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<document ID-DOI="http://doi.org/10.5281/zenodo.5672841" ID-GBIF-Dataset="e7d164f6-b22c-4120-b1f6-6464cfc6de0b" ID-GBIF-Taxon="128090361" ID-ISSN="0005 6219" ID-Zenodo-Dep="5672841" checkinTime="1488976263354" checkinUser="plazi" docAuthor="Ruud, A. Bank, Henk P. M. G Menkhorst &amp; Eike Neubert" docDate="2016" docId="29708788FFD5FFC1FCE1FEA2FE1AF91A" docLanguage="en" docName="basteria.80.1-3.5.2016 Bank et al Lauriidae Enidae Vitrinidae Turkey.pdf" docOrigin="Basteria 80 (1)" docStyle="DocumentStyle{}" docTitle="Multidentula squalina subsp. acutior Lindholm 1922" docType="treatment" docVersion="10" lastPageId="20" lastPageNumber="23" masterDocId="D549FFF0FFC6FFD5FFDDFFA1FFAEFFF7" masterDocTitle="Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)" masterLastPageNumber="30" masterPageNumber="5" pageId="19" pageNumber="22" updateTime="1636598733005" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ruud, A. Bank</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Henk P. M. G Menkhorst</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eike Neubert</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Basteria</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>80</mods:number>
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<mods:number>1</mods:number>
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<mods:identifier type="DOI">10.5281/zenodo.439745</mods:identifier>
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<mods:identifier type="ISSN">00056219</mods:identifier>
<mods:identifier type="Zenodo-Dep">439745</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5672841" ID-GBIF-Taxon="128090361" ID-Zenodo-Dep="5672841" LSID="urn:lsid:plazi:treatment:29708788FFD5FFC1FCE1FEA2FE1AF91A" httpUri="http://treatment.plazi.org/id/29708788FFD5FFC1FCE1FEA2FE1AF91A" lastPageId="20" lastPageNumber="23" pageId="19" pageNumber="22">
<subSubSection box="[828,1432,259,283]" pageId="19" pageNumber="22" type="nomenclature">
<paragraph blockId="19.[828,1432,259,283]" box="[828,1432,259,283]" pageId="19" pageNumber="22">
<taxonomicName authority="Lindholm, 1922" authorityName="Lindholm" authorityYear="1922" box="[828,1335,259,283]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="acutior">
<emphasis box="[828,1137,259,283]" italics="true" pageId="19" pageNumber="22">Multidentula squalina acutior</emphasis>
(
<bibRefCitation author="LINDHOLM, W. A." box="[1151,1328,259,283]" journalOrPublisher="Annuaire de Musee zoologique de l'Academie des Sciences de Russie" pageId="19" pageNumber="22" pagination="273 - 275" part="23" refString="LINDHOLM, W. A., 1922 a. Description of two Bulimini (Gastropoda Pulmonata) from Russian Central Asia. - Annuaire de Musee zoologique de l'Academie des Sciences de Russie 23 (2): 273 - 275." title="Description of two Bulimini (Gastropoda Pulmonata) from Russian Central Asia" type="journal article" year="1922">Lindholm, 1922</bibRefCitation>
)
</taxonomicName>
(
<figureCitation box="[1349,1424,259,283]" captionStart="Figs 16 - 20" captionStartId="21.[151,191,1858,1877]" captionTargetBox="[151,1436,155,1833]" captionTargetId="figure@21.[151,1436,155,1833]" captionTargetPageId="21" captionText="Figs 16 - 20. Multidentula and Imparietula taxa from Turkey. 16, Multidentula squalina squalina (L. Pfeiffer, 1848), NMBE 548313 (H = 7.8 mm), Turkey, Vil. Karabük, eastern part Eskipazar, leg. Hausdorf, 27. ix. 1987. 17, Multidentula squalina eudoxina (Nägele, 1894), paralectotype SMF 14195 a coll. Kobelt ex Nägele (H = 6.3 mm), Turkey, Vil. Tokat, Tokat. 18, Multidentula squalina acutior (Lindholm, 1922), NMBE 544678 (H = 5.7 mm), Turkey, Vil. Artvin, N. Kınalıçam (= NNW. Cağliyan). 19, Multidentula reducta spec. nov., holotype NMBE 544681 (H = 6.2 mm), Turkey, Vil. Sivas, Zara, 16 km towards Imranlı. 20, Imparietula inflexa spec. nov., holotype NMBE 544683 (H = 12, 4 mm), Turkey, Vil. Giresun, 5 km N. Tamdere (= S. Yavuzkemal). All photographs Bochud &amp; Neubert, × 8." httpUri="https://zenodo.org/record/439752/files/figure.png" pageId="19" pageNumber="22">Fig. 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="22" type="reference_group">
