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<mods:title id="40274767BF055DBBE87BD348FFD49F8F">The Californian clade of Cybaeus (Araneae: Cybaeidae) in the Nearctic: the septatus species group and three unplaced species</mods:title>
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<treatment id="39264D0AFFE5AD0EFF6F658BFD2DFBE3" ID-DOI="http://doi.org/10.5281/zenodo.6313355" ID-GBIF-Taxon="193399505" ID-Zenodo-Dep="6313355" LSID="urn:lsid:plazi:treatment:39264D0AFFE5AD0EFF6F658BFD2DFBE3" httpUri="http://treatment.plazi.org/id/39264D0AFFE5AD0EFF6F658BFD2DFBE3" lastPageId="2" lastPageNumber="191" pageId="1" pageNumber="190">
<subSubSection id="F995AF97FFE5AD0DFF6F658BFE57F90D" box="[151,480,1729,1756]" pageId="1" pageNumber="190" type="nomenclature">
<paragraph id="B130FC1CFFE5AD0DFF6F658BFE57F90D" blockId="1.[151,480,1729,1756]" box="[151,480,1729,1756]" pageId="1" pageNumber="190">
<heading id="EA784B70FFE5AD0DFF6F658BFE57F90D" bold="true" box="[151,480,1729,1756]" fontSize="11" level="1" pageId="1" pageNumber="190" reason="1">
<emphasis id="83FB200EFFE5AD0DFF6F658BFE57F90D" bold="true" box="[151,480,1729,1756]" pageId="1" pageNumber="190">
The
<taxonomicName id="768F879FFFE5AD0DFF376588FE86F90A" ID-CoL="32RHC" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[207,305,1730,1756]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE5AD0DFF376588FE86F90A" bold="true" box="[207,305,1730,1756]" italics="true" pageId="1" pageNumber="190">septatus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F995AF97FFE5AD0EFF6F6443FCDBFE07" lastPageId="2" lastPageNumber="191" pageId="1" pageNumber="190" type="diagnosis">
<paragraph id="B130FC1CFFE5AD0EFF6F6443FCA3FF03" blockId="1.[151,1436,1801,2008]" lastBlockId="2.[151,1437,151,1077]" lastPageId="2" lastPageNumber="191" pageId="1" pageNumber="190">
<emphasis id="83FB200EFFE5AD0DFF6F6443FEBDF8F5" bold="true" box="[151,266,1801,1827]" pageId="1" pageNumber="190">Diagnosis</emphasis>
. Unique among the members of the Californian clade of
<taxonomicName id="768F879FFFE5AD0DFC596443FBB6F8F5" box="[929,1025,1801,1827]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="83FB200EFFE5AD0DFC596443FBB6F8F5" box="[929,1025,1801,1827]" italics="true" pageId="1" pageNumber="190">Cybaeus</emphasis>
</taxonomicName>
, the species group affiliation of the males of the
<taxonomicName id="768F879FFFE5AD0DFED46464FE3FF891" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[300,392,1838,1863]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE5AD0DFED46464FE3FF891" box="[300,392,1838,1863]" italics="true" pageId="1" pageNumber="190">septatus</emphasis>
</taxonomicName>
group is relatively easily to determine. The males of the
<taxonomicName id="768F879FFFE5AD0DFBE06464FBC3F891" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[1048,1140,1838,1863]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE5AD0DFBE06464FBC3F891" box="[1048,1140,1838,1863]" italics="true" pageId="1" pageNumber="190">septatus</emphasis>
</taxonomicName>
group are diagnosed by a combination of the prolaterally to slightly anteriorly directed patellar apophysis which is as long as or longer than the width of the palpal patella and bears four to 12 peg setae one of which is in a proximal medial position isolated from, or slightly to noticeably larger than, the remainder of the peg setae (
<figureCitation id="29B4E099FFE5AD0DFC2964D3FB82F865" box="[977,1077,1945,1971]" captionStart="FIGURES 14" captionStartId="3.[151,264,1890,1914]" captionTargetBox="[151,1420,340,1866]" captionTargetId="figure-41@3.[151,1436,329,1866]" captionTargetPageId="3" captionText="FIGURES 14. Cybaeus chauliodous, males (1, 34 from Ashland Summit, Oregon; 2 from Lake Almanor, California), left palpus. 12 Patella and tibia (1 retrolateral, 2 dorsal). 34 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate single enlarged peg seta on patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145514" httpUri="https://zenodo.org/record/6145514/files/figure.png" pageId="1" pageNumber="190">Figs 12</figureCitation>
,
<figureCitation id="29B4E099FFE5AD0DFBB964D3FBDAF862" box="[1089,1133,1945,1972]" captionStart="FIGURES 59" captionStartId="4.[151,264,1266,1290]" captionTargetBox="[158,1430,181,1242]" captionTargetId="figure-30@4.[158,1430,181,1242]" captionTargetPageId="4" captionText="FIGURES 59. Cybaeus chauliodous, males (5, 7, 9 from Lake Almanor, California; 6, 8 from Ashland Summit, Oregon), left palpus. 57 Patellar apophysis, dorsal. 89 Proximal arm of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (57 single enlarged peg seta on patellar apophysis, 89 swollen lip-like tip of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.6145520" httpUri="https://zenodo.org/record/6145520/files/figure.png" pageId="1" pageNumber="190">57</figureCitation>
,
<figureCitation id="29B4E099FFE5AD0DFB8164D3FB2FF865" box="[1145,1176,1945,1971]" captionStart="FIGURES 2224" captionStartId="8.[151,264,1907,1931]" captionTargetBox="[151,1436,381,1883]" captionTargetId="figure-136@8.[151,1436,381,1883]" captionTargetPageId="8" captionText="FIGURES 2224. Cybaeus septatus, male from Mt. Shasta, California, left palpus. 22 Patella, dorsal. 2324 Genital bulb (23 ventral, 24 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis. Unlabelled arrows indicate diagnostic characters (22 single slightly enlarged peg seta on patellar apophysis, 23 weakly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145536" httpUri="https://zenodo.org/record/6145536/files/figure.png" pageId="1" pageNumber="190">22</figureCitation>
