treatments-xml/data/03/F2/87/03F287996905FFB2FF34419FFA620809.xml

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<mods:title id="95A5D3D1FA80213468CB7A5F188B575C">Geometridae (Lepidoptera) of the Baikal region: identification keys and annotated catalogue with notes to DNA barcoding. Part 2. Archiearinae, Geometrinae, Sterrhinae</mods:title>
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<subSubSection id="C34165046905FFB1FF34419FFD5008DB" box="[151,686,1205,1232]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BE4368F6905FFB1FF34419FFD5008DB" blockId="20.[151,686,1205,1232]" box="[151,686,1205,1232]" pageId="20" pageNumber="21">
<emphasis id="B92FEA9D6905FFB1FF34419FFD5008DB" bold="true" box="[151,686,1205,1232]" pageId="20" pageNumber="21">
Key to 
<taxonomicName id="4C5B4D0C6905FFB1FF52419FFE4E08DB" ID-CoL="92MKK" ID-ENA="104443" authority="Schrank, 1802" box="[241,432,1205,1232]" class="Insecta" family="Geometridae" genus="Scopula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9D6905FFB1FF52419FFEAD08C4" bold="true" box="[241,339,1205,1232]" italics="true" pageId="20" pageNumber="21">Scopula</emphasis>
species
</taxonomicName>
of the Baikal region
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34165046905FFB1FF3441D7FDC1092F" pageId="20" pageNumber="21" type="description">
<paragraph id="8BE4368F6905FFB1FF3441D7FDC1092F" blockId="20.[151,1436,1277,1340]" pageId="20" pageNumber="21">
Identification key to 
<taxonomicName id="4C5B4D0C6905FFB1FEDD41D7FDCE0903" authority="Schrank, 1802" box="[382,560,1277,1303]" class="Insecta" family="Geometridae" genus="Scopula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9D6905FFB1FEDD41D7FE260903" box="[382,472,1277,1303]" italics="true" pageId="20" pageNumber="21">Scopula</emphasis>
species
</taxonomicName>
given below partly duplicates the key provided by 
<bibRefCitation id="EFCA4B7E6905FFB1FBC641D7FAD8090C" author="Viidalepp, Ya. R." box="[1125,1318,1277,1304]" pageId="20" pageNumber="21" pagination="443 - 470" refId="ref74514" refString="Viidalepp, Ya. R. (2005) Subfam. Sterrhinae (Scopulini). In: Ler, P. A. (Ed.), Key to the insects of Russian Far East. Vol. V. Trichoptera and Lepidoptera. Pt 5. Dal'nauka, Vladivostok, pp. 443 - 470. [in Russian]" type="book chapter" year="2005">Viidalepp (2005)</bibRefCitation>
. However, the present key also includes females.
</paragraph>
</subSubSection>
<subSubSection id="C34165046905FFB2FF34404CFA620809" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21" type="key">
<key id="FFCA61BE6905FFB2FF34404CFA620809" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<keyStep id="30AF2E2A6905FFB1FF34404CFA6209E5" pageId="20" pageNumber="21">
<paragraph id="8BE4368F6905FFB1FF34404CFA6309C0" blockId="20.[151,1437,1382,2020]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF34404CFA6309C0" pageId="20" pageNumber="21">
1. Antenna distinctly chequered with white and dark brown. Forewing with 4–5 wide ochreous brown transverse lines (
<figureCitation id="13602A0A6905FFB1FA99404CFA6F096F" box="[1338,1425,1382,1404]" captionStart="FIGURE 23" captionStartId="68.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@68.[151,1435,181,1836]" captionTargetPageId="68" captionText="FIGURE 23. Adults of Siberian Sterrhinae. A—Holarctias rufinularia; B—Scopula agutsaensis; C—S. immorata; D—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965143" httpUri="https://zenodo.org/record/7965143/files/figure.png" pageId="20" pageNumber="21">Fig. 23C</figureCitation>
). In male genitalia (
<figureCitation id="13602A0A6905FFB1FE3040A9FDD0098D" box="[403,558,1411,1433]" captionStart="FIGURE 42" captionStartId="87.[151,250,1867,1891]" captionTargetBox="[166,1418,193,1841]" captionTargetId="figure-15@87.[158,1430,181,1844]" captionTargetPageId="87" captionText="FIGURE 42. Male genitalia of Siberian Sterrhinae. A–C—Idaea descitaria; D–F—Holarctias rufinaria; G–I—H. rufinularia; J–L—Scopula agutsaensis; M–O—S. immorata. A, E, I, L, N—genital segment, ventral view; B—ditto, valvae shifted; C, D, H, K, O—phallos; G, F, J, M—sternite A8. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965198" httpUri="https://zenodo.org/record/7965198/files/figure.png" pageId="20" pageNumber="21">Figs. 42M–42O</figureCitation>
) vesica with two cornuti, left ceras significantly shorter than right one; in female genitalia (
<figureCitation id="13602A0A6905FFB1FF4A408BFEBF09A3" box="[233,321,1441,1463]" captionStart="FIGURE 51" captionStartId="96.[151,250,1867,1891]" captionTargetBox="[166,1421,193,1840]" captionTargetId="figure-25@96.[158,1429,181,1844]" captionTargetPageId="96" captionText="FIGURE 51. Female genitalia of Siberian Sterrhinae. A—Idaea straminata; B—I. dimidiata; C—I. descitaria; D—Holarctias rufinaria; E—H. rufinularia; F—Scopula agutsaensis; G—S. immorata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965222" httpUri="https://zenodo.org/record/7965222/files/figure.png" pageId="20" pageNumber="21">Fig. 51G</figureCitation>
) sterigma with wide lamella antevaginalis and semicircular clasp around ostium, signum developed as whole drop-shaped laterally spinulose plate.................................................. 
<taxonomicName id="4C5B4D0C6905FFB1FBD14094FA6309C0" ID-CoL="4VW3M" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1138,1437,1470,1492]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="immorata">
<emphasis id="B92FEA9D6905FFB1FBD14094FA6309C0" bold="true" box="[1138,1437,1470,1492]" pageId="20" pageNumber="21">
<emphasis id="B92FEA9D6905FFB1FBD14094FB1209C7" bold="true" box="[1138,1260,1470,1491]" italics="true" pageId="20" pageNumber="21">S. immorata</emphasis>
(Linnaeus, 1758)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6905FFB1FF3440F6FA6209E5" blockId="20.[151,1437,1382,2020]" box="[151,1436,1499,1521]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF3440F6FA6209E5" box="[151,1436,1499,1521]" pageId="20" pageNumber="21">- Antenna not chequered................................................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6905FFB1FF3440D3FA620A3F" pageId="20" pageNumber="21">
<paragraph id="8BE4368F6905FFB1FF3440D3FA620A1A" blockId="20.[151,1437,1382,2020]" box="[151,1436,1529,1551]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF3440D3FA620A1A" box="[151,1436,1529,1551]" pageId="20" pageNumber="21">2. Frons white.......................................................................................... 3</keyLead>
</paragraph>
<paragraph id="8BE4368F6905FFB1FF34433DFA620A3F" blockId="20.[151,1437,1382,2020]" box="[151,1436,1558,1580]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF34433DFA620A3F" box="[151,1436,1558,1580]" pageId="20" pageNumber="21">- Frons black or dark brown.............................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6905FFB1FF34431EFA630B03" pageId="20" pageNumber="21">
<paragraph id="8BE4368F6905FFB1FF34431EFA620AB5" blockId="20.[151,1437,1382,2020]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF34431EFA620AB5" pageId="20" pageNumber="21">
3. Forewing with distinct dark dentate postmedial line which bordered along the entire length by broad dark spots (
<figureCitation id="13602A0A6905FFB1FABD4319FA860A5D" box="[1310,1400,1587,1609]" captionStart="FIGURE 29" captionStartId="74.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@74.[151,1435,181,1836]" captionTargetPageId="74" captionText="FIGURE 29. Adults of Siberian Sterrhinae. A—Scopula aequifasciata; B—S. cajanderi; C—S. albiceraria; D—Cyclophora albipunctata." figureDoi="http://doi.org/10.5281/zenodo.7965161" httpUri="https://zenodo.org/record/7965161/files/figure.png" pageId="20" pageNumber="21">Fig. 29A</figureCitation>
). In male genitalia sacculus strongly sclerotized and pointed, socii almost as long as sacculus; phallos narrow; right ceras longer than left one (
<figureCitation id="13602A0A6905FFB1FEC54344FE000A97" box="[358,510,1646,1668]" captionStart="FIGURE 47" captionStartId="92.[151,250,1858,1882]" captionTargetBox="[155,1432,212,1827]" captionTargetId="figure-25@92.[151,1436,181,1834]" captionTargetPageId="92" captionText="FIGURE 47. Male genitalia of Siberian Sterrhinae. A–C—Scopula subpunctaria; D–F—S. prouti; G–I—S. flaccidaria; J–L— S. floslactata; M–P—S. aequifasciata. A, D, G, J, P—genital segment; B, E, H, L, O—sternite A8; C, F, I, K, M, N—phallos (M—lateral view, N—ventral view). Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965212" httpUri="https://zenodo.org/record/7965212/files/figure.png" pageId="20" pageNumber="21">Figs. 47M–47P</figureCitation>
). In female genitalia lamella antevaginalis without round sclerites, corpus bursae without signum (
<figureCitation id="13602A0A6905FFB1FF4A43A1FEBF0AB5" box="[233,321,1675,1697]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="20" pageNumber="21">Fig. 54G</figureCitation>
)............................................................... 
<taxonomicName id="4C5B4D0C6905FFB1FBE643A1FA620AB5" ID-CoL="4VVTL" authority="(Christoph, 1881)" baseAuthorityName="Christoph" baseAuthorityYear="1881" box="[1093,1436,1675,1697]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aequifasciata">
<emphasis id="B92FEA9D6905FFB1FBE643A1FA620AB5" bold="true" box="[1093,1436,1675,1697]" pageId="20" pageNumber="21">
<emphasis id="B92FEA9D6905FFB1FBE643A1FB1C0AB5" bold="true" box="[1093,1250,1675,1697]" italics="true" pageId="20" pageNumber="21">S. aequifasciata</emphasis>
(Christoph, 1881)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6905FFB1FF344383FA630B03" blockId="20.[151,1437,1382,2020]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF344383FA630B03" pageId="20" pageNumber="21">
- Forewing without distinct dark postmedial line and broad dark spots (
<figureCitation id="13602A0A6905FFB1FC0E4383FBF40AAA" box="[941,1034,1705,1727]" captionStart="FIGURE 24" captionStartId="69.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@69.[151,1435,181,1836]" captionTargetPageId="69" captionText="FIGURE 24. Adults of Siberian Sterrhinae. A—Scopula virginalis; B—S. dignata; C—S. nemoraria; D—S. umbelaria." figureDoi="http://doi.org/10.5281/zenodo.7965145" httpUri="https://zenodo.org/record/7965145/files/figure.png" pageId="20" pageNumber="21">Fig. 24C</figureCitation>
). In male genitalia sacculus moderately sclerotized, not pointed, socii shorter than sacculus; phallos robust; left ceras longer than right one (
<figureCitation id="13602A0A6905FFB1FB3543ECFACD0ACF" box="[1174,1331,1734,1756]" captionStart="FIGURE 43" captionStartId="88.[151,250,1862,1886]" captionTargetBox="[159,1430,186,1825]" captionTargetId="figure-25@88.[153,1434,181,1837]" captionTargetPageId="88" captionText="FIGURE 43. Male genitalia of Siberian Sterrhinae.A–C—Scopula corrivalaria; D–F—S. virginalis; G–J—S. dignata; K–M— S. nemoraria. A, E, G, K—genital segment; B, D, H, M—sternite A8; C, F, I, L—phallos; J—ditto, vesica everted. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965202" httpUri="https://zenodo.org/record/7965202/files/figure.png" pageId="20" pageNumber="21">Figs. 43K–43M</figureCitation>
). In female genitalia (
<figureCitation id="13602A0A6905FFB1FEE143C9FE670AED" box="[322,409,1763,1785]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="20" pageNumber="21">Fig. 52D</figureCitation>
) lamella antevaginalis anteriorly with two separate round sclerites, corpus bursae with signum..................................................................................... 
<taxonomicName id="4C5B4D0C6905FFB1FBD8422BFA630B03" ID-CoL="4VW7C" authority="(Hubner, 1799)" baseAuthorityName="Hubner" baseAuthorityYear="1799" box="[1147,1437,1793,1815]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nemoraria">
<emphasis id="B92FEA9D6905FFB1FBD8422BFA630B03" bold="true" box="[1147,1437,1793,1815]" pageId="20" pageNumber="21">
<emphasis id="B92FEA9D6905FFB1FBD8422BFB020B02" bold="true" box="[1147,1276,1793,1814]" italics="true" pageId="20" pageNumber="21">S. nemoraria</emphasis>
(Hübner, 1799)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6905FFB1FF344234FA620B7B" pageId="20" pageNumber="21">
<paragraph id="8BE4368F6905FFB1FF344234FA620B45" blockId="20.[151,1437,1382,2020]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF344234FA620B45" pageId="20" pageNumber="21">4. Ground colour of wings clear white, postmedial line narrow, contrast (almost black), terminal area of both wings with brown and/or blue-gray spots................................................................................. 5</keyLead>
</paragraph>
<paragraph id="8BE4368F6905FFB1FF344273FA620B7B" blockId="20.[151,1437,1382,2020]" box="[151,1436,1881,1903]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF344273FA620B7B" box="[151,1436,1881,1903]" pageId="20" pageNumber="21">- Wing pattern different.................................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6905FFB0FF34425CFA630CFC" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<paragraph id="8BE4368F6905FFB1FF34425CFA630BD3" blockId="20.[151,1437,1382,2020]" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB1FF34425CFA630BD3" pageId="20" pageNumber="21">