<paragraph blockId="19.[824,1428,332,491]" pageId="19" pageNumber="22">
<taxonomicName authority="Lindholm, 1922b: 358" authorityName="Lindholm" authorityPageNumber="358" authorityYear="1922" box="[824,1191,332,352]" class="Gastropoda" family="Enidae" genus="Chondrula" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="acutior">
<emphasis box="[824,982,332,351]" italics="true" pageId="19" pageNumber="22">Chondrula acutior</emphasis>
<bibRefCitation author="LINDHOLM, W. A." box="[988,1149,332,352]" journalOrPublisher="Annuaire du Musee Zoologique de l'Academie des Sciences de Russie" pageId="19" pageNumber="22" pagination="355 - 361" part="23" refString="LINDHOLM, W. A., 1922 b. Beschreibung einiger kaukasischer Landschnecken (Gastropoda Pulmonata). - Annuaire du Musee Zoologique de l'Academie des Sciences de Russie 23 (3 / 4): 355 - 361." title="Beschreibung einiger kaukasischer Landschnecken (Gastropoda Pulmonata)" type="journal article" year="1922" yearSuffix="b">Lindholm, 1922b</bibRefCitation>
: 358
</taxonomicName>
-359. Type locality: &quot;Von mir im Genist des Fl. Tschoroch bei Kapandidi (unweit Batum) .... in einem tadellos erhaltenen leeren Stücke gefunden (coll. m.)&quot;. Holotype in ZIN (
<bibRefCitation author="SYSOEV, A. &amp; SCHILEYKO, A. A." bookContentInfo="142 plates" journalOrPublisher="Pensoft, Sofia / Moscow. [Pensoft Series Faunistica No 87]" pageId="19" pageNumber="22" pagination="1 - 312" refString="SYSOEV, A. &amp; SCHILEYKO, A. A., 2009. Land snails and slugs of Russia and adjacent countries: 1 - 312, 142 plates. Pensoft, Sofia / Moscow. [Pensoft Series Faunistica No 87]." title="Land snails and slugs of Russia and adjacent countries" type="book chapter" year="2009">Sysoev &amp; Schileyko, 2009: 85, 259, fig. 38E</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="22" type="description">
<paragraph blockId="19.[824,1430,537,2051]" pageId="19" pageNumber="22">Description. Shell dextral, oval in outline, with a rather wide, open, slit-like umbilicus. The 6.5-7.7</paragraph>
<paragraph blockId="19.[824,1430,537,2051]" pageId="19" pageNumber="22">whorls are rather convex with a rather deep suture. Teleoconch with irregular, oblique striae which are mostly rather fine; there are no spiral striae. Shell</paragraph>
<paragraph blockId="19.[824,1430,537,2051]" pageId="19" pageNumber="22">
solid, not or hardly translucent, rather dark brownish coloured, with a whitish band behind the peristome. The last whorl has an impression at the position of the infrapalatalis; the infrapalatalis and
<taxonomicName box="[1229,1323,814,838]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior are visible outside as blurred white stripes. Peristome somewhat reflected, thickened by a prominent labial callus, the columellar and palatal insertion connected by a clearly visible callus (but the callus is mostly not extra thickened near the insertions). The
<taxonomicName box="[1280,1425,987,1011]" class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="subangularis">subangularis</taxonomicName>
is well developed and fused with the palatal peristome. The
<taxonomicName box="[942,1087,1057,1081]" class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="subangularis">subangularis</taxonomicName>
is connected to the parietalis by a more or less thickened callus. The very prominent parietalis is curved and deeply recessed. A spiralis is missing. Columellaris perpendicular to the columellar peristome, deeply recessed. Basalis present, well developed. Infrapalatalis and
<taxonomicName box="[1260,1354,1230,1254]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior strongly developed; the infrapalatalis is the most prominent one. A suprapalatalis is also present, but is less prominent than the
<taxonomicName box="[1094,1188,1334,1358]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior. A weak suturalis is available.
</paragraph>
<paragraph blockId="19.[824,1430,537,2051]" pageId="19" pageNumber="22">Measurements (n = 13). H = 5.0-6.3 (mean 5.5); LWH = 2.6-3.1 (mean 2.9); MH = 1.7-2.1 (mean 1.8); LWD = 2.0-2.6 (mean 2.3); LWM = 2.3-2.9 (mean 2.6); MD = 1.5-1.8 (mean 1.6); NW = 6.5-7.7 (mean 7.0).</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="23" pageId="19" pageNumber="22" type="distribution">
<paragraph blockId="19.[824,1430,537,2051]" lastBlockId="20.[151,755,155,1774]" lastPageId="20" lastPageNumber="23" pageId="19" pageNumber="22">Localities. Vilayet Artvin: crossing Artvin/Şavşat/Demirkent, 17 km towards Yusufeli, suspension bridge Çoruh Nehri (41.0925°N 41.8146°E) (NMBE/2); 5 km N. Erkinis (= castle N. Demirkent) (40.9258°N 41.7619°E) (HMT/1); N. Kınalıçam (= NNW. Cağliyan) (40.7567°N 41.6449°E) (HMT/41; NMBE 544678/1); Yusufeli (40.8215°N 41.5434°E) (HMT/3); 16 km SE. Yusufeli (40.7826°N 41.6967°E) (HMT/14); 17 km SE. Yusufeli, 5 km N. province boundary Erzurum/Artvin (40.7361°N 41.6651°E) (HMT/47). Vilayet Erzurum: 6 km SE. Çamlıyamaç (40.5854°N 41.5952°E) (HMT/81); Ispir, 38 km towards Yusufeli (40.6517°N 41.2299°E) (NMBE/4); deposition Tortum Gölü (40.6361°N 41.6312°E) (HMT/59); 9 km SW. Uzundere, 2 km S. exit to Dikmen (40.4698°N 41.5019°E) (HMT/117). Vilayet Gümüşhane: 6 km N. Arpalı (= 22 km NW. Bayburt) (40.5792°N 39.1334°E) (HMT/14); Salyazı, deposition Sulama Göletı (40.2446°N 39.8070°E) (HMT/29).</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="23" type="diagnosis">
<paragraph blockId="20.[151,755,155,1774]" pageId="20" pageNumber="23">
Differentiation.
<taxonomicName box="[386,639,294,318]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="squalina">
<emphasis box="[386,639,294,318]" italics="true" pageId="20" pageNumber="23">Multidentula s. squalina</emphasis>
</taxonomicName>
(
<figureCitation box="[653,728,294,318]" captionStart="Figs 16 - 20" captionStartId="21.[151,191,1858,1877]" captionTargetBox="[151,1436,155,1833]" captionTargetId="figure@21.[151,1436,155,1833]" captionTargetPageId="21" captionText="Figs 16 - 20. Multidentula and Imparietula taxa from Turkey. 16, Multidentula squalina squalina (L. Pfeiffer, 1848), NMBE 548313 (H = 7.8 mm), Turkey, Vil. Karabük, eastern part Eskipazar, leg. Hausdorf, 27. ix. 1987. 17, Multidentula squalina eudoxina (Nägele, 1894), paralectotype SMF 14195 a coll. Kobelt ex Nägele (H = 6.3 mm), Turkey, Vil. Tokat, Tokat. 18, Multidentula squalina acutior (Lindholm, 1922), NMBE 544678 (H = 5.7 mm), Turkey, Vil. Artvin, N. Kınalıçam (= NNW. Cağliyan). 19, Multidentula reducta spec. nov., holotype NMBE 544681 (H = 6.2 mm), Turkey, Vil. Sivas, Zara, 16 km towards Imranlı. 20, Imparietula inflexa spec. nov., holotype NMBE 544683 (H = 12, 4 mm), Turkey, Vil. Giresun, 5 km N. Tamdere (= S. Yavuzkemal). All photographs Bochud &amp; Neubert, × 8." httpUri="https://zenodo.org/record/439752/files/figure.png" pageId="20" pageNumber="23">Fig. 16</figureCitation>
) differs from
<taxonomicName box="[291,422,329,352]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="acutior">
<emphasis box="[291,422,329,352]" italics="true" pageId="20" pageNumber="23">M. s. acutior</emphasis>
</taxonomicName>
by the more densily packed and more pronounced oblique striae, the more pointed apex, the wider umbilicus, the mostly absent suturalis, the more reflected peristome, the less slen- der suprapalatalis, and the
<taxonomicName box="[456,550,467,491]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior and infrapalatalis are generally more deeply situated in the aperture (i.e. they are situated further away from the peristome).
<taxonomicName box="[284,544,571,595]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="eudoxina">
<emphasis box="[284,544,571,595]" italics="true" pageId="20" pageNumber="23">Multidentula s. eudoxina</emphasis>
</taxonomicName>
(
<figureCitation box="[558,633,571,595]" captionStart="Figs 16 - 20" captionStartId="21.[151,191,1858,1877]" captionTargetBox="[151,1436,155,1833]" captionTargetId="figure@21.[151,1436,155,1833]" captionTargetPageId="21" captionText="Figs 16 - 20. Multidentula and Imparietula taxa from Turkey. 16, Multidentula squalina squalina (L. Pfeiffer, 1848), NMBE 548313 (H = 7.8 mm), Turkey, Vil. Karabük, eastern part Eskipazar, leg. Hausdorf, 27. ix. 1987. 17, Multidentula squalina eudoxina (Nägele, 1894), paralectotype SMF 14195 a coll. Kobelt ex Nägele (H = 6.3 mm), Turkey, Vil. Tokat, Tokat. 18, Multidentula squalina acutior (Lindholm, 1922), NMBE 544678 (H = 5.7 mm), Turkey, Vil. Artvin, N. Kınalıçam (= NNW. Cağliyan). 19, Multidentula reducta spec. nov., holotype NMBE 544681 (H = 6.2 mm), Turkey, Vil. Sivas, Zara, 16 km towards Imranlı. 20, Imparietula inflexa spec. nov., holotype NMBE 544683 (H = 12, 4 mm), Turkey, Vil. Giresun, 5 km N. Tamdere (= S. Yavuzkemal). All photographs Bochud &amp; Neubert, × 8." httpUri="https://zenodo.org/record/439752/files/figure.png" pageId="20" pageNumber="23">Fig. 17</figureCitation>
) has a different dentition compared to
<taxonomicName box="[479,610,607,630]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="acutior">
<emphasis box="[479,610,607,630]" italics="true" pageId="20" pageNumber="23">M. s. acutior</emphasis>
</taxonomicName>
. The former lacks a suprapalatalis, the infrapalatalis is rather broad and seems slightly bifid, the
<taxonomicName box="[544,638,675,699]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior is more prominent and the angularis is not fused with the palatal peristome, but instead connected to it by a thin callus.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="23" type="discussion">
<paragraph blockId="20.[151,755,155,1774]" pageId="20" pageNumber="23">
Remarks. The type locality &quot;Kapandidi“ of
<taxonomicName class="Gastropoda" family="Enidae" genus="Chondrula" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="acutior">
<emphasis italics="true" pageId="20" pageNumber="23">acutior</emphasis>
</taxonomicName>
could not be traced. It is likely that Maradidi is meant, which is located in Georgia along the river Chorochi (= Çoruh Nehri) a few km north of the Turkish border (41.5030N 41.7123E).
</paragraph>
<paragraph blockId="20.[151,755,155,1774]" pageId="20" pageNumber="23">
An isolated locality from
<taxonomicName box="[473,596,987,1011]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="squalina">
<emphasis box="[473,596,987,1011]" italics="true" pageId="20" pageNumber="23">M. squalina</emphasis>
</taxonomicName>
has been reported by
<bibRefCitation author="FEHER, Z. &amp; EROSS, Z. P." box="[267,695,1022,1046]" journalOrPublisher="Schriften zur Malakozoologie" pageId="20" pageNumber="23" pagination="3 - 21" part="25" refString="FEHER, Z. &amp; EROSS, Z. P., 2009. Contribution to the Mollusca fauna of Albania. Results of the field trips of the Hungarian Natural History Museum between 1992 and 2007. - Schriften zur Malakozoologie 25: 3 - 21." title="Contribution to the Mollusca fauna of Albania. Results of the field trips of the Hungarian Natural History Museum between 1992 and 2007" type="journal article" year="2009">Fehér &amp; Erőss (2009: 12, pl. 1 figs 8-10)</bibRefCitation>
from Albania. All other localities of
<taxonomicName box="[551,673,1056,1080]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="squalina">
<emphasis box="[551,673,1056,1080]" italics="true" pageId="20" pageNumber="23">M. squalina</emphasis>
</taxonomicName>
are restricted to Turkey (except one: the type locality of
<taxonomicName class="Gastropoda" family="Enidae" genus="Chondrula" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="acutior">
<emphasis italics="true" pageId="20" pageNumber="23">acutior</emphasis>
</taxonomicName>
). The shells of Albania (as interpreted from the mentioned figures by Fehér &amp; Erőss as well as those published by Welter-Schultes [2012: 194]) do not belong to the nominotypical subspecies, as the
<taxonomicName class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="subangularis">subangularis</taxonomicName>
is not fused with the palatal peristome, the
</paragraph>
<paragraph blockId="20.[151,755,155,1774]" pageId="20" pageNumber="23">
presence of a suturalis, the oblique striae are less pronounced, the whorls are more convex, and the apex is less pointed. Furthermore, in
<taxonomicName box="[482,627,1368,1392]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="squalina">
<emphasis box="[482,627,1368,1392]" italics="true" pageId="20" pageNumber="23">M. s. squalina</emphasis>
</taxonomicName>
the basalis is shifted to a more subcolumellar position, and the infrapalatalis and
<taxonomicName box="[354,448,1438,1462]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior are more deeply situated in the aperture. The Albanian subspecies differs from
<taxonomicName box="[260,391,1508,1531]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="subSpecies" species="squalina" subSpecies="acutior">
<emphasis box="[260,391,1508,1531]" italics="true" pageId="20" pageNumber="23">M. s. acutior</emphasis>
</taxonomicName>
by the less pronounced
<taxonomicName class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="subangularis">subangularis</taxonomicName>
which is non-fused.
</paragraph>
<paragraph blockId="20.[151,755,155,1774]" pageId="20" pageNumber="23">
Another isolated locality of
<taxonomicName box="[500,622,1576,1600]" class="Gastropoda" family="Enidae" genus="Multidentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="23" phylum="Mollusca" rank="species" species="squalina">
<emphasis box="[500,622,1576,1600]" italics="true" pageId="20" pageNumber="23">M. squalina</emphasis>
</taxonomicName>
is within Turkey itself, namely Çivril and Senirkent (both in the Göller Bölgesi area) (
<bibRefCitation author="YILDIRIM, M. Z. &amp; SCHUTT, H." box="[426,691,1646,1670]" journalOrPublisher="Club Conchylia Informationen" pageId="20" pageNumber="23" pagination="47 - 55" part="29" refString="YILDIRIM, M. Z. &amp; SCHUTT, H., 1997. Verbreitung der Enidae im Seengebiet SW-Anatoliens (Gastropoda Pulmonata: Buliminoidea). - Club Conchylia Informationen 29 (1 / 2): 47 - 55." title="Verbreitung der Enidae im Seengebiet SW-Anatoliens (Gastropoda Pulmonata: Buliminoidea)" type="journal article" year="1997">Yıldırım &amp; Schütt, 1997</bibRefCitation>
: 52, fig. 16;
<bibRefCitation author="SCHUTT, H. &amp; SESEN, R." box="[232,369,1680,1704]" journalOrPublisher="Turkish Journal of Zoology" pageId="20" pageNumber="23" pagination="427 - 429" part="25" refString="SCHUTT, H. &amp; SESEN, R., 2001. Rediscovery of Sesteria gallandi Bourguignat, 1884 (Mollusca, Pulmonata, Buliminoidea, Enidae). - Turkish Journal of Zoology 25 (4): 427 - 429." title="Rediscovery of Sesteria gallandi Bourguignat, 1884 (Mollusca, Pulmonata, Buliminoidea, Enidae)" type="journal article" year="2001">Schütt, 2001</bibRefCitation>
: 148, fig. c;
<bibRefCitation author="SCHUTT, H." box="[496,634,1680,1704]" journalOrPublisher="Verlag Natur &amp; Wissenschaft, Solingen" pageId="20" pageNumber="23" pagination="1 - 559" refString="SCHUTT, H., 2010. Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos: 1 - 559. Verlag Natur &amp; Wissenschaft, Solingen." title="Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos" type="book chapter" year="2010">Schütt, 2010</bibRefCitation>
: 146, fig. c). Unfortunately, the figures are too vague to determine its subspecific state.
</paragraph>
</subSubSection>
</treatment>
</document>