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<mods:title id="04CADFA2DF20863A1A414EAC11CB14A6">Five new species of Bradina Lederer (Lepidoptera, Crambidae) from China, with remarks on the morphology of the genus</mods:title>
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<mods:namePart id="B81B049F60A543B1DEF22B6346C019AB">Guo, Jia-Ming</mods:namePart>
<mods:affiliation id="E3BFCFA071D541685B0135A5503A0221">College of Plant Protection, Southwest University, Chongqing, China</mods:affiliation>
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<mods:namePart id="20D0B0F20F9C61460C77AA3AB6B16511">Du, Xi-Cui</mods:namePart>
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<mods:affiliation id="D557340801D4C8B368EAFD535A7D4B94">College of Plant Protection, Southwest University, Chongqing, China</mods:affiliation>
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<treatment id="289655852DCA5E719B33F3EFEEF2D287" ID-GBIF-Taxon="209563574" LSID="urn:lsid:plazi:treatment:289655852DCA5E719B33F3EFEEF2D287" httpUri="http://treatment.plazi.org/id/289655852DCA5E719B33F3EFEEF2D287" lastPageNumber="49" pageId="0" pageNumber="49" scope_family="Crambidae" scope_order="Lepidoptera">
<subSubSection id="C2396AE642D730BEDBFA40ACF6413E37" pageId="0" pageNumber="49" type="nomenclature">
<paragraph id="9FEED6AEB85D611AB2314E8B3FDA17A3" pageId="0" pageNumber="49">
<taxonomicName id="0E3B435155D6EFD3C6AEA79F027FE3F5" ID-CoL="9242Z" ID-ENA="581787" LSID="28965585-2DCA-5E71-9B33-F3EFEEF2D287" authority="Lederer, 1863" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Bradina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradina" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Bradina Lederer, 1863</taxonomicName>
</paragraph>
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<subSubSection id="FA2D1B1B686AEC3D24F73096F5ABF40E" pageId="0" pageNumber="49" type="reference_group">
<paragraph id="9E6A3346176124F903A97F6D408DF215" pageId="0" pageNumber="49">
<taxonomicName id="A2F1C1D023AFCE1838AB85D5D149EB25" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Bradina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradina" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Bradina</taxonomicName>
Lederer, 1863: 424. Type species:
<taxonomicName id="D898388178E6D5B7A6A6E3D553B8FA32" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Bradina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradina impressalis" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="impressalis">Bradina impressalis</taxonomicName>
Lederer, 1863; subsequent designation by
<bibRefCitation id="C0882C50CB0B1B7275F9E40BAB3B7695" author="Hampson, GF" journalOrPublisher="The Fauna of British India, including Ceylon and Burma, London, Vol. 4" pageId="0" pageNumber="49" refId="B5" refString="Hampson, GF, 1896. Moths. The Fauna of British India, including Ceylon and Burma, London, Vol. 4" title="Moths." year="1896">Hampson 1896</bibRefCitation>
.
</paragraph>
<paragraph id="4F698971BD71E5A2722BC19BCFDF3414" pageId="0" pageNumber="49">
<taxonomicName id="AB22D4BA38D4106F75C8223DA12F682D" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Erilita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erilita" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Erilita</taxonomicName>
Lederer, 1863: 426. Type species:
<taxonomicName id="FDDA261F83557B7B3EF7AC1CF6D1DD41" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Erilita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erilita modestalis" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="modestalis">Erilita modestalis</taxonomicName>
Lederer, 1863, by monotypy.
</paragraph>
<paragraph id="FCBE108EF1689E48D0B2E1DB74B80BE9" pageId="0" pageNumber="49">
<taxonomicName id="2CCC7BA181561AC33688D09290B1684E" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Pleonectusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pleonectusa" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Pleonectusa</taxonomicName>
Lederer, 1863: 426. Type species:
<taxonomicName id="33148A87F25FDDBEECAABFC465855F24" authorityName="Walker" authorityYear="1859" class="Insecta" family="Crambidae" genus="Botys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Botys admixtalis" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="admixtalis">Botys admixtalis</taxonomicName>
Walker, 1859; subsequent designation by
<bibRefCitation id="1544A7BBD74826471C3FA75987013A54" author="Moore, F" journalOrPublisher="Reeve L and Co., London" pageId="0" pageNumber="49" refId="B17" refString="Moore, F, 1884. The Lepidoptera of Ceylon, Vol. 3. Reeve L and Co., London" title="The Lepidoptera of Ceylon, Vol. 3." year="1884">Moore 1884</bibRefCitation>
.
