226 lines
32 KiB
XML
226 lines
32 KiB
XML
<document id="0A151FF62EA82AB989C936B8D683EAC6" ID-CLB-Dataset="39055" ID-DOI="10.11646/zootaxa.4111.3.4" ID-GBIF-Dataset="ea1baba4-76d0-4d30-8543-ff83c08a489c" ID-ISSN="1175-5326" ID-Zenodo-Dep="271466" ID-ZooBank="474D4226-1F84-4412-A680-5DEF7B3FD58B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463468557192" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2016" docId="43620847FFD1FF86FF64F959FDAF5A3F" docLanguage="en" docName="zootaxa.4111.3.4.pdf.imf" docOrigin="Zootaxa 4111 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speodromia Barnard 1946" docType="treatment" docVersion="9" lastPageNumber="262" masterDocId="BF5B703FFFD1FF87FFF3FFCEFFA65D3B" masterDocTitle="The taxonomy of Speodromia platyarthrodes (Stebbing, 1905) (Crustacea: Brachyura), an unusual dromiid crab endemic to South Africa" masterLastPageNumber="275" masterPageNumber="261" pageNumber="261" updateTime="1698662528666" updateUser="ExternalLinkService">
|
||
<mods:mods id="5D27BFF3B15DBCEB3DED962F971F81B2" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A99C1FDDF2CD0590B3666BC98C9A4D70">
|
||
<mods:title id="42F9017045F18DB16B11984C95899D8B">The taxonomy of Speodromia platyarthrodes (Stebbing, 1905) (Crustacea: Brachyura), an unusual dromiid crab endemic to South Africa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6844F5531BE7033296E69B8D8279D984" type="personal">
|
||
<mods:role id="D883FDE4E64DEAA7FE868B854C110C3A">
|
||
<mods:roleTerm id="9A7A7F51534475B33CF36CB3730ED86A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="954F5A6F31DD1F24E5B5FD2FCA7018A1">Ng, Peter K. L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E2F999F3AAD4E23CDF04F256DD83A52F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="38B1ABDC3EF25EDEA5B51EE9FC89BAE0" type="host">
|
||
<mods:titleInfo id="5CBBBB46297000823EF4F605BC3245A7">
|
||
<mods:title id="644C60FB5A304A3F532065575D2BE273">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="5F5A8B6838B4BE3A4C460F6B226FC3CE">
|
||
<mods:date id="2B9430DAC61673445A0F84FC1623C517">2016</mods:date>
|
||
<mods:detail id="62E8191B81744F96EC243A9F1E24E067" type="volume">
|
||
<mods:number id="BD3BCDE6A7970BD18B569E3A8A9FEB6F">4111</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C83E8BB503BB95C4F910DE147A41F7EB" type="issue">
|
||
<mods:number id="8B98DEEE8CC9269C23E8A97788EEF46F">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8AB5737377656476CF4671313F0232B1" unit="page">
|
||
<mods:start id="99017F093AA4338CC37D7467A08B488B">261</mods:start>
|
||
<mods:end id="205E1933424A6F60FAD6348B0D065962">275</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="CD17C561AEDCA0A6875CC171598335A9">journal article</mods:classification>
|
||
<mods:identifier id="0DA86E9490FDE8CFDC6FACC0B040E6AA" type="CLB-Dataset">39055</mods:identifier>
|
||
<mods:identifier id="780A15E282C7C87BBE695A494A282CD6" type="DOI">10.11646/zootaxa.4111.3.4</mods:identifier>
|
||
<mods:identifier id="F1797F715F241A3059E2D0FF87B42F34" type="GBIF-Dataset">ea1baba4-76d0-4d30-8543-ff83c08a489c</mods:identifier>
|
||
<mods:identifier id="68C93BCE1C5E9716EB59269B48211AC5" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="0EC485CC7C5B7FA5EF511E269F0AB8E5" type="Zenodo-Dep">271466</mods:identifier>
|
||
<mods:identifier id="3B328F355A4B8E750A2597EA31C21408" type="ZooBank">474D4226-1F84-4412-A680-5DEF7B3FD58B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="43620847FFD1FF86FF64F959FDAF5A3F" ID-DOI="http://doi.org/10.5281/zenodo.6056973" ID-GBIF-Taxon="120678432" ID-Zenodo-Dep="6056973" LSID="urn:lsid:plazi:treatment:43620847FFD1FF86FF64F959FDAF5A3F" httpUri="http://treatment.plazi.org/id/43620847FFD1FF86FF64F959FDAF5A3F" lastPageId="1" lastPageNumber="262" pageId="0" pageNumber="261">