,
<figureCitation id="29B4E099FFE5AD0DFB5B64D3FB76F862" box="[1187,1217,1945,1972]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1618,1642]" captionTargetBox="[151,1436,813,1595]" captionTargetId="figure-264@9.[151,1436,813,1595]" captionTargetPageId="9" captionText="FIGURES 2526. Cybaeus septatus, male from Mt. Shasta, California, left palpus. 25 Patellar apophysis, dorsal. 26 Proximal arm of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (25 single slightly enlarged peg seta on patellar apophysis; 26 weakly concave posterior margin of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.6145542" httpUri="https://zenodo.org/record/6145542/files/figure.png" pageId="1" pageNumber="190">25</figureCitation>
,
<figureCitation id="29B4E099FFE5AD0DFB3564D3FB5CF865" box="[1229,1259,1945,1971]" captionStart="FIGURES 3336" captionStartId="11.[151,264,1799,1823]" captionTargetBox="[151,1436,181,1770]" captionTargetId="figure-19@11.[151,1436,181,1775]" captionTargetPageId="11" captionText="FIGURES 3336. Cybaeus somesbar, male from Slide Creek, California, left palpus. 33 Patella, tibia, and genital bulb, ventral. 34 Patella and tibia, dorsal. 3536 Genital bulb (35 ventral, 36 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate diagnostic characters (34 single isolated peg seta medially on PTA, 35 strongly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145552" httpUri="https://zenodo.org/record/6145552/files/figure.png" pageId="1" pageNumber="190">34</figureCitation>
,
<figureCitation id="29B4E099FFE5AD0DFB0E64D3FAA2F865" box="[1270,1301,1945,1971]" captionStart="FIGURES 4346" captionStartId="14.[151,264,1860,1884]" captionTargetBox="[151,1426,181,1816]" captionTargetId="figure-28@14.[151,1436,181,1836]" captionTargetPageId="14" captionText="FIGURES 4346. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 4344 Patella and tibia (43 retrolateral, 44 dorsal). 4546 Genital bulb (45 ventral, 46 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145558" httpUri="https://zenodo.org/record/6145558/files/figure.png" pageId="1" pageNumber="190">44</figureCitation>
,
<figureCitation id="29B4E099FFE5AD0DFAD864D3FAF7F865" box="[1312,1344,1945,1971]" captionStart="FIGURES 4748" captionStartId="15.[151,264,977,1001]" captionTargetBox="[177,1410,181,953]" captionTargetId="figure-19@15.[177,1410,181,953]" captionTargetPageId="15" captionText="FIGURES 4748. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 47 Patellar apophysis, dorsal. 48 Proximal arm of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (47 pointed angular tip of patellar apophysis with three peg setae, single isolated peg seta medially; 48 angular posterior margin of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.6145560" httpUri="https://zenodo.org/record/6145560/files/figure.png" pageId="1" pageNumber="190">47</figureCitation>
). To our knowledge, no males of other Californian clade
<taxonomicName id="768F879FFFE5AD0DFD4F64F7FCC7F801" box="[695,880,1981,2007]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="83FB200EFFE5AD0DFD4F64F7FCA0F801" box="[695,791,1981,2007]" italics="true" pageId="1" pageNumber="190">Cybaeus</emphasis>
species
</taxonomicName>
groups (
<taxonomicName id="768F879FFFE5AD0DFC2D64F7FB94F801" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[981,1059,1981,2007]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="adenes">
<emphasis id="83FB200EFFE5AD0DFC2D64F7FB94F801" box="[981,1059,1981,2007]" italics="true" pageId="1" pageNumber="190">adenes</emphasis>
</taxonomicName>
,
<taxonomicName id="768F879FFFE5AD0DFBC964F7FB75F801" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[1073,1218,1981,2007]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="aspenicolens">
<emphasis id="83FB200EFFE5AD0DFBC964F7FB75F801" box="[1073,1218,1981,2007]" italics="true" pageId="1" pageNumber="190">aspenicolens</emphasis>
</taxonomicName>
,
<taxonomicName id="768F879FFFE5AD0DFB2964F4FA88F801" authorityName="Gertsch &amp; Ivie" authorityYear="1936" box="[1233,1343,1982,2007]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="consocius">
<emphasis id="83FB200EFFE5AD0DFB2964F4FA88F801" box="[1233,1343,1982,2007]" italics="true" pageId="1" pageNumber="190">consocius</emphasis>
</taxonomicName>
,
<taxonomicName id="768F879FFFE5AD0DFAB664F7FA21F801" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[1358,1430,1981,2007]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="83FB200EFFE5AD0DFAB664F7FA21F801" box="[1358,1430,1981,2007]" italics="true" pageId="1" pageNumber="190">devius</emphasis>
</taxonomicName>
, and
<taxonomicName id="768F879FFFE6AD0EFF3E63DDFE95FF67" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[198,290,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="83FB200EFFE6AD0EFF3E63DDFE95FF67" box="[198,290,151,177]" italics="true" pageId="2" pageNumber="191">tardatus</emphasis>
</taxonomicName>
) or unplaced Californian species (
<taxonomicName id="768F879FFFE6AD0EFD5A63DDFB95FF67" authority="Chamberlin &amp; Ivie" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[674,1058,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="cribelloides">
