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<document id="31EAEB3D1EBAB2CA8C95E9AAC26A4B9F" ID-CLB-Dataset="282050" ID-DOI="10.11646/zootaxa.5383.1.2" ID-GBIF-Dataset="066cb5f5-49e9-4141-b1e8-c794ed35aac7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10301486" ID-ZooBank="75E27F9C-3ACC-4409-8105-A39648FF2025" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1702023926292" checkinUser="plazi" docAuthor="Khalaji-Pirbalouty, Valiallah &amp; Al-Kandari, Manal Abdulrahman" docDate="2023" docId="325D6174086BFFE435F5FD8A59F1F803" docLanguage="en" docName="zootaxa.5383.1.2.pdf" docOrigin="Zootaxa 5383 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.1.2/52437" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elaphognathia alshumaimah Khalaji-Pirbalouty &amp; Al-Kandari 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="CE64190C086DFFEC3562FFAD5900FFF5" masterDocTitle="Three new species of temporary fish parasite, Gnathiid Isopods (Crustacea: Isopoda: Cymothoida) from Kuwait" masterLastPageNumber="38" masterPageNumber="24" pageNumber="30" updateTime="1732044582731" updateUser="admin" zenodo-license-document="CLOSED">
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<mods:title id="2CED7D006F3431A004AD6F1972AEFD0F">Three new species of temporary fish parasite, Gnathiid Isopods (Crustacea: Isopoda: Cymothoida) from Kuwait</mods:title>
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<mods:name id="0EF96113666ED29883B567A0B4D9A430" type="personal">
<mods:role id="6CA295EFEFDC89B51868CF46CFB94E8E">
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<mods:namePart id="B5DB2814DE909CE361527EACA16F03B5">Khalaji-Pirbalouty, Valiallah</mods:namePart>
<mods:affiliation id="24C182CB3A3577359302C61AD2555138">Department of Biology, Faculty of Basic Science, Shahrekord University, Shahrekord, Iran</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="0B2520E53A1D52888BFFFDEDB9BB4126">Al-Kandari, Manal Abdulrahman</mods:namePart>
<mods:affiliation id="12A97D95A0D02058553175CFD00C9ABB">Ecosystem-Based Management of Marine Resources, Environment and Life Sciences Research Center, Kuwait Institute of Scientific Research, Hamad Al-Mubarak Street, Building 900004, Area 1, Raas Salmiya, Kuwait</mods:affiliation>
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<mods:title id="1123DBF7775CF9501843F74B810F6D7C">Zootaxa</mods:title>
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<mods:part id="5FB8F9E0BD2B4499C42093B3FEC26727">
<mods:date id="706726BF0D2DF16D59625B5957D48BC7">2023</mods:date>
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<mods:number id="3A9E0F932D5EE7B1C20DBBADAC45D252">2023-12-08</mods:number>
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<treatment id="325D6174086BFFE435F5FD8A59F1F803" ID-DOI="http://doi.org/10.5281/zenodo.10391010" ID-GBIF-Taxon="219974362" ID-Zenodo-Dep="10391010" LSID="urn:lsid:plazi:treatment:325D6174086BFFE435F5FD8A59F1F803" httpUri="http://treatment.plazi.org/id/325D6174086BFFE435F5FD8A59F1F803" lastPageId="8" lastPageNumber="32" pageId="6" pageNumber="30">
<subSubSection id="F2EE83E9086BFFEA35F5FD8A5B5FFDB7" box="[151,607,551,578]" pageId="6" pageNumber="30" type="nomenclature">
<paragraph id="BA4BD062086BFFEA35F5FD8A5B5FFDB7" blockId="6.[151,974,551,651]" box="[151,607,551,578]" pageId="6" pageNumber="30">
<heading id="E103670E086BFFEA35F5FD8A5B5FFDB7" bold="true" box="[151,607,551,578]" fontSize="11" level="1" pageId="6" pageNumber="30" reason="1">
<emphasis id="88800C70086BFFEA35F5FD8A5B5FFDB7" bold="true" box="[151,607,551,578]" pageId="6" pageNumber="30">
<taxonomicName id="7DF4ABE1086BFFEA35F5FD8A58F9FDB7" ID-CoL="BXPCL" authority="Khalaji-Pirbalouty &amp; Al-Kandari, 2023" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2023" box="[151,505,551,578]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="30" phylum="Arthropoda" rank="species" species="alshumaimah" status="sp. nov.">
<emphasis id="88800C70086BFFEA35F5FD8A58F9FDB7" bold="true" box="[151,505,551,578]" italics="true" pageId="6" pageNumber="30">Elaphognathia alshumaimah</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93B3B10B086BFFEA3763FD855B5FFDB7" box="[513,607,552,578]" pageId="6" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E9086BFFEA35F5FDE159F9FD7E" pageId="6" pageNumber="30" type="discussion">
<paragraph id="BA4BD062086BFFEA35F5FDE15ACEFD92" blockId="6.[151,974,551,651]" box="[151,974,588,615]" pageId="6" pageNumber="30">
<uri id="CE65DC60086BFFEA35F5FDE15ACEFD92" box="[151,974,588,615]" pageId="6" pageNumber="30">
urn:lsid:zoobank.org:act:
<uuid id="CE52EAB7086BFFEA34C4FDE15ACEFD92" box="[422,974,588,615]" pageId="6" pageNumber="30">AF06EC94-3A95-4562-9E4D-D4ACBB022176</uuid>
</uri>
</paragraph>
<paragraph id="BA4BD062086BFFEA35F5FDDD59F9FD7E" blockId="6.[151,974,551,651]" box="[151,249,624,651]" pageId="6" pageNumber="30">
