213 lines
17 KiB
XML
213 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.363.6482" ID-GBIF-Dataset="36e5551d-3881-45a2-810a-386e517c86fc" ID-PMC="PMC3867181" ID-Pensoft-Pub="1313-2970-363-1" ID-PubMed="24363601" ID-ZBK="780695A9CB2E4FECA0560BA6F456185E" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-363-1" ModsDocOrigin="ZooKeys 363" ModsDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" checkinTime="1451246560986" checkinUser="pensoft" docAuthor="Mathis, Wayne N. & Zatwarnicki, Tadeusz" docDate="2013" docId="10165F5A3A9C3E63E85325E91852829E" docLanguage="en" docName="ZooKeys 363: 1-161" docOrigin="ZooKeys 363" docSource="http://dx.doi.org/10.3897/zookeys.363.6482" docTitle="Hydrochasma kaieteur Mathis & Zatwarnicki, 2013, sp. n." docType="treatment" docUuid="2DFE442B-EA52-4264-9A1B-F2BE46D0EFFB" docUuidSource="ZooBank" docVersion="4" lastPageNumber="59" masterDocId="FFF2FFCA0333765855689824FFCDB839" masterDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="57" updateTime="1668157177207" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>363</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>161</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.363.6482</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.363.6482</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-363-1</mods:identifier>
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<mods:identifier type="ZBK">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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<mods:identifier type="ZooBank">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050168" LSID="urn:lsid:zoobank.org:act:2DFE442B-EA52-4264-9A1B-F2BE46D0EFFB" httpUri="http://treatment.plazi.org/id/10165F5A3A9C3E63E85325E91852829E" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57">
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<subSubSection pageId="56" pageNumber="57" type="nomenclature">
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<paragraph pageId="56" pageNumber="57">
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17.
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<taxonomicName LSID="http://zoobank.org/2DFE442B-EA52-4264-9A1B-F2BE46D0EFFB" class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma kaieteur" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="kaieteur">Hydrochasma kaieteur</taxonomicName>
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<taxonomicNameLabel pageId="56" pageNumber="57">sp. n.</taxonomicNameLabel>
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Figs 96-100
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="diagnosis">
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<paragraph pageId="56" pageNumber="57">Diagnosis.</paragraph>
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<paragraph lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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This species is distinguished from other congeners by the following combination of characters: Small shore flies, body length 1.25-1.80 mm.
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<pageBreakToken pageId="57" pageNumber="58" start="start">Head</pageBreakToken>
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: Antenna mostly dark gray; parafacial silvery white, concolorous with facial coloration; gena-to-eye ratio 0.17-0.18. Thorax: Wing with costal vein ratio 0.65-0.67; M vein ratio 0.54-0.58. Forecoxa mostly gray to silvery gray, some yellowish coloration at ventral apex; forefemur lacking row of spine-like setulae along anteroventral surface; tibiae mostly gray; hindtibia lacking a long, spur-like seta ventroapically. Abdomen: Tergites 3-4 with moderately deep, gray to silvery gray wedges at lateral margins; tergite 5 of male mostly to entirely gray, sometimes with posterior margin darkened. Male terminalia (Figs 96-99): Epandrium generally elongate, almost 3 times longer than wide, and setulose, in posterior view (Fig. 96) with dorsal arch attenuate, not connected, dorsal 1/3 somewhat rectangular, ventral 2/3 tapered with lateral margins shallowly sinuous to a pointed ventral margin, ventromedial apex incised, narrowly V-shaped, in lateral view (Fig. 97) very narrowly developed dorsally laterad of cerci, thereafter ventrally becoming slightly wider to midlength, then becoming wider at ventral 1/3, before becoming tapered to pointed ventral margin; cerci short, length less than twice width, in posterior view (Fig. 96) hemispherical with pointed dorsal apex and truncate ventral margin, in lateral view (Fig. 97) almost evenly semicircular; aedeagus in lateral view (Fig. 99) relatively simple, narrowly tubular except for expanded, mostly membranous apex, length 6
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<normalizedToken originalValue="–7×">-7x</normalizedToken>
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width, in ventral view (Fig. 98) tubular; phallapodeme in lateral view (Fig. 99) spatulate with width of keel equal to width of phallapodeme, keel or spatulate end at end toward hypandrium, rest of phallapodeme tapered, curved subapically toward aedeagus, in ventral view (Fig. 98) linear, with slightly and narrowly developed T-shaped extensions; gonite in lateral view (Fig. 99) bar-like, narrowly linear, sinuous, in ventral view (Fig. 98) short, somewhat shallowly zig-zag; hypandrium in lateral view (Fig. 99) robustly bar-like width about equal to aedeagus, in ventral view (Fig. 98) somewhat rectangular with broadly and deeply emarginate posterior margin and with anterior margin as 2 blunted rounded processes and a moderately deep and narrowly incised medial portion.
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</paragraph>
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<caption pageId="57" pageNumber="58">
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<paragraph pageId="57" pageNumber="58">
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Figures 96-99.
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma kaieteur" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="kaieteur">Hydrochasma kaieteur</taxonomicName>
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sp. n. (Guyana: Dubalay Ranch, Berbice River) 96 epandrium and cerci, posterior view 97 same, lateral view 98 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 99 same, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="57" pageNumber="58" type="type material">
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<paragraph pageId="57" pageNumber="58">Type material.</paragraph>
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<paragraph pageId="57" pageNumber="58">
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The holotype male of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma kaieteur" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="kaieteur">Hydrochasma kaieteur</taxonomicName>
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is labeled "GUYANA. Kaieteur Fal[l]s
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="5.175">5°10.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.448334">59°26.9'W</geoCoordinate>
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[,] 21-24 August 1997[,] Wayne N. Mathis/USNM ENT 00080329 [plastic bar code label]/HOLOTYPE ♂
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma kaieteur" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="kaieteur">Hydrochasma kaieteur</taxonomicName>
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Mathis & Zatwarnicki, USNM [red]." The holotype is double mounted (minuten in a block of plastic), is in excellent condition, and is deposited in the USNM. Thirty paratypes (20♂, 10♀; USNM) bear the same label data as the holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="57" pageNumber="58" type="type locality">
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<paragraph pageId="57" pageNumber="58">Type locality.</paragraph>
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<paragraph pageId="57" pageNumber="58">
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Guyana. Kaieteur Falls (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="5.175">05°10.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.448334">59°26.9'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="other specimens examined">
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<paragraph pageId="57" pageNumber="58">Other specimens examined.</paragraph>
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<paragraph pageId="57" pageNumber="58">
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Neotropical. ARGENTINA. Formosa: Ingeniero
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<normalizedToken originalValue="Juárez">Juarez</normalizedToken>
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(
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-23.911667">23°54.7'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-61.863335">61°51.8'W</geoCoordinate>
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), 2 Jan 1949, R. Golbach (1♂; USNM).
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</paragraph>
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<paragraph lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
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BRAZIL.
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<normalizedToken originalValue="Paraná">Parana</normalizedToken>
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: Antonina (
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-25.473333">25°28.4'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-48.681667">48°40.9'W</geoCoordinate>
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; beach/mangal), 9 Apr 2010, D. and W. N. Mathis (3♂; DZUP, USNM); Antonina (
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-25.451666">25°27.1'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-48.685">48°41.1'W</geoCoordinate>
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; beach; Ponta da Pita), 15 Feb 2010, D. and W. N. Mathis (3♂; DZUP, USNM); Matinhos (N.;
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-25.79">25°47.4'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-48.52667">48°31.6'W</geoCoordinate>
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; 1 m; beach/estuary), 30 Jan-9 Apr 2010, D. and W. N.
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<pageBreakToken pageId="58" pageNumber="59" start="start">Mathis</pageBreakToken>
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(3♂; DZUP, USNM); Morretes (
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-25.466667">25°28'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-43.985">43°59.1'W</geoCoordinate>
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), 29 Aug 2000, D. and W. N. Mathis (14♂, 7♀; USNM);
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<normalizedToken originalValue="Paranaguá">Paranagua</normalizedToken>
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(Rio
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<normalizedToken originalValue="Itiberê">Itibere</normalizedToken>
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;
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-25.523333">25°31.4'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-48.505">48°30.3'W</geoCoordinate>
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; 3 m), 23 Jan 2010, D. and W. N. Mathis (12♂, 2♀; DZUP, USNM); Prainha (5 km S Matinhos;
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-25.853333">25°51.2'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-48.56">48°33.6'W</geoCoordinate>
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; beach), 15 Nov 2010, D. and W. N. Mathis (11♂; DZUP, USNM). Santa Catarina: Barra Velha (
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-26.633333">26°38'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-48.681667">48°40.9'W</geoCoordinate>
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; beach), 29 Apr 2010, D. and W. N. Mathis (2♂; DZUP, USNM).
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</paragraph>
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<paragraph pageId="58" pageNumber="59">
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GUYANA. Conservation of Ecological Interactions and Biotic Associations (CEIBA; ca. 40 km S Georgetown;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="6.4983335">06°29.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-58.218334">58°13.1'W</geoCoordinate>
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), 13-21 Apr 1994, 1995, W. N. Mathis (1♀; USNM); Karanambo (Rupununi River;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="3.7516665">03°45.1'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.31">59°18.6'W</geoCoordinate>
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; 85m), 1-3 Dec 2010, W. N. Mathis (20♂; USNM); Karanambo, Rupununi River (ox bow;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="3.75">03°45'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.291668">59°17.5'W</geoCoordinate>
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; 85m), 2 Dec 2010, W. N. Mathis (1♂; USNM); Kato, Chiung River (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="4.661667">04°39.7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.833332">59°50.0'W</geoCoordinate>
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), 1 May 1995, W. N. Mathis (8♂, 2♀; USNM) Kumo Pond (20 km S Lethem;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="3.2783334">03°16.7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.725">59°43.5'W</geoCoordinate>
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), 30 Apr 1995, W. N. Mathis (2♀; USNM); Kumu River and Falls (25 km SE Lethem in Kanuku Mountains;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="3.2649999">03°15.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.726665">59°43.6'W</geoCoordinate>
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), 28-30 Apr 1995, W. N. Mathis (2♂, 5♀; USNM); Mahaica (3 km E;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="6.725">06°43.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-57.943333">57°56.6'W</geoCoordinate>
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), 14 Apr 1994, W. N. Mathis (1♂, 5♀; USNM); Mahaica (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="6.713333">06°42.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-57.926666">57°55.6'W</geoCoordinate>
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), 14-22 Apr 1994, 1995, W. N. Mathis (8♂, 14; USNM); Menzies Landing (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="5.1683335">05°10.1'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.491665">59°29.5'W</geoCoordinate>
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), 23 Aug 1997, W. N. Mathis (7♂, 4♀; USNM); Paramakatoi (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.7">04°42'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-59.713333">59°42.8'W</geoCoordinate>
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), 24-25 Aug 1997, W. N. Mathis (6♂, 2♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="distribution">
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<paragraph pageId="58" pageNumber="59">Distribution</paragraph>
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<paragraph pageId="58" pageNumber="59">
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(Fig. 100). Neotropical: Argentina (Formosa), Brazil (
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<normalizedToken originalValue="Paraná">Parana</normalizedToken>
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, Santa Catarina), Guyana.
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</paragraph>
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<caption pageId="58" pageNumber="59">
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<paragraph pageId="58" pageNumber="59">
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Figure 100. Distribution of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma kaieteur" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="kaieteur">Hydrochasma kaieteur</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma lineatum" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="lineatum">Hydrochasma lineatum</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="etymology">
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<paragraph pageId="58" pageNumber="59">Etymology.</paragraph>
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<paragraph pageId="58" pageNumber="59">The species epithet, kaieteur, is the name of the world-famous falls in Guyana where this species was collected. The name is a noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="remarks">
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<paragraph pageId="58" pageNumber="59">Remarks.</paragraph>
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<paragraph pageId="58" pageNumber="59">This species is distinguished from congeners, especially those of the incisum group, by the shape of structures of the male terminalia, in particular the epandrium in posterior view and the robust, V-shaped hypandrium in ventral view.</paragraph>
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</subSubSection>
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</treatment>
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</document> |