<emphasis id="83FB200EFFE6AD0EFD5A63DDFCF0FF67" box="[674,839,151,177]" italics="true" pageId="2" pageNumber="191">C. cribelloides</emphasis>
Chamberlin &amp; Ivie
</taxonomicName>
,
<taxonomicName id="768F879FFFE6AD0EFBD563DDFAB7FF67" authority="Bennett" authorityName="Bennett" authorityYear="2009" box="[1069,1280,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="gidneyi">
<emphasis id="83FB200EFFE6AD0EFBD563DDFB28FF67" box="[1069,1183,151,177]" italics="true" pageId="2" pageNumber="191">C. gidneyi</emphasis>
Bennett
</taxonomicName>
,
<taxonomicName id="768F879FFFE6AD0EFAF363DDFF59FF03" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2022" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="irreverens" status="sp. nov.">
<emphasis id="83FB200EFFE6AD0EFAF363DDFA2BFF66" box="[1291,1436,151,177]" italics="true" pageId="2" pageNumber="191">C. irreverens</emphasis>
Bennett
</taxonomicName>
<emphasis id="83FB200EFFE6AD0EFF0D63F1FEDFFF03" bold="true" box="[245,360,187,213]" pageId="2" pageNumber="191">
<taxonomicNameLabel id="98C89D75FFE6AD0EFF0D63F1FEDFFF03" box="[245,360,187,213]" pageId="2" pageNumber="191" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
) show this combination of characters.
</paragraph>
<paragraph id="B130FC1CFFE6AD0EFF3F6395FCDBFE07" blockId="2.[151,1437,151,1077]" pageId="2" pageNumber="191">
The females of the
<taxonomicName id="768F879FFFE6AD0EFE506395FDB3FF2E" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[424,516,223,248]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE6AD0EFE506395FDB3FF2E" box="[424,516,223,248]" italics="true" pageId="2" pageNumber="191">septatus</emphasis>
</taxonomicName>
group are diagnosed by a combination of features of the epigynum, atrium, and copulatory ducts. The epigynum is marked anteriorly by one or more “wrinkles” in the integument (
<figureCitation id="29B4E099FFE6AD0EFB146249FADCFECB" box="[1260,1387,259,285]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 10, 13</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFA8F6249FA23FECB" box="[1399,1428,259,285]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">27</figureCitation>
) and the atrium features paired longitudinal atrial openings medially (
<figureCitation id="29B4E099FFE6AD0EFC5F626DFB9CFE97" box="[935,1067,295,321]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 10, 13</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFBC1626DFBE0FE97" box="[1081,1111,295,321]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1934,1958]" captionTargetBox="[151,1435,452,1910]" captionTargetId="figure-95@6.[151,1435,452,1910]" captionTargetPageId="6" captionText="FIGURES 1621. Cybaeus lockeae spec. nov., females from Castro Valley, California, copulatory organ. 16 Epigynum, ventral. 1721 Vulva (1719 ventral, 2021 dorsal). AT—Atrial opening, BG—Bennetts gland, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145530" httpUri="https://zenodo.org/record/6145530/files/figure.png" pageId="2" pageNumber="191">16</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB9C626DFB35FE97" box="[1124,1154,295,321]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">27</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB68626DFB6BFE97" box="[1168,1244,295,321]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">37, 40</figureCitation>
). The copulatory ducts (
<figureCitation id="29B4E099FFE6AD0EFF196201FE03FEB3" box="[225,436,330,357]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 1112, 1415</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFE476201FDB0FEB3" box="[447,519,331,357]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1934,1958]" captionTargetBox="[151,1435,452,1910]" captionTargetId="figure-95@6.[151,1435,452,1910]" captionTargetPageId="6" captionText="FIGURES 1621. Cybaeus lockeae spec. nov., females from Castro Valley, California, copulatory organ. 16 Epigynum, ventral. 1721 Vulva (1719 ventral, 2021 dorsal). AT—Atrial opening, BG—Bennetts gland, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145530" httpUri="https://zenodo.org/record/6145530/files/figure.png" pageId="2" pageNumber="191">1819</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFDEB6201FDEBFEB3" box="[531,604,331,357]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">2829</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD9E6201FD1AFEB3" box="[614,685,331,357]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">38, 41</figureCitation>
) are short and lead from the atrial openings medially to the midline of the vulva where they are contiguous prior to diverging and leading laterally to the spermathecal heads. As with the
<taxonomicName id="768F879FFFE6AD0EFF3A62D9FEA9FE7A" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[194,286,403,428]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE6AD0EFF3A62D9FEA9FE7A" box="[194,286,403,428]" italics="true" pageId="2" pageNumber="191">septatus</emphasis>
</taxonomicName>
group males, no females of other Californian clade
<taxonomicName id="768F879FFFE6AD0EFC8E62D9FB86FE7B" box="[886,1073,403,429]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="83FB200EFFE6AD0EFC8E62D9FC61FE7B" box="[886,982,403,429]" italics="true" pageId="2" pageNumber="191">Cybaeus</emphasis>
species
</taxonomicName>
groups or unplaced Californian species are known which possess this combination of characters.
