275 lines
26 KiB
XML
275 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne & Haring, Elisabeth" docDate="2021" docId="418C14069E195FBCB8E7E2B00CAC0B07" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Charpentieria dyodon subsp. thomasiana" docType="treatment" docVersion="5" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1732485878053" updateUser="ExternalLinkService">
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<mods:titleInfo id="64A3B1976B4FE8388EFADB9F7DE540A5">
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<mods:title id="49F13C2567779DB79EE12B8588C6C597">Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:namePart id="11B390339ABE9B74B059D55AE61BF0F8">De Mattia, Willy</mods:namePart>
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<mods:nameIdentifier id="C4DEFF6BA88A6843F94AFACEB282C0BA" type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
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<mods:affiliation id="03ACD06922F78E6B796D2970291C124A">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier id="A14A2D8AE6F2109D470EC99EDE1E4E62" type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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<mods:roleTerm id="18E93B17E0203EAE4CBF96AEE8427745">Author</mods:roleTerm>
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<mods:namePart id="F55436667B42474BD9E4749B9FB27FD2">Reier, Susanne</mods:namePart>
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<mods:affiliation id="D31551BC7124CDDB8E19C16E57A27573">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:role>
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<mods:namePart id="B8DEC40157B3FE123F47F97781908159">Haring, Elisabeth</mods:namePart>
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<mods:affiliation id="5EA9ABA8432651F954D1D414E632308B">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:typeOfResource id="BACFC351C99E177FCDEE4F73FF694757">text</mods:typeOfResource>
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<mods:title id="7FB90FC91E419436DFCDF63E41FE81AC">ZooKeys</mods:title>
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<mods:date id="872A9FA7A9DAF0D6B093BC8D04F4912B">2021</mods:date>
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<mods:number id="5A1567FF0034C6D9C3852B5297AB0EC0">2021-12-14</mods:number>
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<mods:number id="FBCB3AC95287F29E5FA16ECE1C3A0E0D">1077</mods:number>
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<mods:url id="C5A3C1E05BA13BD31BE9FB6D7AF4A0F4">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
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<mods:classification id="3D96E41DB30BE876DCF25F89A206A94A">journal article</mods:classification>
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<mods:identifier id="25C2D6B0443EAC039D13775CA00E068B" type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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</mods:mods>
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<treatment id="418C14069E195FBCB8E7E2B00CAC0B07" ID-GBIF-Taxon="191645955" LSID="urn:lsid:plazi:treatment:418C14069E195FBCB8E7E2B00CAC0B07" httpUri="http://treatment.plazi.org/id/418C14069E195FBCB8E7E2B00CAC0B07" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Clausiliidae" scope_infraOrder="Clausilioidei" scope_order="Stylommatophora" scope_phylum="Mollusca" scope_superFamily="Clausilioidea">
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<subSubSection id="87178758BD6D5F09B30BB4A5D34AF35F" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="464B9221B397C2EF06E4AC0FA50A2FE6" pageId="0" pageNumber="1">
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<taxonomicName id="A753043FCE46AB3FAD390A7BA9698785" ID-CoL="7GNWT" LSID="418C1406-9E19-5FBC-B8E7-E2B00CAC0B07" authority="(Kuester, 1850)" baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
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Charpentieria dyodon thomasiana (
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<normalizedToken id="AAC32D5580203A2C9EB47B51F8525174" originalValue="Küster">Kuester</normalizedToken>
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, 1850)
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</taxonomicName>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="943FC6577A364376D6ABAC79B75D2168" pageId="0" pageNumber="1" type="description">
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<paragraph id="8B539181D6B778F896B76E15812FAFA8" pageId="0" pageNumber="1">
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<figureCitation id="A1AF7D50ADE84A325B968EDE74BFFCEA" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">Figs 41.1-41.4, 42.3, 42.4</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="6FFEF4F7A312D711E4C65266EC34C20B" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="FF78745CE7D0A04A8834BD982C7DF543" pageId="0" pageNumber="1">Specimens examined.</paragraph>
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<paragraph id="67AFE25ACC169B06A92E066CE5B8A946" pageId="0" pageNumber="1">
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<materialsCitation id="FD2864A51EE78F5CAD7E03D2DCA343E6" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. De Mattia, J. Macor" country="Italy" elevation="1100" latitude="45.63028" location="Santuario di Oropa." longLatPrecision="1" longitude="7.978353" municipality="Biella" specimenCount="1" stateProvince="Piedmont">
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<collectingCountry id="4BBA7469C2CD19A442D72B3564C979EC" name="Italy">Italy</collectingCountry>
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,
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<collectingRegion id="768F486DD167A92DB8C7EC0FAB99819D" country="Italy" name="Piemonte">Piedmont</collectingRegion>
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,
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<collectingMunicipality id="77942B055E10A4713D991A7ADA4C0713">Biella</collectingMunicipality>
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,
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<location id="F8278824FC4FBF69DDBAFBF741B434BB" LSID="urn:lsid:plazi:treatment:418C14069E195FBCB8E7E2B00CAC0B07:F8278824FC4FBF69DDBAFBF741B434BB" country="Italy" latitude="45.63028" longLatPrecision="1" longitude="7.978353" municipality="Biella" name="Santuario di Oropa." stateProvince="Piedmont">Santuario di Oropa.</location>
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<elevation id="5E27424C454ED158A8B98904ED0401AE" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
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<quantity id="D81B52AC9AC9D691F1F6A577B1481CA5" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</quantity>
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asl
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</elevation>
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<geoCoordinate id="12BBB82DE2856400A2ADA49E26597580" degrees="45" direction="north" minutes="37" orientation="latitude" precision="1" seconds="49.01" value="45.63028">45°37'49.01"N</geoCoordinate>
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,
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<geoCoordinate id="1781AE5616D14ECB770CAA26B24479C8" degrees="07" direction="east" minutes="58" orientation="longitude" precision="1" seconds="42.07" value="7.978353">07°58'42.07"E</geoCoordinate>
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,
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<collectorName id="28B694734AB651052593280C5042C5FF">W. De Mattia</collectorName>
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and
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<collectorName id="DC20840704DF9DE9D4B704944B272F90">J. Macor</collectorName>
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leg. and det., 3 dissected spm
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</materialsCitation>
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.
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||
<materialsCitation id="03EA9AF7C1EC747F5A6F5C53A413FA8B" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Italy" county="Biella" elevation="1200" latitude="45.692326" location="Pra del Vecia" longLatPrecision="1" longitude="7.967325" municipality="Sentiero" specimenCount="1" stateProvince="Piedmont">
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<collectingCountry id="30DB0AFD7894EE9C07CDE82F500F2C9B" name="Italy">Italy</collectingCountry>
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,
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<collectingRegion id="DCB72C18EAF45942141DD813A21D2BD1" country="Italy" name="Piemonte">Piedmont</collectingRegion>
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,
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<collectingCounty id="87F9D68EA15A7EF41C9C1115FE866F01">Biella</collectingCounty>
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,
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<collectingMunicipality id="A62CAE1CE4E695704C65076E8BDE5EE8">Sentiero</collectingMunicipality>
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per
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<location id="B358FEC85D15F47DFEC16D06C3D01684" LSID="urn:lsid:plazi:treatment:418C14069E195FBCB8E7E2B00CAC0B07:B358FEC85D15F47DFEC16D06C3D01684" country="Italy" county="Biella" latitude="45.692326" longLatPrecision="1" longitude="7.967325" municipality="Sentiero" name="Pra del Vecia" stateProvince="Piedmont">
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<normalizedToken id="CE92D30C6F4E0AF3C4A26153DB654258" originalValue="Prà">Pra</normalizedToken>
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del Vecia
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</location>
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||
, Piedicavallo.
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<elevation id="42CD05B867D38F1C3A7C56607F820D2A" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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||
<quantity id="85AEF8ADEBE52C2ECD1FF896549994C9" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
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||
asl
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</elevation>
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||
,
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||
<geoCoordinate id="3FB27BCCB9B8403FFC88C8834A0D826D" degrees="45" direction="north" minutes="41" orientation="latitude" precision="1" seconds="32.37" value="45.692326">45°41'32.37"N</geoCoordinate>
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,
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<geoCoordinate id="64303FB465A6E12F9485618B8FFF7F70" degrees="07" direction="east" minutes="58" orientation="longitude" precision="1" seconds="2.37" value="7.967325">07°58'2.37"E</geoCoordinate>
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, leg. and det., 2 dissected spm
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A33434BCC19DDD6DEBE745E83555E0A1" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="7FE79E7F962023CBC14951BB91403AF6" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="9EA414FE6C4D7B575975559B39C981C0" pageId="0" pageNumber="1">
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<taxonomicName id="1A97B4C47DF9EAB9183BD6A79ACC3609" baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
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<emphasis id="43D1F7CC41EEAB7145DF427193609746" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
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</taxonomicName>
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is found in the surroundings of Oropa and Piedicavallo, surroundings of Biella (Piedmont).
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</paragraph>
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<caption id="611C874864CFA84E99B79A7C53BA654E" doi="10.3897/zookeys.1077.67081.figure40" httpUri="https://binary.pensoft.net/fig/622832" pageId="0" pageNumber="1" start="Figure 40" startId="F40">
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<paragraph id="4C091D686AE679F7FC1E6D057B878978" pageId="0" pageNumber="1">
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<emphasis id="0F5A4C6BA659AE292657E46878119BCA" bold="true" pageId="0" pageNumber="1">Figure 40.</emphasis>
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<taxonomicName id="E5437E94133D3BAF6372827592AF8538" authority="dyodon" authorityName="dyodon (S. Studer" authorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="dyodon">
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<emphasis id="B815C6BED86EAB9236BFC1E6F1012933" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon dyodon</emphasis>
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</taxonomicName>
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(S. Studer, 1820), Verbano
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<emphasis id="D26914AA6FA222347E4CEFC9460A1389" bold="true" pageId="0" pageNumber="1">40.1</emphasis>
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whole distal genital organs
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<emphasis id="47997FC7D091886652F8CF1B7ECCCAB7" bold="true" pageId="0" pageNumber="1">40.2</emphasis>
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internal distal part of genital organs.
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||
<taxonomicName id="41DB0C1D42FF1B53449C3AC79E3BAFF9" baseAuthorityName="Stabile" baseAuthorityYear="1859" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. alpina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="alpina">
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<emphasis id="DEF1AB6C1DAE28084F0A682D8B239253" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon alpina</emphasis>
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</taxonomicName>
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(Stabile, 1859). Val di Lanzo, Usseglio
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<emphasis id="FF2220DAADCE1F627247E9801B3E1576" bold="true" pageId="0" pageNumber="1">40.3</emphasis>
|
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whole distal genital organs
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<emphasis id="6B0BB33498B69D1AAE7FF7C958D6159B" bold="true" pageId="0" pageNumber="1">40.4</emphasis>
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internal distal part of genital organs.
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</paragraph>
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</caption>
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||
</subSubSection>
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<subSubSection id="8D0EA8B88E8EBCC4AEEAC6B3C6D95BF1" pageId="0" pageNumber="1" type="external morphology of the genital organs">
|
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<paragraph id="3682F2E5C60F19445BB8C27596B60F4B" pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
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<paragraph id="013A47B0987C60262D91C69D4122BF88" pageId="0" pageNumber="1">
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<emphasis id="752C49F7821CF06BD85D42D38572AA9F" bold="true" pageId="0" pageNumber="1">
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(Fig.
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<figureCitation id="1A0855C296E279A43FE855A07BB3CBB1" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
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.1, 41.3).
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</emphasis>
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The FO is half as long as the V. The FDBC is shorter than the BC+SDBC (SDBC+BC). The BC+SDBC is club-like and longer than the V with slight distinction between the SDBC and the BC. The apex is pointed. The D is slightly longer than the BC+SDBC, thinner with a round apex. The V is long and cylindrical. The PC is longer than the V. The PR is long and robust. The E is slightly shorter than the P and thin.
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</paragraph>
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</subSubSection>
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<subSubSection id="9E2D47D16843DE69F7EB02192696A400" pageId="0" pageNumber="1" type="internal morphology of the genital organs">
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||
<paragraph id="86A04EC2CECA56101F4D1213F45C8954" pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
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||
<paragraph id="8DCC2035E1DE077DEC529FAC9BBAB30B" pageId="0" pageNumber="1">
|
||
<emphasis id="B15C8BBDF50C389F9E7E62570EE1DD78" bold="true" pageId="0" pageNumber="1">
|
||
(Fig.
