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<document id="2D600302DD84F6A713F3A129DAC356BE" ID-CLB-Dataset="4048" ID-DOI="10.11646/zootaxa.5048.4.2" ID-GBIF-Dataset="ad9f7049-4351-4deb-945a-409f9bbb5076" ID-ISSN="1175-5326" ID-Zenodo-Dep="5556598" ID-ZooBank="6A450390-1C69-42CD-9682-2A766CB31338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633693424633" checkinUser="plazi" docAuthor="Monastyrskii, Alexander L. &amp; Lien, Vu Van" docDate="2021" docId="36019464FFE9B612B7F2FE19FC1FFE2E" docLanguage="en" docName="zootaxa.5048.4.2.pdf" docOrigin="Zootaxa 5048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Faunis indistincta subsp. luctus Monastyrskii &amp; Vu 2021, subspec. nov." docType="treatment" docVersion="8" lastPageNumber="507" masterDocId="CA38EC1CFFFAB607B765FF8AFFE2FFDB" masterDocTitle="New taxa and new records of butterflies from Vietnam (Lepidoptera, Papilionoidea)" masterLastPageNumber="510" masterPageNumber="486" pageNumber="505" updateTime="1699459186830" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B631C7210BC57148884788CCCED20187" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="377E5234114DC4D6F2699F30E8365B67">
<mods:title id="7500BE9D69C1001CDAE485D2C011A66D">New taxa and new records of butterflies from Vietnam (Lepidoptera, Papilionoidea)</mods:title>
</mods:titleInfo>
<mods:name id="3802E507905F0A8263DCC4D8BA7E46F9" type="personal">
<mods:role id="B7255AADAE15D3370D29EB8BE602F65D">
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<mods:namePart id="832B91624FD7AD26E66B120B72784B6D">Monastyrskii, Alexander L.</mods:namePart>
<mods:nameIdentifier id="5E3182A05E5D470B982CEB7CB962C247" type="ORCID">0000-0001-7645-2458</mods:nameIdentifier>
<mods:affiliation id="7143D79372207E30FCF09B81D684731E">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam https: // orcid. org / 0000 - 0001 - 7645 - 2458 &amp; Fauna &amp; Flora International, Vietnam Programme, Ha Noi, Vietnam</mods:affiliation>
</mods:name>
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<mods:role id="7B9C4E2F8A57229EBF5733C77AB6C9A9">
<mods:roleTerm id="1CF841F92C8FB1EE864A85BDDC6E79B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C753255E69BA7CF3DCDD8F024AE3FC7A">Lien, Vu Van</mods:namePart>
<mods:nameIdentifier id="972E36A1F06AFA861E160C2CABFC5256" type="ORCID">0000-0001-7645-2458</mods:nameIdentifier>
<mods:affiliation id="D712527D0334D1178D879458FE6EA8E4">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam https: // orcid. org / 0000 - 0001 - 7645 - 2458 &amp; vulien @ gmail. com; https: // orcid. org / 0000 - 0003 - 2389 - 9402</mods:affiliation>
<mods:nameIdentifier id="A64717B1231337B5D4C9AD54C2D9FAB5" type="email">vulien@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="3982E806FFE0AF209F9E1FD567440E7B">Zootaxa</mods:title>
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<mods:part id="0CC951B837C10DF412CF8CA6150189C4">
<mods:date id="1046BAD4F4BDE8758FC03825CD262CE7">2021</mods:date>
<mods:detail id="B10757A546585D171BB27D3AF9FE4191" type="pubDate">
<mods:number id="13C41EEC35D93FCDEA7DFE67DDF3A165">2021-10-08</mods:number>
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<mods:detail id="0DA259599AA31E8EE7D5A30FE0CE12B1" type="volume">
<mods:number id="04489C890918A97A0081AF75C83350F0">5048</mods:number>
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<mods:number id="A9215D29FCDD65E5EC3DF28013FC1736">4</mods:number>
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<mods:start id="8967571C82B97343D5227CBC6FF89E2F">486</mods:start>
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<mods:classification id="A6E9BA6CD2DD12C1B33B379F0F758C66">journal article</mods:classification>
<mods:identifier id="7E9F61775E46E9DF83BCF3C437280523" type="CLB-Dataset">4048</mods:identifier>