</paragraph>
</subSubSection>
<subSubSection id="F995AF97FFE6AD0EFF3F6290FD2DFBE3" pageId="2" pageNumber="191" type="description">
<paragraph id="B130FC1CFFE6AD0EFF3F6290FBD4FDE8" blockId="2.[151,1437,151,1077]" pageId="2" pageNumber="191">
<emphasis id="83FB200EFFE6AD0EFF3F6290FEEEFE22" bold="true" box="[199,345,474,500]" pageId="2" pageNumber="191">Description.</emphasis>
As in diagnosis. Medium-sized spiders: carapace lengths averaging
<quantity id="767751F9FFE6AD0EFBB16290FB6BFE23" box="[1097,1244,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.325" metricValueMax="2.5" metricValueMin="2.15" pageId="2" pageNumber="191" unit="mm" value="2.325" valueMax="2.5" valueMin="2.15">2.152.5 mm</quantity>
(females). Males subequal. Abdomen usually patterned. Legs unbanded (femora rarely lightly banded). Two or occasionally three complete pairs of ventral tibia I macrosetae (distal pair present, absent, or incomplete).
</paragraph>
<paragraph id="B130FC1CFFE6AD0EFF3F610CFE65FD27" blockId="2.[151,1437,151,1077]" pageId="2" pageNumber="191">
<emphasis id="83FB200EFFE6AD0EFF3F610CFEB5FDB7" bold="true" box="[199,258,582,609]" italics="true" pageId="2" pageNumber="191">Male</emphasis>
: Retrolateral tibial apophysis (
<figureCitation id="29B4E099FFE6AD0EFD9A610DFD1DFDB7" box="[610,682,583,609]" captionStart="FIGURES 14" captionStartId="3.[151,264,1890,1914]" captionTargetBox="[151,1420,340,1866]" captionTargetId="figure-41@3.[151,1436,329,1866]" captionTargetPageId="3" captionText="FIGURES 14. Cybaeus chauliodous, males (1, 34 from Ashland Summit, Oregon; 2 from Lake Almanor, California), left palpus. 12 Patella and tibia (1 retrolateral, 2 dorsal). 34 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate single enlarged peg seta on patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145514" httpUri="https://zenodo.org/record/6145514/files/figure.png" pageId="2" pageNumber="191">Figs 1</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD42610DFD6FFDB7" box="[698,728,583,609]" captionStart="FIGURES 3336" captionStartId="11.[151,264,1799,1823]" captionTargetBox="[151,1436,181,1770]" captionTargetId="figure-19@11.[151,1436,181,1775]" captionTargetPageId="11" captionText="FIGURES 3336. Cybaeus somesbar, male from Slide Creek, California, left palpus. 33 Patella, tibia, and genital bulb, ventral. 34 Patella and tibia, dorsal. 3536 Genital bulb (35 ventral, 36 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate diagnostic characters (34 single isolated peg seta medially on PTA, 35 strongly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145552" httpUri="https://zenodo.org/record/6145552/files/figure.png" pageId="2" pageNumber="191">33</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD1E610DFCB4FDB7" box="[742,771,583,609]" captionStart="FIGURES 4346" captionStartId="14.[151,264,1860,1884]" captionTargetBox="[151,1426,181,1816]" captionTargetId="figure-28@14.[151,1436,181,1836]" captionTargetPageId="14" captionText="FIGURES 4346. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 4344 Patella and tibia (43 retrolateral, 44 dorsal). 4546 Genital bulb (45 ventral, 46 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145558" httpUri="https://zenodo.org/record/6145558/files/figure.png" pageId="2" pageNumber="191">43</figureCitation>
) carinate, nearly as long as tibia. Embolus relatively short and thin, describing a compound (
<figureCitation id="29B4E099FFE6AD0EFDE06121FDD6FD53" box="[536,609,619,645]" captionStart="FIGURES 14" captionStartId="3.[151,264,1890,1914]" captionTargetBox="[151,1420,340,1866]" captionTargetId="figure-41@3.[151,1436,329,1866]" captionTargetPageId="3" captionText="FIGURES 14. Cybaeus chauliodous, males (1, 34 from Ashland Summit, Oregon; 2 from Lake Almanor, California), left palpus. 12 Patella and tibia (1 retrolateral, 2 dorsal). 34 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate single enlarged peg seta on patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145514" httpUri="https://zenodo.org/record/6145514/files/figure.png" pageId="2" pageNumber="191">Figs 3</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD966121FD3BFD53" box="[622,652,619,645]" captionStart="FIGURES 2224" captionStartId="8.[151,264,1907,1931]" captionTargetBox="[151,1436,381,1883]" captionTargetId="figure-136@8.[151,1436,381,1883]" captionTargetPageId="8" captionText="FIGURES 2224. Cybaeus septatus, male from Mt. Shasta, California, left palpus. 22 Patella, dorsal. 2324 Genital bulb (23 ventral, 24 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis. Unlabelled arrows indicate diagnostic characters (22 single slightly enlarged peg seta on patellar apophysis, 23 weakly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145536" httpUri="https://zenodo.org/record/6145536/files/figure.png" pageId="2" pageNumber="191">23</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD616120FD01FD53" box="[665,694,618,645]" captionStart="FIGURES 4346" captionStartId="14.[151,264,1860,1884]" captionTargetBox="[151,1426,181,1816]" captionTargetId="figure-28@14.[151,1436,181,1836]" captionTargetPageId="14" captionText="FIGURES 4346. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 4344 Patella and tibia (43 retrolateral, 44 dorsal). 4546 Genital bulb (45 ventral, 46 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145558" httpUri="https://zenodo.org/record/6145558/files/figure.png" pageId="2" pageNumber="191">45</figureCitation>
) or simple (
<figureCitation id="29B4E099FFE6AD0EFCB86121FC25FD53" box="[832,914,618,645]" captionStart="FIGURES 3336" captionStartId="11.[151,264,1799,1823]" captionTargetBox="[151,1436,181,1770]" captionTargetId="figure-19@11.[151,1436,181,1775]" captionTargetPageId="11" captionText="FIGURES 3336. Cybaeus somesbar, male from Slide Creek, California, left palpus. 33 Patella, tibia, and genital bulb, ventral. 34 Patella and tibia, dorsal. 3536 Genital bulb (35 ventral, 36 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate diagnostic characters (34 single isolated peg seta medially on PTA, 35 strongly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145552" httpUri="https://zenodo.org/record/6145552/files/figure.png" pageId="2" pageNumber="191">Fig. 35</figureCitation>
) curve. Tegular apophysis (
<figureCitation id="29B4E099FFE6AD0EFB336121FAA3FD53" box="[1227,1300,619,645]" captionStart="FIGURES 14" captionStartId="3.[151,264,1890,1914]" captionTargetBox="[151,1420,340,1866]" captionTargetId="figure-41@3.[151,1436,329,1866]" captionTargetPageId="3" captionText="FIGURES 14. Cybaeus chauliodous, males (1, 34 from Ashland Summit, Oregon; 2 from Lake Almanor, California), left palpus. 12 Patella and tibia (1 retrolateral, 2 dorsal). 34 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate single enlarged peg seta on patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145514" httpUri="https://zenodo.org/record/6145514/files/figure.png" pageId="2" pageNumber="191">Figs 3</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFAD96121FA88FD53" box="[1313,1343,619,645]" captionStart="FIGURES 2224" captionStartId="8.[151,264,1907,1931]" captionTargetBox="[151,1436,381,1883]" captionTargetId="figure-136@8.[151,1436,381,1883]" captionTargetPageId="8" captionText="FIGURES 2224. Cybaeus septatus, male from Mt. Shasta, California, left palpus. 22 Patella, dorsal. 2324 Genital bulb (23 ventral, 24 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis. Unlabelled arrows indicate diagnostic characters (22 single slightly enlarged peg seta on patellar apophysis, 23 weakly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145536" httpUri="https://zenodo.org/record/6145536/files/figure.png" pageId="2" pageNumber="191">23</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFAB46120FADDFD53" box="[1356,1386,618,645]" captionStart="FIGURES 3336" captionStartId="11.[151,264,1799,1823]" captionTargetBox="[151,1436,181,1770]" captionTargetId="figure-19@11.[151,1436,181,1775]" captionTargetPageId="11" captionText="FIGURES 3336. Cybaeus somesbar, male from Slide Creek, California, left palpus. 33 Patella, tibia, and genital bulb, ventral. 34 Patella and tibia, dorsal. 3536 Genital bulb (35 ventral, 36 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate diagnostic characters (34 single isolated peg seta medially on PTA, 35 strongly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145552" httpUri="https://zenodo.org/record/6145552/files/figure.png" pageId="2" pageNumber="191">35</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFA8F6120FA23FD53" box="[1399,1428,618,645]" captionStart="FIGURES 4346" captionStartId="14.[151,264,1860,1884]" captionTargetBox="[151,1426,181,1816]" captionTargetId="figure-28@14.[151,1436,181,1836]" captionTargetPageId="14" captionText="FIGURES 4346. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 4344 Patella and tibia (43 retrolateral, 44 dorsal). 4546 Genital bulb (45 ventral, 46 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145558" httpUri="https://zenodo.org/record/6145558/files/figure.png" pageId="2" pageNumber="191">45</figureCitation>
) with distal arm short, unmodified, with rounded or angular terminus; proximal arm prolaterally directed, slightly swollen basally, and tip slightly twisted and swollen (
<figureCitation id="29B4E099FFE6AD0EFD1F61F9FCFBFD1B" box="[743,844,691,717]" captionStart="FIGURES 14" captionStartId="3.[151,264,1890,1914]" captionTargetBox="[151,1420,340,1866]" captionTargetId="figure-41@3.[151,1436,329,1866]" captionTargetPageId="3" captionText="FIGURES 14. Cybaeus chauliodous, males (1, 34 from Ashland Summit, Oregon; 2 from Lake Almanor, California), left palpus. 12 Patella and tibia (1 retrolateral, 2 dorsal). 34 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate single enlarged peg seta on patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145514" httpUri="https://zenodo.org/record/6145514/files/figure.png" pageId="2" pageNumber="191">Figs 34</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFCAF61F9FC33FD1B" box="[855,900,691,717]" captionStart="FIGURES 59" captionStartId="4.[151,264,1266,1290]" captionTargetBox="[158,1430,181,1242]" captionTargetId="figure-30@4.[158,1430,181,1242]" captionTargetPageId="4" captionText="FIGURES 59. Cybaeus chauliodous, males (5, 7, 9 from Lake Almanor, California; 6, 8 from Ashland Summit, Oregon), left palpus. 57 Patellar apophysis, dorsal. 89 Proximal arm of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (57 single enlarged peg seta on patellar apophysis, 89 swollen lip-like tip of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.6145520" httpUri="https://zenodo.org/record/6145520/files/figure.png" pageId="2" pageNumber="191">89</figureCitation>
), bluntly acuminate (
<figureCitation id="29B4E099FFE6AD0EFB9661F9FB58FD1B" box="[1134,1263,691,717]" captionStart="FIGURES 2224" captionStartId="8.[151,264,1907,1931]" captionTargetBox="[151,1436,381,1883]" captionTargetId="figure-136@8.[151,1436,381,1883]" captionTargetPageId="8" captionText="FIGURES 2224. Cybaeus septatus, male from Mt. Shasta, California, left palpus. 22 Patella, dorsal. 2324 Genital bulb (23 ventral, 24 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis. Unlabelled arrows indicate diagnostic characters (22 single slightly enlarged peg seta on patellar apophysis, 23 weakly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145536" httpUri="https://zenodo.org/record/6145536/files/figure.png" pageId="2" pageNumber="191">Figs 2324</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB0261F9FAAEFD18" box="[1274,1305,691,718]" captionStart="FIGURES 2526" captionStartId="9.[151,264,1618,1642]" captionTargetBox="[151,1436,813,1595]" captionTargetId="figure-264@9.[151,1436,813,1595]" captionTargetPageId="9" captionText="FIGURES 2526. Cybaeus septatus, male from Mt. Shasta, California, left palpus. 25 Patellar apophysis, dorsal. 26 Proximal arm of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (25 single slightly enlarged peg seta on patellar apophysis; 26 weakly concave posterior margin of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.6145542" httpUri="https://zenodo.org/record/6145542/files/figure.png" pageId="2" pageNumber="191">26</figureCitation>
), or sharply acuminate (
<figureCitation id="29B4E099FFE6AD0EFEE1619DFEC7FD27" box="[281,368,726,753]" captionStart="FIGURES 3336" captionStartId="11.[151,264,1799,1823]" captionTargetBox="[151,1436,181,1770]" captionTargetId="figure-19@11.[151,1436,181,1775]" captionTargetPageId="11" captionText="FIGURES 3336. Cybaeus somesbar, male from Slide Creek, California, left palpus. 33 Patella, tibia, and genital bulb, ventral. 34 Patella and tibia, dorsal. 3536 Genital bulb (35 ventral, 36 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate diagnostic characters (34 single isolated peg seta medially on PTA, 35 strongly concave posterior margin of PA)." figureDoi="http://doi.org/10.5281/zenodo.6145552" httpUri="https://zenodo.org/record/6145552/files/figure.png" pageId="2" pageNumber="191">Figs 35</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFE84619CFE2DFD27" box="[380,410,726,753]" captionStart="FIGURES 4346" captionStartId="14.[151,264,1860,1884]" captionTargetBox="[151,1426,181,1816]" captionTargetId="figure-28@14.[151,1436,181,1836]" captionTargetPageId="14" captionText="FIGURES 4346. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 4344 Patella and tibia (43 retrolateral, 44 dorsal). 4546 Genital bulb (45 ventral, 46 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.6145558" httpUri="https://zenodo.org/record/6145558/files/figure.png" pageId="2" pageNumber="191">45</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFE5E619DFE71FD27" box="[422,454,727,753]" captionStart="FIGURES 4748" captionStartId="15.[151,264,977,1001]" captionTargetBox="[177,1410,181,953]" captionTargetId="figure-19@15.[177,1410,181,953]" captionTargetPageId="15" captionText="FIGURES 4748. Cybaeus viator spec. nov., male from Fouts Springs, California, left palpus. 47 Patellar apophysis, dorsal. 48 Proximal arm of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (47 pointed angular tip of patellar apophysis with three peg setae, single isolated peg seta medially; 48 angular posterior margin of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.6145560" httpUri="https://zenodo.org/record/6145560/files/figure.png" pageId="2" pageNumber="191">48</figureCitation>
).
</paragraph>
<paragraph id="B130FC1CFFE6AD0EFF3F61B0FCFDFC57" blockId="2.[151,1437,151,1077]" pageId="2" pageNumber="191">
<emphasis id="83FB200EFFE6AD0EFF3F61B0FEA9FCC3" bold="true" box="[199,286,762,789]" italics="true" pageId="2" pageNumber="191">Female</emphasis>
: Atrial openings (
<figureCitation id="29B4E099FFE6AD0EFE1761B1FDC3FCC3" box="[495,628,763,789]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 10, 13</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD7B61B1FD16FCC3" box="[643,673,763,789]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1934,1958]" captionTargetBox="[151,1435,452,1910]" captionTargetId="figure-95@6.[151,1435,452,1910]" captionTargetPageId="6" captionText="FIGURES 1621. Cybaeus lockeae spec. nov., females from Castro Valley, California, copulatory organ. 16 Epigynum, ventral. 1721 Vulva (1719 ventral, 2021 dorsal). AT—Atrial opening, BG—Bennetts gland, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145530" httpUri="https://zenodo.org/record/6145530/files/figure.png" pageId="2" pageNumber="191">16</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD5761B1FD7AFCC3" box="[687,717,763,789]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">27</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFD2361B1FC92FCC3" box="[731,805,763,789]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">37, 40</figureCitation>
) conspicuous or not. Spermathecal heads (
<figureCitation id="29B4E099FFE6AD0EFAEA61B1FA20FCC3" box="[1298,1431,762,789]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 12, 15</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFF6F6055FF57FCEF" box="[151,224,799,825]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1934,1958]" captionTargetBox="[151,1435,452,1910]" captionTargetId="figure-95@6.[151,1435,452,1910]" captionTargetPageId="6" captionText="FIGURES 1621. Cybaeus lockeae spec. nov., females from Castro Valley, California, copulatory organ. 16 Epigynum, ventral. 1721 Vulva (1719 ventral, 2021 dorsal). AT—Atrial opening, BG—Bennetts gland, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145530" httpUri="https://zenodo.org/record/6145530/files/figure.png" pageId="2" pageNumber="191">2021</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFF146055FE82FCEF" box="[236,309,799,825]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">2930</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFEB86055FE6BFCEF" box="[320,476,799,825]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">3839, 4142</figureCitation>
) small dorsolateral lobes; spermathecal stalks short (
<figureCitation id="29B4E099FFE6AD0EFBD76055FB07FCEF" box="[1071,1200,799,825]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 12, 14</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB456055FB6BFCEF" box="[1213,1244,799,825]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1934,1958]" captionTargetBox="[151,1435,452,1910]" captionTargetId="figure-95@6.[151,1435,452,1910]" captionTargetPageId="6" captionText="FIGURES 1621. Cybaeus lockeae spec. nov., females from Castro Valley, California, copulatory organ. 16 Epigynum, ventral. 1721 Vulva (1719 ventral, 2021 dorsal). AT—Atrial opening, BG—Bennetts gland, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145530" httpUri="https://zenodo.org/record/6145530/files/figure.png" pageId="2" pageNumber="191">20</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB1F6055FAB3FCEF" box="[1255,1284,799,825]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">38</figureCitation>
) or relatively long (
<figureCitation id="29B4E099FFE6AD0EFF206009FEEEFC8B" box="[216,345,835,861]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">Figs 2930</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFE9C6009FE33FC8B" box="[356,388,835,861]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">42</figureCitation>
); Bennetts glands near junction with spermathecal bases (
<figureCitation id="29B4E099FFE6AD0EFBE96009FB25FC8B" box="[1041,1170,835,861]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1945,1969]" captionTargetBox="[151,1436,309,1922]" captionTargetId="figure-96@5.[151,1436,309,1922]" captionTargetPageId="5" captionText="FIGURES 1015. Cybaeus chauliodous, females (1012 from Lake Almanor, California; 1315 from Applegate River, Oregon), copulatory organ. 10, 13 Epigynum, ventral. 1112, 1415 Vulva (11, 14 ventral, 12, 15 dorsal). AT—Atrial opening, BG—Bennetts gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145526" httpUri="https://zenodo.org/record/6145526/files/figure.png" pageId="2" pageNumber="191">Figs 12, 14</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB666009FB51FC8B" box="[1182,1254,835,861]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1934,1958]" captionTargetBox="[151,1435,452,1910]" captionTargetId="figure-95@6.[151,1435,452,1910]" captionTargetPageId="6" captionText="FIGURES 1621. Cybaeus lockeae spec. nov., females from Castro Valley, California, copulatory organ. 16 Epigynum, ventral. 1721 Vulva (1719 ventral, 2021 dorsal). AT—Atrial opening, BG—Bennetts gland, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145530" httpUri="https://zenodo.org/record/6145530/files/figure.png" pageId="2" pageNumber="191">2021</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFB096009FAB8FC8B" box="[1265,1295,835,861]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">39</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFAE36009FA8EFC8B" box="[1307,1337,835,861]" captionStart="FIGURES 2732" captionStartId="10.[151,264,1832,1856]" captionTargetBox="[151,1436,418,1809]" captionTargetId="figure-129@10.[151,1436,417,1809]" captionTargetPageId="10" captionText="FIGURES 2732. Cybaeus septatus, females from California (27 from Mt. Shasta, 2829 holotype, 3032 from McCloud), copulatory organ. 27 Epigynum, ventral. 2829, 32 Vulva (28, 32 ventral, 29 dorsal). 30 Subadult vulva, dorsal. 31 Portion of teneral vulva (Bennetts gland and base of spermatheca), dorsal. AT—Atrial opening, BG—Bennetts gland, BS—base of spermatheca, HS—head of spermatheca, W—epigynal “wrinkle”." figureDoi="http://doi.org/10.5281/zenodo.6145546" httpUri="https://zenodo.org/record/6145546/files/figure.png" pageId="2" pageNumber="191">32</figureCitation>
,
<figureCitation id="29B4E099FFE6AD0EFABD6009FA38FC8B" box="[1349,1423,835,861]" captionStart="FIGURES 3742" captionStartId="13.[151,264,1905,1929]" captionTargetBox="[151,1436,454,1878]" captionTargetId="figure-133@13.[151,1436,453,1882]" captionTargetPageId="13" captionText="FIGURES 3742. Cybaeus spp., females, copulatory organ (3739 C. viator spec. nov. from Fouts Springs, California; 4042 holotype of C. somesbar). 37, 40 Epigynum, ventral. 3839, 4142 Vulva (38, 41 ventral, 39, 42 dorsal). AT—Atrial openings, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6145556" httpUri="https://zenodo.org/record/6145556/files/figure.png" pageId="2" pageNumber="191">39, 42</figureCitation>
); bases large, rounded; fertilization ducts exit bases posteriorly.
</paragraph>
<paragraph id="B130FC1CFFE6AD0EFF3F60C0FD2DFBE3" blockId="2.[151,1437,151,1077]" pageId="2" pageNumber="191">
<emphasis id="83FB200EFFE6AD0EFF3F60C0FD86FC72" bold="true" box="[199,561,906,933]" pageId="2" pageNumber="191">Composition and distribution</emphasis>
.
<taxonomicName id="768F879FFFE6AD0EFDBB60C1FC2BFC73" authority="Bennett" authorityName="Bennett" authorityYear="2009" box="[579,924,907,933]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="chauliodous">
<emphasis id="83FB200EFFE6AD0EFDBB60C1FC80FC73" box="[579,823,907,933]" italics="true" pageId="2" pageNumber="191">Cybaeus chauliodous</emphasis>
Bennett
</taxonomicName>
,
<taxonomicName id="768F879FFFE6AD0EFC5360C1FB3FFC73" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2022" box="[939,1160,907,933]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="lockeae" status="sp. nov.">
<emphasis id="83FB200EFFE6AD0EFC5360C1FB91FC73" box="[939,1062,907,933]" italics="true" pageId="2" pageNumber="191">C. lockeae</emphasis>
Bennett
</taxonomicName>
<emphasis id="83FB200EFFE6AD0EFB6B60C1FABDFC73" bold="true" box="[1171,1290,907,933]" pageId="2" pageNumber="191">
<taxonomicNameLabel id="98C89D75FFE6AD0EFB6B60C1FABDFC73" box="[1171,1290,907,933]" pageId="2" pageNumber="191" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="768F879FFFE6AD0EFAE460C1FEC7FC1F" authority="Chamberlin &amp; Ivie" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE6AD0EFAE460C1FA2BFC72" box="[1308,1436,907,933]" italics="true" pageId="2" pageNumber="191">C. septatus</emphasis>
Chamberlin &amp; Ivie
</taxonomicName>
,
<taxonomicName id="768F879FFFE6AD0EFE8560E5FDDAFC1F" authority="Bennett" authorityName="Bennett" authorityYear="2009" box="[381,621,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="somesbar">
<emphasis id="83FB200EFFE6AD0EFE8560E5FDBDFC1F" box="[381,522,943,969]" italics="true" pageId="2" pageNumber="191">C. somesbar</emphasis>
Bennett
</taxonomicName>
, and
<taxonomicName id="768F879FFFE6AD0EFD5260E5FCD8FC1F" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2022" box="[682,879,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="viator" status="sp. nov.">
<emphasis id="83FB200EFFE6AD0EFD5260E5FCB8FC1E" box="[682,783,943,969]" italics="true" pageId="2" pageNumber="191">C. viator</emphasis>
Bennett
</taxonomicName>
<emphasis id="83FB200EFFE6AD0EFC8F60E5FC5CFC1F" bold="true" box="[887,1003,943,969]" pageId="2" pageNumber="191">
<taxonomicNameLabel id="98C89D75FFE6AD0EFC8F60E5FC5CFC1F" box="[887,1003,943,969]" pageId="2" pageNumber="191" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="768F879FFFE6AD0EFC0B60E5FB0FFC1E" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[1011,1208,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="septatus">
<emphasis id="83FB200EFFE6AD0EFC0B60E5FB0FFC1E" box="[1011,1208,943,969]" italics="true" pageId="2" pageNumber="191">Cybaeus septatus</emphasis>
</taxonomicName>
and
<taxonomicName id="768F879FFFE6AD0EFB0960E5FA2BFC1F" authorityName="Bennett" authorityYear="2009" box="[1265,1436,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="chauliodous">
<emphasis id="83FB200EFFE6AD0EFB0960E5FA2BFC1F" box="[1265,1436,943,969]" italics="true" pageId="2" pageNumber="191">C. chauliodous</emphasis>
</taxonomicName>
are relatively common within their known distributions; the other species are uncommon. All have restricted ranges within an area encompassing southwestern
<collectingRegion id="734B32FEFFE6AD0EFD8260BDFD79FBC7" box="[634,718,1015,1041]" country="United States of America" name="Oregon" pageId="2" pageNumber="191">Oregon</collectingRegion>
(Jackson and Josephine Counties) and northern
<collectingRegion id="734B32FEFFE6AD0EFB1060BDFAEDFBC7" box="[1256,1370,1015,1041]" country="United States of America" name="California" pageId="2" pageNumber="191">California</collectingRegion>
south to Alameda and Sierra Counties (
<figureCitation id="29B4E099FFE6AD0EFDF26751FD39FBE3" box="[522,654,1050,1077]" captionStart-0="FIGURE 49" captionStart-1="FIGURE 50" captionStartId-0="16.[151,250,1944,1968]" captionStartId-1="17.[151,250,1943,1967]" captionTargetBox-0="[160,1427,181,1920]" captionTargetBox-1="[160,1427,181,1919]" captionTargetId-0="figure-28@16.[160,1427,181,1920]" captionTargetId-1="figure-19@17.[160,1427,181,1919]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 49. Distributions of Cybaeus chauliodous, C. septatus, and C. somesbar. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California and Oregon county names capitalized." captionText-1="FIGURE 50. Distributions of Cybaeus lockeae spec. nov. and C. viator spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi-0="http://doi.org/10.5281/zenodo.6145566" figureDoi-1="http://doi.org/10.5281/zenodo.6145573" httpUri-0="https://zenodo.org/record/6145566/files/figure.png" httpUri-1="https://zenodo.org/record/6145573/files/figure.png" pageId="2" pageNumber="191">Figs 4950</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>