169 lines
18 KiB
XML
169 lines
18 KiB
XML
<document id="33D11A93C751671CBADBDCB53ED81A50" ID-ISSN="11755334" ID-ZooBank="C1FA90AF-1C31-45D6-9CB6-C7D3058E501C" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629820076988" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2012" docId="3601F35E9E1D937DA3F8FBDAFC17DDB4" docLanguage="en" docName="zt03531p095.pdf" docOrigin="Zootaxa 3531" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Epimetopus lanceolatus Perkins 2012, new species" docType="treatment" docVersion="2" lastPageNumber="69" masterDocId="CA388B269E599339A36FFFE4FFC2DA6D" masterDocTitle="3531" masterLastPageNumber="95" masterPageNumber="1" pageNumber="69" updateTime="1699116377101" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AC0127BBEC0D2047760EE7081A631BE9">3531</mods:title>
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<mods:namePart id="339CA60FC1C00D9E4317ED3955851458">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="44545FE322838A31405B61A6147AC8AB">text</mods:typeOfResource>
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<mods:title id="4FE23B5F9BCD04D77C5D85981A80F072">Zootaxa</mods:title>
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<mods:part id="00879312BFB3DE5491555BD4A560E6F6">
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<mods:date id="9E2A70C5CE95F7A03158ED884EFFDC86">2012</mods:date>
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<mods:number id="0BF56D35FD1CE41030987DA138AC9DDA">2012-10-30</mods:number>
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<mods:number id="44DA4C8AE1D5E9978099AFD4D6288DB3">3531</mods:number>
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<treatment id="3601F35E9E1D937DA3F8FBDAFC17DDB4" LSID="urn:lsid:plazi:treatment:3601F35E9E1D937DA3F8FBDAFC17DDB4" httpUri="http://treatment.plazi.org/id/3601F35E9E1D937DA3F8FBDAFC17DDB4" lastPageNumber="69" pageId="68" pageNumber="69">
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<subSubSection id="F6B211C39E1D937DA3F8FBDAFD19DE14" pageId="68" pageNumber="69" type="nomenclature">
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<paragraph id="BE1742489E1D937DA3F8FBDAFD94DE35" blockId="68.[151,731,1086,1145]" box="[151,598,1086,1112]" pageId="68" pageNumber="69">
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<heading id="E55FF5249E1D937DA3F8FBDAFD94DE35" bold="true" box="[151,598,1086,1112]" fontSize="11" level="1" pageId="68" pageNumber="69" reason="1">
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<emphasis id="8CDC9E5A9E1D937DA3F8FBDAFD94DE35" bold="true" box="[151,598,1086,1112]" pageId="68" pageNumber="69">
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<taxonomicName id="79A839CB9E1D937DA3F8FBDAFE75DE35" ID-CoL="6G5XL" authority="Perkins, 2012" authorityName="Perkins" authorityYear="2012" box="[151,439,1086,1112]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lanceolatus" status="sp. nov.">
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<emphasis id="8CDC9E5A9E1D937DA3F8FBDAFE75DE35" bold="true" box="[151,439,1086,1112]" italics="true" pageId="68" pageNumber="69">Epimetopus lanceolatus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="97EF23219E1D937DA2A9FBDAFD94DE35" box="[454,598,1086,1112]" pageId="68" pageNumber="69" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BE1742489E1D937DA3F8FB85FD19DE14" blockId="68.[151,731,1086,1145]" box="[151,731,1121,1145]" pageId="68" pageNumber="69">
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Figs.
