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<mods:title id="7AEB19DA62E41F7919E7E9892AA2DE91">A monographic catalogue on the systematics and phylogeny of the South American iguanian lizard family Liolaemidae (Squamata, Iguania)</mods:title>
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Complex
<taxonomicName id="7BB630F1FF9F61376D62E918FE5F8B78" ID-CoL="3V6DB" authorityName="Koslowsky" authorityYear="1898" box="[270,400,519,545]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="8EC29760FF9F61376D62E918FE5F8B78" bold="true" box="[270,400,519,545]" italics="true" pageId="47" pageNumber="48">boulengeri</emphasis>
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An appreciable amount of research has been devoted to understand the diversity, biogeography and phylogenetic relationships of the
<taxonomicName id="7BB630F1FF9F61376DBDE965FD658BCD" box="[465,682,634,660]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="8EC29760FF9F61376DBDE965FD848BCA" box="[465,587,634,659]" italics="true" pageId="47" pageNumber="48">Liolaemus</emphasis>
species
</taxonomicName>
belonging to the complex
<taxonomicName id="7BB630F1FF9F61376F96E965FBB68BCA" authorityName="Koslowsky" authorityYear="1898" box="[1018,1145,634,659]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="8EC29760FF9F61376F96E965FBB68BCA" box="[1018,1145,634,659]" italics="true" pageId="47" pageNumber="48">boulengeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="D8273683FF9F613768E1E965FA868BCD" author="Etheridge" box="[1165,1353,634,660]" firstAuthor="Etheridge" pageId="47" pageNumber="48" pagination="137 - 199" refId="ref59190" refString="Etheridge, R. (1993) Lizards of the Liolaemus darwinii Complex (Squamata: Iguania: Tropiduridae) in Northern Argentina. Bollettino del Museo Regionale di Scienze Naturali di Torino, 11, 137 - 199." type="journal article" year="1993">Etheridge 1993</bibRefCitation>
,
<bibRefCitation id="D8273683FF9F61376936E965FA588BCD" author="Etheridge" box="[1370,1431,634,660]" firstAuthor="Etheridge" pageId="47" pageNumber="48" pagination="1 - 34" refId="ref59230" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">1995</bibRefCitation>
,
<bibRefCitation id="D8273683FF9F61376CFBE9BDFF1A8BE5" author="Etheridge, R." box="[151,213,674,700]" pageId="47" pageNumber="48" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">2000</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F61376C8DE9BDFE1A8BE5" author="Morando" box="[225,469,674,700]" etAl="et al." firstAuthor="Morando" pageId="47" pageNumber="48" pagination="842 - 861" refId="ref66517" refString="Morando, M., Avila, L. J., Baker, J. &amp; Sites, J. W. (2004) Phylogeny and phylogeography of the Liolaemus darwinii complex (Squamata: Liolaemidae): evidence for introgression and incomplete lineage sorting. Evolution, 58, 842 - 861." type="journal article" year="2004">
Morando
<emphasis id="8EC29760FF9F61376D38E9BDFE408BE2" box="[340,399,674,699]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F61376D8DE9BDFDB38BE5" author="Abdala" box="[481,636,674,700]" firstAuthor="Abdala" pageId="47" pageNumber="48" pagination="1 - 84" refId="ref50199" refString="Abdala, C. S. (2007) Phylogeny of the boulengeri group (Iguania: Liolaemidae, Liolaemus) based on morphological and molecular characters. Zootaxa, 1538, 1 - 84." type="journal article" year="2007">Abdala 2007</bibRefCitation>
; see also
<bibRefCitation id="D8273683FF9F61376E80E9BDFC0B8BE5" author="Halloy" box="[748,964,674,700]" etAl="et al." firstAuthor="Halloy" pageId="47" pageNumber="48" pagination="1 - 37" refId="ref61046" refString="Halloy, M., Etheridge, R. &amp; Burghardt, G. M. (1998) To bury in the sand: phylogenetic relationships among lizard species of the boulengeri group, Liolaemus (Reptilia: Squamata: Tropiduridae), based on behavioral characters. Herpetological Monographs, 12, 1 - 37." type="journal article" year="1998">
Halloy
<emphasis id="8EC29760FF9F61376F2FE9BDFCB18BE2" box="[835,894,674,699]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F61376FBCE9BDFB5C8BE5" author="Cruz" box="[976,1171,674,700]" etAl="et al." firstAuthor="Cruz" pageId="47" pageNumber="48" pagination="1559 - 1574" refId="ref56953" refString="Cruz, F. B., Fitzgerald, L. A., Espinoza, R. E. &amp; Schulte II, J. A. (2005). The importance of phylogenetic scale in tests of Bergmann's and Rapoport's rules: lessons from a clade of South American lizards. Journal of Evolutionary Biology, 18, 1559 - 1574." type="journal article" year="2005">
Cruz
<emphasis id="8EC29760FF9F6137687DE9BDFB838BE2" box="[1041,1100,674,699]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2005
</bibRefCitation>
). Indeed, this group of
<taxonomicName id="7BB630F1FF9F61376CFBE9D5FEDE8BBA" authorityName=", Pincheira-Donoso &amp; Nunez" authorityYear="2005" box="[151,273,714,739]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="8EC29760FF9F61376CFBE9D5FEDE8BBA" box="[151,273,714,739]" italics="true" pageId="47" pageNumber="48">Liolaemus</emphasis>
</taxonomicName>
is perhaps the best known phylogenetically within the entire family
<taxonomicName id="7BB630F1FF9F61376857E9D5FB1E8BBD" authorityName="Frost &amp; Etheridge" authorityYear="1989" box="[1083,1233,714,740]" class="Reptilia" family="Liolaemidae" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="family">Liolaemidae</taxonomicName>
. In general, most phylogenetic hypotheses support the idea that two major clades form this large complex of species, one recognized as
<taxonomicName id="7BB630F1FF9F61376C93E805FE478A6A" authorityName="Etheridge" authorityYear="2000" box="[255,392,794,819]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="wiegmannii">
<emphasis id="8EC29760FF9F61376C93E805FE478A6A" box="[255,392,794,819]" italics="true" pageId="47" pageNumber="48">wiegmannii</emphasis>
</taxonomicName>
clade or subgenus
<taxonomicName id="7BB630F1FF9F61376E00E805FCCA8A6A" authorityName="Girard" authorityYear="1858" box="[620,773,794,819]" class="Reptilia" family="Liolaemidae" genus="Ortholaemus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="8EC29760FF9F61376E00E805FCCA8A6A" box="[620,773,794,819]" italics="true" pageId="47" pageNumber="48">Ortholaemus</emphasis>
</taxonomicName>
(
<bibRefCitation id="D8273683FF9F61376F79E805FC498A6D" author="Cei" box="[789,902,794,820]" firstAuthor="Cei" pageId="47" pageNumber="48" refId="ref54875" refString="Cei, J. M. (1993) Reptiles del noroeste, nordeste y este de la Argentina. Herpetofauna de las selvas subtropicales, Puna y Pampas. Museo Regionale di Scienze Naturali di Torino, Monografie 14, 949 pp." type="book" year="1993">Cei 1993</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F61376FFEE805FB858A6D" author="Etheridge" box="[914,1098,794,820]" firstAuthor="Etheridge" pageId="47" pageNumber="48" pagination="1 - 34" refId="ref59230" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995</bibRefCitation>
,
<bibRefCitation id="D8273683FF9F6137683BE805FB5A8A6D" author="Etheridge, R." box="[1111,1173,794,820]" pageId="47" pageNumber="48" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">2000</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F613768CEE805FA588A6D" author="Espinoza" box="[1186,1431,794,820]" etAl="et al." firstAuthor="Espinoza" pageId="47" pageNumber="48" pagination="16819 - 16824" refId="ref59093" refString="Espinoza, R. E., Wiens, J. J. &amp; Tracy, C. R. (2004) Recurrent evolution of herbivory in small, cold-climate lizards: Breaking the ecophysiological rules or reptilian herbivory. Proceedings of the National Academy of Sciences, USA, 101, 16819 - 16824." type="journal article" year="2004">
Espinoza
<emphasis id="8EC29760FF9F61376979E805FA9E8A6A" box="[1301,1361,794,819]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F61376CFBE85DFE908A05" author="Cruz" box="[151,351,834,860]" etAl="et al." firstAuthor="Cruz" pageId="47" pageNumber="48" pagination="1559 - 1574" refId="ref56953" refString="Cruz, F. B., Fitzgerald, L. A., Espinoza, R. E. &amp; Schulte II, J. A. (2005). The importance of phylogenetic scale in tests of Bergmann's and Rapoport's rules: lessons from a clade of South American lizards. Journal of Evolutionary Biology, 18, 1559 - 1574." type="journal article" year="2005">
Cruz
<emphasis id="8EC29760FF9F61376CB6E85DFED88A02" box="[218,279,834,859]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2005
</bibRefCitation>
), and the other as
<taxonomicName id="7BB630F1FF9F61376E2CE85DFD7D8A02" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[576,690,834,859]" class="Reptilia" family="Liolaemidae" genus="Proctotretus" kingdom="Animalia" order="Squamata" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="fitzingerii">
<emphasis id="8EC29760FF9F61376E2CE85DFD7D8A02" box="[576,690,834,859]" italics="true" pageId="47" pageNumber="48">fitzingerii</emphasis>
</taxonomicName>
clade (
<bibRefCitation id="D8273683FF9F61376F67E85DFB6F8A05" author="Pincheira-Donoso &amp; Nunez" box="[779,1184,834,860]" firstAuthor="Pincheira-Donoso" pageId="47" pageNumber="48" refId="ref69923" refString="Pincheira-Donoso, D. &amp; Nunez, H. (2005) Las especies chilenas del genero Liolaemus (Iguania, Tropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion, Publicacion Ocasional del Museo Nacional de Historia Natural de Chile, Santiago, 486 pp." type="book" year="2005">Pincheira-Donoso &amp; Núñez 2005</bibRefCitation>
; see
<bibRefCitation id="D8273683FF9F613768B1E85DFA588A05" author="Etheridge" box="[1245,1431,834,860]" firstAuthor="Etheridge" pageId="47" pageNumber="48" pagination="1 - 34" refId="ref59230" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995</bibRefCitation>
;
<bibRefCitation id="D8273683FF9F61376CFBE875FEBA8ADD" author="Schulte" box="[151,373,874,900]" etAl="et al." firstAuthor="Schulte" pageId="47" pageNumber="48" pagination="75 - 102" refId="ref71803" refString="Schulte II, J. A., Macey, J. R., Espinoza, R. E. &amp; Larson, A. (2000) Phylogenetic relationships in the iguanid lizard genus Liolaemus: multiple origins of viviparous reproduction and evidence for recurring Andean vicariance and dispersal. Biological Journal of the Linnean Society, 69, 75 - 102." type="journal article" year="2000">
Schulte
<emphasis id="8EC29760FF9F61376C9AE875FEFE8ADA" box="[246,305,874,899]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2000
</bibRefCitation>
, for phylogenetic details).
</paragraph>
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