|
||
<subSubSection id="83D1EADAFFD1FF87FF64F959FE425B89" box="[151,484,1687,1714]" pageId="0" pageNumber="261" type="nomenclature">
|
||
<paragraph id="CB74B951FFD1FF87FF64F959FE425B89" blockId="0.[151,484,1687,1714]" box="[151,484,1687,1714]" pageId="0" pageNumber="261">
|
||
<heading id="903C0E3DFFD1FF87FF64F959FE425B89" bold="true" box="[151,484,1687,1714]" fontSize="11" level="1" pageId="0" pageNumber="261" reason="1">
|
||
<taxonomicName id="0CCBC2D2FFD1FF87FF64F959FE425B89" ID-CoL="7Q2B4" authority="Barnard, 1946" authorityName="Barnard" authorityYear="1946" box="[151,484,1687,1714]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD1FF87FF64F959FE425B89" bold="true" box="[151,484,1687,1714]" pageId="0" pageNumber="261">
|
||
<emphasis id="F9BF6543FFD1FF87FF64F959FE8E5B8A" bold="true" box="[151,296,1687,1713]" italics="true" pageId="0" pageNumber="261">Speodromia</emphasis>
|
||
<bibRefCitation id="AF5AC4A0FFD1FF87FEDCF956FE425B89" author="Barnard" box="[303,484,1688,1714]" pageId="0" pageNumber="261" refString="Barnard, K. H. (1946) Descriptions of new species of South African decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, Series 11, 13 (102), 361 - 392. http: // dx. doi. org / 10.1080 / 00222934608654562" type="journal article" year="1946">Barnard, 1946</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83D1EADAFFD1FF87FF64F910FA245BCF" box="[151,1410,1758,1781]" pageId="0" pageNumber="261" type="reference_group">
|
||
<paragraph id="CB74B951FFD1FF87FF64F910FA245BCF" blockId="0.[151,1410,1758,1781]" box="[151,1410,1758,1781]" pageId="0" pageNumber="261">
|
||
<treatmentCitationGroup id="EBDB9E7FFFD1FF87FF64F910FA245BCF" box="[151,1410,1758,1781]" pageId="0" pageNumber="261">
|
||
<taxonomicName id="0CCBC2D2FFD1FF87FF64F910FE715BCF" ID-CoL="7Q2B4" authority="Barnard 1946: 370" authorityName="Barnard" authorityPageNumber="370" authorityYear="1946" box="[151,471,1758,1780]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD1FF87FF64F910FEB75BCF" box="[151,273,1758,1780]" italics="true" pageId="0" pageNumber="261">Speodromia</emphasis>
|
||
<treatmentCitation id="4A6A9F40FFD1FF87FEE5F910FE715BCF" author="Barnard" box="[278,471,1758,1780]" page="370" pageId="0" pageNumber="261" year="1946">
|
||
<bibRefCitation id="AF5AC4A0FFD1FF87FEE5F910FE025BCF" author="Barnard" box="[278,420,1758,1780]" pageId="0" pageNumber="261" refString="Barnard, K. H. (1946) Descriptions of new species of South African decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, Series 11, 13 (102), 361 - 392. http: // dx. doi. org / 10.1080 / 00222934608654562" type="journal article" year="1946">Barnard 1946</bibRefCitation>
|
||
: 370
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="4A6A9F40FFD1FF87FE12F910FD045BCF" author="Barnard" box="[481,674,1758,1780]" page="333" pageId="0" pageNumber="261" year="1950">
|
||
<bibRefCitation id="AF5AC4A0FFD1FF87FE12F910FDC95BCF" author="Barnard" box="[481,623,1758,1780]" pageId="0" pageNumber="261" refString="Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38, 1 - 837, figs. 1 - 154." type="journal article" year="1950">Barnard 1950</bibRefCitation>
|
||
: 333
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="4A6A9F40FFD1FF87FD5FF911FCC35BCF" author="McLay" box="[684,869,1758,1780]" page="182" pageId="0" pageNumber="261" year="1993">
|
||
<bibRefCitation id="AF5AC4A0FFD1FF87FD5FF911FC945BCF" author="McLay" box="[684,818,1758,1780]" pageId="0" pageNumber="261" refString="McLay, C. L. (1993) Crustacea: Decapoda: The Sponge Crabs (Dromiidae) of New Caledonia and the Philippines with a review of the genera. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, 10. Memoires du Museum national d'Histoire naturelle, 156, 111 - 251." type="journal article" year="1993">McLay 1993</bibRefCitation>
|
||
: 182
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="4A6A9F40FFD1FF87FC9CF910FB205BCF" author="Guinot" box="[879,1158,1758,1780]" page="74" pageId="0" pageNumber="261" year="2003">
|
||
<bibRefCitation id="AF5AC4A0FFD1FF87FC9CF910FBC65BCF" author="Guinot" box="[879,1120,1758,1780]" pageId="0" pageNumber="261" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae De Haan, 1833, with diagnoses and description of new genera and species (Crustacea, Decapoda, Brachyura). Zoosystema 25 (1), 43 - 129." type="journal article" year="2003">Guinot & Tavares 2003</bibRefCitation>
|
||
: 74
|
||
</treatmentCitation>
|
||
, 113; Ng
|
||
<emphasis id="F9BF6543FFD1FF87FB1BF92EFABA5BCF" box="[1256,1308,1758,1781]" italics="true" pageId="0" pageNumber="261">et al.</emphasis>
|
||
2008: 35.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83D1EADAFFD1FF87FF64F8EFFC125A01" box="[151,948,1825,1850]" pageId="0" pageNumber="261" type="materials_examined">
|
||
<paragraph id="CB74B951FFD1FF87FF64F8EFFC125A01" blockId="0.[151,1436,1825,2030]" box="[151,948,1825,1850]" pageId="0" pageNumber="261">
|
||
<emphasis id="F9BF6543FFD1FF87FF64F8EFFE8D5A01" bold="true" box="[151,299,1825,1850]" pageId="0" pageNumber="261">
|
||
<typeStatus id="147007F3FFD1FF87FF64F8EFFF745A01" box="[151,210,1825,1850]" pageId="0" pageNumber="261">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="0CCBC2D2FFD1FF87FECAF8ECFCA85A01" authority="Stebbing, 1905" authorityName="Stebbing" authorityYear="1905" box="[313,782,1825,1850]" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="261" phylum="Arthropoda" rank="species" species="platyarthrodes">
|
||
<emphasis id="F9BF6543FFD1FF87FECAF8ECFDFD5A02" box="[313,603,1825,1849]" italics="true" pageId="0" pageNumber="261">Dynomene platyarthrodes</emphasis>
|
||
<bibRefCitation id="AF5AC4A0FFD1FF87FD91F8EFFCA85A01" author="Stebbing" box="[610,782,1825,1850]" pageId="0" pageNumber="261" refString="Stebbing, T. R. R. (1905) South African Crustacea. Part III. Marine Investigations in South Africa, Cape of Good Hope, Department of Agriculture, Cape Town 4, 21 - 123, pls. 17 - 21, 21 bis, 22 - 26." type="journal article" year="1905">Stebbing, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83D1EADAFFD1FF86FF34F88BFC5F5FCB" lastPageId="1" lastPageNumber="262" pageId="0" pageNumber="261" type="diagnosis">
|
||
<paragraph id="CB74B951FFD1FF86FF34F88BFC5F5FCB" blockId="0.[151,1436,1825,2030]" lastBlockId="1.[151,1437,151,1797]" lastPageId="1" lastPageNumber="262" pageId="0" pageNumber="261">
|
||
<emphasis id="F9BF6543FFD1FF87FF34F88BFE9C5A65" bold="true" box="[199,314,1861,1886]" pageId="0" pageNumber="261">Diagnosis</emphasis>
|
||