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<figureCitation id="26935ECD9E1D937DA3A1FB85FEF4DE14" box="[206,310,1121,1145]" captionStart="FIGURE 107" captionStartId="69.[151,250,1728,1751]" captionTargetBox="[227,1355,213,1689]" captionTargetId="figure-102@69.[203,1384,195,1707]" captionTargetPageId="69" captionText="FIGURE 107. Epimetopus lanceolatus holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="68" pageNumber="69">Figs. 107</figureCitation>
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(habitus), 109 (aedeagus), 128 (map)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F6B211C39E1D937DA3F8FB4DFF3EDF64" pageId="68" pageNumber="69" type="materials_examined">
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<paragraph id="BE1742489E1D937DA3F8FB4DFF3EDF64" blockId="68.[151,1436,1193,2010]" pageId="68" pageNumber="69">
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<emphasis id="8CDC9E5A9E1D937DA3F8FB4DFE89DEAF" bold="true" box="[151,331,1193,1218]" pageId="68" pageNumber="69">Type Material.</emphasis>
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<materialsCitation id="0EC048159E1D937DA239FB4DFEA2DE88" collectingDate="1955-12-01" collectingDateMax="1955-12-30" collectingDateMin="1955-12-01" collectionCode="MSP" collectorName="M. Alvarenga" country="Brazil" latitude="-16.066668" location="Caceres" longLatPrecision="1283" longitude="-57.683334" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
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<emphasis id="8CDC9E5A9E1D937DA239FB4DFE01DEAC" bold="true" box="[342,451,1193,1217]" pageId="68" pageNumber="69">
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<typeStatus id="6113FCEA9E1D937DA239FB4DFE01DEAC" box="[342,451,1193,1217]" pageId="68" pageNumber="69" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingCountry id="C6BF02D89E1D937DA146FB4DFDB0DEAC" box="[553,626,1193,1217]" name="Brazil" pageId="68" pageNumber="69">Brazil</collectingCountry>
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:
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<collectingRegion id="7C6C8CAA9E1D937DA110FB4EFCD5DEAC" box="[639,791,1193,1217]" country="Brazil" name="Mato Grosso" pageId="68" pageNumber="69">Mato Grosso</collectingRegion>
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,
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<location id="BB7714939E1D937DA049FB4DFC40DEAC" LSID="urn:lsid:plazi:treatment:3601F35E9E1D937DA3F8FBDAFC17DDB4:BB7714939E1D937DA049FB4DFC40DEAC" box="[806,898,1193,1217]" country="Brazil" latitude="-16.066668" longLatPrecision="1283" longitude="-57.683334" name="Caceres" pageId="68" pageNumber="69" stateProvince="Mato Grosso">Caceres</location>
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, MT.,
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<geoCoordinate id="DB9C248F9E1D937DA0BEFB4DFBF3DEAC" box="[977,1073,1193,1217]" degrees="16" direction="south" minutes="4" orientation="latitude" pageId="68" pageNumber="69" precision="925" value="-16.066668">16° 4' S</geoCoordinate>
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,
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||
<geoCoordinate id="DB9C248F9E1D937DA72EFB4DFB78DEAC" box="[1089,1210,1193,1217]" degrees="57" direction="west" minutes="41" orientation="longitude" pageId="68" pageNumber="69" precision="925" value="-57.683334">57° 41' W</geoCoordinate>
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,
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<date id="CA1664889E1D937DA7A7FB4DFAAFDEAC" box="[1224,1389,1193,1217]" pageId="68" pageNumber="69" value="1955-12-01" valueMax="1955-12-30" valueMin="1955-12-01">
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<collectingDate id="DA529D609E1D937DA7A7FB4DFAAFDEAC" box="[1224,1389,1193,1217]" pageId="68" pageNumber="69" value="1955-12-01" valueMax="1955-12-30" valueMin="1955-12-01">1–30 xii 1955</collectingDate>
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</date>
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,
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<collectorName id="135D279E9E1D937DA613FB4EFECEDE88" pageId="68" pageNumber="69">M. Alvarenga</collectorName>
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||
(
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||
<collectionCode id="D8B9DA8D9E1D937DA274FB29FE9BDE88" box="[283,345,1229,1253]" pageId="68" pageNumber="69">MSP</collectionCode>
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||
)
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</materialsCitation>
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.
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<materialsCitation id="0EC048159E1D937DA203FB29FF3ADF64" collectingDate="1955-12-02" collectingDateMax="1955-12-04" collectingDateMin="1955-12-02" collectionCode="CAS" collectorName="M. Alvarenga" country="Brazil" latitude="-16.066668" location="Caceres" longLatPrecision="1283" longitude="-57.683334" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
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||
<emphasis id="8CDC9E5A9E1D937DA203FB29FE20DE88" bold="true" box="[364,482,1229,1253]" pageId="68" pageNumber="69">
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||
<typeStatus id="6113FCEA9E1D937DA203FB29FE1FDE88" box="[364,477,1229,1253]" pageId="68" pageNumber="69" type="paratype">Paratype</typeStatus>
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||
:
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||
</emphasis>
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||
<collectingCountry id="C6BF02D89E1D937DA286FB29FDF0DE88" box="[489,562,1229,1253]" name="Brazil" pageId="68" pageNumber="69">Brazil</collectingCountry>
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:
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<collectingRegion id="7C6C8CAA9E1D937DA152FB2AFD13DE88" box="[573,721,1229,1253]" country="Brazil" name="Mato Grosso" pageId="68" pageNumber="69">Mato Grosso</collectingRegion>
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,
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||
<location id="BB7714939E1D937DA1B2FB29FCFADE88" LSID="urn:lsid:plazi:treatment:3601F35E9E1D937DA3F8FBDAFC17DDB4:BB7714939E1D937DA1B2FB29FCFADE88" box="[733,824,1229,1253]" country="Brazil" latitude="-16.066668" longLatPrecision="1283" longitude="-57.683334" name="Caceres" pageId="68" pageNumber="69" stateProvince="Mato Grosso">Caceres</location>
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, MT.,
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||
<geoCoordinate id="DB9C248F9E1D937DA0EEFB29FC19DE88" box="[897,987,1229,1253]" degrees="16" direction="south" minutes="4" orientation="latitude" pageId="68" pageNumber="69" precision="925" value="-16.066668">16° 4' S</geoCoordinate>
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||
,
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||
<geoCoordinate id="DB9C248F9E1D937DA088FB29FB9BDE88" box="[999,1113,1229,1253]" degrees="57" direction="west" minutes="41" orientation="longitude" pageId="68" pageNumber="69" precision="925" value="-57.683334">57° 41' W</geoCoordinate>
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,
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<date id="CA1664889E1D937DA70BFB29FB36DE88" box="[1124,1268,1229,1253]" pageId="68" pageNumber="69" value="1955-12-02" valueMax="1955-12-04" valueMin="1955-12-02">
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||
<collectingDate id="DA529D609E1D937DA70BFB29FB36DE88" box="[1124,1268,1229,1253]" pageId="68" pageNumber="69" value="1955-12-02" valueMax="1955-12-04" valueMin="1955-12-02">2–4 xii 1955</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="135D279E9E1D937DA66FFB2AFA59DE88" box="[1280,1435,1229,1253]" pageId="68" pageNumber="69">M. Alvarenga</collectorName>
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||
(1
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||
<collectionCode id="D8B9DA8D9E1D937DA3DAFB15FF32DF64" box="[181,240,1265,1289]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="68" pageNumber="69" type="Museum">CAS</collectionCode>
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)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F6B211C39E1D937DA3A8FAF1FEEBDC44" pageId="68" pageNumber="69" type="diagnosis">
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||
<paragraph id="BE1742489E1D937DA3A8FAF1FEEBDC44" blockId="68.[151,1436,1193,2010]" pageId="68" pageNumber="69">
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||
<emphasis id="8CDC9E5A9E1D937DA3A8FAF1FE24DF43" bold="true" box="[199,486,1301,1326]" pageId="68" pageNumber="69">Differential Diagnosis.</emphasis>
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Differentiated from
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<taxonomicName id="79A839CB9E1D937DA18BFAF2FC6ADF40" authorityName="Perkins" authorityYear="2012" box="[740,936,1301,1325]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lanceolatulus">
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<emphasis id="8CDC9E5A9E1D937DA18BFAF2FC6ADF40" box="[740,936,1301,1325]" italics="true" pageId="68" pageNumber="69">E. lanceolatulus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="26935ECD9E1D937DA0D4FAF1FBF2DF40" box="[955,1072,1301,1325]" captionStart="FIGURE 107" captionStartId="69.[151,250,1728,1751]" captionTargetBox="[227,1355,213,1689]" captionTargetId="figure-102@69.[203,1384,195,1707]" captionTargetPageId="69" captionText="FIGURE 107. Epimetopus lanceolatus holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="68" pageNumber="69">Figs. 107</figureCitation>
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,
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<figureCitation id="26935ECD9E1D937DA751FAF1FBA9DF40" box="[1086,1131,1301,1325]" captionStart="FIGURE 108" captionStartId="70.[151,250,1870,1893]" captionTargetBox="[268,1336,356,1835]" captionTargetId="figure-141@70.[237,1351,337,1849]" captionTargetPageId="70" captionText="FIGURE 108. Epimetopus lanceolatulus holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="68" pageNumber="69">108</figureCitation>
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) by the much larger size (ca. 3.63 vs.
|
||
<quantity id="7950EFAD9E1D937DA254FADDFE69DF3C" box="[315,427,1337,1361]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" pageId="68" pageNumber="69" unit="mm" value="2.61">2.61 mm</quantity>
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), the longer and more sharply arcuate pronotal hood, the different spearhead-like shape formed by the pronotal carinae, the presence of a pronotal ridge leading posteriorly from the posterior margin of the pronotal hood, and more robust elytra. In addition, the pronotal hood carinae are anteriorly confluent in
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||
<taxonomicName id="79A839CB9E1D937DA2DFFA42FD9FDFD0" authorityName="Perkins" authorityYear="2012" box="[432,605,1445,1469]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lanceolatus">
|
||
<emphasis id="8CDC9E5A9E1D937DA2DFFA42FD9FDFD0" box="[432,605,1445,1469]" italics="true" pageId="68" pageNumber="69">E. lanceolatus</emphasis>
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||
</taxonomicName>
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||
, and separated in
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||
<taxonomicName id="79A839CB9E1D937DA052FA42FBC3DFD0" authorityName="Perkins" authorityYear="2012" box="[829,1025,1445,1469]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lanceolatulus">
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||
<emphasis id="8CDC9E5A9E1D937DA052FA42FBC3DFD0" box="[829,1025,1445,1469]" italics="true" pageId="68" pageNumber="69">E. lanceolatulus</emphasis>
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||
</taxonomicName>
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||
. The aedeagus is notably broader than that of
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||
<taxonomicName id="79A839CB9E1D937DA243FA2EFE33DF8C" authorityName="Perkins" authorityYear="2012" box="[300,497,1481,1505]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lanceolatulus">
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||
<emphasis id="8CDC9E5A9E1D937DA243FA2EFE33DF8C" box="[300,497,1481,1505]" italics="true" pageId="68" pageNumber="69">E. lanceolatulus</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<figureCitation id="26935ECD9E1D937DA16BFA2DFD74DF8C" box="[516,694,1481,1505]" captionStart="FIGURES 109–110" captionStartId="71.[151,264,1990,2013]" captionTargetBox="[484,1105,472,1951]" captionTargetId="figure-136@71.[472,1115,457,1969]" captionTargetPageId="71" captionText="FIGURES 109–110. Epimetopus, aedeagi of holotypes. 109. E. lanceolatus. 110. E. lanceolatulus." pageId="68" pageNumber="69">Figs. 109, 110</figureCitation>
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), but despite the great difference in size of adults of the two species the aedeagi are of approximately the same length. Also, the parameres are shaped differently in the two species.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="F6B211C39E1D937DA3A8F9D1FEDFDDFC" pageId="68" pageNumber="69" type="description">
|
||
<paragraph id="BE1742489E1D937DA3A8F9D1FEDFDDFC" blockId="68.[151,1436,1193,2010]" pageId="68" pageNumber="69">
|
||
<emphasis id="8CDC9E5A9E1D937DA3A8F9D1FEA3DC20" box="[199,353,1589,1614]" inLineHeading="true" pageId="68" pageNumber="69" reason="1">
|
||
<emphasis id="8CDC9E5A9E1D937DA3A8F9D1FE98DC23" bold="true" box="[199,346,1589,1614]" pageId="68" pageNumber="69">Description</emphasis>
|
||
.
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||
</emphasis>
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||
Size:
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||
<typeStatus id="6113FCEA9E1D937DA2D9F9D1FDDCDC20" box="[438,542,1589,1613]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
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||
(length/width, mm): body (length from anterior margin of pronotum to elytral apices) 3.63/2.03; head (width) 0.94; pronotum 1.34/1.44; elytra 2.44/2.03. Habitus as illustrated (
|
||
<figureCitation id="26935ECD9E1D937DA3CFF999FECFDCF8" box="[160,269,1661,1685]" captionStart="FIGURE 107" captionStartId="69.[151,250,1728,1751]" captionTargetBox="[227,1355,213,1689]" captionTargetId="figure-102@69.[203,1384,195,1707]" captionTargetPageId="69" captionText="FIGURE 107. Epimetopus lanceolatus holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="68" pageNumber="69">Fig. 107</figureCitation>
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||
). Head dark brown to piceous, maxillary palpi light brown, dorsum dark brown except light brown narrow transverse fascia on elytra, venter and coxae dark brown to piceous, remainder of legs brown, abdominal ventrites brown, margin slightly lighter. Pronotum with primary lateral lobes large, angulate, secondary lobes small; hood anterior margin rather sharply arcuate; hood carinae anteriorly converging, confluent at anterior tip, together forming very sharp angle; pronotal ridges strong. Elytra with carinae strong, except third carina absent in short area subbasally; non-carinate intervals each with an irregular row of small granules. Metaventral depression moderately deep and wide, ca. eight large granules along base.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F6B211C39E1D937DA3A8F879FCB9DDD8" box="[199,891,1949,1973]" pageId="68" pageNumber="69" type="etymology">
|
||
<paragraph id="BE1742489E1D937DA3A8F879FCB9DDD8" blockId="68.[151,1436,1193,2010]" box="[199,891,1949,1973]" pageId="68" pageNumber="69">
|
||
<emphasis id="8CDC9E5A9E1D937DA3A8F879FE8CDDD8" bold="true" box="[199,334,1949,1973]" inLineHeading="true" pageId="68" pageNumber="69" reason="1">Etymology.</emphasis>
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Named in reference to the shape of the aedeagus.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F6B211C39E1D937DA3A8F825FC17DDB4" box="[199,981,1985,2010]" pageId="68" pageNumber="69" type="distribution">
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||
<paragraph id="BE1742489E1D937DA3A8F825FC17DDB4" blockId="68.[151,1436,1193,2010]" box="[199,981,1985,2010]" pageId="68" pageNumber="69">
|
||
<emphasis id="8CDC9E5A9E1D937DA3A8F825FEA2DDB7" bold="true" box="[199,352,1985,2010]" inLineHeading="true" pageId="68" pageNumber="69" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="6113FCEA9E1D937DA1A3F826FD3EDDB7" box="[716,764,1986,2010]" pageId="68" pageNumber="69">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="26935ECD9E1D937DA009F825FC0BDDB4" box="[870,969,1985,2009]" captionStart="FIGURE 128" captionStartId="88.[151,250,2001,2024]" captionTargetBox="[283,1304,469,1981]" captionTargetId="figure-67@88.[283,1304,469,1981]" captionTargetPageId="88" captionText="FIGURE 128. Geographical distributions of Epimetopus species." pageId="68" pageNumber="69">Fig. 128</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |