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<document ID-DOI="http://doi.org/10.5281/zenodo.5672811" ID-GBIF-Dataset="e7d164f6-b22c-4120-b1f6-6464cfc6de0b" ID-GBIF-Taxon="128090371" ID-ISSN="0005 6219" ID-Zenodo-Dep="5672811" checkinTime="1488976263354" checkinUser="plazi" docAuthor="Ruud, A. Bank, Henk P. M. G Menkhorst &amp; Eike Neubert" docDate="2016" docId="29708788FFCFFFDCFF72FE57FC1FFD4C" docLanguage="en" docName="basteria.80.1-3.5.2016 Bank et al Lauriidae Enidae Vitrinidae Turkey.pdf" docOrigin="Basteria 80 (1)" docStyle="DocumentStyle{}" docTitle="Turanena (Turanena) elegantula Ruud, Menkhorst &amp; Neubert, 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="12" masterDocId="D549FFF0FFC6FFD5FFDDFFA1FFAEFFF7" masterDocTitle="Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)" masterLastPageNumber="30" masterPageNumber="5" pageId="9" pageNumber="12" updateTime="1636598733005" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ruud, A. Bank</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Henk P. M. G Menkhorst</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eike Neubert</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Basteria</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>5</mods:start>
<mods:end>30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.439745</mods:identifier>
<mods:identifier type="GBIF-Dataset">e7d164f6-b22c-4120-b1f6-6464cfc6de0b</mods:identifier>
<mods:identifier type="ISSN">00056219</mods:identifier>
<mods:identifier type="Zenodo-Dep">439745</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5672811" ID-GBIF-Taxon="128090371" ID-Zenodo-Dep="5672811" LSID="urn:lsid:plazi:treatment:29708788FFCFFFDCFF72FE57FC1FFD4C" httpUri="http://treatment.plazi.org/id/29708788FFCFFFDCFF72FE57FC1FFD4C" lastPageNumber="12" pageId="9" pageNumber="12">
<subSubSection box="[175,739,502,526]" pageId="9" pageNumber="12" type="nomenclature">
<paragraph blockId="9.[175,739,502,526]" box="[175,739,502,526]" pageId="9" pageNumber="12">
<heading bold="true" box="[175,739,502,526]" centered="true" fontSize="10" level="4" pageId="9" pageNumber="12" reason="0">
<emphasis bold="true" box="[175,739,502,526]" pageId="9" pageNumber="12">
<taxonomicName box="[175,534,502,526]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="elegantula" status="spec. nov." subGenus="Turanena">
<emphasis bold="true" box="[175,280,502,525]" italics="true" pageId="9" pageNumber="12">Turanena</emphasis>
(
<emphasis bold="true" box="[296,401,502,525]" italics="true" pageId="9" pageNumber="12">Turanena</emphasis>
)
<emphasis bold="true" box="[416,534,502,526]" italics="true" pageId="9" pageNumber="12">elegantula</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[540,654,502,526]" pageId="9" pageNumber="12" rank="species">spec. nov.</taxonomicNameLabel>
(
<figureCitation box="[667,731,502,526]" captionStart="Figs 5 - 7" captionStartId="8.[151,191,1684,1703]" captionTargetBox="[151,1436,155,1649]" captionTargetId="figure@8.[151,1436,155,1649]" captionTargetPageId="8" captionText="Figs 5 - 7. Turanena taxa from Turkey. 5, Turanena (Turanena) elegantula spec. nov., holotype NMBE 544664 (H = 9.0 mm), Turkey, Vil. Van, 4 km SE. Göründü. 6, Turanena (Turanena) zilchi E. Gittenberger &amp; Menkhorst 1993, NMBE 546171 (H = 6.1 mm), Turkey, Vil. Hakkâri, 7 km S. Hakkâri. 7, Turanena (Turanena) andronakii salpinx subspec. nov., holotype NMBE 546173 (H = 8.1 mm), Turkey, Vil. Artvin, 1 km S. Oruçlu. All photographs Bochud &amp; Neubert, × 8." httpUri="https://zenodo.org/record/439748/files/figure.png" pageId="9" pageNumber="12">Fig. 5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="materials_examined">
<paragraph blockId="9.[151,753,575,699]" pageId="9" pageNumber="12">
Type locality &amp; type specimens.
<materialsCitation ID-GBIF-Occurrence="1455386809" collectingDate="1992-08-21" collectionCode="NMBE" collectorName="H. P. M. G. Menkhorst" country="Turkey" elevation="1650" latitude="38.3271" location="Vilayet Van" longLatPrecision="6" longitude="42.9331" pageId="9" pageNumber="12" specimenCode="NMBE 544664" specimenCount="1" stateProvince="Vilayet Van" typeStatus="holotype">
<collectingCountry box="[473,540,575,595]" name="Turkey" pageId="9" pageNumber="12">Turkey</collectingCountry>
,
<collectingRegion box="[550,659,575,594]" pageId="9" pageNumber="12">Vilayet Van</collectingRegion>
,
<locationDeviation pageId="9" pageNumber="12">
<quantity box="[670,717,575,594]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="12" unit="km" value="4.0">4 km</quantity>
SE. Göründü
</locationDeviation>
,
<quantity box="[249,316,610,629]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="12" unit="m" value="1650.0">
<elevation box="[249,316,610,629]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="12" unit="m" value="1650.0">1650 m</elevation>
</quantity>
(
<geoCoordinate box="[327,424,610,629]" direction="north" orientation="latitude" pageId="9" pageNumber="12" precision="5" value="38.3271">38.3271°N</geoCoordinate>
<geoCoordinate box="[429,521,610,630]" direction="east" orientation="longitude" pageId="9" pageNumber="12" precision="5" value="42.9331">42.9331°E</geoCoordinate>
),
<collectorName box="[538,735,610,629]" pageId="9" pageNumber="12">H.P.M.G. Menkhorst</collectorName>
leg.,
<date box="[195,303,644,663]" pageId="9" pageNumber="12" value="1992-08-21">
<collectingDate box="[195,303,644,663]" pageId="9" pageNumber="12" value="1992-08-21">21.viii.1992</collectingDate>
</date>
.
<typeStatus box="[311,400,644,663]" pageId="9" pageNumber="12">Holotype</typeStatus>
<specimenCode box="[406,539,644,664]" pageId="9" pageNumber="12">NMBE 544664</specimenCode>
</materialsCitation>
, paratypes NMBE 544665/1, HMK/&gt;50, RBA/2, RMNH/2, ZMH/2.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="diagnosis">
<paragraph blockId="9.[151,764,745,2051]" pageId="9" pageNumber="12">
Diagnosis. A slender, spindle-shaped light hornyyellowish coloured
<taxonomicName box="[371,469,780,803]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis box="[371,469,780,803]" italics="true" pageId="9" pageNumber="12">Turanena</emphasis>
</taxonomicName>
species with a well developed parietal callus, a slightly thickened, not reflected, peristome, and a juxtaposed columellar and palatal insertion of the peristome.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="description">
<paragraph blockId="9.[151,764,745,2051]" pageId="9" pageNumber="12">
Description. Shell dextral, elongated spindle-like in outline, with an open, relatively wide, slit-like umbilicus. The 6.5-7.6 whorls are quite convex with a deep suture. Teleoconch with irregular, fine, oblique striae; there are no spiral striae. Shell moderately solid, somewhat translucent, glossy, yellowish to horny-yellowish coloured; there is no whitish band behind the peristome. Aperture elliptical-rounded, peristome not reflected (only slightly at the columellar part) and only slightly thickened. Columellar and palatal insertion connected by a well developed callus (which is often thickened near its ends).
<taxonomicName class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="subangularis">A subangularis</taxonomicName>
is not present (the thickened parietal callus does not form
<taxonomicName box="[255,419,1369,1393]" class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="subangularis">a subangularis</taxonomicName>
but normally fuses with the palatal insertion of the peristome). The somewhat oblique columellar ledge reaches halfway to above the middle of the columellar side of the aperture.
</paragraph>
<paragraph blockId="9.[151,764,745,2051]" pageId="9" pageNumber="12">Measurements (n = 6). H = 7.3-9.1 (mean 8.4); LWH = 4.1-4.7 (mean 4.4); MH = 2.2-2.7 (mean 2.4); LWD = 2.9-3.1 (mean 3.0); LWM = 3.0-3.4 (mean 3.2); MD = 1.7-1.9 (mean 1.8); NW = 6.5-7.6 (mean 7.1).</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="distribution">
<paragraph blockId="9.[151,764,745,2051]" pageId="9" pageNumber="12">Localities. Known from the locus typicus only (see above).</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="etymology">
<paragraph blockId="9.[151,764,745,2051]" pageId="9" pageNumber="12">
Derivatio nominis. Derived from
<emphasis box="[543,618,1715,1739]" italics="true" pageId="9" pageNumber="12">elegans</emphasis>
(graceful, elegant), which relates to the slenderness of the shell.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="diagnosis">
<paragraph blockId="9.[151,764,745,2051]" pageId="9" pageNumber="12">
Differentiation.
<taxonomicName box="[386,596,1784,1808]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="elegantula">
<emphasis box="[386,596,1784,1808]" italics="true" pageId="9" pageNumber="12">Turanena elegantula</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="cochlicopoides">
<emphasis italics="true" pageId="9" pageNumber="12">T. cochlicopoides</emphasis>
</taxonomicName>
by its larger size, the more elongated shape, the higher number of whorls, the more prominent parietal callus, and the more clearly thickened peristome.
<taxonomicName box="[276,424,1923,1947]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="conelongata">
<emphasis box="[276,424,1923,1947]" italics="true" pageId="9" pageNumber="12">T. conelongata</emphasis>
</taxonomicName>
is less slender, the whorls are more convex, the peristome is sharp, the parietal callus is weaker, and the peristome is clearly more reflected at its columellar insertion.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="12" type="discussion">
<paragraph blockId="9.[824,1432,155,699]" pageId="9" pageNumber="12">
Remarks. Only a single locality is known for
<taxonomicName class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="elegantula">
<emphasis italics="true" pageId="9" pageNumber="12">T. elegantula</emphasis>
</taxonomicName>
, which is located at the southern shore of Van Gölü. From the same area,
<taxonomicName box="[1125,1273,224,248]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="conelongata">
<emphasis box="[1125,1273,224,248]" italics="true" pageId="9" pageNumber="12">T. conelongata</emphasis>
</taxonomicName>
and
<taxonomicName box="[1329,1411,224,248]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="zilchi">
<emphasis box="[1329,1411,224,248]" italics="true" pageId="9" pageNumber="12">T. zilchi</emphasis>
</taxonomicName>
has been described, originally also from a single locality. However, the latter two species have now been recorded from a wider area:
<taxonomicName box="[1143,1291,328,352]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="conelongata">
<emphasis box="[1143,1291,328,352]" italics="true" pageId="9" pageNumber="12">T. conelongata</emphasis>
</taxonomicName>
from Kozluk (vil. Batman), Baykan (vil. Siirt) and Tatvan (vil. Bitlis) (
<bibRefCitation author="SCHUTT, H." box="[902,1175,398,422]" journalOrPublisher="Verlag Natur &amp; Wissenschaft, Solingen" pageId="9" pageNumber="12" pagination="1 - 559" refString="SCHUTT, H., 2010. Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos: 1 - 559. Verlag Natur &amp; Wissenschaft, Solingen." title="Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos" type="book chapter" year="2010" yearSuffix="e">Schütt, 2010: 87, 89 fig. e</bibRefCitation>
), and
<taxonomicName box="[1245,1327,398,422]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="zilchi">
<emphasis box="[1245,1327,398,422]" italics="true" pageId="9" pageNumber="12">T. zilchi</emphasis>
</taxonomicName>
from Baykan and Siirt (vil. Siirt), as well as Silvan, Ergani, Eğil, Çatakköprü and Kulp (vil. Diyarbakır) (
<bibRefCitation author="SESEN, R. &amp; SCHUTT, H." journalOrPublisher="Zoology in the Middle East" pageId="9" pageNumber="12" pagination="77 - 84" part="20" refString="SESEN, R. &amp; SCHUTT, H., 2000. Land snails from Sason Mountains in Siirt province in eastern Anatolia. - Zoology in the Middle East 20: 77 - 84." title="Land snails from Sason Mountains in Siirt province in eastern Anatolia" type="journal article" year="2000">Şeşen &amp; Schütt, 2000</bibRefCitation>
: 80, fig. 8;
<bibRefCitation author="SCHUTT, H." box="[1077,1214,502,526]" journalOrPublisher="Verlag Natur &amp; Wissenschaft, Solingen" pageId="9" pageNumber="12" pagination="1 - 559" refString="SCHUTT, H., 2010. Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos: 1 - 559. Verlag Natur &amp; Wissenschaft, Solingen." title="Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos" type="book chapter" year="2010">Schütt, 2010</bibRefCitation>
: 88, 89 figs j+k). We can now add another record of
<taxonomicName box="[1175,1257,536,560]" class="Gastropoda" family="Enidae" genus="Turanena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="zilchi">
<emphasis box="[1175,1257,536,560]" italics="true" pageId="9" pageNumber="12">T. zilchi</emphasis>
</taxonomicName>
, namely Vilayet Hakkâri, 7 km S. Hakkâri, along the road D400, 1230 m. We here provide a picture (
<figureCitation box="[1220,1282,605,630]" captionStart="Figs 5 - 7" captionStartId="8.[151,191,1684,1703]" captionTargetBox="[151,1436,155,1649]" captionTargetId="figure@8.[151,1436,155,1649]" captionTargetPageId="8" captionText="Figs 5 - 7. Turanena taxa from Turkey. 5, Turanena (Turanena) elegantula spec. nov., holotype NMBE 544664 (H = 9.0 mm), Turkey, Vil. Van, 4 km SE. Göründü. 6, Turanena (Turanena) zilchi E. Gittenberger &amp; Menkhorst 1993, NMBE 546171 (H = 6.1 mm), Turkey, Vil. Hakkâri, 7 km S. Hakkâri. 7, Turanena (Turanena) andronakii salpinx subspec. nov., holotype NMBE 546173 (H = 8.1 mm), Turkey, Vil. Artvin, 1 km S. Oruçlu. All photographs Bochud &amp; Neubert, × 8." httpUri="https://zenodo.org/record/439748/files/figure.png" pageId="9" pageNumber="12">Fig. 6</figureCitation>
) of this littleknown species, so as to become acquainted with its variability.
</paragraph>
</subSubSection>
</treatment>
</document>