</paragraph>
<paragraph id="A9DC92D4551E12D037BD2931BCB7BADB" pageId="0" pageNumber="49">
<taxonomicName id="AE76D6E47BB25076A9E2593A0B52B6C9" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Trematarcha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trematarcha" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Trematarcha</taxonomicName>
Meyrick, 1886: 233. Type species:
<taxonomicName id="33DF1CF6579DB39EC56D4A6DBDE4883D" authorityName="C.Felder, R.Felder &amp; Rogenhofer" authorityYear="1875" class="Insecta" family="Crambidae" genus="Marasmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Marasmia erilitalis" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="erilitalis">Marasmia erilitalis</taxonomicName>
Felder, Felder &amp; Rogenhofer, 1875; subsequent designation by
<bibRefCitation id="EB39BAD5C8D427C6E6CF220F0061A6C6" DOI="https://doi.org/10.5962/bhl.title.124187" author="Klima, A" journalOrPublisher="Junk, ' s-Gravenhage" pageId="0" pageNumber="49" refId="B9" refString="Klima, A, 1937. Pyralidae: Subfam.: Scopariinae et Nymphulinae. In: Bryk F (Ed.) Lepidopterorum Catalogus 84. Dr. W. Junk, 's-Gravenhage, DOI: https://doi.org/10.5962/bhl.title.124187" title="Pyralidae: Subfam.: Scopariinae et Nymphulinae. In: Bryk F (Ed.) Lepidopterorum Catalogus 84. Dr. W." url="https://doi.org/10.5962/bhl.title.124187" year="1937">Klima 1937</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71FEB888F9FF8FD5446BA1D366871072" pageId="0" pageNumber="49" type="diagnosis">
<paragraph id="473FD478168138518D8E25AA27D04A74" pageId="0" pageNumber="49">Diagnosis.</paragraph>
<paragraph id="7665877D12FA33B72A1165DA06878E29" pageId="0" pageNumber="49">
<emphasis id="E92B5C636D9160A66E06A13FFDB6631D" bold="true" italics="true" pageId="0" pageNumber="49">Head</emphasis>
(Fig.
<figureCitation id="611595BE1EC092CCF6D65A31DF92255F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bradina megesalis, male A head B wing venation, wing slide no. GJM 21001. Scale bars: 0.5 mm (A); 1.0 mm (B)." figureDoi="10.3897/zookeys.1158.99411.figure1" httpUri="https://binary.pensoft.net/fig/839996" pageId="0" pageNumber="49">1A</figureCitation>
). Frons rounded. Antenna annulated, male with short cilia ventrally. Labial palpus obliquely upturned, second segment with broad scales ventrally, third joint minute and forward, apex blunt. Maxillary palpus filiform.
<emphasis id="5F512703B77B61DA4C689F984D57FCE6" bold="true" pageId="0" pageNumber="49">
<emphasis id="5A1ADC000DD0994D225C8835CD535FB2" italics="true" pageId="0" pageNumber="49">Thorax</emphasis>
.