<figureCitation id="22CFCCE7086BFFEA35F5FDDD59F9FD7E" box="[151,249,624,651]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1649,1673]" captionStartId-1="8.[151,250,1474,1498]" captionTargetBox-0="[159,1428,183,1617]" captionTargetBox-1="[166,1422,245,1443]" captionTargetId-0="figure-22@7.[151,1436,181,1625]" captionTargetId-1="figure-40@8.[158,1430,237,1451]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." captionText-1="FIGURE 6. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively." figureDoi-0="http://doi.org/10.5281/zenodo.10301498" figureDoi-1="http://doi.org/10.5281/zenodo.10301502" httpUri-0="https://zenodo.org/record/10301498/files/figure.png" httpUri-1="https://zenodo.org/record/10301502/files/figure.png" pageId="6" pageNumber="30">Figs 5–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E9086BFFEA35F5FD1D5BCEFD3C" box="[151,718,688,713]" pageId="6" pageNumber="30" type="reference_group">
<paragraph id="BA4BD062086BFFEA35F5FD1D5BCEFD3C" blockId="6.[151,718,688,713]" box="[151,718,688,713]" pageId="6" pageNumber="30">
<treatmentCitationGroup id="9AE4F74C086BFFEA35F5FD1D5BCEFD3C" box="[151,718,688,713]" pageId="6" pageNumber="30">
<taxonomicName id="7DF4ABE1086BFFEA35F5FD1D5851FD3D" box="[151,337,688,712]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="30" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="88800C70086BFFEA35F5FD1D582EFD3D" box="[151,302,688,712]" italics="true" pageId="6" pageNumber="30">Elaphognathia</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="3B55F673086BFFEA343EFD1D5B76FD3C" author="Al-Kandari, M. A. &amp; Khalaji-Pirbalouty, V. &amp; Abdulkhaliq, H. &amp; Chen, W." box="[348,630,688,713]" page="118" pageId="6" pageNumber="30" year="2022">
<bibRefCitation id="DE65AD93086BFFEA343EFD1D5B76FD3C" author="Al-Kandari, M. A. &amp; Khalaji-Pirbalouty, V. &amp; Abdulkhaliq, H. &amp; Chen, W." box="[348,630,688,713]" pageId="6" pageNumber="30" pagination="107 - 133" refId="ref6210" refString="Al-Kandari, M. A., Khalaji-Pirbalouty, V., Abdulkhaliq, H. &amp; Chen, W. (2022) Diversity and distribution of the Isopoda (Crustacea, Peracarida) of Kuwait, with an updated checklist. ZooKeys, 1080, 107 - 133. https: // doi. org / 10.3897 / zookeys. 1080.71370" type="journal article" year="2022">
Al-Kandari
<emphasis id="88800C70086BFFEA34B4FD1C5B03FD3D" box="[470,515,688,712]" italics="true" pageId="6" pageNumber="30">et al</emphasis>
. 2022: 118
</bibRefCitation>
</treatmentCitation>
, fig. 3B.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E9086BFFEA35F5FD5E585DFCA0" pageId="6" pageNumber="30" type="materials_examined">
<paragraph id="BA4BD062086BFFEA35F5FD5E585DFCA0" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35F5FD5E58C4FCF8" bold="true" box="[151,452,754,781]" pageId="6" pageNumber="30">
<typeStatus id="654F6EC0086BFFEA35F5FD5E59D2FCF8" box="[151,210,755,781]" pageId="6" pageNumber="30">Type</typeStatus>
material.
<materialsCitation id="0A9CDA3F086BFFEA3437FD5F5A28FCC4" ID-GBIF-Occurrence="4464606309" collectingDate="2014-11-24" collectionCode="ZMH-K" latitude="29.656717" location="Al-Shumaimah" longLatPrecision="1" longitude="48.130833" municipality="Khor Al-Subiyah" pageId="6" pageNumber="30" specimenCode="63976" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="88800C70086BFFEA3437FD5F58C4FCF8" bold="true" box="[341,452,754,781]" italics="true" pageId="6" pageNumber="30">
<typeStatus id="654F6EC0086BFFEA3437FD5F58C0FCF8" box="[341,448,754,781]" pageId="6" pageNumber="30" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="ACF21BEB086BFFEA34B2FD5E58FFFCF8" box="[464,511,755,781]" count="1" pageId="6" pageNumber="30" type="male">1 ♂</specimenCount>
(
<quantity id="7D0C7D87086BFFEA3771FD5E5B73FCF8" box="[531,627,755,781]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="30" unit="mm" value="2.1">2.1 mm</quantity>
);
<collectingMunicipality id="5A2F4A18086BFFEA37E9FD5E5A50FCF8" box="[651,848,755,781]" pageId="6" pageNumber="30">Khor Al-Subiyah</collectingMunicipality>
(
<location id="BF2B86B9086BFFEA3601FD5E5D15FCF8" LSID="urn:lsid:plazi:treatment:325D6174086BFFE435F5FD8A59F1F803:BF2B86B9086BFFEA3601FD5E5D15FCF8" box="[867,1045,755,781]" latitude="29.656717" longLatPrecision="1" longitude="48.130833" municipality="Khor Al-Subiyah" name="Al-Shumaimah" pageId="6" pageNumber="30">Al-Shumaimah</location>
);
<geoCoordinate id="DFC0B6A5086BFFEA314FFD5E5DCEFCF8" box="[1069,1230,755,781]" degrees="29" direction="north" minutes="39.403" orientation="latitude" pageId="6" pageNumber="30" precision="1" value="29.656717">29° 39.403’N</geoCoordinate>
;
<geoCoordinate id="DFC0B6A5086BFFEA31BCFD5F5C70FCF8" box="[1246,1392,754,781]" degrees="48" direction="east" minutes="07.850" orientation="longitude" pageId="6" pageNumber="30" precision="1" value="48.130833">48°07.850’E</geoCoordinate>
;
<date id="CE4AF6A2086BFFEA30E2FD5E5852FCC4" pageId="6" pageNumber="30" value="2014-11-24">
<collectingDate id="DE0E0F4A086BFFEA30E2FD5E5852FCC4" pageId="6" pageNumber="30" value="2014-11-24">24 November 2014</collectingDate>
</date>
, dead coral,
<quantity id="7D0C7D87086BFFEA3484FCBB5B59FCC4" box="[486,601,790,817]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="6" pageNumber="30" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 m</quantity>
(
<collectionCode id="DCE548A7086BFFEA3708FCBA5BD3FCC4" box="[618,723,791,817]" pageId="6" pageNumber="30">ZMH-K-</collectionCode>
<specimenCode id="EA527819086BFFEA37B1FCBA5A1CFCC4" box="[723,796,791,817]" pageId="6" pageNumber="30">63976</specimenCode>
).
</materialsCitation>
</emphasis>
<materialsCitation id="0A9CDA3F086BFFEA3652FCBA5859FCA0" ID-GBIF-Occurrence="4464606302" collectingDate="2014-11-24" collectionCode="ZMH-K" latitude="29.656717" location="Al-Shumaimah" longLatPrecision="1" longitude="48.130833" municipality="Khor Al-Subiyah" pageId="6" pageNumber="30" specimenCode="63977" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="88800C70086BFFEA3652FCBA5A9EFCC4" bold="true" box="[816,926,791,817]" italics="true" pageId="6" pageNumber="30">
<typeStatus id="654F6EC0086BFFEA3652FCBA5A99FCC4" box="[816,921,791,817]" pageId="6" pageNumber="30" type="paratype">Paratype</typeStatus>
.
</emphasis>
1 Praniza (
<quantity id="7D0C7D87086BFFEA3146FCBA5D82FCC4" box="[1060,1154,791,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="30" unit="mm" value="1.4">1.4 mm</quantity>
); same data as holotype, (
<collectionCode id="DCE548A7086BFFEA35FDFC965809FCA0" box="[159,265,827,853]" pageId="6" pageNumber="30">ZMH-K-</collectionCode>
<specimenCode id="EA527819086BFFEA346BFC965851FCA0" box="[265,337,827,853]" pageId="6" pageNumber="30">63977</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E9086BFFEA35A5FCF35D12FC34" pageId="6" pageNumber="30" type="diagnosis">
<paragraph id="BA4BD062086BFFEA35A5FCF35D12FC34" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5FCF35842FC8D" bold="true" box="[199,322,862,888]" pageId="6" pageNumber="30">Diagnosis.</emphasis>
Frontal margin with deep excavation; a conical and acute mediofrontal process, bearing two fine long setae on each side. Mandible elongate, distally curved, with a single pronounced conical internal lobe. Lobi laterales absent on pereonite 6. Pylopod with three articles, third article minute.
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E9086BFFE435A5FC6758FDFF20" lastPageId="8" lastPageNumber="32" pageId="6" pageNumber="30" type="description">
<paragraph id="BA4BD062086BFFEA35A5FC675D70FC10" blockId="6.[151,1437,754,2041]" box="[199,1136,970,997]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5FC6758B7FC10" bold="true" box="[199,439,970,997]" pageId="6" pageNumber="30">Description of male.</emphasis>
<emphasis id="88800C70086BFFEA34DCFC6658F8FC10" box="[446,504,971,997]" italics="true" pageId="6" pageNumber="30">Body</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3765FC665B5FFC10" box="[519,607,970,997]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA372BFC675B5FFC10" box="[585,607,970,997]" pageId="6" pageNumber="30">A</collectionCode>
</figureCitation>
) length
<quantity id="7D0C7D87086BFFEA37D9FC665A13FC10" box="[699,787,971,997]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="30" unit="mm" value="2.1">2.1 mm</quantity>
, about 3 times as long as wide.
</paragraph>
<paragraph id="BA4BD062086BFFEA35A5FC425AF7FA84" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5FC42585FFBFC" box="[199,351,1007,1033]" italics="true" pageId="6" pageNumber="30">Cephalosome</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3412FC4258EBFBFC" box="[368,491,1006,1033]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA34D0FC4358C9FBFC" box="[434,457,1006,1033]" pageId="6" pageNumber="30">A</collectionCode>
,
<collectionCode id="DCE548A7086BFFEA34B4FC4258EBFBFC" box="[470,491,1007,1033]" pageId="6" pageNumber="30">B</collectionCode>
</figureCitation>
), about 0.7 times as long as wide.
<emphasis id="88800C70086BFFEA3611FC425D37FBFC" box="[883,1079,1007,1033]" italics="true" pageId="6" pageNumber="30">Supraocular lobe</emphasis>
conical, rounded.
<emphasis id="88800C70086BFFEA306BFC4259ECFBD9" italics="true" pageId="6" pageNumber="30">Mediofrontal process</emphasis>
present, conical, acute, each side with 2 fine long setae.
<emphasis id="88800C70086BFFEA36F1FBBE5DE3FBD9" box="[915,1251,1043,1069]" italics="true" pageId="6" pageNumber="30">Inferior frontolateral process</emphasis>
small, serrated.
<emphasis id="88800C70086BFFEA35F5FB9A58EFFBA5" box="[151,495,1079,1105]" italics="true" pageId="6" pageNumber="30">Superior frontolateral process</emphasis>
present, strong, with 4 fine long setae. Dorsal sulcus wide, deep, triangular. Eyes lateral, about 0.2 times as long as cephalosome length.
<emphasis id="88800C70086BFFEA3795FBF65A9EFB81" box="[759,926,1114,1140]" italics="true" pageId="6" pageNumber="30">Pereonites 1–6</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA36CEFBF65D03FB80" box="[940,1027,1114,1141]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA368FFBF75D03FB80" box="[1005,1027,1114,1141]" pageId="6" pageNumber="30">A</collectionCode>
</figureCitation>
) without tubercles on dorsal surface, lateral margins smooth. Pereonites 1–2 subequal.
<emphasis id="88800C70086BFFEA37BEFBD25A60FB6C" box="[732,864,1151,1177]" italics="true" pageId="6" pageNumber="30">Pereonite 1</emphasis>
dorsally reaching lateral margins, divided into 3 regions by posterior margin of cephalosome. Pereonite 3 smaller then the previous ones.
<emphasis id="88800C70086BFFEA31FBFB0E5C1BFB48" box="[1177,1307,1187,1213]" italics="true" pageId="6" pageNumber="30">Pereonite 4</emphasis>
longer and narrower than pereonite 3, with anterior constriction.
<emphasis id="88800C70086BFFEA3667FB6A5A89FB14" box="[773,905,1223,1249]" italics="true" pageId="6" pageNumber="30">Pereonite 5</emphasis>
with areae laterales.
<emphasis id="88800C70086BFFEA31E1FB6A5C07FB15" box="[1155,1287,1222,1248]" italics="true" pageId="6" pageNumber="30">Pereonite 6</emphasis>
without lobi laterales; with weak lobuii.
<emphasis id="88800C70086BFFEA34B3FB465B53FAF0" box="[465,595,1259,1285]" italics="true" pageId="6" pageNumber="30">Pereonite 7</emphasis>
overlapping pleon, longer than pleonite 1, with rounded posterior margin.
<emphasis id="88800C70086BFFEA35F5FAA25830FADC" box="[151,304,1295,1321]" italics="true" pageId="6" pageNumber="30">Pleonites 1–5</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3422FAA25899FADC" box="[320,409,1294,1321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA34E0FAA35899FADC" box="[386,409,1294,1321]" pageId="6" pageNumber="30">A</collectionCode>
</figureCitation>
), epimera visible dorsally, a row of 3 fine setae posteriorly on all pleonites.
<emphasis id="88800C70086BFFEA318BFAA25C5FFADC" box="[1257,1375,1295,1321]" italics="true" pageId="6" pageNumber="30">Pleotelson</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA300DFAA259B9FAB8" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA35C7FA9F59B9FAB8" box="[165,185,1330,1357]" pageId="6" pageNumber="30">C</collectionCode>
</figureCitation>
) 0.95 times as long as anterior width; lateral margins smooth; mid-dorsal surface with 2 sub-median simple setae, posterolateral margins with 1 submarginal seta, apex with 2 long setae.
</paragraph>
<paragraph id="BA4BD062086BFFEA35A5FAD658B0F9F4" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5FAD6583AFA60" box="[199,314,1403,1429]" italics="true" pageId="6" pageNumber="30">Antennula</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3428FAD658A1FA60" box="[330,417,1402,1429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA34E9FAD758A1FA60" box="[395,417,1402,1429]" pageId="6" pageNumber="30">D</collectionCode>
</figureCitation>
) short, peduncle article 2 0.9 times as long as article 1; article 3 1.3 times as long as article 2; flagellum with 5 articles, about 1.25 times as long as peduncle article 3, articles 3–5 each bearing an aesthetasc.
<emphasis id="88800C70086BFFEA35F5FA6E59F4FA28" box="[151,244,1475,1501]" italics="true" pageId="6" pageNumber="30">Antenna</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3467FA6E585DFA28" box="[261,349,1474,1501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA3425FA6F585DFA28" box="[327,349,1474,1501]" pageId="6" pageNumber="30">D</collectionCode>
</figureCitation>
) peduncle article 4 1.25 times as long as article 3, with 2 small sensory palmate setae ventrally; flagellum with 7 articles.
</paragraph>
<paragraph id="BA4BD062086BFFEA35A5F9A65AECF9BC" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5F9A65832F9D0" box="[199,306,1547,1573]" italics="true" pageId="6" pageNumber="30">Mandible</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3421F9A65898F9D0" box="[323,408,1546,1573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA34E7F9A75898F9D0" box="[389,408,1546,1573]" pageId="6" pageNumber="30">E</collectionCode>
</figureCitation>
) elongate, distally curved; about 1.3 times as long as cephalosome; with no obvious blade; with a single pronounced conical internal lobe, mandibular seta at midpoint.
</paragraph>
<paragraph id="BA4BD062086BFFEA35A5F9FE59EFF940" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5F9FE5840F998" box="[199,320,1619,1645]" italics="true" pageId="6" pageNumber="30">Maxilliped</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3430F9FE58A6F998" box="[338,422,1618,1645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA34F7F9FF58A6F998" box="[405,422,1618,1645]" pageId="6" pageNumber="30">F</collectionCode>
</figureCitation>
) article 1 lateral margin fringed with fine, small setae; endite extended beyond article 2; external margins of articles 2–5 bearing 2, 8, 5 and 7 stout plumose setae respectively; article 5 with 5 simple setae distally.
</paragraph>
<paragraph id="BA4BD062086BFFEA35A5F91258B0F8B0" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5F9125824F92C" box="[199,292,1727,1753]" italics="true" pageId="6" pageNumber="30">Pylopod</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3451F9125889F92C" box="[307,393,1726,1753]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="6" pageNumber="30">
Fig. 5
<collectionCode id="DCE548A7086BFFEA3411F9135889F92C" box="[371,393,1726,1753]" pageId="6" pageNumber="30">G</collectionCode>
</figureCitation>
) with 3 articles; article 1 about 1.4 times as long as wide, lateral margin of article 1 with about 27 plumose setae; mesial and proximal margins with continuous fine setae, mesial margin with 2 submarginal long setae; distal end with 4 setae; article 2, 1.8 times as long as wide, fringed with dense fine setae, 6 long simple setae distally; article 3 minute.
</paragraph>
<paragraph id="BA4BD062086BFFEA35A5F8E25DD4F80C" blockId="6.[151,1437,754,2041]" pageId="6" pageNumber="30">
<emphasis id="88800C70086BFFEA35A5F8E2584BF89C" box="[199,331,1871,1897]" italics="true" pageId="6" pageNumber="30">Pereopod 2</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA343CF8E258BAF89C" box="[350,442,1871,1897]" captionStart="FIGURE 6" captionStartId="8.[151,250,1474,1498]" captionTargetBox="[166,1422,245,1443]" captionTargetId="figure-40@8.[158,1430,237,1451]" captionTargetPageId="8" captionText="FIGURE 6. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively." figureDoi="http://doi.org/10.5281/zenodo.10301502" httpUri="https://zenodo.org/record/10301502/files/figure.png" pageId="6" pageNumber="30">
Fig. 6
<collectionCode id="DCE548A7086BFFEA34C6F8E258BAF89C" box="[420,442,1871,1897]" pageId="6" pageNumber="30">A</collectionCode>
</figureCitation>
) basis 2.9 times as long as greatest width, superior margin with 2 long and 2sensory palmate setae, inferior margin 6 long setae; ischium 0.7 times as long as basis,
<emphasis id="88800C70086BFFEA314CF8DE5D72F879" box="[1070,1138,1907,1932]" italics="true" pageId="6" pageNumber="30">merus</emphasis>
as long as carpus; carpus inferior margin with 1 biserrate robust seta distally; propodus 3.4 times as long as wide, superior margin with 1 simple and 1 sensory palmate setae, inferior margin with 2 robust serrate setae.
<emphasis id="88800C70086BFFEA317FF8165DA9F820" box="[1053,1193,1979,2005]" italics="true" pageId="6" pageNumber="30">Pereopods 3</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA31D8F8165C10F820" box="[1210,1296,1979,2005]" captionStart="FIGURE 6" captionStartId="8.[151,250,1474,1498]" captionTargetBox="[166,1422,245,1443]" captionTargetId="figure-40@8.[158,1430,237,1451]" captionTargetPageId="8" captionText="FIGURE 6. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively." figureDoi="http://doi.org/10.5281/zenodo.10301502" httpUri="https://zenodo.org/record/10301502/files/figure.png" pageId="6" pageNumber="30">
Fig. 6
<collectionCode id="DCE548A7086BFFEA319FF8165C10F820" box="[1277,1296,1979,2005]" pageId="6" pageNumber="30">B</collectionCode>
</figureCitation>
) and
<emphasis id="88800C70086BFFEA3033F8165C5FF820" box="[1361,1375,1979,2005]" italics="true" pageId="6" pageNumber="30">4</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA300DF81659B9F80C" captionStart="FIGURE 6" captionStartId="8.[151,250,1474,1498]" captionTargetBox="[166,1422,245,1443]" captionTargetId="figure-40@8.[158,1430,237,1451]" captionTargetPageId="8" captionText="FIGURE 6. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively." figureDoi="http://doi.org/10.5281/zenodo.10301502" httpUri="https://zenodo.org/record/10301502/files/figure.png" pageId="6" pageNumber="30">
Fig. 6
<collectionCode id="DCE548A7086BFFEA35C7F87259B9F80C" box="[165,185,2015,2041]" pageId="6" pageNumber="30">C</collectionCode>
</figureCitation>
) similar to pereopod 2. As illustrated
<emphasis id="88800C70086BFFEA3702F8725BE1F80C" box="[608,737,2015,2041]" italics="true" pageId="6" pageNumber="30">Pereopod 5</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3793F8725A48F80C" box="[753,840,2015,2041]" captionStart="FIGURE 6" captionStartId="8.[151,250,1474,1498]" captionTargetBox="[166,1422,245,1443]" captionTargetId="figure-40@8.[158,1430,237,1451]" captionTargetPageId="8" captionText="FIGURE 6. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively." figureDoi="http://doi.org/10.5281/zenodo.10301502" httpUri="https://zenodo.org/record/10301502/files/figure.png" pageId="6" pageNumber="30">
Fig. 6
<collectionCode id="DCE548A7086BFFEA3651F8725A48F80C" box="[819,840,2015,2041]" pageId="6" pageNumber="30">D</collectionCode>
</figureCitation>
) is similar to
<emphasis id="88800C70086BFFEA3681F8725D60F80D" box="[995,1120,2014,2041]" italics="true" pageId="6" pageNumber="30">pereopod 6</emphasis>
(
<figureCitation id="22CFCCE7086BFFEA3112F8725DC7F80C" box="[1136,1223,2015,2041]" captionStart="FIGURE 6" captionStartId="8.[151,250,1474,1498]" captionTargetBox="[166,1422,245,1443]" captionTargetId="figure-40@8.[158,1430,237,1451]" captionTargetPageId="8" captionText="FIGURE 6. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively." figureDoi="http://doi.org/10.5281/zenodo.10301502" httpUri="https://zenodo.org/record/10301502/files/figure.png" pageId="6" pageNumber="30">
Fig. 6
<collectionCode id="DCE548A7086BFFEA31D1F8725DC7F80C" box="[1203,1223,2015,2041]" pageId="6" pageNumber="30">E</collectionCode>
</figureCitation>
).
</paragraph>
<caption id="EE8B80EA086AFFEB35F5F9DC5889F927" ID-DOI="http://doi.org/10.5281/zenodo.10301498" ID-Zenodo-Dep="10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="7" pageNumber="31" startId="7.[151,250,1649,1673]" targetBox="[159,1428,183,1617]" targetPageId="7" targetType="figure">
<paragraph id="BA4BD062086AFFEB35F5F9DC5889F927" blockId="7.[151,1436,1649,1746]" pageId="7" pageNumber="31">
<emphasis id="88800C70086AFFEB35F5F9DC5819F97F" bold="true" box="[151,281,1649,1674]" pageId="7" pageNumber="31">FIGURE 5.</emphasis>
<taxonomicName id="7DF4ABE1086AFFEB3447F9DC5B50F97C" authority="Khalaji-Pirbalouty &amp; Al-Kandari, 2023" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2023" box="[293,592,1649,1673]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="31" phylum="Arthropoda" rank="species" species="alshumaimah" status="sp. nov.">
<emphasis id="88800C70086AFFEB3447F9DC5B50F97C" box="[293,592,1649,1673]" italics="true" pageId="7" pageNumber="31">Elaphognathia alshumaimah</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93B3B10B086AFFEB373EF9DF5BB0F97C" box="[604,688,1650,1673]" pageId="7" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2.
</paragraph>
</caption>
<paragraph id="BA4BD062086AFFEB35A5F8895B48F85F" blockId="7.[151,1436,1827,2034]" pageId="7" pageNumber="31">
<emphasis id="88800C70086AFFEB35A5F889583BF8CB" box="[199,315,1828,1854]" italics="true" pageId="7" pageNumber="31">Pleopod 1</emphasis>
(
<figureCitation id="22CFCCE7086AFFEB3429F889589DF8CB" box="[331,413,1827,1854]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="7" pageNumber="31">
Fig 5
<collectionCode id="DCE548A7086AFFEB34E5F88E589DF8CB" box="[391,413,1827,1854]" pageId="7" pageNumber="31">H</collectionCode>
</figureCitation>
) endopod and exopod with 8 plumose marginal setae; endopod longer than exopod, about 2.1 times as long as wide; exopod 2.3 times as long as wide; sympodite mesial margin with 2 coupling hooks, lateral margin with 1 stout simple seta.
<emphasis id="88800C70086AFFEB3760F8C15B74F873" box="[514,628,1900,1926]" italics="true" pageId="7" pageNumber="31">Pleopod 2</emphasis>
(
<figureCitation id="22CFCCE7086AFFEB37E6F8C15BD0F873" box="[644,720,1899,1926]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="7" pageNumber="31">
Fig.
<date id="CE4AF6A2086AFFEB37DAF8C65BD0F873" box="[696,720,1899,1926]" pageId="7" pageNumber="31">
5
<collectionCode id="DCE548A7086AFFEB37A4F8C65BD0F873" box="[710,720,1899,1926]" pageId="7" pageNumber="31">I</collectionCode>
</date>
</figureCitation>
) endopod and exopod shape as in pleopod 1, distally with 7 and 9 plumose marginal setae respectively.
</paragraph>
<paragraph id="BA4BD062086AFFE435A5F81958FDFF20" blockId="7.[151,1436,1827,2034]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="32" pageId="7" pageNumber="31">
<emphasis id="88800C70086AFFEB35A5F8195833F83B" box="[199,307,1972,1998]" italics="true" pageId="7" pageNumber="31">Uropodal</emphasis>
(
<figureCitation id="22CFCCE7086AFFEB3426F8195896F83B" box="[324,406,1971,1998]" captionStart="FIGURE 5" captionStartId="7.[151,250,1649,1673]" captionTargetBox="[159,1428,183,1617]" captionTargetId="figure-22@7.[151,1436,181,1625]" captionTargetPageId="7" captionText="FIGURE 5. Elaphognathia alshumaimah sp. nov. (ZMH-K-63976), male holotype: A, dorsal view; B, ventral view of cephalosome; C, dorsal view of pleotelson and uropod; D, antennula and antenna; E, mandible; F, maxilliped; G, pylopod; H, pleopod 1; I, pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.10301498" httpUri="https://zenodo.org/record/10301498/files/figure.png" pageId="7" pageNumber="31">
Fig 5
<collectionCode id="DCE548A7086AFFEB34E0F81E5896F83B" box="[386,406,1971,1998]" pageId="7" pageNumber="31">C</collectionCode>
</figureCitation>
) rami extending beyond pleotelson apex, endopod and exopod nearly subequal in length, endopod wider than exopod. Endopod lateral margin fringed with small fine setae between 3 long setae; mesial margin weakly convex, with 7 long plumose setae. Exopod mesial and lateral margins fringed with small fine setae, between 10 long plumose setae.
</paragraph>
</subSubSection>
<caption id="EE8B80EA0865FFE435F5FA6F5C45FA2F" ID-DOI="http://doi.org/10.5281/zenodo.10301502" ID-Zenodo-Dep="10301502" box="[151,1349,1474,1499]" httpUri="https://zenodo.org/record/10301502/files/figure.png" pageId="8" pageNumber="32" startId="8.[151,250,1474,1498]" targetBox="[166,1422,245,1443]" targetPageId="8" targetType="figure">
<paragraph id="BA4BD0620865FFE435F5FA6F5C45FA2F" blockId="8.[151,1349,1474,1499]" box="[151,1349,1474,1499]" pageId="8" pageNumber="32">
<emphasis id="88800C700865FFE435F5FA6F5813FA2F" bold="true" box="[151,275,1474,1498]" pageId="8" pageNumber="32">FIGURE 6.</emphasis>
<taxonomicName id="7DF4ABE10865FFE4347BFA6F5B3FFA2F" authority="Khalaji-Pirbalouty &amp; Al-Kandari, 2023" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2023" box="[281,575,1474,1498]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="alshumaimah" status="sp. nov.">
<emphasis id="88800C700865FFE4347BFA6F5B3FFA2F" box="[281,575,1474,1498]" italics="true" pageId="8" pageNumber="32">Elaphognathia alshumaimah</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93B3B10B0865FFE43727FA6E5B94FA2F" box="[581,660,1475,1498]" pageId="8" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
(ZMH-K-63976), male holotype, A–E: pereopods 2–6 respectively.
</paragraph>
</caption>
<subSubSection id="F2EE83E90865FFE435A5F9815BF1F9B3" box="[199,753,1580,1607]" pageId="8" pageNumber="32" type="distribution">
<paragraph id="BA4BD0620865FFE435A5F9815BF1F9B3" blockId="8.[151,1437,1580,2038]" box="[199,753,1580,1607]" pageId="8" pageNumber="32">
<emphasis id="88800C700865FFE435A5F9815861F9B3" bold="true" box="[199,353,1580,1606]" pageId="8" pageNumber="32">Distribution.</emphasis>
Only known from the
<typeStatus id="654F6EC00865FFE43702F9805B90F9B2" box="[608,656,1581,1607]" pageId="8" pageNumber="32">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E90865FFE435A5F9FC5D8CF85B" pageId="8" pageNumber="32" type="discussion">
<paragraph id="BA4BD0620865FFE435A5F9FC5DC8F90E" blockId="8.[151,1437,1580,2038]" pageId="8" pageNumber="32">
<emphasis id="88800C700865FFE435A5F9FC583BF99E" bold="true" box="[199,315,1617,1643]" pageId="8" pageNumber="32">Remarks.</emphasis>
Eight species of the genus
<taxonomicName id="7DF4ABE10865FFE437FDF9FD5A45F99F" authorityName="Monod" authorityYear="1926" box="[671,837,1616,1642]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="88800C700865FFE437FDF9FD5A45F99F" box="[671,837,1616,1642]" italics="true" pageId="8" pageNumber="32">Elaphognathia</emphasis>
</taxonomicName>
have been described from the Indian Ocean:
<taxonomicName id="7DF4ABE10865FFE430E6F9FC5B82F97A" authority="Kensley, Schotte &amp; Poore, 2009" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="aldabrae">
<emphasis id="88800C700865FFE430E6F9FC59FCF97B" italics="true" pageId="8" pageNumber="32">E. aldabrae</emphasis>
<bibRefCitation id="DE65AD930865FFE43461F9D95B82F97A" author="Kensley, B. &amp; Schotte, M. &amp; Poore, G. C. B." box="[259,642,1652,1679]" pageId="8" pageNumber="32" pagination="32 - 51" refId="ref6652" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (2), 32 - 51. https: // doi. org / 10.2988 / 07 - 16.1" type="journal article" year="2009">Kensley, Schotte &amp; Poore, 2009</bibRefCitation>
</taxonomicName>
from Picard Island,
<collectingCountry id="C2E390F20865FFE4361DF9D95AFAF97B" box="[895,1018,1652,1678]" name="Seychelles" pageId="8" pageNumber="32">Seychelles</collectingCountry>
;
<taxonomicName id="7DF4ABE10865FFE43169F9D85C59F97A" authority="(Cals, 1978)" baseAuthorityName="Cals" baseAuthorityYear="1978" box="[1035,1369,1652,1679]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="88800C700865FFE43169F9D85DBCF97B" box="[1035,1212,1652,1678]" italics="true" pageId="8" pageNumber="32">E. amboinensis</emphasis>
(Cals, 1978)
</taxonomicName>
from
<collectingCountry id="C2E390F20865FFE435F5F9355808F947" box="[151,264,1688,1714]" name="Indonesia" pageId="8" pageNumber="32">Indonesia</collectingCountry>
;
<taxonomicName id="7DF4ABE10865FFE43470F9345BE3F946" authority="Kensley, Schotte &amp; Poore, 2009" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[274,739,1688,1715]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="gladia">
<emphasis id="88800C700865FFE43470F9345879F947" box="[274,377,1688,1714]" italics="true" pageId="8" pageNumber="32">E. gladia</emphasis>
<bibRefCitation id="DE65AD930865FFE4341DF9355BE3F946" author="Kensley, B. &amp; Schotte, M. &amp; Poore, G. C. B." box="[383,739,1688,1715]" pageId="8" pageNumber="32" pagination="32 - 51" refId="ref6652" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (2), 32 - 51. https: // doi. org / 10.2988 / 07 - 16.1" type="journal article" year="2009">Kensley, Schotte &amp; Poore, 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7DF4ABE10865FFE4367AF9345DF6F946" authority="Kensley, Schotte &amp; Poore, 2009" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[792,1270,1688,1715]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="88800C700865FFE4367AF9345A8BF947" box="[792,907,1689,1714]" italics="true" pageId="8" pageNumber="32">E. ramosa</emphasis>
<bibRefCitation id="DE65AD930865FFE436F3F9355DF6F946" author="Kensley, B. &amp; Schotte, M. &amp; Poore, G. C. B." box="[913,1270,1688,1715]" pageId="8" pageNumber="32" pagination="32 - 51" refId="ref6652" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (2), 32 - 51. https: // doi. org / 10.2988 / 07 - 16.1" type="journal article" year="2009">Kensley, Schotte &amp; Poore, 2009</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="C2E390F20865FFE4305BF9355C98F947" box="[1337,1432,1688,1714]" name="Somalia" pageId="8" pageNumber="32">Somalia</collectingCountry>
;
<taxonomicName id="7DF4ABE10865FFE435F5F91058D3F922" authority="(Stebbing, 1905)" baseAuthorityName="Stebbing" baseAuthorityYear="1905" box="[151,467,1724,1751]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="88800C700865FFE435F5F910580CF923" box="[151,268,1724,1750]" italics="true" pageId="8" pageNumber="32">E. insolita</emphasis>
(
<bibRefCitation id="DE65AD930865FFE4347FF91158CBF922" author="Stebbing, T. R. R." box="[285,459,1724,1751]" pageId="8" pageNumber="32" pagination="1 - 64" refId="ref7130" refString="Stebbing, T. R. R. (1905) Report on the Isopoda collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A., Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar. Supplemental report, 23, 1 - 64." type="journal article" year="1905">Stebbing, 1905</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="C2E390F20865FFE43779F9115B8EF923" box="[539,654,1724,1750]" name="Sri Lanka" pageId="8" pageNumber="32">Sri Lanka</collectingCountry>
;
<taxonomicName id="7DF4ABE10865FFE437F9F9105DC2F922" authority="Svavarsson &amp; Gisladottir, 2002" authorityName="Svavarsson &amp; Gisladottir" authorityYear="2002" box="[667,1218,1724,1751]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="korachaensis">
<emphasis id="88800C700865FFE437F9F9105A50F923" box="[667,848,1724,1750]" italics="true" pageId="8" pageNumber="32">E. korachaensis</emphasis>
<bibRefCitation id="DE65AD930865FFE4363AF9105DC2F922" author="Svavarsson, J. &amp; Gisladottir, E." box="[856,1218,1724,1751]" pageId="8" pageNumber="32" pagination="157 - 164" refId="ref7282" refString="Svavarsson, J. &amp; Gisladottir, E. (2002) Elaphognathia korachaensis sp. Nov., a new gnathiid species (Crustacea, Isopoda) from Thailand. Phuket Marine Biological Center Special Publication, 23 (10), 157 - 164." type="journal article" year="2002">Svavarsson &amp; Gisladottir, 2002</bibRefCitation>
</taxonomicName>
: Andaman Sea;
<taxonomicName id="7DF4ABE10865FFE430E6F91058A4F90F" authority="(Monod, 1926)" baseAuthorityName="Monod" baseAuthorityYear="1926" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="rangifer">
<emphasis id="88800C700865FFE430E6F91059F3F90E" italics="true" pageId="8" pageNumber="32">E. rangifer</emphasis>
(
<bibRefCitation id="DE65AD930865FFE43460F94D589BF90F" author="Monod, T." box="[258,411,1760,1786]" pageId="8" pageNumber="32" pagination="1 - 668" refId="ref6823" refString="Monod, T. (1926) Les Gnathiidae. Memories de la Societe des Sciences Naturelles du Maroc, 13, 1 - 668. [in French]" type="journal article" year="1926">Monod, 1926</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="C2E390F20865FFE43485F94D5B4BF90F" box="[487,587,1760,1786]" name="Thailand" pageId="8" pageNumber="32">Thailand</collectingCountry>
and
<collectingCountry id="C2E390F20865FFE437E3F94D5BF3F90F" box="[641,755,1760,1786]" name="Singapore" pageId="8" pageNumber="32">Singapore</collectingCountry>
and
<taxonomicName id="7DF4ABE10865FFE4364BF94C5D33F90E" authority="(Muller, 1989)" baseAuthorityName="Muller" baseAuthorityYear="1989" box="[809,1075,1760,1787]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="wolffi">
<emphasis id="88800C700865FFE4364BF94C5A88F90F" box="[809,904,1760,1786]" italics="true" pageId="8" pageNumber="32">E. wolffi</emphasis>
(
<bibRefCitation id="DE65AD930865FFE436F5F94D5D2BF90E" author="Muller, H. G." box="[919,1067,1760,1787]" pageId="8" pageNumber="32" pagination="25 - 30" refId="ref6906" refString="Muller, H. G. (1989) Redescription of Gnathia sugashimaensis Nunomura (Crustacea: Isopoda) from Japan. Publications of the Seto Marine Biological Laboratory, 34, 25 - 30. https: // doi. org / 10.5134 / 176161" type="journal article" year="1989">Müller, 1989</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="C2E390F20865FFE43115F94C5DC4F90E" box="[1143,1220,1761,1787]" name="Kenya" pageId="8" pageNumber="32">Kenya</collectingCountry>
.
</paragraph>
<paragraph id="BA4BD0620865FFE435A5F8A95AF2F892" blockId="8.[151,1437,1580,2038]" pageId="8" pageNumber="32">
The new species is most similar to
<taxonomicName id="7DF4ABE10865FFE43737F8A85D2EF8EA" authority="Kensley, Schotte &amp; Poore, 2009" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[597,1070,1796,1823]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="gladia">
<emphasis id="88800C700865FFE43737F8A85BBCF8EB" box="[597,700,1796,1822]" italics="true" pageId="8" pageNumber="32">E. gladia</emphasis>
<bibRefCitation id="DE65AD930865FFE437A6F8A95D2EF8EA" author="Kensley, B. &amp; Schotte, M. &amp; Poore, G. C. B." box="[708,1070,1796,1823]" pageId="8" pageNumber="32" pagination="32 - 51" refId="ref6652" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (2), 32 - 51. https: // doi. org / 10.2988 / 07 - 16.1" type="journal article" year="2009">
Kensley, Schotte &amp; Poore,
<quantity id="7D0C7D870865FFE43694F8A85D4BF8EB" box="[1014,1099,1796,1823]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="8" pageNumber="32" unit="in" value="2009.0">
<quantity id="7D0C7D870865FFE43694F8A85D2EF8EA" box="[1014,1070,1797,1823]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="8" pageNumber="32" unit="in" value="2009.0">2009</quantity>
in
</quantity>
</bibRefCitation>
</taxonomicName>
having a long, thin saber-like mandible. However, it differs from
<taxonomicName id="7DF4ABE10865FFE43747F8845B8CF8B7" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[549,652,1832,1858]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="gladia">
<emphasis id="88800C700865FFE43747F8845B8CF8B7" box="[549,652,1832,1858]" italics="true" pageId="8" pageNumber="32">E. gladia</emphasis>
</taxonomicName>
in having a mandible with only one conical lobe at its base (vs two in
<taxonomicName id="7DF4ABE10865FFE435F5F8E059FEF893" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[151,254,1868,1894]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="gladia">
<emphasis id="88800C700865FFE435F5F8E059FEF893" box="[151,254,1868,1894]" italics="true" pageId="8" pageNumber="32">E. gladia</emphasis>
</taxonomicName>
) and having an acute mediofrontal process (vs absent in
<taxonomicName id="7DF4ABE10865FFE43619F8E05AE3F893" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[891,995,1868,1894]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="gladia">
<emphasis id="88800C700865FFE43619F8E05AE3F893" box="[891,995,1868,1894]" italics="true" pageId="8" pageNumber="32">E. gladia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="BA4BD0620865FFE435A5F8DD5D8CF85B" blockId="8.[151,1437,1580,2038]" pageId="8" pageNumber="32">
<taxonomicName id="7DF4ABE10865FFE435A5F8DD5B9EF87F" authority="(Kussakin, 1969)" baseAuthorityName="Kussakin" baseAuthorityYear="1969" box="[199,670,1904,1930]" class="Malacostraca" family="Gnathiidae" genus="Elaphognathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="bacescoi">
<emphasis id="88800C700865FFE435A5F8DD58D6F87F" box="[199,470,1904,1930]" italics="true" pageId="8" pageNumber="32">Elaphognathia bacescoi</emphasis>
(
<bibRefCitation id="DE65AD930865FFE43487F8DD5B95F87F" author="Kussakin, O. G." box="[485,661,1904,1930]" pageId="8" pageNumber="32" pagination="408 - 439" refId="ref6776" refString="Kussakin, O. G. (1969) K Opredelitel fauni Cerngo i Azooskogo Morei. Isopoda. [Keys for the identification of fauna of the Black and Azov Seas. Isopoda]. Trudy Nauk Dumky, Kiev 3, 408 - 439." type="journal article" year="1969">Kussakin, 1969</bibRefCitation>
)
</taxonomicName>
from the Black Sea is also similar to the new species, but the former has a mandible with a bigger conical internal lobe, more setosed antenna and uropod rami.
</paragraph>
</subSubSection>
<subSubSection id="F2EE83E90865FFE435A5F81459F1F803" pageId="8" pageNumber="32" type="etymology">
<paragraph id="BA4BD0620865FFE435A5F81459F1F803" blockId="8.[151,1437,1580,2038]" pageId="8" pageNumber="32">
<emphasis id="88800C700865FFE435A5F814584EF826" bold="true" box="[199,334,1977,2003]" pageId="8" pageNumber="32">Etymology.</emphasis>
The name refers to the
<typeStatus id="654F6EC00865FFE43703F8145B91F826" box="[609,657,1977,2003]" pageId="8" pageNumber="32">type</typeStatus>
locality in the intertidal area of Al-Shumaimah at Khor Al-Subiyah,
<collectingCountry id="C2E390F20865FFE435F5F87159EDF803" box="[151,237,2012,2038]" name="Kuwait" pageId="8" pageNumber="32">Kuwait</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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