<mods:identifier id="0C399C0CD14F26167435B6B3E2721EFE" type="DOI">10.11646/zootaxa.5048.4.2</mods:identifier>
<mods:identifier id="62080E4828F508A70BE7A3A19C15E42B" type="GBIF-Dataset">ad9f7049-4351-4deb-945a-409f9bbb5076</mods:identifier>
<mods:identifier id="3EE71C6AEDE96A9AA2BB0A0330AF8465" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E9E4CD53F36E469520DB42E155BFE258" type="Zenodo-Dep">5556598</mods:identifier>
<mods:identifier id="E6D1864A3812D6C8F1473A1260CC8CBB" type="ZooBank">6A450390-1C69-42CD-9682-2A766CB31338</mods:identifier>
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<treatment id="36019464FFE9B612B7F2FE19FC1FFE2E" ID-DOI="http://doi.org/10.5281/zenodo.5570243" ID-GBIF-Taxon="188639125" ID-Zenodo-Dep="5570243" LSID="urn:lsid:plazi:treatment:36019464FFE9B612B7F2FE19FC1FFE2E" httpUri="http://treatment.plazi.org/id/36019464FFE9B612B7F2FE19FC1FFE2E" lastPageId="21" lastPageNumber="507" pageId="19" pageNumber="505">
<subSubSection id="F6B276F9FFE9B614B7F2FE19FC85FE76" box="[151,871,402,429]" pageId="19" pageNumber="505" type="nomenclature">
<paragraph id="BE172572FFE9B614B7F2FE19FC85FE76" blockId="19.[151,871,402,465]" box="[151,871,402,429]" pageId="19" pageNumber="505">
<heading id="E55F921EFFE9B614B7F2FE19FC85FE76" bold="true" box="[151,871,402,429]" fontSize="11" level="1" pageId="19" pageNumber="505" reason="1">
<emphasis id="8CDCF960FFE9B614B7F2FE19FC85FE76" bold="true" box="[151,871,402,429]" pageId="19" pageNumber="505">
<taxonomicName id="79A85EF1FFE9B614B7F2FE19FD5EFE76" authority="Monastyrskii &amp; Vu" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[151,700,402,429]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="505" phylum="Arthropoda" rank="subSpecies" species="indistincta" status="subsp. nov." subSpecies="luctus">
<emphasis id="8CDCF960FFE9B614B7F2FE19FE26FE76" bold="true" box="[151,452,402,429]" italics="true" pageId="19" pageNumber="505">Faunis indistincta luctus</emphasis>
Monastyrskii &amp; Vu
</taxonomicName>
<taxonomicNameLabel id="97EF441BFFE9B614B5A6FE19FC85FE76" box="[707,871,403,429]" pageId="19" pageNumber="505" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFE9B613B7F2FE3DFE75FE52" lastPageId="20" lastPageNumber="506" pageId="19" pageNumber="505" type="description">
<paragraph id="BE172572FFE9B614B7F2FE3DFEA2FE0A" blockId="19.[151,871,402,465]" box="[151,320,439,465]" pageId="19" pageNumber="505">
(
<figureCitation id="269339F7FFE9B614B7FAFE3DFEEDFE0A" box="[159,271,439,465]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="19" pageNumber="505">Figure 13</figureCitation>
,
<figureCitation id="269339F7FFE9B614B67EFE3DFEDAFE0A" box="[283,312,439,465]" captionStart="FIGURE 14" captionStartId="20.[151,250,1657,1681]" captionTargetBox="[169,1424,421,1629]" captionTargetId="figure-116@20.[161,1426,417,1633]" captionTargetPageId="20" captionText="FIGURE 14. The male genitalia of Faunis aerope-group: A, C–E—Faunis indistincta luctus Monastyrskii &amp; Vu subspec. nov., ♂PT: A—general view of male genitalia in lateral aspect; C–E—ditto, apex of right claspers; B—general view of male genitalia of F. caelestis Monastyrskii &amp; Lang, 2016; F—F. aerope aerope (Leech, 1890)—right clasper (after Monastyrskii &amp; Lang, 2016); G—F. aerope yunnanensis Brooks, 1933 (after Monastyrskii &amp; Lang, 2016)." figureDoi="http://doi.org/10.5281/zenodo.5556640" httpUri="https://zenodo.org/record/5556640/files/figure.png" pageId="19" pageNumber="505">14</figureCitation>
)
</paragraph>
<paragraph id="BE172572FFE9B614B7F2FE74FA7AFDC2" blockId="19.[151,1437,510,825]" box="[151,1432,510,537]" pageId="19" pageNumber="505">
<emphasis id="8CDCF960FFE9B614B7F2FE74FEC9FDC3" bold="true" box="[151,299,510,536]" pageId="19" pageNumber="505">Description:</emphasis>
Male: length of forewing: HT: 45.0 mm; PTs: 46.0; 45.0; 46.0 mm; antennae dark brown (22.0 mm).
</paragraph>
<paragraph id="BE172572FFE9B614B7A2FDA8FEFDFD72" blockId="19.[151,1437,510,825]" pageId="19" pageNumber="505">
<emphasis id="8CDCF960FFE9B614B7A2FDA8FE54FDE7" bold="true" box="[199,438,546,572]" pageId="19" pageNumber="505">Forewing upperside</emphasis>
(
<figureCitation id="269339F7FFE9B614B6A3FDA9FDACFDE6" box="[454,590,547,573]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="19" pageNumber="505">Figure 13A</figureCitation>
): ground colour uniformly grey; area from termen and apex to postmedian and subcostal penetrating into discal cell uniformly blackish brown; there is a smooth transition from greyish black to bright grey in the postmedian area; series of small whitish subterminal spots is lined from cell R
<subScript id="222C2737FFE9B614B398FDF7FAE7FD50" attach="left" box="[1277,1285,637,651]" fontSize="6" pageId="19" pageNumber="505">4</subScript>
to Cu
<subScript id="222C2737FFE9B614B228FDF7FABEFD50" attach="left" box="[1357,1372,637,651]" fontSize="6" pageId="19" pageNumber="505">1a</subScript>
; cilia dark brown.
</paragraph>
<paragraph id="BE172572FFE9B614B7A2FD38FD9EFCE2" blockId="19.[151,1437,510,825]" pageId="19" pageNumber="505">
<emphasis id="8CDCF960FFE9B614B7A2FD38FE58FD17" bold="true" box="[199,442,690,716]" pageId="19" pageNumber="505">Hindwing upperside</emphasis>
(
<figureCitation id="269339F7FFE9B614B6ADFD39FDACFD16" box="[456,590,691,717]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="19" pageNumber="505">Figure 13A</figureCitation>
): ground colour uniformly grey with blackish brown bordering from termen to subterminal area and smooth transition to greyish colour; there is a series of small and subtle whitish subterminal spots from cell Rs to Cu
<subScript id="222C2737FFE9B614B6C0FC87FE56FCC0" attach="left" box="[421,436,781,795]" fontSize="6" pageId="19" pageNumber="505">1a</subScript>
; cilia dark brown. At base of cubitus there is hair tuft touching a small spot of specialised scent scales on underside of forewing base.
</paragraph>
<caption id="EAD775FAFFE9B614B7F2F8F0FEAEF86E" ID-DOI="http://doi.org/10.5281/zenodo.5556638" ID-Zenodo-Dep="5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="19" pageNumber="505" startId="19.[151,250,1914,1938]" targetBox="[153,1428,850,1885]" targetPageId="19">
<paragraph id="BE172572FFE9B614B7F2F8F0FEAEF86E" blockId="19.[151,1437,1914,1974]" pageId="19" pageNumber="505">
<emphasis id="8CDCF960FFE9B614B7F2F8F0FEBBF849" bold="true" box="[151,345,1914,1938]" pageId="19" pageNumber="505">FIGURE 13. A–B</emphasis>
—
<taxonomicName id="79A85EF1FFE9B614B616F8F1FCA7F84A" authority="Monastyrskii &amp; Vu" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[371,837,1914,1938]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="505" phylum="Arthropoda" rank="subSpecies" species="indistincta" status="subsp. nov." subSpecies="luctus">
<emphasis id="8CDCF960FFE9B614B616F8F1FD91F849" box="[371,627,1914,1938]" italics="true" pageId="19" pageNumber="505">Faunis indistincta luctus</emphasis>
Monastyrskii &amp; Vu
</taxonomicName>
,
<emphasis id="8CDCF960FFE9B614B437F8F0FC03F849" bold="true" box="[850,993,1914,1938]" pageId="19" pageNumber="505">
<taxonomicNameLabel id="97EF441BFFE9B614B437F8F0FC03F849" box="[850,993,1914,1938]" pageId="19" pageNumber="505" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
, ♂HT;
<emphasis id="8CDCF960FFE9B614B351F8F0FB87F849" bold="true" box="[1076,1125,1914,1938]" pageId="19" pageNumber="505">C–D</emphasis>
—
<taxonomicName id="79A85EF1FFE9B614B31AF8F1FA62F849" authority="Monastyrskii" authorityName="Monastyrskii" box="[1151,1408,1914,1938]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="505" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="8CDCF960FFE9B614B31AF8F1FB13F849" box="[1151,1265,1914,1938]" italics="true" pageId="19" pageNumber="505">F. caelestis</emphasis>
Monastyrskii
</taxonomicName>
&amp; Lang, 2016, ♂HT
</paragraph>
</caption>
<paragraph id="BE172572FFEEB613B7A2FF1CFE3EFF22" blockId="20.[151,1437,150,393]" pageId="20" pageNumber="506">
<emphasis id="8CDCF960FFEEB613B7A2FF1CFE54FF6B" bold="true" box="[199,438,150,176]" pageId="20" pageNumber="506">Forewing underside</emphasis>
(
<figureCitation id="269339F7FFEEB613B6A0FF1DFDA8FF6A" box="[453,586,151,177]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="20" pageNumber="506">Figure 13B</figureCitation>
): Ground colour dark brown with three darker brown fasciae located within discal cell (submedian) and at postmedian and subterminal areas; series of subterminal small whitish spots located in a row from cell R
<subScript id="222C2737FFEEB613B61CFF7BFE63FF24" attach="left" box="[377,385,241,255]" fontSize="6" pageId="20" pageNumber="506">4</subScript>
to Cu
<subScript id="222C2737FFEEB613B6A3FF7BFE37FF24" attach="left" box="[454,469,241,255]" fontSize="6" pageId="20" pageNumber="506">1a</subScript>
.