</emphasis>
Forewing long and narrow usually; length of cell c. half of wing; discocellulars incurved; R from cell at c. four fifths above; Rs1 anastomosed with Rs2+s3 at base and with a long stalk c. two fifths of Rs3; Rs2 and Rs3 stalked c. three fifths of Rs3; basal half of Rs4 straight and clearly separated from Rs1+s2+s3; M2, M3 and CuA1 uniformly from posterior angle of cell at base (except for
<taxonomicName id="02EF76AC4246672E15138BF9B9E2A2B2" lsidName="B. diagonalis" pageId="0" pageNumber="49" rank="species" species="diagonalis">
<emphasis id="AAA0D76BF903FF88969C6A222B35090C" italics="true" pageId="0" pageNumber="49">B. diagonalis</emphasis>
</taxonomicName>
Hampson, 1896); CuA2 from cell at three quarters below. Hindwing with length of cell c. one third of wing; discocellulars incurved; Sc+R and Rs stalked c. one third of Rs; Rs and M1 from anterior angle of cell; M2, M3 and CuA1 uniformly from posterior angle of cell; CuA2 from cell at four fifths below (Fig.
<figureCitation id="A3CE49F54BDE5DCCBB6CCC9B9B7D1D3A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bradina megesalis, male A head B wing venation, wing slide no. GJM 21001. Scale bars: 0.5 mm (A); 1.0 mm (B)." figureDoi="10.3897/zookeys.1158.99411.figure1" httpUri="https://binary.pensoft.net/fig/839996" pageId="0" pageNumber="49">1B</figureCitation>
). Legs long and slender; middle tibia with outer distal spur c. half-length of inner spur.
<emphasis id="79F05C428C1C123BEEE31600855A91C1" bold="true" pageId="0" pageNumber="49">
<emphasis id="8D0843FF3500127C642C39DD00D005C1" italics="true" pageId="0" pageNumber="49">Abdomen</emphasis>
.
</emphasis>
Male abdomen long and slender (except for
<taxonomicName id="4626F90783AF27E4D6DA368D6FB7CCE9" lsidName="B. melanoperas" pageId="0" pageNumber="49" rank="species" species="melanoperas">
<emphasis id="63F38CA124E9D9139F4AEE05224258DD" italics="true" pageId="0" pageNumber="49">B. melanoperas</emphasis>
</taxonomicName>
Hampson, 1896).
</paragraph>
<caption id="8F38AA4F5B0A3B09733AE7D6D9C22890" doi="10.3897/zookeys.1158.99411.figure1" httpUri="https://binary.pensoft.net/fig/839996" pageId="0" pageNumber="49" start="Figure 1" startId="F1">
<paragraph id="BE7855AC2F213F1AD2E69854B9ACD541" pageId="0" pageNumber="49">
<emphasis id="CE1B21D3251A1F61B7BC5AD9757B7DBE" bold="true" pageId="0" pageNumber="49">Figure 1.</emphasis>
<taxonomicName id="2904A4163B8AF59A6CBE28D3B2C2EEA4" baseAuthorityName="Walker" baseAuthorityYear="1859" class="Insecta" family="Crambidae" genus="Bradina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradina megesalis" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="megesalis">
<emphasis id="CA4C82C306621F2573CC31385C798D66" italics="true" pageId="0" pageNumber="49">Bradina megesalis</emphasis>
</taxonomicName>
, male
<emphasis id="E823A9C1F57A650FA7F3B6B5A2272575" bold="true" pageId="0" pageNumber="49">A</emphasis>
head
<emphasis id="F357C065E5CB979C2C9369117F2F9C6E" bold="true" pageId="0" pageNumber="49">B</emphasis>
wing venation, wing slide no. GJM21001. Scale bars: 0.5 mm (
<emphasis id="26D62A314AD9DB0A001BFB240FD107FF" bold="true" pageId="0" pageNumber="49">A</emphasis>
); 1.0 mm (
<emphasis id="8F32A1EFBD9CAB6F5719E49C3393544A" bold="true" pageId="0" pageNumber="49">B</emphasis>
).
</paragraph>
</caption>
<paragraph id="E4069D67BD87FCBD9A46591BED0F4BC7" pageId="0" pageNumber="49">
<emphasis id="665757C4BEFDF618701F1A628ADAE521" bold="true" pageId="0" pageNumber="49">
<emphasis id="2B91D742DBD0E2908F1657A968AA3906" italics="true" pageId="0" pageNumber="49">Male genitalia</emphasis>
.