. Carapace transversely subovate, with anterior part produced; gastric, branchial regions swollen, other regions well demarcated; branchial regions with large, deep cavity on sub-branchial side, opening to subbranchial cavity joins shallow grooved area on remaining sub-branchial part of carapace. Front trilobate; supraorbital margin with median spine; anterolateral margin arcuate, anteriorly curving downwards towards pterygostomial region, with 6–8 low, triangular teeth. Epistome with posterior margin separated into 2 truncate lobes by deep median fissure. Antenna with article 1 subrectangular, inner margin with prominent cleft, article 2 subquadrate, distal margin with cleft. Third maxilliped relatively long, inner margin of ischium lined with prominent pectinate crista dentata. Chelipeds closely coapted to lateral sides of carapace when retracted; dorsal margin of merus with large expanded crista on distal half, that on ventral half forming C-shaped concavity; chela stout, outer surface with longitudinal rows of short setae; inner surface of fingers excavate, bracketed by strong teeth. P2 and P3 similar in form; coxa with short ledge-like projection on inner margin of posterior part which locks onto margin of pleonal somites when closed; merus with strong dorsal crista; carpus with median ridge, strong dorsal ridge, 2 low ventral ridges; propodus with median ridge, well developed dorsal ridge, 2 low ventral ridges; dactylo-propodal lock well developed. P4 prominently reduced; dorsal crista of merus distinct; flexor side of distal margin with long fixed spine which opposes strongly recurved, hook-like dactylus. P5 slender, short; merus slender, elongate; carpus, propodus elongate, unarmed; distal margin of propodus with long fixed spine on flexor side, relatively shorter spine on extensor side, dactylus hook-like. Thoracic sternite 4 semicircular; penis not calcified, long, mobile, reaching to median part of coxa of P3. P1–P5 without epipods or podobranchs. Male telson triangular, tip sharp; uropod ventral, ovate, mobile, not visible from dorsal view; no pleopods on somites 3–5. G2 without endopod. Female spermathecal groove (sternal suture 7/8) long, prominent, reaching to sternite 4; aperture to spermatheca on each side of rounded tip; coxae of P2, P3 without locking mechanism as in male; female telson semicircular, uropod relatively more longitudinally ovate compared to males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83D1EADAFFD0FF86FF34FD32FDAF5A3F" pageId="1" pageNumber="262" type="discussion">
|
||
<paragraph id="CB74B951FFD0FF86FF34FD32FCE75EF3" blockId="1.[151,1437,151,1797]" pageId="1" pageNumber="262">
|
||
<emphasis id="F9BF6543FFD0FF86FF34FD32FE925E2E" bold="true" box="[199,308,764,789]" pageId="1" pageNumber="262">Remarks</emphasis>
|
||
. The morphology of
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FDCFFD32FCDC5E2E" authority="Barnard, 1946" authorityName="Barnard" authorityYear="1946" box="[572,890,764,789]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FDCFFD32FD645E2F" box="[572,706,764,788]" italics="true" pageId="1" pageNumber="262">Speodromia</emphasis>
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FD23FD32FCDC5E2E" author="Barnard" box="[720,890,764,789]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1946) Descriptions of new species of South African decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, Series 11, 13 (102), 361 - 392. http: // dx. doi. org / 10.1080 / 00222934608654562" type="journal article" year="1946">Barnard, 1946</bibRefCitation>
|
||
</taxonomicName>
|
||
, is quite unlike any known dromiid, with its transversely ovate carapace and anteriorly produced frontal region. In fact, in its carapace form, it superficially resembles species of