</paragraph>
<paragraph id="BE172572FFEEB613B7A2FE88FAB4FEB0" blockId="20.[151,1437,150,393]" pageId="20" pageNumber="506">
<emphasis id="8CDCF960FFEEB613B7A2FE88FE5FFEC7" bold="true" box="[199,445,258,284]" pageId="20" pageNumber="506">Hindwing underside</emphasis>
(
<figureCitation id="269339F7FFEEB613B6ABFE89FDB7FEC6" box="[462,597,259,285]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="20" pageNumber="506">Figure 13B</figureCitation>
): ground colour is similar to the forewing; the hindwing pattern includes submedian straight fascia extending from Sc+R
<subScript id="222C2737FFEEB613B5A5FEB3FD2AFE9C" attach="left" box="[704,712,313,327]" fontSize="6" pageId="20" pageNumber="506">1</subScript>
to Cu
<subScript id="222C2737FFEEB613B475FEB3FCC2FE9C" attach="left" box="[784,800,313,327]" fontSize="6" pageId="20" pageNumber="506">1b</subScript>
; postmedian fascia extending from Sc+R
<subScript id="222C2737FFEEB613B261FEB3FAEEFE9C" attach="left" box="[1284,1292,313,327]" fontSize="6" pageId="20" pageNumber="506">1</subScript>
to Cu
<subScript id="222C2737FFEEB613B23CFEB3FA8BFE9C" attach="left" box="[1369,1385,313,327]" fontSize="6" pageId="20" pageNumber="506">1b</subScript>
and subterminal fascia from Rs to Cu1b; series of small whitish subterminal spots extending from cell Rs to Cu
<subScript id="222C2737FFEEB613B222FED7FAB4FEB0" attach="left" box="[1351,1366,349,363]" fontSize="6" pageId="20" pageNumber="506">1a</subScript>
</paragraph>
<paragraph id="BE172572FFEEB613B7A2FEE5FE75FE52" blockId="20.[151,1437,150,393]" box="[199,407,367,393]" pageId="20" pageNumber="506">
<emphasis id="8CDCF960FFEEB613B7A2FEE5FECAFE52" bold="true" box="[199,296,367,393]" pageId="20" pageNumber="506">Female:</emphasis>
unknown
</paragraph>
</subSubSection>
<caption id="EAD775FAFFEEB613B7F2F9F3FBC9F926" ID-DOI="http://doi.org/10.5281/zenodo.5556640" ID-Zenodo-Dep="5556640" httpUri="https://zenodo.org/record/5556640/files/figure.png" pageId="20" pageNumber="506" startId="20.[151,250,1657,1681]" targetBox="[169,1424,421,1629]" targetPageId="20">
<paragraph id="BE172572FFEEB613B7F2F9F3FBC9F926" blockId="20.[151,1437,1657,1790]" pageId="20" pageNumber="506">
<emphasis id="8CDCF960FFEEB613B7F2F9F3FEC0F94A" bold="true" box="[151,290,1657,1681]" pageId="20" pageNumber="506">FIGURE 14.</emphasis>
The male genitalia of
<taxonomicName id="79A85EF1FFEEB613B571F9F0FD4BF94A" authorityName="Leech" authorityYear="1890" box="[532,681,1658,1681]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="aerope">
<emphasis id="8CDCF960FFEEB613B571F9F0FD4BF94A" box="[532,681,1658,1681]" italics="true" pageId="20" pageNumber="506">Faunis aerope</emphasis>
</taxonomicName>
-group:
<emphasis id="8CDCF960FFEEB613B598F9F3FC85F94A" bold="true" box="[765,871,1657,1681]" pageId="20" pageNumber="506">A, C–E—</emphasis>
<taxonomicName id="79A85EF1FFEEB613B402F9F0FADAF94A" authority="Monastyrskii &amp; Vu" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[871,1336,1657,1682]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="subSpecies" species="indistincta" status="subsp. nov." subSpecies="luctus">
<emphasis id="8CDCF960FFEEB613B402F9F0FB8AF94A" box="[871,1128,1657,1681]" italics="true" pageId="20" pageNumber="506">Faunis indistincta luctus</emphasis>
Monastyrskii &amp; Vu
</taxonomicName>
<emphasis id="8CDCF960FFEEB613B224F9F0FF21F96D" bold="true" pageId="20" pageNumber="506">
<taxonomicNameLabel id="97EF441BFFEEB613B224F9F0FF21F96D" pageId="20" pageNumber="506" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
, ♂PT:
<emphasis id="8CDCF960FFEEB613B66AF917FEC3F96E" bold="true" box="[271,289,1693,1717]" pageId="20" pageNumber="506">A</emphasis>
—general view of male genitalia in lateral aspect;
<emphasis id="8CDCF960FFEEB613B44CF917FCBBF96E" bold="true" box="[809,857,1693,1717]" pageId="20" pageNumber="506">C–E</emphasis>
—ditto, apex of right claspers;
<emphasis id="8CDCF960FFEEB613B3FFF914FB49F96D" bold="true" box="[1178,1195,1694,1718]" pageId="20" pageNumber="506">B</emphasis>
—general view of male genitalia of
<taxonomicName id="79A85EF1FFEEB613B677F948FDF2F902" authority="Monastyrskii" authorityName="Monastyrskii" box="[274,528,1729,1753]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="8CDCF960FFEEB613B677F948FE60F902" box="[274,386,1729,1753]" italics="true" pageId="20" pageNumber="506">F. caelestis</emphasis>
Monastyrskii
</taxonomicName>
&amp; Lang, 2016;
<emphasis id="8CDCF960FFEEB613B5D1F948FD21F901" bold="true" box="[692,707,1730,1754]" pageId="20" pageNumber="506">F</emphasis>
—
<taxonomicName id="79A85EF1FFEEB613B5B8F948FBFEF902" authority="aerope (Leech, 1890)" authorityName="aerope (Leech" authorityYear="1890" baseAuthorityName="Leech" baseAuthorityYear="1890" box="[733,1052,1729,1753]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="subSpecies" species="aerope" subSpecies="aerope">
<emphasis id="8CDCF960FFEEB613B5B8F948FC6AF902" box="[733,904,1730,1753]" italics="true" pageId="20" pageNumber="506">F. aerope aerope</emphasis>
(Leech, 1890)
</taxonomicName>
—right clasper (after Monastyrskii &amp; Lang, 2016);
<emphasis id="8CDCF960FFEEB613B645F96FFED6F926" bold="true" box="[288,308,1765,1789]" pageId="20" pageNumber="506">G</emphasis>
—
<taxonomicName id="79A85EF1FFEEB613B628F96CFD5CF925" authority="Brooks, 1933" authorityName="Brooks" authorityYear="1933" box="[333,702,1765,1790]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="subSpecies" species="aerope" subSpecies="yunnanensis">
<emphasis id="8CDCF960FFEEB613B628F96CFDCDF926" box="[333,559,1766,1789]" italics="true" pageId="20" pageNumber="506">F. aerope yunnanensis</emphasis>
Brooks, 1933
</taxonomicName>
(after Monastyrskii &amp; Lang, 2016).
</paragraph>
</caption>
<subSubSection id="F6B276F9FFEEB612B7A2F8A1FD37FEC6" lastPageId="21" lastPageNumber="507" pageId="20" pageNumber="506" type="diagnosis">
<paragraph id="BE172572FFEEB612B7A2F8A1FD37FEC6" blockId="20.[151,1436,1835,2042]" lastBlockId="21.[151,1437,151,501]" lastPageId="21" lastPageNumber="507" pageId="20" pageNumber="506">
<emphasis id="8CDCF960FFEEB613B7A2F8A1FEA0F89E" bold="true" box="[199,322,1835,1861]" pageId="20" pageNumber="506">Diagnosis.</emphasis>
The new taxon collected in Lao Cai Province resembles members of the
<taxonomicName id="79A85EF1FFEEB613B3F0F8A7FADBF89D" authorityName="Leech" authorityYear="1890" box="[1173,1337,1837,1862]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="aerope">
<emphasis id="8CDCF960FFEEB613B3F0F8A7FADBF89D" box="[1173,1337,1837,1862]" italics="true" pageId="20" pageNumber="506">Faunis aerope</emphasis>
</taxonomicName>
speciesgroup distributed in southern
<collectingCountry id="C6BF65E2FFEEB613B68BF8DAFDD3F8B1" box="[494,561,1872,1898]" name="China" pageId="20" pageNumber="506">China</collectingCountry>
and northern
<collectingCountry id="C6BF65E2FFEEB613B5B0F8DAFCDAF8B1" box="[725,824,1872,1898]" name="Vietnam" pageId="20" pageNumber="506">Vietnam</collectingCountry>
. The most distinctive character of
<taxonomicName id="79A85EF1FFEEB613B3ACF8DBFA7EF8B1" authority="Mell" authorityName="Mell" box="[1225,1436,1872,1898]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="indistincta">
<emphasis id="8CDCF960FFEEB613B3ACF8DBFABFF8B1" box="[1225,1373,1872,1898]" italics="true" pageId="20" pageNumber="506">F. indistincta</emphasis>
Mell
</taxonomicName>
is the male genitalia structure which vaguely resembles that of
<taxonomicName id="79A85EF1FFEEB613B402F8FFFC33F855" authorityName="Leech" authorityYear="1890" box="[871,977,1909,1934]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="aerope">
<emphasis id="8CDCF960FFEEB613B402F8FFFC33F855" box="[871,977,1909,1934]" italics="true" pageId="20" pageNumber="506">F. aerope</emphasis>
</taxonomicName>
: relatively short and curved valvae with apex covered by dense spines (
<figureCitation id="269339F7FFEEB613B567F812FD2DF869" box="[514,719,1944,1970]" captionStart="FIGURE 14" captionStartId="20.[151,250,1657,1681]" captionTargetBox="[169,1424,421,1629]" captionTargetId="figure-116@20.[161,1426,417,1633]" captionTargetPageId="20" captionText="FIGURE 14. The male genitalia of Faunis aerope-group: A, C–E—Faunis indistincta luctus Monastyrskii &amp; Vu subspec. nov., ♂PT: A—general view of male genitalia in lateral aspect; C–E—ditto, apex of right claspers; B—general view of male genitalia of F. caelestis Monastyrskii &amp; Lang, 2016; F—F. aerope aerope (Leech, 1890)—right clasper (after Monastyrskii &amp; Lang, 2016); G—F. aerope yunnanensis Brooks, 1933 (after Monastyrskii &amp; Lang, 2016)." figureDoi="http://doi.org/10.5281/zenodo.5556640" httpUri="https://zenodo.org/record/5556640/files/figure.png" pageId="20" pageNumber="506">Figure 14A, C–E</figureCitation>
).
<taxonomicName id="79A85EF1FFEEB613B580F813FBE2F869" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[741,1024,1944,1970]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="subSpecies" species="indistincta" subSpecies="luctus">
<emphasis id="8CDCF960FFEEB613B580F813FBE2F869" box="[741,1024,1944,1970]" italics="true" pageId="20" pageNumber="506">Faunis indistincta luctus</emphasis>
</taxonomicName>
is similar to recently the described species
<taxonomicName id="79A85EF1FFEEB613B795F837FDA1F80D" authority="Monastyrskii" authorityName="Monastyrskii" box="[240,579,1980,2006]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="8CDCF960FFEEB613B795F837FE44F80D" box="[240,422,1980,2006]" italics="true" pageId="20" pageNumber="506">Faunis caelestis</emphasis>
Monastyrskii
</taxonomicName>
&amp; Lang, 2016 (
<figureCitation id="269339F7FFEEB613B59CF836FC43F80D" box="[761,929,1980,2006]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="20" pageNumber="506">Figure 13C–D</figureCitation>
). Both species have similar wing patterns on the undersides of both wings, though in
<taxonomicName id="79A85EF1FFEEB613B530F86BFD2DF821" box="[597,719,2016,2042]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="8CDCF960FFEEB613B530F86BFD2DF821" box="[597,719,2016,2042]" italics="true" pageId="20" pageNumber="506">F. caelestis</emphasis>
</taxonomicName>
the ground colour is paler. The upperside of
<taxonomicName id="79A85EF1FFEEB613B3A7F86BFA7EF821" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[1218,1436,2016,2042]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="506" phylum="Arthropoda" rank="subSpecies" species="indistincta" subSpecies="luctus">
<emphasis id="8CDCF960FFEEB613B3A7F86BFA7EF821" box="[1218,1436,2016,2042]" italics="true" pageId="20" pageNumber="506">F. indistincta luctus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="79A85EF1FFEFB612B641FF1DFD95FF6A" authority="Brooks" authorityName="Brooks" authorityYear="1933" box="[292,631,151,177]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="507" phylum="Arthropoda" rank="subSpecies" species="aerope" subSpecies="yunnanensis">
<emphasis id="8CDCF960FFEFB612B641FF1DFDFCFF6B" box="[292,542,151,176]" italics="true" pageId="21" pageNumber="507">F. aerope yunnanensis</emphasis>
Brooks
</taxonomicName>
and
<taxonomicName id="79A85EF1FFEFB612B5CBFF1DFCC8FF6A" box="[686,810,151,177]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="507" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="8CDCF960FFEFB612B5CBFF1DFCC8FF6A" box="[686,810,151,177]" italics="true" pageId="21" pageNumber="507">F. caelestis</emphasis>
</taxonomicName>
, though the new taxon has broad black borders on both wings (
<figureCitation id="269339F7FFEFB612B78CFF31FE8EFF0E" box="[233,364,187,213]" captionStart="FIGURE 13" captionStartId="19.[151,250,1914,1938]" captionTargetBox="[153,1428,850,1885]" captionTargetId="figure-244@19.[153,1434,849,1891]" captionTargetPageId="19" captionText="FIGURE 13. A–B—Faunis indistincta luctus Monastyrskii &amp; Vu, subspec. nov., ♂HT; C–D—F. caelestis Monastyrskii &amp; Lang, 2016, ♂HT" figureDoi="http://doi.org/10.5281/zenodo.5556638" httpUri="https://zenodo.org/record/5556638/files/figure.png" pageId="21" pageNumber="507">Figure 13A</figureCitation>
) and distinctive male genitalia (
<figureCitation id="269339F7FFEFB612B5B1FF31FC79FF0E" box="[724,923,187,213]" captionStart="FIGURE 14" captionStartId="20.[151,250,1657,1681]" captionTargetBox="[169,1424,421,1629]" captionTargetId="figure-116@20.[161,1426,417,1633]" captionTargetPageId="20" captionText="FIGURE 14. The male genitalia of Faunis aerope-group: A, C–E—Faunis indistincta luctus Monastyrskii &amp; Vu subspec. nov., ♂PT: A—general view of male genitalia in lateral aspect; C–E—ditto, apex of right claspers; B—general view of male genitalia of F. caelestis Monastyrskii &amp; Lang, 2016; F—F. aerope aerope (Leech, 1890)—right clasper (after Monastyrskii &amp; Lang, 2016); G—F. aerope yunnanensis Brooks, 1933 (after Monastyrskii &amp; Lang, 2016)." figureDoi="http://doi.org/10.5281/zenodo.5556640" httpUri="https://zenodo.org/record/5556640/files/figure.png" pageId="21" pageNumber="507">Figure 14A, C–E</figureCitation>
). The female genitalia structures in species of
<taxonomicName id="79A85EF1FFEFB612B7F2FF55FEE2FF23" authorityName="Leech" authorityYear="1890" box="[151,256,223,248]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="507" phylum="Arthropoda" rank="species" species="aerope">
<emphasis id="8CDCF960FFEFB612B7F2FF55FEE2FF23" box="[151,256,223,248]" italics="true" pageId="21" pageNumber="507">F. aerope</emphasis>
</taxonomicName>
group are also distinctive; however, this character cannot be used in the current study because the female of
<taxonomicName id="79A85EF1FFEFB612B7D3FE89FE73FEC6" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[182,401,259,285]" class="Insecta" family="Nymphalidae" genus="Faunis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="507" phylum="Arthropoda" rank="subSpecies" species="indistincta" subSpecies="luctus">
<emphasis id="8CDCF960FFEFB612B7D3FE89FE73FEC6" box="[182,401,259,285]" italics="true" pageId="21" pageNumber="507">F. indistincta luctus</emphasis>
</taxonomicName>
has not yet been discovered.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFEFB612B7A2FEADFDEBFE75" pageId="21" pageNumber="507" type="materials_examined">
<paragraph id="BE172572FFEFB612B7A2FEADFDEBFE75" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="507">
<emphasis id="8CDCF960FFEFB612B7A2FEADFE95FE9B" bold="true" box="[199,375,294,321]" pageId="21" pageNumber="507">Type material.</emphasis>
<materialsCitation id="0EC02F2FFFEFB612B61AFEADFB78FEBE" ID-GBIF-Occurrence="3383539308" collectingDate="2011-05-20" collectionCode="VVL" collectorName="Hoang Lien N. P. &amp; Cong Troi" country="Vietnam" elevation="2" location="Lao Cai province" pageId="21" pageNumber="507" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<typeStatus id="61139BD0FFEFB612B61AFEADFE0EFE9A" box="[383,492,295,321]" pageId="21" pageNumber="507" type="holotype">Holotype</typeStatus>
: NORTHERN
<collectingCountry id="C6BF65E2FFEFB612B5F2FEADFCF9FE9A" box="[663,795,295,321]" name="Vietnam" pageId="21" pageNumber="507">VIETNAM</collectingCountry>
,
<specimenCount id="A8AEEEFBFFEFB612B44DFEADFCDDFE9A" box="[808,831,295,321]" pageId="21" pageNumber="507" type="male">♂</specimenCount>
,
<collectingRegion id="7C6CEB90FFEFB612B429FEADFBF7FE9A" box="[844,1045,295,321]" country="Vietnam" name="Lao Cai" pageId="21" pageNumber="507">Lao Cai province</collectingRegion>
,
<collectorName id="135D40A4FFEFB612B347FEADFB00FE9A" box="[1058,1250,295,321]" pageId="21" pageNumber="507">Hoang Lien N.P.</collectorName>
,
<collectorName id="135D40A4FFEFB612B38AFEADFA81FE9A" box="[1263,1379,295,321]" pageId="21" pageNumber="507">Cong Troi</collectorName>
pass (Tram Ton ranger station),
<date id="CA1603B2FFEFB612B6B1FEC1FDABFEBE" box="[468,585,331,357]" pageId="21" pageNumber="507" value="2011-05-20">
<collectingDate id="DA52FA5AFFEFB612B6B1FEC1FDABFEBE" box="[468,585,331,357]" pageId="21" pageNumber="507" value="2011-05-20">20.V.2011</collectingDate>
</date>
; evergreen mountain forest at ~
<quantity id="79508897FFEFB612B4D9FEC1FBC2FEBE" box="[956,1056,331,357]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="507" unit="m" value="1.9">
<elevation id="3585C241FFEFB612B4D9FEC1FBC2FEBE" box="[956,1056,331,357]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="507" unit="m" value="1.9">1,900 m</elevation>
</quantity>
;
<collectionCode id="D8B9BDB7FFEFB612B34AFEC1FB8AFEBE" box="[1071,1128,331,357]" pageId="21" pageNumber="507">VVL</collectionCode>
leg
</materialsCitation>
.;
<materialsCitation id="0EC02F2FFFEFB612B3CBFEC1FDC5FE52" ID-GBIF-Occurrence="3383539317" collectingDate="2011-05-20" collectionCode="VVL" collectorName="Hoang Lien N. P. &amp; Cong Troi" country="Vietnam" elevation="2" location="The" pageId="21" pageNumber="507" specimenCount="3" specimenCount-male="3" stateProvince="Lao Cai" typeStatus="paratype">
<typeStatus id="61139BD0FFEFB612B3CBFEC1FAFDFEBE" box="[1198,1311,331,357]" pageId="21" pageNumber="507" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A8AEEEFBFFEFB612B248FEC1FAB6FEBE" box="[1325,1364,331,357]" pageId="21" pageNumber="507" type="male">3♂</specimenCount>
: same data as holotype.
<location id="BB7773A9FFEFB612B605FEE5FE6EFE52" LSID="urn:lsid:plazi:treatment:36019464FFE9B612B7F2FE19FC1FFE2E:BB7773A9FFEFB612B605FEE5FE6EFE52" box="[352,396,367,393]" name="The" pageId="21" pageNumber="507">The</location>
<typeStatus id="61139BD0FFEFB612B6F0FEE5FE15FE52" box="[405,503,367,393]" pageId="21" pageNumber="507" type="holotype">holotype</typeStatus>
and
</materialsCitation>
<materialsCitation id="0EC02F2FFFEFB612B554FEE5FB44FE51" ID-GBIF-Occurrence="3383539320" box="[561,1190,367,394]" collectionCode="VNMN" country="Vietnam" location="Hanoi" pageId="21" pageNumber="507" specimenCount="1" stateProvince="Ha Noi" typeStatus="paratype">
<specimenCount id="A8AEEEFBFFEFB612B554FEE5FD2DFE52" box="[561,719,367,393]" pageId="21" pageNumber="507" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
will be deposited in the
<collectionCode id="D8B9BDB7FFEFB612B488FEE5FBA1FE52" box="[1005,1091,367,393]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="21" pageNumber="507">VNMN</collectionCode>
(
<collectingRegion id="7C6CEB90FFEFB612B336FEE5FB7FFE51" box="[1107,1181,367,394]" country="Vietnam" name="Ha Noi" pageId="21" pageNumber="507">Hanoi</collectingRegion>
)
</materialsCitation>
;
<materialsCitation id="0EC02F2FFFEFB612B3D6FEE5FDE7FE75" ID-GBIF-Occurrence="3383539324" country="Vietnam" location="London" pageId="21" pageNumber="507" specimenCount="1" stateProvince="Ha Noi" typeStatus="paratype">
<specimenCount id="A8AEEEFBFFEFB612B3D6FEE5FAA6FE52" box="[1203,1348,367,393]" pageId="21" pageNumber="507" type="generic" typeStatus="paratype">one paratype</specimenCount>
will be deposited in the NHM (
<location id="BB7773A9FFEFB612B6C5FE19FE1CFE75" LSID="urn:lsid:plazi:treatment:36019464FFE9B612B7F2FE19FC1FFE2E:BB7773A9FFEFB612B6C5FE19FE1CFE75" box="[416,510,403,430]" country="Vietnam" name="London" pageId="21" pageNumber="507" stateProvince="Ha Noi">London</location>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFEFB612B7A2FE3CFC1FFE2E" pageId="21" pageNumber="507" type="distribution">
<paragraph id="BE172572FFEFB612B7A2FE3CFC1FFE2E" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="507">
<emphasis id="8CDCF960FFEFB612B7A2FE3CFE1EFE0B" bold="true" box="[199,508,438,465]" pageId="21" pageNumber="507">Distribution and habitats.</emphasis>
The new taxon has been recorded only at Hoang Lien National Park in the vicinity of Fan Si Pan peak and the forest ranger station at elevations around
<quantity id="79508897FFEFB612B4FFFE51FC1BFE2E" box="[922,1017,475,501]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="507" unit="m" value="2.0">2,000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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