</emphasis>
Uncus diverse, apex with setae dorsally. Valva narrow or broad, some with well-developed setal cluster. Saccus developed. Phallus long and cylindrical.
</paragraph>
<paragraph id="3ED4A766208734827285478DF5135E48" pageId="0" pageNumber="49">
<emphasis id="CD610B7A4E592CFC4C0F986F197722D6" bold="true" pageId="0" pageNumber="49">
<emphasis id="866F7F242F23B4E7721DC92C7AFC1874" italics="true" pageId="0" pageNumber="49">Female genitalia</emphasis>
.
</emphasis>
Papillae anales densely setose. Apophyses anteriores c. twice length of apophyses posteriores. Ostium bursae well developed. Corpus bursae rounded or oval, inside usually densely studded with tiny spines; signum crescent and densely spinose, spines on concave side developed.
</paragraph>
</subSubSection>
<subSubSection id="9066BFDD95BC40C0D4819E2AAC4AA5B4" pageId="0" pageNumber="49" type="remarks">
<paragraph id="C3896751BAD5ECFC6FE2CFC8941F4FBF" pageId="0" pageNumber="49">Remarks.</paragraph>
<paragraph id="A0017248E20F95D7D26D831CD315B504" pageId="0" pageNumber="49">
The generic characteristics were summarized by
<bibRefCitation id="5302EF5E9BA748B5D654692202417A62" author="Khan, ZH" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="0" pageNumber="49" publicationUrl="http://hdl.handle.net/10603/52039" refId="B8" refString="Khan, ZH, 2000. Biosystematic Studies on Pyralid Pests of Rice Crop in India (Pyralidae: Lepidoptera). PhD Thesis, Aligarh Muslim University, Aligarh, India. http://hdl.handle.net/10603/52039" title="Biosystematic Studies on Pyralid Pests of Rice Crop in India (Pyralidae: Lepidoptera). PhD Thesis, Aligarh Muslim University, Aligarh, India." url="http://hdl.handle.net/10603/52039" year="2000">Khan (2000)</bibRefCitation>
, but the description about wing venation was incomplete. In addition, the lengths of the maxillary palpus and labial palpus were identical in
<normalizedToken id="8FCC8E1DACB817E0D70B209DA23DC4C2" originalValue="Khan’s">Khan's</normalizedToken>
description, while we found that both have their ends at the same height but they are of different lengths. Therefore, the generic characteristics are revised in the present study.
</paragraph>
<paragraph id="3CEEB23E2018B2967864BC78A9F2E116" pageId="0" pageNumber="49">
The bodies of
<taxonomicName id="453F5B8B1B502B4A00177CF51CE8A86B" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Bradina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradina" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="9805D7DB7BA15A7C1311B110AF311542" italics="true" pageId="0" pageNumber="49">Bradina</emphasis>
</taxonomicName>
species are usually brown, yellowish brown, or pale brown, except for a few species with white bodies, and have nearly identical wing markings. The male genitalia, on the other hand, are very diverse among species of this genus. We found that the male genitalia can be divided into three types according to the morphological characteristics of the valvae. The first type, represented by
<taxonomicName id="9371302A4282FE43236D01BA9FBCBC22" lsidName="B. admixtalis" pageId="0" pageNumber="49" rank="species" species="admixtalis">
<emphasis id="C9CD4692AF891FFB63DB46683D99F388" italics="true" pageId="0" pageNumber="49">B. admixtalis</emphasis>
</taxonomicName>
, have long and narrow valvae; the second type, represented by
<taxonomicName id="D152E3032287759FA8597D6EB2C1E800" lsidName="B. melanoperas" pageId="0" pageNumber="49" rank="species" species="melanoperas">
<emphasis id="0AA1052AEC31E32F24DF8620FB3DC886" italics="true" pageId="0" pageNumber="49">B. melanoperas</emphasis>
</taxonomicName>
, have short and broad valvae; the third type, represented by
<taxonomicName id="D6759268AA2FBB6332C2D0EBB66C7960" lsidName="B. megesalis" pageId="0" pageNumber="49" rank="species" species="megesalis">
<emphasis id="010AC4F5EC1D786495528BB12944D4C0" italics="true" pageId="0" pageNumber="49">B. megesalis</emphasis>
</taxonomicName>
, have broad valvae in which the costa is arched near the base or middle. Species in the present study have the third valva type, accompanied by the following common characteristics: body brown of various shades. Uncus broad, distal part bilobed, with dense short setae dorsally. Valva broad; costa arched near base or middle and accompanied by a cluster of long setae, usually followed by a depression. Saccus nearly trapezoidal, slightly concaved terminally. Juxta nearly rounded, split posteriorly. Phallus distinctly inflated at anterior end. Apophyses anteriores expanded at c. one third from base.
</paragraph>
</subSubSection>
<subSubSection id="34A6A3423311850B19514FFBB237A3DD" pageId="0" pageNumber="49" type="key to bradina species in the present study based on genitalia">
<paragraph id="72DB7C40B0630A54B3B2EEC95CABE53A" pageId="0" pageNumber="49">
Key to
<taxonomicName id="B6A1D766352B4334C12005F83A061985" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Bradina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bradina" order="Lepidoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="8EBB807A3870F38B8A9F724D2D417089" italics="true" pageId="0" pageNumber="49">Bradina</emphasis>
</taxonomicName>
species in the present study based on genitalia
</paragraph>
<paragraph id="E09DDFF42674E09F5FD4ED37F2CEBD8D" pageId="0" pageNumber="49">
<table id="8EF4FF1FF9F0B84DD518D7381FF9594D" inLine="true" pageId="0" pageNumber="49">
<tr id="A2EC8DD3EAF444D7E66C8A530EE41F25" pageId="0" pageNumber="49">
<td id="9FDDDD116F0EA4482001ECC3EE1AEAF0" colspan="1" pageId="0" pageNumber="49" rowspan="1">1</td>
<td id="5D9459AF96E3193F8F225A422BB78205" colspan="1" pageId="0" pageNumber="49" rowspan="1">Sacculus with a cluster of long setae near middle</td>
<td id="0ECD5673CE1D9C00E1355A1674BA848D" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="4CCA680FDB779EDF529836C2AE98BD59" bold="true" pageId="0" pageNumber="49">2</emphasis>
</td>
</tr>
<tr id="8FAB4AEEBA38689E0EC98815CD671726" pageId="0" pageNumber="49">
<td id="07620E83000A156810CEED54F66011D2" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="4266BCFC397E950E59AF7A9B82918745" colspan="1" pageId="0" pageNumber="49" rowspan="1">Sacculus without long setal cluster</td>
<td id="CA97F07FF49C617E8207FAAE56E0F826" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="C3F1094AE64B54AD82FB4CB9CFB8C8AC" bold="true" pageId="0" pageNumber="49">5</emphasis>
</td>
</tr>
<tr id="A400026BB481F9541218D3CA2FEFCE1D" pageId="0" pageNumber="49">
<td id="4C4534D57480EE4A12EE23FD1BCD4DD7" colspan="1" pageId="0" pageNumber="49" rowspan="1">2</td>
<td id="2CE4B317D99078537069D7A12A4B16DB" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with one fusiform cornutus composed of short and blunt spines, and one subcircular sclerotized cornutus</td>
<td id="985C49F265ED2AAD2734BCDAFFD9AF4F" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="1C189C95ACF5A2BF69B25A7FB756A76E" bold="true" pageId="0" pageNumber="49">
<taxonomicName id="333D5D774290E265E0647C0430A458A8" lsidName="B. fusoidea" pageId="0" pageNumber="49" rank="species" species="fusoidea">
<emphasis id="8520453CDABEAF9F8D562132970603BD" italics="true" pageId="0" pageNumber="49">B. fusoidea</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr id="343260CE36759D3290C62D55DF5F447E" pageId="0" pageNumber="49">
<td id="8DD4704F8FA5475CCDE668991B556AF6" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="13A93C5896D4B8FB355838FBA829CC18" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with various cornuti, but not as above</td>
<td id="06764280B2D0F02D43E39A75A657CEA5" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="AAA3CDA72302C15254A0FBE744A312B7" bold="true" pageId="0" pageNumber="49">3</emphasis>
</td>
</tr>
<tr id="3C2849C0405B0DAECD6AFF053EBE70B5" pageId="0" pageNumber="49">
<td id="D8F3209A6EB08EA4214C9E4E3F368F81" colspan="1" pageId="0" pageNumber="49" rowspan="1">3</td>
<td id="6CA7015B5C7D14EB3883D2DE6845367F" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with three long leaf-like cornuti tapered apically</td>
<td id="5BEE46893458D76D74D68A13605FC038" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="88B308BED07E71D3D842F94D408C1AAD" bold="true" pageId="0" pageNumber="49">
<taxonomicName id="2D7EB370DC2B4EFEA019B8D829683FE9" lsidName="B. ternifolia" pageId="0" pageNumber="49" rank="species" species="ternifolia">
<emphasis id="4C064E70D63C7EDCCB68244AABFE92B4" italics="true" pageId="0" pageNumber="49">B. ternifolia</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr id="A219ED604E44C670A957E9A241318395" pageId="0" pageNumber="49">
<td id="1A3758F8E1CF2C8FBFB6ABBF5B3E063E" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="663A645D4002E5858A1430B883CE9394" colspan="1" pageId="0" pageNumber="49" rowspan="1">Cornuti not as above</td>
<td id="CE31BA9B5AC6C5ACB238677615BCECA0" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="CAA29DC5B8BE1F0922C649385BDF09C2" bold="true" pageId="0" pageNumber="49">4</emphasis>
</td>
</tr>
<tr id="9B1428D4A34DB826A4E141739456C1DF" pageId="0" pageNumber="49">
<td id="03A18E2F667FDA71811E8D25634A3F8F" colspan="1" pageId="0" pageNumber="49" rowspan="1">4</td>
<td id="41EF8BF482433B1C90CFFADBD2330452" colspan="1" pageId="0" pageNumber="49" rowspan="1">Posterior phallus with two developed spear-like cornuti</td>
<td id="2D40917361C12D9D03F59ADDC2156A25" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<taxonomicName id="8BD0B591604F8F392D47FF258EED4751" lsidName="B. megesalis" pageId="0" pageNumber="49" rank="species" species="megesalis">
<emphasis id="80815B6E4AB4835A9F579D5D682B8C07" bold="true" italics="true" pageId="0" pageNumber="49">B. megesalis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="0148D4DCEEB5B6A814CF2C62C1C8F949" pageId="0" pageNumber="49">
<td id="A32018431BD1AB72588243DE7E6DEB76" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="3689C42C8B4F96B3AFA827D3A0D28C24" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with one short and spiral band-like cornutus tapered at posterior end, along with two lamellar cornuti in posterior half</td>
<td id="3597C286509B497B740F223360A062FC" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="2025D5FDBB05B6188FD8F40F4DD75F1F" bold="true" pageId="0" pageNumber="49">
<taxonomicName id="087CB9F34D81839FC65A484AABC95835" lsidName="B. spirella" pageId="0" pageNumber="49" rank="species" species="spirella">
<emphasis id="912376FC4D4EEF6502FA5AFE13A0AB61" italics="true" pageId="0" pageNumber="49">B. spirella</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr id="B7BB492D9E9C4F0D0404EF6BA33955CB" pageId="0" pageNumber="49">
<td id="11EDE0BF86818F3D9D3FE59BB75A318B" colspan="1" pageId="0" pageNumber="49" rowspan="1">5</td>
<td id="CF8F34ECCEC9AE6F0F7D6582781A6611" colspan="1" pageId="0" pageNumber="49" rowspan="1">Valva nearly rectangular, broad distally</td>
<td id="FFA137895EC4CB0A65EA44CE800866E3" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<taxonomicName id="17B2A12C2608A197A20D05E0256B1A5D" lsidName="B. subpurpurescens" pageId="0" pageNumber="49" rank="species" species="subpurpurescens">
<emphasis id="B513662CCFA071FA4E1F847EBA0C6DD4" bold="true" italics="true" pageId="0" pageNumber="49">B. subpurpurescens</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="1885EA7C37999E09311810C6DDE3839E" pageId="0" pageNumber="49">
<td id="B9C2926FB996A73E617A138F6F419A4C" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="5BE7D52D7795B5499AC23811D25A89F3" colspan="1" pageId="0" pageNumber="49" rowspan="1">Valva nearly elliptical or narrowed distally</td>
<td id="7B1BB0A11E05E3DE440F5437C267B24A" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="B4D377461F7F0DD4AA54F792713A9E04" bold="true" pageId="0" pageNumber="49">6</emphasis>
</td>
</tr>
<tr id="C3F7B09F92FA81E8833A88EBD35623FF" pageId="0" pageNumber="49">
<td id="FD4D768A3F14E93A208F9F86EAB4D4B4" colspan="1" pageId="0" pageNumber="49" rowspan="1">6</td>
<td id="448A50A3DA022A8C29454956D55CFBEF" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with one broad and spiral band-like cornutus</td>
<td id="55C2D10C3AC5C16D846315A42E21AF27" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="4A0A803F55B3F4BCA15369E9FF225901" bold="true" pageId="0" pageNumber="49">
<taxonomicName id="F88B664F7F6369512DE71212521AA547" lsidName="B. torsiva" pageId="0" pageNumber="49" rank="species" species="torsiva">
<emphasis id="7192F7D4638594447075DCE01EE3F2D4" italics="true" pageId="0" pageNumber="49">B. torsiva</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr id="434A162443EE5B58B7274C538A7F10D6" pageId="0" pageNumber="49">
<td id="30AC5E26523AC083C393FF96AC5A6F54" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="74B0F4B1B3612D7FE65B1886868A46BA" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with cornutus not as above</td>
<td id="16073531AF6269CBB70F87A28479BA13" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="3A29B4112D9CB6BE0175F656B74D3466" bold="true" pageId="0" pageNumber="49">7</emphasis>
</td>
</tr>
<tr id="F9C3D5DB1205D433A1CD558E6F0DF66E" pageId="0" pageNumber="49">
<td id="F2F6DFC77DECEB061BE35258039D6848" colspan="1" pageId="0" pageNumber="49" rowspan="1">7</td>
<td id="3745FA7EDD412BB83F8CD1530C3F32A9" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with one fusiform cornutus medially, and with two lamellar cornuti posteriorly. Ductus bursae slender, membranous</td>
<td id="467151D20E978C0A344F27EABD5A436E" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<taxonomicName id="885C85F851BD7198E4AAFD94EAE63E39" lsidName="B. translinealis" pageId="0" pageNumber="49" rank="species" species="translinealis">
<emphasis id="029336B0C166F265F548E8A285BBC994" bold="true" italics="true" pageId="0" pageNumber="49">B. translinealis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="52677A928EF74CD9A50A0F5CB30BBA6C" pageId="0" pageNumber="49">
<td id="C8E331648E9A7D2298FB2C73395073AB" colspan="1" pageId="0" pageNumber="49" rowspan="1">-</td>
<td id="1A8101A0A42D9FC80B5FBCF4505E5E17" colspan="1" pageId="0" pageNumber="49" rowspan="1">Phallus with one small fusiform cornutus anteriorly, and with two sickle-shaped cornuti posteriorly. Ductus bursae relatively thick, slightly sclerotized medially</td>
<td id="9B155FEC3A8244348F6D03D36B32D03B" colspan="1" pageId="0" pageNumber="49" rowspan="1">
<emphasis id="716080CED1BBF263E91A330674FD0EB3" bold="true" pageId="0" pageNumber="49">
<taxonomicName id="AA467F7064D9835EA087695D1BBC0456" lsidName="B. falciculata" pageId="0" pageNumber="49" rank="species" species="falciculata">
<emphasis id="B5ED316D3DF2D5DBAF599E9FD835B938" italics="true" pageId="0" pageNumber="49">B. falciculata</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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