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FE77FC8AFD075E67" authority="Stimpson, 1871" authorityName="Stimpson" authorityYear="1871" box="[388,673,836,861]" class="Malacostraca" family="Aethridae" genus="Osachila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FE77FC8AFE4F5E67" box="[388,489,836,860]" italics="true" pageId="1" pageNumber="262">Osachila</emphasis>
|
||
Stimpson, 1871
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FD5EFC8AFB9F5E67" authority="Manning & Holthuis, 1981" authorityName="Manning & Holthuis" authorityYear="1981" box="[685,1081,836,861]" class="Malacostraca" family="Aethridae" genus="Sakaila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FD5EFC8AFCA75E67" box="[685,769,836,860]" italics="true" pageId="1" pageNumber="262">Sakaila</emphasis>
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FCFBFC8AFB9F5E67" author="Manning" box="[776,1081,836,861]" pageId="1" pageNumber="262" refString="Manning, R. B. & Holthuis, L. B. (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology 306, i - xii, 1 - 379, figs. 1 - 88, table 1." type="journal article" year="1981">Manning & Holthuis, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FB87FC8AFF745EBB" authority="Latreille in Cuvier, 1816" authorityName="Latreille in Cuvier" authorityYear="1816" class="Malacostraca" family="Aethridae" genus="Aethra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FB87FC8AFB665E67" box="[1140,1216,836,860]" italics="true" pageId="1" pageNumber="262">Aethra</emphasis>
|
||
Latreille in Cuvier, 1816
|
||
</taxonomicName>
|
||
, genera now placed in
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FE26FCA9FD6A5EBB" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[469,716,871,896]" class="Malacostraca" family="Aethridae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="family">Aethridae Dana, 1851</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FCFBFCA9FC0C5EBB" author="Rathbun" box="[776,938,871,896]" pageId="1" pageNumber="262" refString="Rathbun, M. J. (1916) Description of three species of crabs (Osachila) from the eastern coast of North America. Proceedings of the United States National Museum, 50, 647 - 652. http: // dx. doi. org / 10.5479 / si. 00963801.50 - 2138.647" type="journal article" year="1916">Rathbun 1916</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FC46FCA9FB475EBB" author="Manning" box="[949,1249,871,896]" pageId="1" pageNumber="262" refString="Manning, R. B. & Holthuis, L. B. (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology 306, i - xii, 1 - 379, figs. 1 - 88, table 1." type="journal article" year="1981">Manning & Holthuis 1981</bibRefCitation>
|
||
; Ng 1999; Ng
|
||
<emphasis id="F9BF6543FFD0FF86FA7BFCA7FF125E9F" italics="true" pageId="1" pageNumber="262">et al.</emphasis>
|
||
2008;
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FEFBFC42FE465E9F" author="Martin" box="[264,480,908,933]" pageId="1" pageNumber="262" refString="Martin, J. W., Godwin, S. & Moffitt, R. (2009) Additions to the decapod crustacean fauna of the Hawaiian Islands, I. A review of the genus Sakaila Manning and Holthuis, 1981 (Decapoda, Brachyura, Aethridae), with the description of a new species from the Northwestern Hawaiian Islands. Zootaxa 2018, 25 - 44." type="journal article" year="2009">
|
||
Martin
|
||
<emphasis id="F9BF6543FFD0FF86FEADFC43FE3F5E9F" box="[350,409,908,932]" italics="true" pageId="1" pageNumber="262">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
). The highly reduced last two ambulatory legs with the subcheliform dactylus and propodus (P4 and P5), however, make it clear it is a dromiid.