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||
<figureCitation id="A1610C0523A83E5AF82B2CAE88E99484" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
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.2, 41.4).
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</emphasis>
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The A and the P are smooth, with a very fine granulated sculpturing. The V shows many smooth longitudinal pleats. The smooth simple PP is simple, elongated with a pointed apex. Its base is partially connected to the ELP and transversally extends along the transition wall, narrowing the transition passage. The epiphallar formula is: PP(ELP). The E shows two main longitudinal moderately fringed pleats that proximally fade before the VD.
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</paragraph>
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||
<caption id="1F3851B776BC122AF759F17E1C8040DD" doi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
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||
<paragraph id="2D8A65F4106B8A56D0E1F9BB96FB24DF" pageId="0" pageNumber="1">
|
||
<emphasis id="69A0507050A9F4245871A941039B3198" bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
|
||
<taxonomicName id="86C7B52123B078B64B4F192754740F81" baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
|
||
<emphasis id="F34F6B2908D205F242932E1A123F7B5D" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="FC344F125DA2FB6F7F379C903AAB5956" originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850), Piedicavallo
|
||
<emphasis id="C1A4975A293E1E455F33EE7FBE7A0346" bold="true" pageId="0" pageNumber="1">41.1</emphasis>
|
||
whole distal genital organs
|
||
<emphasis id="EAEEADBBBD00D8E36C968DA1D9FC3012" bold="true" pageId="0" pageNumber="1">41.2</emphasis>
|
||
internal distal part of genital organs. Oropa, Biella
|
||
<emphasis id="92CCF7AB049A4D6DE25D1E59705BC001" bold="true" pageId="0" pageNumber="1">41.3</emphasis>
|
||
whole distal genital organs
|
||
<emphasis id="EDB61D2EB717E3F9AEFFE623466BCD6D" bold="true" pageId="0" pageNumber="1">41.4</emphasis>
|
||
internal distal part of genital organs.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="F0CAFB1BB0C0B53915C3C4563ACB6561" pageId="0" pageNumber="1" type="brief genital anatomical description of the subspecific taxa of charpentieria ornata and charpentieria itala">
|
||
<paragraph id="DA80464756F872EA1290A28871BBABCF" pageId="0" pageNumber="1">
|
||
Brief genital anatomical description of the subspecific taxa of
|
||
<taxonomicName id="62FC7CFDD6985A7DF31F61879B3675EF" baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria ornata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ornata">
|
||
<emphasis id="3E811E7CFFAECFCDE947DF68B193F588" italics="true" pageId="0" pageNumber="1">Charpentieria ornata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B8DFB1B818B598195A229CF8025089C1" baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis id="5C83070B091989810D705E22845EDAB6" italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="A456EF337E2AB972F4108A1B07A22EF2" pageId="0" pageNumber="1">
|
||
The Southern Alpine
|
||
<taxonomicName id="A7A9149B3F7D50E177FE281361EFD75B" lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis id="81C359F7D677B2B4458A93A71FD6AC76" italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
is distributed from Como Lake to the Garda Lake, Trentino, Veneto (including Colli Berici and Colli Euganei) and the Pre-Alps of Friuli.
|
||
<bibRefCitation id="3162271EAD6D0B4F0565B13266322D1E" author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">Nordsieck (1963b)</bibRefCitation>
|
||
provided, for the eastern border of its distribution, the axis Pordenone-Ampezzo. Recent collecting extended its eastern distribution as far Udine and Cividale in Friuli (WDM, personal unpublished data). Its eastern distribution range overlaps with
|
||
<taxonomicName id="784D459D02D6D0120E86BE6588E85BB2" lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis id="08DB93D824233E83ECE973356D99B5ED" italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
and in few localities they are found syntopic (e.g., San Romedio, Trento, where
|
||
<taxonomicName id="524CCF29AAE47D3B9E285BAC02877D17" authority="stenzii" lsidName="C. stenzii subsp. stenzii" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="stenzii">
|
||
<emphasis id="4F13266474AAD0052D06250B1EC04A52" italics="true" pageId="0" pageNumber="1">C. stenzii stenzii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4A97F31DA1C8433EA3A3B269EB7C4155" lsidName="C. itala subsp. serravalensis" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="serravalensis">
|
||
<emphasis id="8C0B8D4A84D10D6428934E4BF56FFBEF" italics="true" pageId="0" pageNumber="1">C. itala serravalensis</emphasis>
|
||
</taxonomicName>
|
||
are found syntopic). Concerning the nomenclature of
|
||
<taxonomicName id="73DA25B8BFFED539DDF5580127E68EDC" lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis id="9C230F6CC81B21953C0DE19F372E3183" italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
, we adopted the system by Scheel and Hausdorf (2012) that considered
|
||
<taxonomicName id="BA07B1007421D3C2A90BFC498E55F3F3" lsidName="C. clavata" pageId="0" pageNumber="1" rank="species" species="clavata">
|
||
<emphasis id="E7EE806BC0AC5AA3794774E0312F5954" italics="true" pageId="0" pageNumber="1">C. clavata</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName id="9AC1D13A08EEC3F1E2D05D4DADE9F958" lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis id="8AD8AD60E830E5B12C485C27FC321419" italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="60D202CB6C27C7AC325D5E579D006AFC" doi="10.3897/zookeys.1077.67081.figure42" httpUri="https://binary.pensoft.net/fig/622834" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
|
||
<paragraph id="D5C4E544500D0FD1B49D0B296276C280" pageId="0" pageNumber="1">
|
||
<emphasis id="E69CC009A95EF688B06DDD5721DDA3CC" bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
|
||
<taxonomicName id="A1FB98756E8F6D2A38E154BB1714467D" authority="dyodon" authorityName="dyodon (S. Studer" authorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="dyodon">
|
||
<emphasis id="26304AC86C8A03FE3CB34446EB2386C4" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon dyodon</emphasis>
|
||
</taxonomicName>
|
||
(S. Studer, 1820), Verbano
|
||
<emphasis id="BE8DF775D015373B4849FB210855BC31" bold="true" pageId="0" pageNumber="1">42.1</emphasis>
|
||
shell.
|
||
<taxonomicName id="B45022BC23069794C842B53EC1FF7CB9" baseAuthorityName="Stabile" baseAuthorityYear="1859" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. alpina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="alpina">
|
||
<emphasis id="C3BBC09A36C8A00B0A03FD6607EBCACB" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon alpina</emphasis>
|
||
</taxonomicName>
|
||
(Stabile, 1859). Val di Lanzo, Usseglio
|
||
<emphasis id="6246DF956BBBE843D8B84A07CA6BDE1B" bold="true" pageId="0" pageNumber="1">42.2</emphasis>
|
||
shell.
|
||
<taxonomicName id="712EB6EC2A456319F84769DDE6B110B7" baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
|
||
<emphasis id="3DA16010C30136860757359DC0FDAAAF" italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="A0155460186B6A4D4DE0B97420112469" originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850), Piedicavallo
|
||
<emphasis id="28B2539CF2DEB94F2DBF5D2AAB2420F8" bold="true" pageId="0" pageNumber="1">42.3</emphasis>
|
||
shell. Oropa, Biella
|
||
<emphasis id="87FDCAAB9178B3FC685B026D5AA7B867" bold="true" pageId="0" pageNumber="1">42.4</emphasis>
|
||
shell.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="94E5E9EA5781EA692C5757621281D6B6" pageId="0" pageNumber="1">
|
||
Its shell is weakly striated to frankly ribbed, with evident sutural papillae and a strong lunella, which is remarkably curved. Twelve out of the seventeen currently valid subspecies of
|
||
<taxonomicName id="3D3E14FF07BE1CA93A2616ED768EA2A0" lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis id="CDAD041B1D1FE71DA0F5049B98CFCAAC" italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
were anatomically investigated, revealing a common overall arrangement of the genital organs. The pseudopapilla can be either smooth or markedly wrinkled and occupies almost the whole penial volume.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |