264 lines
27 KiB
XML
264 lines
27 KiB
XML
<document ID-CLB-Dataset="21365" ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. & Hofman, Sebastian" docDate="2020" docId="16F422A81C675FFF9B361C753D667574" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Caucasogeyeria ignidona Grego & Palatov 2020, sp. nov." docType="treatment" docUuid="6DE56E85-4B3D-431D-AE7F-2CB087B421CD" docUuidSource="ZooBank" docVersion="7" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1732936627699" updateUser="ExternalLinkService">
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<mods:mods id="BAD34AE7969E58C06675143A9354AE71" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1C6ACC5AC127813BD9BF3356AC547D0A">
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<mods:title id="224937F35834EBA785E61ADDE9659253">Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
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<mods:name id="E5243A270DC4DE59A72F05E6AB380B69" type="personal">
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<mods:roleTerm id="B677F5DD7EC3C1DE1D44906A6488526C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="31DF06A6AEE181BDDBAE95D62D600465">Grego, Jozef</mods:namePart>
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<mods:affiliation id="BC075DF1CFE7D4BA6782E47809C35E78">Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name id="C9A9D4E1768087AD6F511F2240CA1457" type="personal">
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<mods:roleTerm id="86B2B66F03849D5C994ACEE61367EDD3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="751BA9467823EA5834E54021AC446371">Mumladze, Levan</mods:namePart>
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<mods:affiliation id="4F22C2C4C0796EF8E86900464BB221AA">Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
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<mods:nameIdentifier id="56C9F66AB6BCAA8CF46CD182CAB89BD4" type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
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</mods:name>
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<mods:name id="7BCF5078846930C5ADB3DFCCABFEF01E" type="personal">
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<mods:roleTerm id="619A54D7D818B04E15C3344EF533B64A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D025BB61CBF2DC8E2D5DFF5BABCFB199">Falniowski, Andrzej</mods:namePart>
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<mods:affiliation id="3C8E1521562849CE8B136FBA3DA9C008">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:name id="64075AE6BF56041F3A35BE1717664CE9" type="personal">
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<mods:roleTerm id="D38DC3C5D0AB51D796708A7AF78D9038">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5C6298FCB9830BAF312E5E7AAE68AA85">Osikowski, Artur</mods:namePart>
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<mods:affiliation id="5A97B8A1891A0E593228AA32C4268979">Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
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<mods:nameIdentifier id="9CCFBCC509BD86BE825AA6BA5921A12D" type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
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<mods:nameIdentifier id="93206C339D6DC40BCB972EE0F5BB3513" type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
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<mods:roleTerm id="F8A3C9F4BD3AF7AE4244C36C3E5A6D59">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="240383E9AFA8AF45C65DCACCED386CC9">Rysiewska, Aleksandra</mods:namePart>
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<mods:affiliation id="D93AA2ACDAE113CAC42C0455BE6C40A4">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:namePart id="31AF39BD61CE41D2A857B826C26779A4">Palatov, Dimitry M.</mods:namePart>
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<mods:affiliation id="803F14A835CF87EFCBCF13B6156AC73D">Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
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<mods:name id="9057B18DAA91DF830A823D8B583244BD" type="personal">
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<mods:namePart id="734709D27AC02D42413CA43E32B2E6E3">Hofman, Sebastian</mods:namePart>
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<mods:affiliation id="ABCE7B6A45DDE61F6C5815F2F0D7BEC8">Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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<mods:typeOfResource id="08327A6C3E1A8490C085740211935CCA">text</mods:typeOfResource>
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<mods:title id="28B942FB4E7E8EEFDBD0D349EBA2DCC4">ZooKeys</mods:title>
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<mods:part id="D69276AFDE2A7BC33A3AC5D672FE45A6">
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<mods:date id="050DDD456CBDC5DC08BADFF1C8AC010F">2020</mods:date>
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<mods:detail id="DAB4210B5DD4DDA8A98E7028EC68944C" type="volume">
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<mods:number id="E5CE05C94E205EAA3EC30C1C22861F7C">955</mods:number>
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<mods:start id="C02A46744FBC2169BD32B1CFDBB91CCD">1</mods:start>
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<mods:end id="05B41A40F744D154EC5BA7004D6F52E3">77</mods:end>
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<mods:url id="6478903E12A411C76217AAAEA464018D">http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
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<mods:classification id="41DD5DBD4FEDC64225FC87ECCE767D12">journal article</mods:classification>
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<mods:identifier id="AF3E05544AF3B91E792BEA61F8E7B065" type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
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<mods:identifier id="C2005FF0D87F18B95BDC5AC59E06A199" type="Pensoft-Pub">1313-2970-955-1</mods:identifier>
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<mods:identifier id="55B90745D1687C19A14FD0630383E437" type="ZooBank">C8EF4A4B6F054621B9B5AE47FEE7C217</mods:identifier>
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<mods:identifier id="F5EEB3E53BBEB718A2F0115725B1AC17" type="Pensoft-UUID">F7018CF19C6158309D57C83A2F5E9478</mods:identifier>
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</mods:mods>
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<treatment id="16F422A81C675FFF9B361C753D667574" ID-GBIF-Taxon="166096069" LSID="urn:lsid:zoobank.org:act:6DE56E85-4B3D-431D-AE7F-2CB087B421CD" httpUri="http://treatment.plazi.org/id/16F422A81C675FFF9B361C753D667574" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_phylum="Mollusca" scope_subFamily="Sadlerianinae" scope_superFamily="Truncatelloidea">
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<subSubSection id="AB50690547F41FD057C0FFB044B9D64F" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="277CC975C4A2221761FB4B962360B35C" pageId="0" pageNumber="1">
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<taxonomicName id="44113C090E9B84F09770CBA314B46069" ID-CoL="7QL4S" LSID="16F422A8-1C67-5FFF-9B36-1C753D667574" authority="Grego & Palatov" authorityName="Grego & Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria ignidona" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ignidona" status="sp. nov.">Caucasogeyeria ignidona Grego & Palatov</taxonomicName>
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<taxonomicNameLabel id="D1038B4A152BE589175979E307C5833B" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="21B0ED470376AD855214D90B963FA6C2" captionStart="Plate 15" captionStartId="F28" captionText="Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego." figureDoi="10.3897/zookeys.955.51983.plate15" httpUri="https://binary.pensoft.net/fig/439180" pageId="0" pageNumber="1">Plate 15</figureCitation>
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<figureCitation id="6E395184CC60B3F4D522AF47F896137C" captionStart="Figure 14" captionStartId="F29" captionText="Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska." figureDoi="10.3897/zookeys.955.51983.figure14" httpUri="https://binary.pensoft.net/fig/439181" pageId="0" pageNumber="1">(1); Fig. 14A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6C2A09D6A80FA6A8EFE30F508D894AA6" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="215087E5311EF05EE432E1EB47E5E583" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="1015D1789A8EE1847740AC26926347B6" pageId="0" pageNumber="1">
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Georgia • Imereti, Kumistavi, Prometheus Cave (პრომეთეს
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<normalizedToken id="838907FA03A469058D0A5A63599598EE" originalValue="მღვიმე">მღვიმე</normalizedToken>
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);
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<geoCoordinate id="E7907D41EE007E2C12DB09195F3226B5" degrees="42" direction="north" minutes="22" orientation="latitude" precision="15" seconds="33" value="42.37583">42°22'33"N</geoCoordinate>
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,
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<geoCoordinate id="1F553C7DC1E36C1B34993F3D00B7B87B" degrees="42" direction="east" minutes="36" orientation="longitude" precision="15" seconds="2" value="42.600555">42°36'2"E</geoCoordinate>
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; 175 m a.s.l.; bottom of cave stream.
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</paragraph>
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<caption id="FB4D57647F64B5BFC361987E0FA960E9" doi="10.3897/zookeys.955.51983.plate15" httpUri="https://binary.pensoft.net/fig/439180" pageId="0" pageNumber="1" start="Plate 15" startId="F28">
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<paragraph id="EF000A6F3A5C672D975F76B9458F630A" pageId="0" pageNumber="1">
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<emphasis id="DFE7F93FF896F5D1298749EEE5F47FB8" bold="true" pageId="0" pageNumber="1">Plate 15.</emphasis>
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<emphasis id="919986956985D720FE33A856738D6465" bold="true" pageId="0" pageNumber="1">1</emphasis>
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<taxonomicName id="8561BC2701C6FCDFF42EBE24EC30F046" authorityName="Grego & Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria ignidona" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ignidona">
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<emphasis id="E47D351FF39D51D1AA31C2343A310C69" italics="true" pageId="0" pageNumber="1">Caucasogeyeria ignidona</emphasis>
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</taxonomicName>
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sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype
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<emphasis id="550428096F218A7DA9CFD43F262E0165" bold="true" pageId="0" pageNumber="1">2</emphasis>
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<taxonomicName id="BE14B50F66505F696C0CEAC806DB9A79" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
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<emphasis id="AD3202463F6AC0BC87451378D69C5488" italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype
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<emphasis id="156EEDFD03F61D9717AD242E83531FCC" bold="true" pageId="0" pageNumber="1">3</emphasis>
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<taxonomicName id="B38AD4EC73872883A73BEB2513E94BB1" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
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<emphasis id="78DBE309FAD314B122AE374A7164B30C" italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype
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<emphasis id="0B0BC8CB3BAB14B47E9752ECC1C72C0D" bold="true" pageId="0" pageNumber="1">4</emphasis>
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<taxonomicName id="862A50AB06113295C78F944BEED623E5" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
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<emphasis id="C8E9D03BF5A33D2E5F8677E37E03BE06" italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
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</taxonomicName>
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sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype
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<emphasis id="759EE853CA93D4931373BB965D0E6F9E" bold="true" pageId="0" pageNumber="1">5-8</emphasis>
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<taxonomicName id="259181BA5CAFDEA9AA2609F798B71771" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria pseudocolchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pseudocolchis">
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<emphasis id="B352368FA941DF7114D355D75FF2BB07" italics="true" pageId="0" pageNumber="1">Caucasogeyeria pseudocolchis</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Mukhuri, Shisha Spring
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<emphasis id="48E3E4D51D639FD69765850E056C0C6D" bold="true" pageId="0" pageNumber="1">5</emphasis>
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holotype
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<emphasis id="4B20015324D6E3078E26159FCA0AA7DC" bold="true" pageId="0" pageNumber="1">6-8</emphasis>
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paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="D24A051EBF6654F14D2355C98C1373F6" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="A13C425D60D2EF4F8DB436343D064EEF" pageId="0" pageNumber="1">Material.</paragraph>
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<paragraph id="67D3EAF5BC3BFCA36A490902FDC86E15" pageId="0" pageNumber="1">
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<emphasis id="2407CC34292E565160EA6E0F08BAFE01" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Georgia • adult, dry; type locality; 01 May 2018; J. Grego, L. Mumladze and M.
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<normalizedToken id="F9CF04B7F6C591A2C5A1C41A1F473C87" originalValue="Olšavský">Olsavsky</normalizedToken>
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leg.; ISU FM-T005-H.
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<emphasis id="1F98E9A6C5A312F9B1AB40FAB67E5FF3" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Georgia • same as for holotype; ISU FM-T005-P1/1 dry and 1 wet, coll. JG F0969/1 dry and 2 wet.
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</paragraph>
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</subSubSection>
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<subSubSection id="CB7F5E18F61FAF625F6AC169E537142D" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="75A849C48C1E7E362E0270199A3EF551" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="FBD6D13D0E8DB0F8AA6D3C7CC0822906" pageId="0" pageNumber="1">
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The new species can be distinguished from other members of the genus by the typical shell aperture.
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<taxonomicName id="6E8DDBC7D27C90AFFB97222ED2004F8E" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria gloeri" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gloeri">
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<emphasis id="854DEFD6E63F301A1EADBB78388992B5" italics="true" pageId="0" pageNumber="1">Caucasogeyeria gloeri</emphasis>
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</taxonomicName>
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sp. nov. has a larger, more robust shell with different aperture,
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<taxonomicName id="C1D0E97946A7DDBF9D489A4ABAEC3351" lsidName="C. colchis" pageId="0" pageNumber="1" rank="species" species="colchis">
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<emphasis id="6427182E9EC8685B3BED5990E84C25C6" italics="true" pageId="0" pageNumber="1">C. colchis</emphasis>
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</taxonomicName>
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sp. nov. has more sinuated and more deeply cut labral margin at its columellar side, and
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|
<taxonomicName id="C859FE43DCB315A8C4320B1C6568A8B7" lsidName="C. chrysomallos" pageId="0" pageNumber="1" rank="species" species="chrysomallos">
|
|
<emphasis id="E9217EE50F82B5A49B602021E0AE8201" italics="true" pageId="0" pageNumber="1">C. chrysomallos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. has smaller, more conical and elongate shell with a greater number of whorls and proportionally smaller, differently shaped aperture.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E8027818A4551106CCEF756BC1956B9D" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="6B78B483998672880AD7DA8119BA5C36" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="E661C05F8683409A6F03E3D7E162C5F7" pageId="0" pageNumber="1">
|
|
<emphasis id="DAB213BE30715C62A8B7C59E469FD738" bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
|
: conical with blunt protoconch and with
|
|
<normalizedToken id="E79DA37ADCB3CFDB09042963C34D2CF2" originalValue="3½">31/2</normalizedToken>
|
|
inflated whorls separated by semi-deep suture. Height 1.4-1.7 mm. Shell surface milky white, glossy with occasional rusty brown incrustations. Aperture expanded, proportionally larger, rhomboidal with a weak negative sinuation at labral junction with the body whorl and a weak positive sinuation at columellar margin. Umbilicus slit-like.
|
|
</paragraph>
|
|
<paragraph id="63929745AEF84DFE8A77ECDF1542AF8C" pageId="0" pageNumber="1">
|
|
<emphasis id="A2F379703FC73F7D00017A439EAECF22" bold="true" italics="true" pageId="0" pageNumber="1">Operculum</emphasis>
|
|
: paucispiral, glass-like translucent.
|
|
</paragraph>
|
|
<paragraph id="3B87F2979F297B34830EB9F1F2F844AE" pageId="0" pageNumber="1">
|
|
<emphasis id="F1EEECB1E6FB4AA620046C2D45BEC0DD" bold="true" italics="true" pageId="0" pageNumber="1">Animal body</emphasis>
|
|
: white, without eye spots.
|
|
</paragraph>
|
|
<paragraph id="DF7B18F060CD56A8D99BC2DBBD3A9FFF" pageId="0" pageNumber="1">
|
|
<emphasis id="8E61E6DA7DE69AA5E78F7ED7F9C0AFF8" bold="true" italics="true" pageId="0" pageNumber="1">Holotype measurements</emphasis>
|
|
: H-1.60 mm; W-1.10 mm; BH-1.15 mm; BW-0.9 mm; AH-0.85 mm; AW-0.70 mm; CA: 28°.
|
|
</paragraph>
|
|
<paragraph id="0DC85AFFD3F3AFF99764C1E03E4C4284" pageId="0" pageNumber="1">
|
|
<emphasis id="A8890F114AD183DFF18643133E84601D" bold="true" italics="true" pageId="0" pageNumber="1">Anatomy</emphasis>
|
|
: the penis (Fig.
|
|
<figureCitation id="94F8592770B3FABF97D0FA8BD0E60197" captionStart="Figure 14" captionStartId="F29" captionText="Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska." figureDoi="10.3897/zookeys.955.51983.figure14" httpUri="https://binary.pensoft.net/fig/439181" pageId="0" pageNumber="1">14A-C</figureCitation>
|
|
) bent, simple, narrow, gradually narrowing towards its distal end, vas deferens inside running straight.
|
|
</paragraph>
|
|
<caption id="48D3509AA9873769481CC3044FCAD363" doi="10.3897/zookeys.955.51983.figure14" httpUri="https://binary.pensoft.net/fig/439181" pageId="0" pageNumber="1" start="Figure 14" startId="F29">
|
|
<paragraph id="D9CAAE27E6DA1E9822378124AE10F8FD" pageId="0" pageNumber="1">
|
|
<emphasis id="6657975A957C6BFFD0E86DA55B1BCD86" bold="true" pageId="0" pageNumber="1">Figure 14.</emphasis>
|
|
<emphasis id="CF1397C130ED9B9CC0EAEEBE0C2F812C" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
|
<taxonomicName id="C487D84380CB0523E99E364293A63175" authorityName="Grego & Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria ignidona" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ignidona">
|
|
<emphasis id="1556D59D64EDCA84A311AB4A629BBBC3" italics="true" pageId="0" pageNumber="1">Caucasogeyeria ignidona</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., Imereti, Kumistavi, Prometheus Cave
|
|
<emphasis id="82DE5C1E17AD3468D8EFE312D02CF667" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
<taxonomicName id="763B2BC0E9BC61AE42CACAA9AB35CA07" lsidName="C. chrysomallos" pageId="0" pageNumber="1" rank="species" species="chrysomallos">
|
|
<emphasis id="6F4D2D9C8809326BA49EE6811718BA2E" italics="true" pageId="0" pageNumber="1">C. chrysomallos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="93924EF4FEDDCB1A9473D21E8E49E040" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="B31CA3256AB69654B81CFE935777172A" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="5CF49B57682EC7AC002AA776A2D6F3A0" pageId="0" pageNumber="1">
|
|
Name derived from Latin word
|
|
<taxonomicName id="2ADFD84B7AB8BB7FCCAA17A91A4BC8CA" lsidName="ignidona" pageId="0" pageNumber="1" rank="species" species="ignidona">
|
|
<emphasis id="878C1AAFE017A244732C82E12F7735F9" italics="true" pageId="0" pageNumber="1">ignidona</emphasis>
|
|
</taxonomicName>
|
|
meaning of "donating fire", referring to the gift of Prometheus to the mankind, indirectly indicating the name of type locality in the Prometheus Cave near Kutaisi.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B285C5784FC9F4632C58C8787F5A0D43" pageId="0" pageNumber="1" type="habitat">
|
|
<paragraph id="36A941AE0AC3E04C0F9C5E9AC154667E" pageId="0" pageNumber="1">Habitat.</paragraph>
|
|
<paragraph id="2727E7322D806F55A578EE6D17DB9FF2" pageId="0" pageNumber="1">Stygobiotic species. Live individuals of the new species were found in the cave stream on submerged stones and gravel, covered by a layer of dark brown-black layer of bacterial mats. Empty shells were found in sandy sediment of the cave stream.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="343DBF563939EEFC13EEA366CB99E25F" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="6D45E6F1BE9AFDBF3660FD5AACBFBA68" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="2AD5B3EC25E478DD42974EEEB67F21C4" pageId="0" pageNumber="1">Only known from the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CD1E3BBC253275C3292CAC0077264281" pageId="0" pageNumber="1" type="conservation status">
|
|
<paragraph id="CDDB1CBB517F141793CB1E8E91AA56D1" pageId="0" pageNumber="1">Conservation status.</paragraph>
|
|
<paragraph id="7D72FF7F6956DAA97FF97FF7C3A00155" pageId="0" pageNumber="1">The number of known locations (1) is no more than 5 and EOO is smaller than 20 km2. There is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number or mature individuals are declining however due to its extremely small EOO we assessed as Vulnerable (VU) D2.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6301009632479EB9D95B838C78E85AEE" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="9F8416640487AA5122A256C51C88BC7D" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="E44B91FB6A57F3A73099BAA8047FFC1C" pageId="0" pageNumber="1">
|
|
The sympatric
|
|
<taxonomicName id="F0D472BB7453E009EB342860B2288014" authorityName="Grego & Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis prometheus" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prometheus">
|
|
<emphasis id="85660D178B1EBCC83701A1210C8B6EB5" italics="true" pageId="0" pageNumber="1">Imeretiopsis prometheus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. has more numerous populations throughout the cave stream. It is not clear whether both species share the micro-habitats within the same cave stream.
|
|
</paragraph>
|
|
<caption id="B5BABDF5B46B9C1415A6F2E4F00351A2" doi="10.3897/zookeys.955.51983.plate16" httpUri="https://binary.pensoft.net/fig/439182" pageId="0" pageNumber="1" start="Plate 16" startId="F30">
|
|
<paragraph id="8E64B55E43A0B0FE456B5EA85E3D03B9" pageId="0" pageNumber="1">
|
|
<emphasis id="85E740908F6222B922AD4BC9137A1D99" bold="true" pageId="0" pageNumber="1">Plate 16.</emphasis>
|
|
<emphasis id="DFACF189FA6EAEC8550143A5D4781337" bold="true" pageId="0" pageNumber="1">1-6</emphasis>
|
|
<taxonomicName id="10FA48145853DE00B9A51C63B1EB6EAD" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria chrysomallos" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysomallos">
|
|
<emphasis id="8E6FEC06AE3804E12234C66644ED392E" italics="true" pageId="0" pageNumber="1">Caucasogeyeria chrysomallos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring
|
|
<emphasis id="3F1616403214F1330354A64BF2849799" bold="true" pageId="0" pageNumber="1">1</emphasis>
|
|
holotype
|
|
<emphasis id="FEC88FC7D58F1C83B9C3494A5F821652" bold="true" pageId="0" pageNumber="1">2-6</emphasis>
|
|
paratypes
|
|
<emphasis id="66CDE595B1587CA5CB58446DC71809FA" bold="true" pageId="0" pageNumber="1">7, 8</emphasis>
|
|
<taxonomicName id="C5492B248B52095FF0B2096277E7644D" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria chrysomallos" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysomallos">
|
|
<emphasis id="D5CFB87A86BFF0A89B56B6A2B2971C72" italics="true" pageId="0" pageNumber="1">Caucasogeyeria chrysomallos</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring
|
|
<emphasis id="324E60808893609809A922E002AAE88A" bold="true" pageId="0" pageNumber="1">9</emphasis>
|
|
<taxonomicName id="AA2E511A0D27EFF1381201E2C735E38D" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria gloeri" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gloeri">Caucasogeyeria cf. gloeri</taxonomicName>
|
|
, Samegrelo, Mukhuri, Mapeli Cave, paratype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |