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<document id="72856D89AE630EE882A07642E2E9222E" ID-CLB-Dataset="25313" ID-DOI="10.11646/zootaxa.4679.3.7" ID-GBIF-Dataset="0a75e36a-4254-44be-9a1a-2c2bc6f115fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471264" ID-ZooBank="1FF46982-622E-4C27-AB8E-8F8A993A419A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099354004" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03F287B9FFD91B1BFF10FC2D093D278E" docLanguage="en" docName="zootaxa.4679.3.7.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Microscydmus) elytratus Jałoszyński 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="546" masterDocId="FFCBFFC1FFDF1B1CFF87FFF80864257D" masterDocTitle="Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="552" masterPageNumber="539" pageNumber="545" updateTime="1698757590553" updateUser="ExternalLinkService">
<mods:mods id="EF0D4F86E4706DA2160CD967CCAE7062" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B61F67EA51F587B2835BDF35F199CE24">
<mods:title id="15ADF0817F61183ECFDB9ED115A7FDC1">Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="3FEA51F66AC5D2DBFF5683E47C2D42C7" type="personal">
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</mods:role>
<mods:namePart id="9B93C61C18F3E6CAF49018D1DD6E14A1">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="6C547120D371B54984E14AC2CBAD2583">text</mods:typeOfResource>
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<mods:title id="B1EEF12DEA740FEA730CFC32F77642C5">Zootaxa</mods:title>
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<mods:part id="20B694BC94EE2AA700682AA39012344C">
<mods:date id="289E6DA9B286942395A758C1139C8CAD">2019</mods:date>
<mods:detail id="01337DDF162CF0224C0D80BB2550BBB6" type="pubDate">
<mods:number id="4D5881D3531838BABAB655E4B32DB4A2">2019-10-03</mods:number>
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<mods:number id="D0E38FB9DD2D12E0925546F3047C8C37">4679</mods:number>
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<mods:number id="E3B82C6347DE01DA3A930574C7FC83D1">3</mods:number>
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<mods:classification id="B0DBDAFD7ABF704F157C95B885158C0D">journal article</mods:classification>
<mods:identifier id="F6B5BA18D01F3A87521E84E24CA8CFFA" type="CLB-Dataset">25313</mods:identifier>
<mods:identifier id="2F831DFD33710273BD680E46A4B6B14C" type="DOI">10.11646/zootaxa.4679.3.7</mods:identifier>
<mods:identifier id="23E220F7005E015FCE2C086526CC3C8B" type="GBIF-Dataset">0a75e36a-4254-44be-9a1a-2c2bc6f115fa</mods:identifier>
<mods:identifier id="5014701A5ED56F677B595687AE39E939" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03F287B9FFD91B1BFF10FC2D093D278E" ID-DOI="http://doi.org/10.5281/zenodo.5925430" ID-GBIF-Taxon="159628634" ID-Zenodo-Dep="5925430" LSID="urn:lsid:plazi:treatment:03F287B9FFD91B1BFF10FC2D093D278E" httpUri="http://treatment.plazi.org/id/03F287B9FFD91B1BFF10FC2D093D278E" lastPageId="7" lastPageNumber="546" pageId="6" pageNumber="545">
<subSubSection id="C3416524FFD91B1AFF10FC2D0AAF2692" box="[151,715,981,1008]" pageId="6" pageNumber="545" type="nomenclature">
<paragraph id="8BE436AFFFD91B1AFF10FC2D0AAF2692" blockId="6.[151,715,981,1043]" box="[151,715,981,1008]" pageId="6" pageNumber="545">
<heading id="D0AC81C3FFD91B1AFF10FC2D0AAF2692" bold="true" box="[151,715,981,1008]" fontSize="11" level="1" pageId="6" pageNumber="545" reason="1">
<emphasis id="B92FEABDFFD91B1AFF10FC2D0AAF2692" bold="true" box="[151,715,981,1008]" pageId="6" pageNumber="545">
<taxonomicName id="4C5B4D2CFFD91B1AFF10FC2D0AE5268D" ID-CoL="8Y6FC" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,641,981,1008]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="elytratus" status="sp. nov." subGenus="Microscydmus">
<emphasis id="B92FEABDFFD91B1AFF10FC2D0923268D" bold="true" box="[151,327,981,1008]" italics="true" pageId="6" pageNumber="545">Microscydmus</emphasis>
(
<emphasis id="B92FEABDFFD91B1AFEDFFC2D0A6C268D" bold="true" box="[344,520,981,1008]" italics="true" pageId="6" pageNumber="545">Microscydmus</emphasis>
)
<emphasis id="B92FEABDFFD91B1AFD9EFC2D0AE5268D" bold="true" box="[537,641,981,1008]" italics="true" pageId="6" pageNumber="545">elytratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57C6FFD91B1AFD0EFC2D0AAF2692" box="[649,715,981,1007]" pageId="6" pageNumber="545" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1AFF10FC010909216E" box="[151,365,1016,1043]" pageId="6" pageNumber="545" type="description">
<paragraph id="8BE436AFFFD91B1AFF10FC010909216E" blockId="6.[151,715,981,1043]" box="[151,365,1016,1043]" pageId="6" pageNumber="545">
(
<figureCitation id="13602A2AFFD91B1AFF18FC01088C216E" box="[159,232,1016,1043]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="6" pageNumber="545">Figs 5</figureCitation>
,
<figureCitation id="13602A2AFFD91B1AFF73FC010976216E" box="[244,274,1017,1043]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="6" pageNumber="545">13</figureCitation>
,
<figureCitation id="13602A2AFFD91B1AFE99FC010958216E" box="[286,316,1017,1043]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="6" pageNumber="545">21</figureCitation>
,
<figureCitation id="13602A2AFFD91B1AFECFFC010901216E" box="[328,357,1017,1043]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="6" pageNumber="545">29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1AFF10FBB90BD221BA" pageId="6" pageNumber="545" type="materials_examined">
<paragraph id="8BE436AFFFD91B1AFF10FBB90BD221BA" blockId="6.[151,1437,1088,2015]" pageId="6" pageNumber="545">
<emphasis id="B92FEABDFFD91B1AFF10FBB909232126" bold="true" box="[151,327,1088,1115]" pageId="6" pageNumber="545">Type material.</emphasis>
<materialsCitation id="3B333CF2FFD91B1AFEC8FBB90AFC21DE" ID-GBIF-Occurrence="2423812462" collectingDate="1999-05-22" collectingDateMax="1999-05-23" collectingDateMin="1999-05-22" collectionCode="JAPAN, EUMJ" collectorName="Pref. &amp; Tatsuya Kan" country="Japan" location="Kuroson" municipality="Nishitosa-Village" pageId="6" pageNumber="545" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B92FEABDFFD91B1AFEC8FBB90AB02126" bold="true" box="[335,724,1088,1115]" pageId="6" pageNumber="545">
<typeStatus id="54E0880DFFD91B1AFEC8FBB909A42126" box="[335,448,1089,1115]" pageId="6" pageNumber="545" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34C763FFFD91B1AFE4BFBB80A4C2126" box="[460,552,1088,1115]" name="Japan" pageId="6" pageNumber="545">JAPAN</collectingCountry>
(Kôchi
<collectorName id="26AE5379FFD91B1AFD00FBB80AA32126" box="[647,711,1088,1115]" pageId="6" pageNumber="545">Pref.</collectorName>
):
</emphasis>
<specimenCount id="9D5DFD26FFD91B1AFD5BFBB90A972126" box="[732,755,1089,1115]" pageId="6" pageNumber="545" type="male">♂</specimenCount>
, two labels: &quot;(Kôchi:
<collectionCode id="ED4AAE6AFFD91B1AFC6EFBB90C262126" box="[1001,1090,1089,1115]" pageId="6" pageNumber="545">JAPAN</collectionCode>
) /
<location id="8E846074FFD91B1AFBE7FBB90CA42126" LSID="urn:lsid:plazi:treatment:03F287B9FFD91B1BFF10FC2D093D278E:8E846074FFD91B1AFBE7FBB90CA42126" box="[1120,1216,1089,1115]" country="Japan" municipality="Nishitosa-Village" name="Kuroson" pageId="6" pageNumber="545">Kuroson</location>
/
<collectingMunicipality id="6B80ACD5FFD91B1AFB50FBB90DF92126" box="[1239,1437,1089,1115]" pageId="6" pageNumber="545">Nishitosa-Village</collectingMunicipality>
/
<date id="FFE5106FFFD91B1AFF20FB9D09302102" box="[167,340,1125,1151]" pageId="6" pageNumber="545" value="1999-05-22" valueMax="1999-05-23" valueMin="1999-05-22">
<collectingDate id="EFA1E987FFD91B1AFF20FB9D09302102" box="[167,340,1125,1151]" pageId="6" pageNumber="545" value="1999-05-22" valueMax="1999-05-23" valueMin="1999-05-22">22-23. V. 1999</collectingDate>
</date>
/
<collectorName id="26AE5379FFD91B1AFEEAFB9D09992102" box="[365,509,1125,1151]" pageId="6" pageNumber="545">Tatsuya Kan</collectorName>
leg.&quot; [white, printed]; &quot;
<taxonomicName id="4C5B4D2CFFD91B1AFC8AFB9D0B802102" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[781,996,1125,1151]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFD91B1AFC8AFB9D0B802102" box="[781,996,1125,1151]" italics="true" pageId="6" pageNumber="545">MICROSCYDMUS</emphasis>
</taxonomicName>
(s. str.) /
<emphasis id="B92FEABDFFD91B1AFBD2FB9D0CD32102" box="[1109,1207,1125,1151]" italics="true" pageId="6" pageNumber="545">elytratus</emphasis>
m. / P.
<bibRefCitation id="EFCA4B5EFFD91B1AFA93FB9D08AB21DE" author="Jaloszynski, P." pageId="6" pageNumber="545" pagination="205 - 220" refId="ref9055" refString="Jaloszynski, P. (2019) Taxonomy of the &quot; Afroeudesis group &quot; of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4612 (2), 205 - 220. https: // doi. org / 10.11646 / zootaxa. 4612.2.4" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
/
<typeStatus id="54E0880DFFD91B1AFF62FB7109EB21DE" box="[229,399,1161,1187]" pageId="6" pageNumber="545" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED4AAE6AFFD91B1AFDC7FB710AEB21DE" box="[576,655,1161,1187]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="6" pageNumber="545">EUMJ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B333CF2FFD91B1AFD25FB7108DC21BA" ID-GBIF-Occurrence="2423812458" collectingDate="1999-05-22" collectingDateMax="1999-05-23" collectingDateMin="1999-05-22" collectionCode="VI" collectorName="Pref. &amp; Tatsuya Kan" country="India" location="Kuroson" municipality="Nishitosa-Village" pageId="6" pageNumber="545" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="B92FEABDFFD91B1AFD25FB710BE621DE" bold="true" box="[674,898,1160,1187]" pageId="6" pageNumber="545">
<typeStatus id="54E0880DFFD91B1AFD25FB710B7E21DE" box="[674,794,1161,1187]" pageId="6" pageNumber="545" type="paratype">Paratypes</typeStatus>
(3 exx.):
</emphasis>
<specimenCount id="9D5DFD26FFD91B1AFC0FFB710BAC21DE" box="[904,968,1161,1187]" pageId="6" pageNumber="545" type="female">3 ♀♀</specimenCount>
, same data as for holotype, except for
<date id="FFE5106FFFD91B1AFAFEFB7108DC21BA" pageId="6" pageNumber="545" value="1999-06-21">
21.
<collectionCode id="ED4AAE6AFFD91B1AFF10FB5508DC21BA" box="[151,184,1197,1223]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="545">VI</collectionCode>
</date>
</materialsCitation>
. 1999; all
<typeStatus id="54E0880DFFD91B1AFEAAFB5509FC21BA" box="[301,408,1197,1223]" pageId="6" pageNumber="545" type="paratype">paratypes</typeStatus>
with yellow
<materialsCitation id="3B333CF2FFD91B1AFDACFB550BD621BA" ID-GBIF-Occurrence="2423812452" box="[555,946,1197,1223]" collectionCode="EUMJ" country="India" pageId="6" pageNumber="545" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E0880DFFD91B1AFDACFB550AAA21BA" box="[555,718,1197,1223]" pageId="6" pageNumber="545" type="paratype">PARATYPUS</typeStatus>
labels (cPJ,
<collectionCode id="ED4AAE6AFFD91B1AFCDBFB550BCE21BA" box="[860,938,1197,1223]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="6" pageNumber="545">EUMJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1AFF40FB280BBE2006" pageId="6" pageNumber="545" type="diagnosis">
<paragraph id="8BE436AFFFD91B1AFF40FB280BBE2006" blockId="6.[151,1437,1088,2015]" pageId="6" pageNumber="545">
<emphasis id="B92FEABDFFD91B1AFF40FB2809262196" bold="true" box="[199,322,1232,1259]" pageId="6" pageNumber="545">Diagnosis.</emphasis>
Body length ~
<quantity id="4CA39B4AFFD91B1AFE6DFB290A262196" box="[490,578,1233,1259]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="545" unit="mm" value="1.1">1.1 mm</quantity>
; entire frons and vertex with distinct but shallow and diffuse dense punctures; bristles on tempora and vertex moderately long and dense, well-visible; protibia gradually thickened distad, with apical comb of six teeth that occupy distomesal margin; elytra between middle and posterior third with an oval circumsutural patch of long and dense setae directed toward suture; aedeagus with conspicuous spherical vesicular structure in sub-basal region of median lobe; parameres strongly recurved.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1BFF40FA7C0AC62766" lastPageId="7" lastPageNumber="546" pageId="6" pageNumber="545" type="description">
<paragraph id="8BE436AFFFD91B1AFF40FA7C0B4120BE" blockId="6.[151,1437,1088,2015]" pageId="6" pageNumber="545">
<emphasis id="B92FEABDFFD91B1AFF40FA7C093D20E2" bold="true" box="[199,345,1412,1439]" pageId="6" pageNumber="545">Description.</emphasis>
Body of male (
<figureCitation id="13602A2AFFD91B1AFD86FA7D0A2620E2" box="[513,578,1412,1439]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="6" pageNumber="545">Fig. 5</figureCitation>
) moderately slender, slightly flattened, pigmentation moderately dark brown, vestiture light brown; BL
<quantity id="4CA39B4AFFD91B1AFE3DFA510A3920BE" box="[442,605,1449,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.115" metricValueMax="1.13" metricValueMin="1.1" pageId="6" pageNumber="545" unit="mm" value="1.115" valueMax="1.13" valueMin="1.1">1.10–1.13 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFD36FA510B7D20BE" box="[689,793,1449,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="6" pageNumber="545" unit="mm" value="1.11">1.11 mm</quantity>
).
</paragraph>
<paragraph id="8BE436AFFFD91B1AFF40FA350C3E23C2" blockId="6.[151,1437,1088,2015]" pageId="6" pageNumber="545">
Head (
<figureCitation id="13602A2AFFD91B1AFE93FA35090C209A" box="[276,360,1485,1511]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="6" pageNumber="545">Fig. 13</figureCitation>
) broadest at eyes, HL
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, HW
<quantity id="4CA39B4AFFD91B1AFCA4FA340BE9209A" box="[803,909,1484,1511]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="545" unit="mm" value="0.25">0.25 mm</quantity>
; tempora weakly arcuate, slightly longer than eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small and dense but shallow and diffuse punctures; setae sparse, short and suberect, bristles on tempora and vertex relatively short but well-visible; vertex with a pair of long posterolateral setae. Antennae slender, AnL
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(mean
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), antennomeres I and II strongly elongate, III indistinctly transverse, IV–VI each about as long as broad, VII–X each transverse, XI about as long as broad.
</paragraph>
<paragraph id="8BE436AFFFD91B1AFF40F9310A6522C6" blockId="6.[151,1437,1088,2015]" pageId="6" pageNumber="545">
Pronotum in dorsal view nearly circular, broadest slightly in front of middle, PL
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(mean
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), PW
<quantity id="4CA39B4AFFD91B1AFF40F9150949227A" box="[199,301,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="545" unit="mm" value="0.33">0.33 mm</quantity>
; anterior and posterior margins weakly arcuate; anterior corners not marked; sides strongly rounded in anterior half, in posterior half slightly sinuate; posterior pronotal corners distinct, obtuse-angled; pronotal base with a well-defined, short transverse groove deepened at each end, lateral pit near each hind corner present, but barely discernible, very small and obscured by dense punctures. Punctures on disc conspicuously dense, slightly larger than those on head, but very shallow and with diffuse margins; setae sparse, short, suberect; lateral bristles dense and relatively long, well visible.
</paragraph>
<paragraph id="8BE436AFFFD91B1BFF40F83D0DFE2462" blockId="6.[151,1437,1088,2015]" lastBlockId="7.[151,1437,153,755]" lastPageId="7" lastPageNumber="546" pageId="6" pageNumber="545">
Elytra together oval, broadest distinctly in front of middle, EL
<quantity id="4CA39B4AFFD91B1AFC00F83D0C4F22A2" box="[903,1067,1989,2015]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.3" metricValueMin="6.0" pageId="6" pageNumber="545" unit="mm" value="0.615" valueMax="0.63" valueMin="0.6">0.60–0.63 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFBF9F83D0C8322A2" box="[1150,1255,1989,2015]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="6" pageNumber="545" unit="mm" value="0.62">0.62 mm</quantity>
), EW
<quantity id="4CA39B4AFFD91B1BFAABF83C08A725CE" lastPageId="7" lastPageNumber="546" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="6" pageNumber="545" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43–0.45 mm</quantity>
(mean
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), EI 1.39–1.41; humeral calli distinct; elytral base with large and deep basal impression; adsutural area not impressed in anterior half; apices of elytra separately rounded; small oval circumsutural region between middle and posterior half slightly impressed and surrounded by dense, long setae directed toward suture. Surface of elytra finely shagreened, individual punctures not discernible; setae short, sparse and nearly recumbent.
</paragraph>
<paragraph id="8BE436AFFFD81B1BFF40FED10D0B243E" blockId="7.[151,1437,153,755]" box="[199,1391,297,323]" pageId="7" pageNumber="546">
Protibia (
<figureCitation id="13602A2AFFD81B1BFEA8FED109E5243E" box="[303,385,297,323]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="7" pageNumber="546">Fig. 21</figureCitation>
) gradually thickened distad, with apical comb of six teeth that occupy distomesal margin.
</paragraph>
<paragraph id="8BE436AFFFD81B1BFF40FEB5097024D2" blockId="7.[151,1437,153,755]" pageId="7" pageNumber="546">
Aedeagus (
<figureCitation id="13602A2AFFD81B1BFECDFEB509C4241A" box="[330,416,333,359]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="7" pageNumber="546">Fig. 29</figureCitation>
) moderately slender, AeL
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, endophallus with long median tubular component abruptly broadened in its proximal portion to form a spherical vesicle; parameres strongly sinuate, each with one apical seta.
</paragraph>
<paragraph id="8BE436AFFFD81B1BFF40FE410AC62766" blockId="7.[151,1437,153,755]" pageId="7" pageNumber="546">
Female. Externally similar to male, with weakly thickened protibiae lacking apical combs, and elytra lacking posteromesal patch of modified setae; BL
<quantity id="4CA39B4AFFD81B1BFDF5FE250ABE248A" box="[626,730,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="546" unit="mm" value="1.1">1.10 mm</quantity>
; HL
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, HW
<quantity id="4CA39B4AFFD81B1BFC3EFE240C44248A" box="[953,1056,476,503]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="546" unit="mm" value="0.25">0.25 mm</quantity>
, AnL
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; PL
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, PW
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; EL
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, EW
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, EI 1.41.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD81B1BFF40FDDC0AE62742" box="[199,642,548,575]" pageId="7" pageNumber="546" type="distribution">
<paragraph id="8BE436AFFFD81B1BFF40FDDC0AE62742" blockId="7.[151,1437,153,755]" box="[199,642,548,575]" pageId="7" pageNumber="546">
<emphasis id="B92FEABDFFD81B1BFF40FDDC09052742" bold="true" box="[199,353,548,575]" pageId="7" pageNumber="546">Distribution.</emphasis>
<collectingCountry id="F34C763FFFD81B1BFEE0FDDD09CE2742" box="[359,426,549,575]" name="Japan" pageId="7" pageNumber="546">Japan</collectingCountry>
, southern Shikoku.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD81B1BFF40FDB10C67271E" box="[199,1027,585,611]" pageId="7" pageNumber="546" type="etymology">
<paragraph id="8BE436AFFFD81B1BFF40FDB10C67271E" blockId="7.[151,1437,153,755]" box="[199,1027,585,611]" pageId="7" pageNumber="546">
<emphasis id="B92FEABDFFD81B1BFF40FDB1092B271E" bold="true" box="[199,335,585,611]" pageId="7" pageNumber="546">Etymology.</emphasis>
The adjective
<emphasis id="B92FEABDFFD81B1BFE72FDB10A33271E" box="[501,599,585,611]" italics="true" pageId="7" pageNumber="546">elytratus</emphasis>
refers to the modified elytra in males.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD81B1BFF40FD95093D278E" pageId="7" pageNumber="546" type="discussion">
<paragraph id="8BE436AFFFD81B1BFF40FD95093D278E" blockId="7.[151,1437,153,755]" pageId="7" pageNumber="546">
<emphasis id="B92FEABDFFD81B1BFF40FD95095F27FA" bold="true" box="[199,315,621,647]" pageId="7" pageNumber="546">Remarks.</emphasis>
This is a giant among Japanese species of
<taxonomicName id="4C5B4D2CFFD81B1BFC81FD950BCD27FA" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[774,937,621,647]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="546" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFD81B1BFC81FD950BCD27FA" box="[774,937,621,647]" italics="true" pageId="7" pageNumber="546">Microscydmus</emphasis>
</taxonomicName>
, the only one with adults distinctly exceeding
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in length. The elytral patch of modified setae is unique among Palaearctic species of this genus. Similarities and differences in relation to
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<emphasis id="B92FEABDFFD81B1BFE5BFD4E0A1227B2" box="[476,630,693,719]" italics="true" pageId="7" pageNumber="546">M. kochiensis</emphasis>
</taxonomicName>
, a species with the most similar aedeagus, were discussed in remarks for the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>
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