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<document id="5C3A298F0390585153B555F31310F34C" ID-CLB-Dataset="25313" ID-DOI="10.11646/zootaxa.4679.3.7" ID-GBIF-Dataset="0a75e36a-4254-44be-9a1a-2c2bc6f115fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471264" ID-ZooBank="1FF46982-622E-4C27-AB8E-8F8A993A419A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099354004" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03F287B9FFDE1B1EFF10FA09094E20E2" docLanguage="en" docName="zootaxa.4679.3.7.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Microscydmus) okinawanus Jałoszyński 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="541" masterDocId="FFCBFFC1FFDF1B1CFF87FFF80864257D" masterDocTitle="Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="552" masterPageNumber="539" pageNumber="540" updateTime="1698757590553" updateUser="ExternalLinkService">
<mods:mods id="1240196C1F3435493000BC9FE6FE82AA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="75B1EFF70440B6EDA78BAC04A5A67AEB">
<mods:title id="455524475014A555B1640352E5348942">Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="DBDC7F6A5FEBF4F4773E25E2DE231B4F">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="245F9DD6ACA22B09F11F01C40697ACCA">Zootaxa</mods:title>
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<mods:part id="9F7B0FF41B023C6E7A9D4CA7DE114BA4">
<mods:date id="34DFECE3E97C13D1F2894CCF7CCC2DF9">2019</mods:date>
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<mods:number id="AC4139660E8E3FE883CE5F06DB76B86F">2019-10-03</mods:number>
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<mods:number id="D5E3695F78C4F397D7DF5BD7FBB2E99A">4679</mods:number>
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<mods:classification id="B15C0AD964305D24FBC59315EBFB1210">journal article</mods:classification>
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<mods:identifier id="B412D07704CA7F7A291902FB3795BCE0" type="DOI">10.11646/zootaxa.4679.3.7</mods:identifier>
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</mods:mods>
<treatment id="03F287B9FFDE1B1EFF10FA09094E20E2" ID-GBIF-Taxon="159628630" LSID="urn:lsid:plazi:treatment:03F287B9FFDE1B1EFF10FA09094E20E2" httpUri="http://treatment.plazi.org/id/03F287B9FFDE1B1EFF10FA09094E20E2" lastPageId="2" lastPageNumber="541" pageId="1" pageNumber="540">
<subSubSection id="C3416524FFDE1B1DFF10FA090A922376" box="[151,758,1521,1548]" pageId="1" pageNumber="540" type="nomenclature">
<paragraph id="8BE436AFFFDE1B1DFF10FA090A922376" blockId="1.[151,758,1521,1583]" box="[151,758,1521,1548]" pageId="1" pageNumber="540">
<heading id="D0AC81C3FFDE1B1DFF10FA090A922376" bold="true" box="[151,758,1521,1548]" fontSize="11" level="1" pageId="1" pageNumber="540" reason="1">
<emphasis id="B92FEABDFFDE1B1DFF10FA090A922376" bold="true" box="[151,758,1521,1548]" pageId="1" pageNumber="540">
<taxonomicName id="4C5B4D2CFFDE1B1DFF10FA090AC82371" ID-CoL="8Y6FH" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,684,1521,1548]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="species" species="okinawanus" status="sp. nov." subGenus="Microscydmus">
<emphasis id="B92FEABDFFDE1B1DFF10FA0909232371" bold="true" box="[151,327,1521,1548]" italics="true" pageId="1" pageNumber="540">Microscydmus</emphasis>
(
<emphasis id="B92FEABDFFDE1B1DFEDFFA090A6C2371" bold="true" box="[344,520,1521,1548]" italics="true" pageId="1" pageNumber="540">Microscydmus</emphasis>
)
<emphasis id="B92FEABDFFDE1B1DFD9EFA090AC82371" bold="true" box="[537,684,1521,1548]" italics="true" pageId="1" pageNumber="540">okinawanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57C6FFDE1B1DFD34FA090A922376" box="[691,758,1521,1547]" pageId="1" pageNumber="540" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDE1B1DFF10F9ED093B2352" box="[151,351,1556,1583]" pageId="1" pageNumber="540" type="description">
<paragraph id="8BE436AFFFDE1B1DFF10F9ED093B2352" blockId="1.[151,758,1521,1583]" box="[151,351,1556,1583]" pageId="1" pageNumber="540">
(
<figureCitation id="13602A2AFFDE1B1DFF18F9ED08822352" box="[159,230,1557,1583]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="1" pageNumber="540">Figs 1</figureCitation>
,
<figureCitation id="13602A2AFFDE1B1DFF73F9ED09602352" box="[244,260,1557,1583]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="1" pageNumber="540">9</figureCitation>
,
<figureCitation id="13602A2AFFDE1B1DFE97F9ED094A2352" box="[272,302,1557,1583]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="1" pageNumber="540">17</figureCitation>
,
<figureCitation id="13602A2AFFDE1B1DFEBDF9EC09332352" box="[314,343,1556,1583]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="1" pageNumber="540">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDE1B1DFF10F9A50CC923C2" pageId="1" pageNumber="540" type="materials_examined">
<paragraph id="8BE436AFFFDE1B1DFF10F9A50CC923C2" blockId="1.[151,1436,1628,2015]" pageId="1" pageNumber="540">
<emphasis id="B92FEABDFFDE1B1DFF10F9A50927230A" bold="true" box="[151,323,1628,1655]" pageId="1" pageNumber="540">Type material.</emphasis>
<materialsCitation id="3B333CF2FFDE1B1DFECFF9A50CC923C2" ID-GBIF-Occurrence="2423812459" collectingDate="2003-09-06" collectingDateMax="2003-09-09" collectingDateMin="2003-09-06" collectionCode="JAPAN, MICROSCYD-, NSMT" collectorName="S. &amp; K. Arai" country="Japan" location="Okinawa Pref." pageId="1" pageNumber="540" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="B92FEABDFFDE1B1DFECFF9A50A8F230A" bold="true" box="[328,747,1628,1655]" pageId="1" pageNumber="540">
<typeStatus id="54E0880DFFDE1B1DFECFF9A509DD230A" box="[328,441,1629,1655]" pageId="1" pageNumber="540" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34C763FFFDE1B1DFE43F9A40A7A230A" box="[452,542,1628,1655]" name="Japan" pageId="1" pageNumber="540">JAPAN</collectingCountry>
(
<collectingRegion id="499FF84DFFDE1B1DFDADF9A40ABA230A" box="[554,734,1628,1655]" country="Japan" name="Okinawa" pageId="1" pageNumber="540">Okinawa Pref.</collectingRegion>
):
</emphasis>
<specimenCount id="9D5DFD26FFDE1B1DFD77F9A50B63230A" box="[752,775,1629,1655]" pageId="1" pageNumber="540" type="male">♂</specimenCount>
, two labels: &quot;
<collectionCode id="ED4AAE6AFFDE1B1DFC10F9A50B96230A" box="[919,1010,1629,1655]" pageId="1" pageNumber="540">JAPAN</collectionCode>
,
<collectingRegion id="499FF84DFFDE1B1DFC7BF9A50CDD230A" box="[1020,1209,1629,1655]" country="Japan" name="Okinawa" pageId="1" pageNumber="540">OKINAWA Pref.</collectingRegion>
/
<collectingRegion id="499FF84DFFDE1B1DFB4CF9A50D29230A" box="[1227,1357,1629,1655]" country="Japan" name="Okinawa" pageId="1" pageNumber="540">OKINAWA</collectingRegion>
Is. FIT / OKINAWASHI,
<date id="FFE5106FFFDE1B1DFEE0F979099623E6" box="[359,498,1665,1691]" pageId="1" pageNumber="540" value="2003-09-06" valueMax="2003-09-09" valueMin="2003-09-06">
<collectingDate id="EFA1E987FFDE1B1DFEE0F979099623E6" box="[359,498,1665,1691]" pageId="1" pageNumber="540" value="2003-09-06" valueMax="2003-09-09" valueMin="2003-09-06">6-9 IX 2003</collectingDate>
</date>
/ CHIBANAGUSUKU-ATO /
<collectorName id="26AE5379FFDE1B1DFCD1F9790B8223E6" box="[854,998,1665,1691]" pageId="1" pageNumber="540">S. &amp; K. Arai</collectorName>
leg.&quot; [white, printed]; &quot;
<emphasis id="B92FEABDFFDE1B1DFB71F97908B523C2" italics="true" pageId="1" pageNumber="540">
<collectionCode id="ED4AAE6AFFDE1B1DFB71F9790DF823E6" box="[1270,1436,1665,1691]" pageId="1" pageNumber="540">MICROSCYD-</collectionCode>
MUS
</emphasis>
(s. str.) /
<emphasis id="B92FEABDFFDE1B1DFEBCF95D09A523C2" box="[315,449,1701,1727]" italics="true" pageId="1" pageNumber="540">okinawanus</emphasis>
m. / P.
<bibRefCitation id="EFCA4B5EFFDE1B1DFD93F95D0ABF23C2" author="Jaloszynski, P." box="[532,731,1701,1727]" pageId="1" pageNumber="540" pagination="205 - 220" refId="ref9055" refString="Jaloszynski, P. (2019) Taxonomy of the &quot; Afroeudesis group &quot; of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4612 (2), 205 - 220. https: // doi. org / 10.11646 / zootaxa. 4612.2.4" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
/
<typeStatus id="54E0880DFFDE1B1DFD76F95D0BFF23C2" box="[753,923,1701,1727]" pageId="1" pageNumber="540" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED4AAE6AFFDE1B1DFBCAF95D0CC523C2" box="[1101,1185,1701,1727]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="1" pageNumber="540">NSMT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDE1B1DFF40F93009DE22EA" pageId="1" pageNumber="540" type="diagnosis">
<paragraph id="8BE436AFFFDE1B1DFF40F93009DE22EA" blockId="1.[151,1436,1628,2015]" pageId="1" pageNumber="540">
<emphasis id="B92FEABDFFDE1B1DFF40F9300926239E" bold="true" box="[199,322,1736,1763]" pageId="1" pageNumber="540">Diagnosis.</emphasis>
Body length ~
<quantity id="4CA39B4AFFDE1B1DFE6BF9310A20239E" box="[492,580,1737,1763]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="540" unit="mm" value="0.9">0.9 mm</quantity>
; posteromedian portion of vertex with punctures larger and denser than those on surrounding areas; bristles on tempora and vertex short and sparse, barely discernible; protibia strongly thickened in apical region, with apical comb of ten robust spines that extend from apical onto mesal margin of tibia, the most proximal spine longer and of different shape than the penultimate one; aedeagus with a pair of subapical arcuate structures connected at middle and with strongly elongate lateral components flanking a long tubular structure; parameres nearly straight.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDE1B1EFF40F85809FA2692" lastPageId="2" lastPageNumber="541" pageId="1" pageNumber="540" type="description">
<paragraph id="8BE436AFFFDE1B1DFF40F858096622A2" blockId="1.[151,1436,1628,2015]" pageId="1" pageNumber="540">
<emphasis id="B92FEABDFFDE1B1DFF40F858093D22C6" bold="true" box="[199,345,1952,1979]" pageId="1" pageNumber="540">Description.</emphasis>
Body of male (
<figureCitation id="13602A2AFFDE1B1DFD8CF8590A2822C6" box="[523,588,1953,1979]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="1" pageNumber="540">Fig. 1</figureCitation>
) slender, slightly flattened, pigmentation brown, vestiture light brown; BL
<quantity id="4CA39B4AFFDE1B1DFF10F83D089A22A2" box="[151,254,1989,2015]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="540" unit="mm" value="0.88">0.88 mm</quantity>
.
</paragraph>
<paragraph id="8BE436AFFFDD1B1EFF40FF610AD924F6" blockId="2.[151,1437,153,1439]" pageId="2" pageNumber="541">
Head (
<figureCitation id="13602A2AFFDD1B1EFE92FF61093825CE" box="[277,348,153,179]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="2" pageNumber="541">Fig. 9</figureCitation>
) broadest at eyes, HL
<quantity id="4CA39B4AFFDD1B1EFDECFF610AB125CE" box="[619,725,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="2" pageNumber="541" unit="mm" value="0.18">0.18 mm</quantity>
, HW
<quantity id="4CA39B4AFFDD1B1EFC9AFF610BEC25CE" box="[797,904,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="541" unit="mm" value="0.2">0.20 mm</quantity>
; tempora weakly arcuate, slightly longer than eyes; posterior margin of vertex straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex covered with small and shallow but dense and distinct punctures, which on posteromesal portion of vertex are distinctly larger than those on surrounding areas; setae sparse, short and suberect, bristles on tempora and vertex short and sparse; vertex with a pair of long posterolateral setae. Antennae slender, AnL
<quantity id="4CA39B4AFFDD1B1EFCE7FEB40BA3241A" box="[864,967,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="541" unit="mm" value="0.35">0.35 mm</quantity>
, antennomeres I and II strongly elongate, III–X each transverse, XI about as long as broad.
</paragraph>
<paragraph id="8BE436AFFFDD1B1EFF40FE6D099C2742" blockId="2.[151,1437,153,1439]" pageId="2" pageNumber="541">
Pronotum in dorsal view nearly circular, broadest at middle, PL
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, PW
<quantity id="4CA39B4AFFDD1B1EFBBEFE6D0CC424D2" box="[1081,1184,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="541" unit="mm" value="0.23">0.23 mm</quantity>
; anterior and posterior margins weakly arcuate, sides strongly rounded, anterior and posterior pronotal corners indistinct, obtuse-angled; pronotal base with a well-defined, short transverse groove, additionally with one tiny pit near each hind corner. Punctures on disc fine and inconspicuous, smaller and shallower than those on head; setae sparse, short, suberect; lateral bristles short and sparse.
</paragraph>
<paragraph id="8BE436AFFFDD1B1EFF40FDB10A4127B2" blockId="2.[151,1437,153,1439]" pageId="2" pageNumber="541">
Elytra together oval, broadest slightly in front of middle, EL
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, EW
<quantity id="4CA39B4AFFDD1B1EFBAAFDB10CF1271E" box="[1069,1173,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="541" unit="mm" value="0.33">0.33 mm</quantity>
, EI 1.46; humeral calli distinct; elytral base with large and deep basal impression; narrow adsutural region in anterior half of each elytron slightly impressed; apices of elytra separately rounded. Punctures about as indistinct as those on pronotum; setae short, sparse and nearly recumbent.
</paragraph>
<paragraph id="8BE436AFFFDD1B1EFF40FD210A4B2646" blockId="2.[151,1437,153,1439]" pageId="2" pageNumber="541">
Protibia (
<figureCitation id="13602A2AFFDD1B1EFEB5FD2109E3278E" box="[306,391,729,755]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="2" pageNumber="541">Fig. 17</figureCitation>
) strongly thickened in apical region, broadest at apex, with apical comb composed of ten robust teeth that occupy apical margin and extend onto mesal margin, the most basal spine strongly differs in length and shape from the penultimate one.
</paragraph>
<paragraph id="8BE436AFFFDD1B1EFF40FCBD092026B6" blockId="2.[151,1437,153,1439]" pageId="2" pageNumber="541">
Aedeagus (
<figureCitation id="13602A2AFFDD1B1EFEC3FCBD09F02622" box="[324,404,836,863]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="2" pageNumber="541">Fig. 25</figureCitation>
) relatively stout, AeL
<quantity id="4CA39B4AFFDD1B1EFD0BFCBD0A952622" box="[652,753,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="541" unit="mm" value="0.14">0.14 mm</quantity>
, endophallus with long median tubular component flanked by a pair of shorter elongate and nearly symmetrical sclerites in submedian region of median lobe, in subapical region the tubular component connected to a pair of long, arcuate, transverse sclerites; parameres nearly straight, each with one apical seta.
</paragraph>
<paragraph id="8BE436AFFFDD1B1EFF40FC2D09FA2692" blockId="2.[151,1437,153,1439]" box="[199,414,981,1007]" pageId="2" pageNumber="541">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDD1B1EFF40FC000A39216E" box="[199,605,1016,1043]" pageId="2" pageNumber="541" type="distribution">
<paragraph id="8BE436AFFFDD1B1EFF40FC000A39216E" blockId="2.[151,1437,153,1439]" box="[199,605,1016,1043]" pageId="2" pageNumber="541">
<emphasis id="B92FEABDFFDD1B1EFF40FC000905216E" bold="true" box="[199,353,1016,1043]" pageId="2" pageNumber="541">Distribution.</emphasis>
<collectingCountry id="F34C763FFFDD1B1EFEE0FC0109CE216E" box="[359,426,1017,1043]" name="Japan" pageId="2" pageNumber="541">Japan</collectingCountry>
, Okinawa-jima.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDD1B1EFF40FBE50BE2214A" box="[199,902,1053,1079]" pageId="2" pageNumber="541" type="etymology">
<paragraph id="8BE436AFFFDD1B1EFF40FBE50BE2214A" blockId="2.[151,1437,153,1439]" box="[199,902,1053,1079]" pageId="2" pageNumber="541">
<emphasis id="B92FEABDFFDD1B1EFF40FBE5092B214A" bold="true" box="[199,335,1053,1079]" pageId="2" pageNumber="541">Etymology.</emphasis>
Locotypical, after
<collectingRegion id="499FF84DFFDD1B1EFDA3FBE50AED214A" box="[548,649,1053,1079]" country="Japan" name="Okinawa" pageId="2" pageNumber="541">Okinawa</collectingRegion>
Island and Prefecture.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDD1B1EFF40FBB9094E20E2" pageId="2" pageNumber="541" type="discussion">
<paragraph id="8BE436AFFFDD1B1EFF40FBB9094E20E2" blockId="2.[151,1437,153,1439]" pageId="2" pageNumber="541">
<emphasis id="B92FEABDFFDD1B1EFF40FBB9095F2126" bold="true" box="[199,315,1089,1115]" pageId="2" pageNumber="541">Remarks.</emphasis>
Among Japanese species,
<taxonomicName id="4C5B4D2CFFDD1B1EFDE0FBBA0B762126" authorityName="Jałoszyński" authorityYear="2019" box="[615,786,1089,1115]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="B92FEABDFFDD1B1EFDE0FBBA0B762126" box="[615,786,1089,1115]" italics="true" pageId="2" pageNumber="541">M. okinawanus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C5B4D2CFFDD1B1EFC64FBBA0CF02126" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[995,1172,1089,1115]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="nishimeanus" status="sp. nov.">
<emphasis id="B92FEABDFFDD1B1EFC64FBBA0CF02126" box="[995,1172,1089,1115]" italics="true" pageId="2" pageNumber="541">M. nishimeanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57C6FFDD1B1EFB1CFBB90CB32126" box="[1179,1239,1089,1115]" pageId="2" pageNumber="541" rank="species">sp. n.</taxonomicNameLabel>
described below. They share the protibiae strongly and abruptly thickened in distal region (either not thickened or thickened gradually, not abruptly, in all remaining Far Eastern
<taxonomicName id="4C5B4D2CFFDD1B1EFDFDFB710B7921DE" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[634,797,1161,1187]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFDD1B1EFDFDFB710B7921DE" box="[634,797,1161,1187]" italics="true" pageId="2" pageNumber="541">Microscydmus</emphasis>
</taxonomicName>
); they also have a similar general body form, relatively indistinct punctures on the pronotum and elytra, but distinct on the vertex, short and sparse bristles on the tempora and vertex, and similar endophallic structures. The apical protibial comb in
<taxonomicName id="4C5B4D2CFFDD1B1EFC5EFB2A0CE72196" authorityName="Jałoszyński" authorityYear="2019" box="[985,1155,1233,1259]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="B92FEABDFFDD1B1EFC5EFB2A0CE72196" box="[985,1155,1233,1259]" italics="true" pageId="2" pageNumber="541">M. okinawanus</emphasis>
</taxonomicName>
is composed of 10 robust teeth that occupy not only the apical margin, but extend onto the mesal margin, and the most basal spine or tooth of the comb strongly differs in length and shape from the penultimate one. In
<taxonomicName id="4C5B4D2CFFDD1B1EFC65FAE20CF0204E" authorityName="Jałoszyński" authorityYear="2019" box="[994,1172,1305,1331]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="nishimeanus">
<emphasis id="B92FEABDFFDD1B1EFC65FAE20CF0204E" box="[994,1172,1305,1331]" italics="true" pageId="2" pageNumber="541">M. nishimeanus</emphasis>
</taxonomicName>
, the comb is composed of only six teeth that do not extend onto the mesal tibial margin, and the most basal tooth is inconspicuous, of the same shape and length as the penultimate one.
<taxonomicName id="4C5B4D2CFFDD1B1EFD61FA990C7D2006" authorityName="Jałoszyński" authorityYear="2019" box="[742,1049,1377,1403]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="B92FEABDFFDD1B1EFD61FA990C7D2006" box="[742,1049,1377,1403]" italics="true" pageId="2" pageNumber="541">Microscydmus okinawanus</emphasis>
</taxonomicName>
is also distinctly smaller than
<taxonomicName id="4C5B4D2CFFDD1B1EFAF9FA9A094720E2" authorityName="Jałoszyński" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="nishimeanus">
<emphasis id="B92FEABDFFDD1B1EFAF9FA9A094720E2" italics="true" pageId="2" pageNumber="541">M. nishimeanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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