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<document ID-CLB-Dataset="24202" ID-DOI="http://dx.doi.org/10.3897/zse.96.48312" ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0" ID-Pensoft-Pub="1860-0743-1-25" ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF" ID-ZooBank="F2C8585A1268443693348B64AE20F6EE" ModsDocID="1860-0743-1-25" checkinTime="1580744411276" checkinUser="pensoft" docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian &amp; Neubauer, Thomas A" docDate="2020" docId="0B43E9776B3A5F6A926554482B2B4A6F" docLanguage="en" docName="ZoosystEvol 96(1): 25-66" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.48312" docTitle="Theodoxus gloeri Odabasi &amp; Arslan 2015" docType="treatment" docVersion="8" id="4EDA9B5B777551B7BBB0152DD188E5EF" lastPageNumber="25" masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF" masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species" masterLastPageNumber="66" masterPageNumber="25" pageNumber="25" updateTime="1732731517510" updateUser="ExternalLinkService">
<mods:mods id="C06FAA3D7DD0AEECB4D056D1D04DAE19" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="07CECEC7D75881338AA531DCD929E173">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:name id="C6EE8D7315780C753038733D4D31A0CF" type="personal">
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<mods:namePart id="7774B85F951792585150EC22055437AE">Sands, Arthur F</mods:namePart>
<mods:affiliation id="BCCC12913BD2389B6BDEB34FB2D73580">Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="2D31C1685DDEE6510D8D704AD3109830">Gloeer, Peter</mods:namePart>
<mods:affiliation id="A6B1DFB2CFFAFF82559ADE80C395518C">Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:namePart id="84E0B56E390B3A4D35E8BC11E19C3214">Guerlek, Mustafa E</mods:namePart>
<mods:affiliation id="F567A791BCBBDEE8DBEB8E8DEA3BEFAB">Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:namePart id="785F61B290450F9CF5008E2B37AB8F39">Albrecht, Christian</mods:namePart>
<mods:affiliation id="016186CE0F9B01C31AED557AE72C1914">Justus Liebig University Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="BF55332E43D43D63B31BDC8AFA788E91">Neubauer, Thomas A</mods:namePart>
<mods:affiliation id="BA51864ED7AC7DC6C8B3AFDA41C4AB05">Justus Liebig University Giessen, Giessen, Germany &amp; Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<mods:date id="A2D5694A094F51ACA09CFA89FF96AC35">2020</mods:date>
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<mods:number id="801A5EFD2203FD9261AF08854D16622E">96</mods:number>
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<treatment id="0B43E9776B3A5F6A926554482B2B4A6F" ID-GBIF-Taxon="161939053" LSID="urn:lsid:plazi:treatment:0B43E9776B3A5F6A926554482B2B4A6F" httpUri="http://treatment.plazi.org/id/0B43E9776B3A5F6A926554482B2B4A6F" lastPageNumber="25" pageId="0" pageNumber="25" scope_class="Gastropoda" scope_family="Neritidae" scope_subFamily="Neritininae" scope_superFamily="Neritoidea">
<subSubSection id="AEB8F9022F034C3F3770556C2D507B97" pageId="0" pageNumber="25" type="nomenclature">
<paragraph id="4D154060EC4473533E360317BC17572C" pageId="0" pageNumber="25">
<taxonomicName id="02BF521EEFD537C3E0CFD7196011EACE" ID-CoL="56C5Y" LSID="0B43E977-6B3A-5F6A-9265-54482B2B4A6F" authority="Odabasi &amp; Arslan, 2015" authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">
Theodoxus gloeri
<normalizedToken id="4F11FE4F051B54AC52C5A4CF57EC545A" originalValue="Odabaşı">Odabasi</normalizedToken>
&amp; Arslan, 2015
</taxonomicName>
<figureCitation id="AF0580F69DC3002DCEF829C972A636B4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi &amp; Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">Figure 11A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D48F8839B2C8493B98955F967BD5E332" pageId="0" pageNumber="25" type="reference_group">
<paragraph id="ED6F4449654237DE9712262B0967C819" pageId="0" pageNumber="25">
<taxonomicName id="7BC2D0EECAADB3E1F83DFCE7BAE86FC0" authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">Theodoxus gloeri</taxonomicName>
<bibRefCitation id="D05833CC06C74F6EAD7852ABD9898B45" author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken id="F79FC383584F78E68D708D2E1B118419" originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
: 329-330, fig. 2;
<bibRefCitation id="C76BBB113AE38A8600E3398A587F36B0" author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken id="DFC5D2927B673C5B1B7C79F21542D6FE" originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993;
<bibRefCitation id="0AB915EE2D59086969B30E95AD5D9319" author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken id="0D5179874268852B190CA5E3CF556C05" originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
: 44, fig. 29.
</paragraph>
</subSubSection>
<subSubSection id="D69C38EDD6D79AA02826A75B0E604F68" pageId="0" pageNumber="25" type="type locality">
<paragraph id="BD0EA9779BD70F7F19305B7D37551985" pageId="0" pageNumber="25">
<typeStatus id="E93ABA7F897D7B6E0051B25C46807E9A">Type</typeStatus>
locality.
</paragraph>
<paragraph id="E57D7BA69A72616911A9CAF8669FEC87" pageId="0" pageNumber="25">
<normalizedToken id="EA67A2DB2D718F6D9F9135D606E3097F" originalValue="Balıkdamı">Balikdami</normalizedToken>
Wetland-Sakarya River,
<normalizedToken id="54C912B20C42388B5A3C2BF587569761" originalValue="Eskişehir">Eskisehir</normalizedToken>
, Turkey;
<geoCoordinate id="4F04C72E00E8E27E67609FC3FC9991A2" degrees="39.15277" direction="north" orientation="latitude" precision="1" value="39.15277">39.15277°N</geoCoordinate>
,
<geoCoordinate id="5F12F8986A4EE59BF9BCE125DD137012" degrees="31.61562" direction="east" orientation="longitude" precision="1" value="31.61562">31.61562°E</geoCoordinate>
(Fig.
<figureCitation id="2C5CFDF3CC3C1E000B2932E04F26161C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2749B04EB1C23F9A21E116472803288" pageId="0" pageNumber="25" type="type material">
<paragraph id="84433E1E6A48092776D586F1A8DE336C" pageId="0" pageNumber="25">
<typeStatus id="E2C77E58FB8F9404DD169B7BF0472DFB">Type</typeStatus>
material.
</paragraph>
<paragraph id="2BE9AF64BA8F171A6AA5413A0CD7045A" pageId="0" pageNumber="25">
<materialsCitation id="205FBC61675159B73551B9681606BD85" country="Turkey" typeStatus="Holotype">
<typeStatus id="2BFF16478C82708C8E56965CDB1C1C04">Holotype</typeStatus>
and seven
<typeStatus id="4717BB7E811AD4E55CE1BEB2EC25AD88">paratypes</typeStatus>
are deposited in COMULM (coll. no. COMULM-G 0052-0053)
</materialsCitation>
.
<materialsCitation id="3F741EECFE2DDB70DA189285707C2BFD" collectionCode="OGUHB" country="Turkey" typeStatus="paratypes">
Further nine
<typeStatus id="096D604278D3B48877C13CEBFEA57B3F">paratypes</typeStatus>
are deposited in
<collectionCode id="F5A6A8B3F11B7DD813E63D03BE0BC568" collectionName="OGUHB">OGUHB</collectionCode>
(coll. no. OGUHB-01759)
</materialsCitation>
.
<materialsCitation id="3C78FEFBF0DD6EEC7AF1C93C68B36C95" country="Turkey" typeStatus="paratype">
An additional
<typeStatus id="00AA65B8CB01771C7B3A3F5E91B7E19E">paratype</typeStatus>
is stored in the private collection of one of us (coll.
<normalizedToken id="8C2E32D2FE6D8975DACE9F4AF3E22AB3" originalValue="Glöer">Gloeer</normalizedToken>
; Fig.
<figureCitation id="72D33471D95B0DFA09BA99270B056285" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi &amp; Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">11A-C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7C214BF6E1CF74850FB0C27D8E8415A5" pageId="0" pageNumber="25" type="remarks">
<paragraph id="31AD78D1E53066FDB9F264D345E52572" pageId="0" pageNumber="25">Remarks.</paragraph>
<paragraph id="2C1071080572F5A1557370FAA5744BED" pageId="0" pageNumber="25">
<taxonomicName id="C28F3EE2CC40FFE46A30B62A414078D0" authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">
<emphasis id="9A5E4C92A2E5DC83D591813E45F5F1F2" italics="true" pageId="0" pageNumber="25">Theodoxus gloeri</emphasis>
</taxonomicName>
<normalizedToken id="E6699338633FE38F0EA9AFEF0387299A" originalValue="Odabaşı">Odabasi</normalizedToken>
&amp; Arslan, 2015 shows strong similarity to the fossil species
<taxonomicName id="5C4E93B9F7506C4D4D16E600EA9B8496" lsidName="T. pilidei" pageId="0" pageNumber="25" rank="species" species="pilidei">
<emphasis id="3E0E57153CE30AF7EE63557011B579D5" italics="true" pageId="0" pageNumber="25">T. pilidei</emphasis>
</taxonomicName>
(
<normalizedToken id="94437439EA9115639C27939CF275C4B3" originalValue="Tournouër">Tournouer</normalizedToken>
, 1879) from Romania and
<taxonomicName id="303753123AFACCC2CA5A73AA05255516" lsidName="T. lamelliferus" pageId="0" pageNumber="25" rank="species" species="lamelliferus">
<emphasis id="C99799512F02462DF4FBCDEE54302B7A" italics="true" pageId="0" pageNumber="25">T. lamelliferus</emphasis>
</taxonomicName>
(Milaschewitsch, 1912) from the Black Sea (near Alushta, Crimea) with strongly ribbed shells, but it is smaller than both (
<bibRefCitation id="8040EE46C1CB9799EF49187D66D79CC6" author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken id="6E47868C1E0C65802090216D36841FB7" originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
;
<bibRefCitation id="789B2676EDD31B6D5EC65BD88FBA7FFD" author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken id="B4E58CBF68AA53183F40771AD69408AA" originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
; Fig.
<figureCitation id="B9419BD2F664C34B854598BB44454442" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi &amp; Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">11</figureCitation>
). Although it is expected to be an extant subterranean species, no living material has ever been recorded and its phylogenetic position is presently unknown (
<bibRefCitation id="6F6A1AE9434963BA722518567CD47303" author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken id="EBADB0E4D862653B32CDACFA1F000A0C" originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
).
</paragraph>
<caption id="7776FDA7C189E8FB7748E62AF0231F7E" doi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25" start="Figure 11" startId="F11">
<paragraph id="E4D974A414EE2B60FD741D1A0F0D95EE" pageId="0" pageNumber="25">
<emphasis id="83E6B976743DB9DBC3D134DBCCB34A20" bold="true" pageId="0" pageNumber="25">Figure 11.</emphasis>
<taxonomicName id="02AC09D37E7E2B73FFED5FAB4F82442F" authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">
<emphasis id="808CC6C2160BDA61469B787B181D672A" italics="true" pageId="0" pageNumber="25">Theodoxus gloeri</emphasis>
</taxonomicName>
<normalizedToken id="7B8730809455E2C7193E118DCF82F9BA" originalValue="Odabaşı">Odabasi</normalizedToken>
&amp; Arslan, 2015.
<emphasis id="C1A7210AD74719A26FC94B1B60EF1F88" bold="true" pageId="0" pageNumber="25">A</emphasis>
-
<emphasis id="176D1200B3342FAEAE81FEA19297518E" bold="true" pageId="0" pageNumber="25">C</emphasis>
.
<typeStatus id="382C3B811FEF43FE341E7034C8AA467C">Paratype</typeStatus>
from
<normalizedToken id="941F77FF47F2B66484AB10654925E38F" originalValue="Balıkdamı">Balikdami</normalizedToken>
Wetland-Sakarya River,
<normalizedToken id="D8FCE64391D61331F1D2BBA7B53953ED" originalValue="Eskişehir">Eskisehir</normalizedToken>
, Turkey (coll.
<normalizedToken id="C2E73810DB6DA7E8A675EA86984D93AF" originalValue="Glöer">Gloeer</normalizedToken>
). Scale bar:
<quantity id="DA8C359026D5ED82EBFE72D944518BB5" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8713305EE3E0C315AFE840CC6B0BA09E" pageId="0" pageNumber="25" type="distribution">
<paragraph id="A6BCCB872629E7DE4F239E4F4B081F76" pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph id="B5D13E8A9D3ADC69DCCE5416CAFB0175" pageId="0" pageNumber="25">
Only known from two sites in very close proximity within the
<normalizedToken id="42A10A61DC954FB021920262C6B65C41" originalValue="Balıkdamı">Balikdami</normalizedToken>
Wetland-Sakarya River,
<normalizedToken id="1E1BF9EB74BB38D4B7CCCB7B3A23E7CE" originalValue="Eskişehir">Eskisehir</normalizedToken>
, Turkey (
<bibRefCitation id="1E2C0D2637B4C2E9A0694EA8ADC0618B" author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken id="B58DA56871466B1A2E902558DA3206C8" originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
; Fig.
<figureCitation id="A5EB9027DBFAF1ACF8785C37E972F60E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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