|
||
</paragraph>
|
||
<paragraph id="CB74B951FFD0FF86FF34FC1AFD6159FF" blockId="1.[151,1437,151,1797]" pageId="1" pageNumber="262">
|
||
The characteristic form and density of the setae on the surfaces of
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FC45FC1AFB9A5ED7" box="[950,1084,980,1004]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FC45FC1AFB9A5ED7" box="[950,1084,980,1004]" italics="true" pageId="1" pageNumber="262">Speodromia</emphasis>
|
||
</taxonomicName>
|
||
are not known to be present in any other dromiids and were accurately figured by
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FD09FC39FC00592B" author="Barnard" box="[762,934,1015,1040]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38, 1 - 837, figs. 1 - 154." type="journal article" year="1950">Barnard (1950</bibRefCitation>
|
||
: fig. 64b, c). On the dorsal surface of the carapace, ventral surfaces of the thoracic sternum and pleon, and surfaces of the pereiopods, these setae are short and squamiform, forming clumps which resemble patches of granules but do not completely obscure the surface. Along the margins of the structures, however, these scale-like structures are larger and stalked to various degrees, resulting in distinct clavate setae. These clavate setae are often closely packed, making the cristae and margins higher and more prominent than they actually are.
|
||
</paragraph>
|
||
<paragraph id="CB74B951FFD0FF86FF34FB01FACE58A6" blockId="1.[151,1437,151,1797]" pageId="1" pageNumber="262">
|
||
The deep cavity in the sub-branchial region (
|
||
<figureCitation id="53F0A5D4FFD0FF86FD39FB01FCAB59D3" box="[714,781,1231,1256]" captionStart="FIGURE 5" captionStartId="6.[151,250,1778,1800]" captionTargetBox="[151,1436,193,1755]" captionTargetId="figure@6.[151,1436,193,1756]" captionTargetPageId="6" captionText="FIGURE 5. Speodromia platyarthrodes, male (28.1 × 22.1 mm) (SAM MB-A 000816), South Africa. A, left lateral side of cephalothorax; B, right lateral side of cephalothorax (denuded); C, left side of sub-branchial surface showing cavity; D, left side of sub-branchial cavity showing groove towards posterior carapace margin (denuded); E, F, posterior view of cephalothorax." httpUri="https://zenodo.org/record/271471/files/figure.png" pageId="1" pageNumber="262">Fig. 5</figureCitation>
|
||
A–D) is on the anterior half of the carapace, and forms a chamber in the branchial area which is connected to the gill chamber along the ventral margin (see
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FB08FB3AFA3E5836" author="Barnard" box="[1275,1432,1268,1293]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38, 1 - 837, figs. 1 - 154." type="journal article" year="1950">Barnard 1950</bibRefCitation>
|
||
: pl. 64d). With regards to cavities on the carapace,
|
||
<emphasis id="F9BF6543FFD0FF86FD2CFAD6FC50580B" box="[735,1014,1304,1328]" italics="true" pageId="1" pageNumber="262">Sphaeorodromia brizops</emphasis>
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FC0CFAD7FA84580B" author="McLay" box="[1023,1314,1303,1329]" pageId="1" pageNumber="262" refString="McLay, C. L. & Crosnier, A. (1991) Description of a new and unusual species of Sphaerodromia (Brachyura, Dromiidae) from the Seychelles Islands. Bulletin du Museum national d'histoire naturelle, 4 A (Zoology), 13 (1 - 2), 181 - 188, pl. 1." type="journal article" year="1991">McLay & Crosnier, 1991</bibRefCitation>
|
||
, described from one male specimen from the
|
||
<collectingCountry id="B3DCF9C1FFD0FF86FDE5FAF2FD37586E" box="[534,657,1340,1365]" name="Seychelles" pageId="1" pageNumber="262">Seychelles</collectingCountry>
|
||
, has an unusually deep and prominent orbit but the branchial and subbranchial regions are normal (
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FE0BFAAEFCB95843" author="McLay" box="[504,799,1375,1400]" pageId="1" pageNumber="262" refString="McLay, C. L. & Crosnier, A. (1991) Description of a new and unusual species of Sphaerodromia (Brachyura, Dromiidae) from the Seychelles Islands. Bulletin du Museum national d'histoire naturelle, 4 A (Zoology), 13 (1 - 2), 181 - 188, pl. 1." type="journal article" year="1991">McLay & Crosnier 1991</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FCC5FA91FC465843" author="Barnard" box="[822,992,1375,1400]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1946) Descriptions of new species of South African decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, Series 11, 13 (102), 361 - 392. http: // dx. doi. org / 10.1080 / 00222934608654562" type="journal article" year="1946">Barnard (1946</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FC1CFA91FB8F5843" author="Barnard" box="[1007,1065,1375,1400]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38, 1 - 837, figs. 1 - 154." type="journal article" year="1950">1950</bibRefCitation>
|
||
) also dissected a specimen and showed that all the pereiopods (including the chelipeds) do not possess an epipod (cf.
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FBA9FA4AFB5158A6" author="Barnard" box="[1114,1271,1412,1437]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38, 1 - 837, figs. 1 - 154." type="journal article" year="1950">Barnard 1950</bibRefCitation>
|
||
: fig. 64e).
|
||
</paragraph>
|
||
<paragraph id="CB74B951FFD0FF86FF34FA69FACD5B86" blockId="1.[151,1437,151,1797]" pageId="1" pageNumber="262">
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FF34FA69FE4858FB" author="Guinot" box="[199,494,1447,1472]" pageId="1" pageNumber="262" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae De Haan, 1833, with diagnoses and description of new genera and species (Crustacea, Decapoda, Brachyura). Zoosystema 25 (1), 43 - 129." type="journal article" year="2003">Guinot & Tavares (2003)</bibRefCitation>
|
||
divided the
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FD76FA69FD5B58FB" box="[645,765,1447,1472]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="family">Dromiidae</taxonomicName>
|
||
into three subfamilies:
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FBE4FA69FB2958FB" box="[1047,1167,1447,1472]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Dromiinae">Dromiinae</taxonomicName>
|
||
De Haan, 1833 s. str.,
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FF64FA02FEE158DE" box="[151,327,1484,1509]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Hypoconchinae">Hypoconchinae</taxonomicName>
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FEA1FA02FDD758DF" author="Guinot" box="[338,625,1484,1509]" pageId="1" pageNumber="262" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae De Haan, 1833, with diagnoses and description of new genera and species (Crustacea, Decapoda, Brachyura). Zoosystema 25 (1), 43 - 129." type="journal article" year="2003">Guinot & Tavares, 2003</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FD45FA02FC2558DE" box="[694,899,1484,1509]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Sphaerodromiinae">Sphaerodromiinae</taxonomicName>
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FC7CFA02FB0858DF" author="Guinot" box="[911,1198,1484,1509]" pageId="1" pageNumber="262" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae De Haan, 1833, with diagnoses and description of new genera and species (Crustacea, Decapoda, Brachyura). Zoosystema 25 (1), 43 - 129." type="journal article" year="2003">Guinot & Tavares, 2003</bibRefCitation>
|
||
, on the basis of the structure of the carapace, penis, male pleopodal formula, structure of P4 and P5, form and position of the male and female uropods, relative length of the female thoracic sternal sutures 7/8, position of the spermathecal apertures, and presence of an endopod of the G2.
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FDBDF9F9FCC15B6B" author="Guinot" box="[590,871,1591,1616]" pageId="1" pageNumber="262" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae De Haan, 1833, with diagnoses and description of new genera and species (Crustacea, Decapoda, Brachyura). Zoosystema 25 (1), 43 - 129." type="journal article" year="2003">Guinot & Tavares (2003)</bibRefCitation>
|
||
retained
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FC3CF9F6FBF35B6B" box="[975,1109,1592,1616]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FC3CF9F6FBF35B6B" box="[975,1109,1592,1616]" italics="true" pageId="1" pageNumber="262">Speodromia</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FB8AF9F9FB575B6B" box="[1145,1265,1591,1616]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Dromiinae">Dromiinae</taxonomicName>
|
||
s. str. based on
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FF64F992FEEC5B4E" author="Barnard" box="[151,330,1628,1653]" pageId="1" pageNumber="262" refString="Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38, 1 - 837, figs. 1 - 154." type="journal article" year="1950">Barnard (1950)</bibRefCitation>
|
||
. The study of the present specimens confirms this classification. While the carapace shape of
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FF64F94EFEBB5BA3" box="[151,285,1664,1688]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FF64F94EFEBB5BA3" box="[151,285,1664,1688]" italics="true" pageId="1" pageNumber="262">Speodromia</emphasis>
|
||
</taxonomicName>
|
||
is unusual and the deep sub-branchial cavity is a unique character, all the other features used to define
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FF64F96AFEA95B86" box="[151,271,1700,1725]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Dromiinae">Dromiinae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FECAF96AFDF45B87" author="Guinot" box="[313,594,1700,1725]" pageId="1" pageNumber="262" refString="Guinot, D. & Tavares, M. (2003) A new subfamilial arrangement for the Dromiidae De Haan, 1833, with diagnoses and description of new genera and species (Crustacea, Decapoda, Brachyura). Zoosystema 25 (1), 43 - 129." type="journal article" year="2003">Guinot & Tavares (2003)</bibRefCitation>
|
||
apply to
|
||
<taxonomicName id="0CCBC2D2FFD0FF86FD48F96AFCE75B87" box="[699,833,1700,1724]" class="Malacostraca" family="Dromiidae" genus="Speodromia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9BF6543FFD0FF86FD48F96AFCE75B87" box="[699,833,1700,1724]" italics="true" pageId="1" pageNumber="262">Speodromia</emphasis>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FC42F96AFBD95B87" author="Guinot" box="[945,1151,1700,1725]" pageId="1" pageNumber="262" refString="Guinot, D., Tavares, M. & Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa 3665 (1), 1 - 414. http: // dx. doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
|
||
Guinot
|
||
<emphasis id="F9BF6543FFD0FF86FBF6F96BFB985B87" box="[1029,1086,1700,1724]" italics="true" pageId="1" pageNumber="262">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FB78F96AFAF95B86" author="Davie" box="[1163,1375,1700,1725]" pageId="1" pageNumber="262" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 b) Classification and Systematics and classification of Brachyura, pp. 1049 - 1130. In: Castro, P., Davie, P., Guinot, D., Schram, F. R. & Vaupel Klein, J. C. von (Eds.), Decapoda: Brachyura, Treatise on Zoology - Anatomy, Taxonomy, Biology, 9, part C-II, Brill, Leiden." type="journal article" year="2015" yearSuffix="b">
|
||
Davie
|
||
<emphasis id="F9BF6543FFD0FF86FB26F96BFAAB5B87" box="[1237,1293,1700,1724]" italics="true" pageId="1" pageNumber="262">et al.</emphasis>
|
||
2015b
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CB74B951FFD0FF86FF34F909FDAF5A3F" blockId="1.[151,1437,151,1797]" pageId="1" pageNumber="262">
|
||
The authority of the genus is usually given as “Barnard, 1947”, but the publication was actually released in 1946 (see Ng &
|
||
<bibRefCitation id="AF5AC4A0FFD0FF86FEA3F923FE475A3E" author="Kumar" box="[336,481,1772,1797]" pageId="1" pageNumber="262" refString="Ng, P. K. L. & Kumar, A. B. (2015) The species of Moloha Barnard, 1946, from the western Indian Ocean, with the description of a new species from India (Crustacea: Brachyura: Homolidae). European Journal of Taxonomy, 166, 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.166" type="journal article" year="2015">Kumar 2015</bibRefCitation>
|
||
: 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |