treatments-xml/data/29/70/87/29708788FFC2FFD0FF78F959FE20FE96.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.5672801" ID-GBIF-Dataset="e7d164f6-b22c-4120-b1f6-6464cfc6de0b" ID-GBIF-Taxon="128090357" ID-ISSN="0005 6219" ID-Zenodo-Dep="5672801" checkinTime="1488976263354" checkinUser="plazi" docAuthor="Ruud, A. Bank, Henk P. M. G Menkhorst &amp; Eike Neubert" docDate="2016" docId="29708788FFC2FFD0FF78F959FE20FE96" docLanguage="en" docName="basteria.80.1-3.5.2016 Bank et al Lauriidae Enidae Vitrinidae Turkey.pdf" docOrigin="Basteria 80 (1)" docStyle="DocumentStyle{}" docTitle="Leiostyla paphlagonica subsp. subangulosa Ruud, Menkhorst &amp; Neubert, 2016, subspec. nov." docType="treatment" docVersion="10" lastPageId="5" lastPageNumber="7" masterDocId="D549FFF0FFC6FFD5FFDDFFA1FFAEFFF7" masterDocTitle="Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)" masterLastPageNumber="30" masterPageNumber="5" pageId="4" pageNumber="7" updateTime="1636598733005" updateUser="ExternalLinkService">
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<mods:title>Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ruud, A. Bank</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Henk P. M. G Menkhorst</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eike Neubert</mods:namePart>
</mods:name>
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<mods:titleInfo>
<mods:title>Basteria</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>80</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:identifier type="DOI">10.5281/zenodo.439745</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5672801" ID-GBIF-Taxon="128090357" ID-Zenodo-Dep="5672801" LSID="urn:lsid:plazi:treatment:29708788FFC2FFD0FF78F959FE20FE96" httpUri="http://treatment.plazi.org/id/29708788FFC2FFD0FF78F959FE20FE96" lastPageId="5" lastPageNumber="7" pageId="4" pageNumber="7">
<subSubSection pageId="4" pageNumber="7" type="nomenclature">
<paragraph blockId="4.[165,741,1784,1843]" pageId="4" pageNumber="7">
<heading bold="true" centered="true" fontSize="10" level="4" pageId="4" pageNumber="7" reason="0">
<emphasis bold="true" box="[165,741,1784,1809]" pageId="4" pageNumber="7">
<taxonomicName box="[165,580,1784,1808]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="subSpecies" species="paphlagonica" status="subspec. nov." subSpecies="subangulosa">
<emphasis bold="true" box="[165,580,1784,1808]" italics="true" pageId="4" pageNumber="7">Leiostyla paphlagonica subangulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[586,741,1785,1809]" pageId="4" pageNumber="7" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
<emphasis bold="true" box="[414,493,1819,1843]" pageId="4" pageNumber="7">
(
<figureCitation box="[421,485,1819,1843]" captionStart="Figs 1 - 3" captionStartId="3.[151,191,1164,1183]" captionTargetBox="[151,1436,155,1114]" captionTargetId="figure@3.[151,1436,155,1114]" captionTargetPageId="3" captionText="Figs 1 - 3. Leiostyla taxa from Turkey. 1, Leiostyla paphlagonica paphlagonica Hausdorf 1990, holotype SMF 307492 (4.0 mm), Turkey, Vil. Kastamonu, Kapısuyu. 2, Leiostyla paphlagonica subangulosa subspec. nov., holotype NMBE 544659 (H = 3.7 mm), Turkey, Vil. Amasya, 10.3 km NE. Amasya. 3, Leiostyla eikenboomi spec. nov., holotype NMBE 544682 (H = 3.4 mm), Turkey, Vil. Karabük, Suçatı Tüneli 9 km ESE. Yenice. All photographs Bochud &amp; Neubert, × 15." httpUri="https://zenodo.org/record/439746/files/figure.png" pageId="4" pageNumber="7">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="7" type="materials_examined">
<paragraph blockId="4.[151,746,1892,2051]" pageId="4" pageNumber="7">
Type locality &amp; type specimens. – 
<materialsCitation ID-GBIF-Occurrence="1455386815" collectingDate="1992-08-03" collectionCode="NMBE" collectorName="H. P. M. G. Menkhorst" country="Turkey" elevation="380" latitude="40.7018" location="10.3 km NE. Amasya (in the direction of Yenice / Direkli" longLatPrecision="6" longitude="35.9608" pageId="4" pageNumber="7" specimenCode="NMBE 544656" specimenCount="1" stateProvince="Vilayet Amasya" typeStatus="holotype">
<collectingCountry box="[473,540,1892,1912]" name="Turkey" pageId="4" pageNumber="7">Turkey</collectingCountry>
, 
<collectingRegion box="[550,699,1892,1913]" pageId="4" pageNumber="7">Vilayet Amasya</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:29708788FFC2FFD0FF78F959FE20FE96:A4066045FFC2FFD1FD18F8C4FDDFF86C" country="Turkey" latitude="40.7018" longLatPrecision="6" longitude="35.9608" name="10.3 km NE. Amasya (in the direction of Yenice / Direkli" pageId="4" pageNumber="7" stateProvince="Vilayet Amasya">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="7" unit="km" value="10.3">10.3 km</quantity>
NE. Amasya (in the direction of Yenice/Direkli
</location>
), 
<elevation box="[641,698,1927,1946]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="7" unit="m" value="380.0">
<quantity box="[641,698,1927,1946]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="7" unit="m" value="380.0">380 m</quantity>
</elevation>
(
<geoCoordinate box="[157,253,1962,1981]" direction="north" orientation="latitude" pageId="4" pageNumber="7" precision="5" value="40.7018">40.7018°N</geoCoordinate>
<geoCoordinate box="[259,351,1962,1982]" direction="east" orientation="longitude" pageId="4" pageNumber="7" precision="5" value="35.9608">35.9608°E</geoCoordinate>
), 
<collectorName box="[368,564,1962,1981]" pageId="4" pageNumber="7">H.P.M.G. Menkhorst</collectorName>
leg., 
<date box="[614,711,1962,1981]" pageId="4" pageNumber="7" value="1992-08-03">
<collectingDate box="[614,711,1962,1981]" pageId="4" pageNumber="7" value="1992-08-03">3.viii.1992</collectingDate>
</date>
. 
<typeStatus box="[151,240,1996,2015]" pageId="4" pageNumber="7">Holotype</typeStatus>
<specimenCode box="[245,379,1996,2016]" pageId="4" pageNumber="7">NMBE 544656</specimenCode>
</materialsCitation>
, paratypes NMBE 544657/11, HMK/&gt;50, RBA/3, RMNH/3, ZMH/3.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="7" type="diagnosis">
<paragraph blockId="4.[831,1435,155,2051]" pageId="4" pageNumber="7">
Diagnosis. – A subspecies of 
<taxonomicName box="[1154,1314,155,179]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="paphlagonica">
<emphasis box="[1154,1314,155,179]" italics="true" pageId="4" pageNumber="7">L. paphlagonica</emphasis>
</taxonomicName>
with a basal keel, a small 
<taxonomicName box="[1039,1184,190,214]" class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="subangularis">subangularis</taxonomicName>
or, alternatively, a well developed parietal callus between the angular lamella and the palatal insertion of the peristome.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="7" type="description">
<paragraph blockId="4.[831,1435,155,2051]" pageId="4" pageNumber="7">
Description. – Shell dextral, elongated oviform in outline with conical spire, with closely arranged, distinctly and rather regularly, oblique striae; there are no spiral striae. The 7.4-7.8 whorls are convex with a deep suture. Shell rather solid, somewhat translucent, dark horn-coloured. The last whorl has a lengthy impression below the 
<taxonomicName box="[1049,1143,502,526]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
inferior on the outer wall, resulting in a distinct basal keel. Umbilicus open, deep and narrow. Peristome strongly reflected at right angles to form a flat, thickened, whitish lip. There is a marked thickening below the parieto- palatal angle of the peristome. Columellar and palatal insertion of the peristome connected by a clearly visible parietal callus, which slightly thickens near its ends. Angular lamella prominent, high, without appendages; it does not reach the border of the parietal callus. The angular lamella is fused with the palatal insertion of the peristome by a well developed callus; alternatively, a small subangular thickening is present that fuses on one side with the palatal insertion and on the other side with the angular lamella. Parietalis rather weak, deep inside the aperture; it is not connected with the angular lamella. 
<taxonomicName box="[1238,1332,1056,1080]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="23" phylum="Mollusca" rank="species" species="palatalis">Palatalis</taxonomicName>
inferior very long and well developed; it does not reach the lip. The columellaris is horizontally projected at or above the middle of the columellar side of the aperture.
</paragraph>
<paragraph blockId="4.[831,1435,155,2051]" pageId="4" pageNumber="7">Measurements (n = 6). – H = 3.6-4.0 (mean 3.8); LWH = 1.8-1.9 (mean 1.9); MH = 1.2-1.4 (mean 1.3); LWD = 1.7-1.8 (mean 1.8); LWM = 1.8-1.9 (mean 1.8); MD = 1.1-1.2 (mean 1.1); NW = 7.4-7.8 (mean 7.6).</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="7" type="distribution">
<paragraph blockId="4.[831,1435,155,2051]" pageId="4" pageNumber="7">Localities. – Known from the locus typicus only (see above).</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="7" type="etymology">
<paragraph blockId="4.[831,1435,155,2051]" pageId="4" pageNumber="7">
Derivatio nominis. – Named after the presence of the 
<taxonomicName box="[873,1018,1472,1496]" class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="subangularis">subangularis</taxonomicName>
, which is missing in 
<taxonomicName authority="Hausdorf, 1990" authorityName="Hausdorf" authorityYear="1990" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="subSpecies" species="paphlagonica" subSpecies="incisa">
<emphasis italics="true" pageId="4" pageNumber="7">L. paphlagonica incisa</emphasis>
<bibRefCitation author="HAUSDORF, B." box="[897,1071,1507,1531]" journalOrPublisher="Archiv fur Molluskenkunde 120 (1 / 3)" pageId="4" pageNumber="7" pagination="47 - 56" part="1989" refString="HAUSDORF, B., 1990. Zur Kenntnis einiger Arten der Gattung Leiostyla Lowe aus NW-Anatolien (Gastropoda: Lauriinae). - Archiv fur Molluskenkunde 120 (1 / 3) [1989]: 47 - 56." title="Zur Kenntnis einiger Arten der Gattung Leiostyla Lowe aus NW-Anatolien (Gastropoda: Lauriinae)" type="journal article" year="1990">Hausdorf, 1990</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="7" type="diagnosis">
<paragraph blockId="4.[831,1435,155,2051]" pageId="4" pageNumber="7">
Differentiation. – 
<taxonomicName box="[1066,1316,1542,1566]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="subSpecies" species="paphlagonica" subSpecies="subangulosa">
<emphasis box="[1066,1316,1542,1566]" italics="true" pageId="4" pageNumber="7">Leiostyla p. subangulosa</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[892,1078,1576,1600]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="subSpecies" species="paphlagonica" subSpecies="paphlagonica">
<emphasis box="[892,1078,1576,1600]" italics="true" pageId="4" pageNumber="7">L. p. paphlagonica</emphasis>
</taxonomicName>
(
<figureCitation box="[1093,1152,1576,1600]" captionStart="Figs 1 - 3" captionStartId="3.[151,191,1164,1183]" captionTargetBox="[151,1436,155,1114]" captionTargetId="figure@3.[151,1436,155,1114]" captionTargetPageId="3" captionText="Figs 1 - 3. Leiostyla taxa from Turkey. 1, Leiostyla paphlagonica paphlagonica Hausdorf 1990, holotype SMF 307492 (4.0 mm), Turkey, Vil. Kastamonu, Kapısuyu. 2, Leiostyla paphlagonica subangulosa subspec. nov., holotype NMBE 544659 (H = 3.7 mm), Turkey, Vil. Amasya, 10.3 km NE. Amasya. 3, Leiostyla eikenboomi spec. nov., holotype NMBE 544682 (H = 3.4 mm), Turkey, Vil. Karabük, Suçatı Tüneli 9 km ESE. Yenice. All photographs Bochud &amp; Neubert, × 15." httpUri="https://zenodo.org/record/439746/files/figure.png" pageId="4" pageNumber="7">Fig. 1</figureCitation>
) by the presence of a spiral furrow on the outer wall of the last whorl; this furrow is located below the 
<taxonomicName box="[1144,1238,1646,1670]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
inferior. Furthermore, the parietal callus is clearly less thickened, and the angular lamella and parietalis are more weakly developed. In addition, the radial striae are less regular and less densely packed. 
<taxonomicName box="[1151,1400,1784,1808]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="subSpecies" species="paphlagonica" subSpecies="subangulosa">
<emphasis box="[1151,1400,1784,1808]" italics="true" pageId="4" pageNumber="7">Leiostyla p. subangulosa</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[971,1083,1820,1843]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="subSpecies" species="paphlagonica" subSpecies="incisa">
<emphasis box="[971,1083,1820,1843]" italics="true" pageId="4" pageNumber="7">L. p. incisa</emphasis>
</taxonomicName>
by the presence of 
<taxonomicName class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="12" phylum="Mollusca" rank="species" species="subangularis">a subangularis</taxonomicName>
or, alternatively, the more prominently developed parietal callus (which fuses the angular lamella to the palatal insertion of the peristome) and the less diffuse 
<taxonomicName box="[915,1009,1958,1982]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
inferior.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="8" pageId="4" pageNumber="7" type="discussion">
<paragraph blockId="4.[831,1435,155,2051]" lastBlockId="5.[151,761,155,353]" lastPageId="5" lastPageNumber="8" pageId="4" pageNumber="7">
Remarks. – So far only three populations have been recorded for 
<taxonomicName box="[1036,1197,2027,2051]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="paphlagonica">
<emphasis box="[1036,1197,2027,2051]" italics="true" pageId="4" pageNumber="7">L. paphlagonica</emphasis>
</taxonomicName>
, each population having its own subspecific status (
<taxonomicName authority="Vilayet" authorityName="Vilayet" box="[535,761,155,179]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="paphlagonica">
<emphasis box="[535,669,155,179]" italics="true" pageId="5" pageNumber="8">paphlagonica</emphasis>
: Vilayet
</taxonomicName>
Kastamonu, Kapısuyu, 
<emphasis box="[414,474,191,214]" italics="true" pageId="5" pageNumber="8">incisa</emphasis>
: Vilayet Kastamonu, Azdavay; 
<emphasis box="[232,359,225,249]" italics="true" pageId="5" pageNumber="8">subangulosa</emphasis>
: Vilayet Amasya, Amasya). Much more field work has to be carried out to delineate the precise distribution area and subspecies boundaries of this polytypic species.
</paragraph>
</subSubSection>
</treatment>
</document>