5. Postmedial line of hindwing distally bordered by dark ochreous spots between 
<emphasis id="B92FEA9D6905FFB1FC6F425CFC120B9F" box="[972,1004,1910,1931]" italics="true" pageId="20" pageNumber="21">M1</emphasis>
and 
<emphasis id="B92FEA9D6905FFB1FBBA425CFBC70B9F" box="[1049,1081,1910,1931]" italics="true" pageId="20" pageNumber="21">M3</emphasis>
and in tornal area (
<figureCitation id="13602A0A6905FFB1FB4D425CFABB0B98" box="[1262,1349,1910,1932]" captionStart="FIGURE 25" captionStartId="70.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@70.[151,1435,181,1836]" captionTargetPageId="70" captionText="FIGURE 25. Adults of Siberian Sterrhinae. A—Scopula nigropunctata; B—S. virgulata; C—S. ornata; D—S. decorata." figureDoi="http://doi.org/10.5281/zenodo.7965151" httpUri="https://zenodo.org/record/7965151/files/figure.png" pageId="20" pageNumber="21">Fig. 25C</figureCitation>
). In male genitalia (
<figureCitation id="13602A0A6905FFB1FEE242B9FE280BBD" box="[321,470,1939,1961]" captionStart="FIGURE 45" captionStartId="90.[151,250,1859,1883]" captionTargetBox="[160,1424,191,1829]" captionTargetId="figure-25@90.[153,1434,181,1835]" captionTargetPageId="90" captionText="FIGURE 45. Male genitalia of Siberian Sterrhinae. A–C—Scopula ornata; D–F—S. decorata; G–I—S. permutata; J–L—S. rubiginata; M–O—S. impersonata. A, E, G, K, N—genital segment; B, D, I, J, O—sternite A8; C, F, H, L, M—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965206" httpUri="https://zenodo.org/record/7965206/files/figure.png" pageId="20" pageNumber="21">Figs. 45A–45C</figureCitation>
) fibula tapering, cerata near equal in length. In female genitalia (
<figureCitation id="13602A0A6905FFB1FB9C42B9FB6B0BBD" box="[1087,1173,1939,1961]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="20" pageNumber="21">Fig. 53B</figureCitation>
) lamella antevaginalis with V-shaped central fold, signum developed as numerous small spinules........................ 
<taxonomicName id="4C5B4D0C6905FFB1FB06429BFA630BD3" ID-CoL="93MFG" authority="(Scopoli, 1763)" authorityName=": Viidalepp" authorityYear="1976" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[1189,1437,1969,1991]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="B92FEA9D6905FFB1FB06429BFB010BD2" box="[1189,1279,1969,1991]" italics="true" pageId="20" pageNumber="21">
<emphasis id="B92FEA9D6905FFB1FB06429BFB4C0BD2" bold="true" box="[1189,1202,1969,1990]" italics="true" pageId="20" pageNumber="21">S</emphasis>
. 
<emphasis id="B92FEA9D6905FFB1FB1D429BFB010BD2" bold="true" box="[1214,1279,1969,1990]" pageId="20" pageNumber="21">ornata</emphasis>
</emphasis>
(Scopoli, 1763)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6905FFB0FF3442E5FA630CFC" blockId="20.[151,1437,1382,2020]" lastBlockId="21.[151,1437,151,2021]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<keyLead id="30AA95BA6905FFB0FF3442E5FA630CFC" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
- Postmedial line of hindwing distally bordered by bluish spots between 
<emphasis id="B92FEA9D6905FFB1FC0542E4FC3B0BF7" box="[934,965,1998,2019]" italics="true" pageId="20" pageNumber="21">M1</emphasis>
and 
<emphasis id="B92FEA9D6905FFB1FC5942E4FBE40BF7" box="[1018,1050,1998,2019]" italics="true" pageId="20" pageNumber="21">M3</emphasis>
and in tornal area (
<figureCitation id="13602A0A6905FFB1FB4242E4FAC00BF0" box="[1249,1342,1998,2020]" captionStart="FIGURE 25" captionStartId="70.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@70.[151,1435,181,1836]" captionTargetPageId="70" captionText="FIGURE 25. Adults of Siberian Sterrhinae. A—Scopula nigropunctata; B—S. virgulata; C—S. ornata; D—S. decorata." figureDoi="http://doi.org/10.5281/zenodo.7965151" httpUri="https://zenodo.org/record/7965151/files/figure.png" pageId="20" pageNumber="21">Fig. 25D</figureCitation>
). In male genitalia (
<figureCitation id="13602A0A6904FFB0FEE045BDFE260CB9" box="[323,472,151,173]" captionStart="FIGURE 45" captionStartId="90.[151,250,1859,1883]" captionTargetBox="[160,1424,191,1829]" captionTargetId="figure-25@90.[153,1434,181,1835]" captionTargetPageId="90" captionText="FIGURE 45. Male genitalia of Siberian Sterrhinae. A–C—Scopula ornata; D–F—S. decorata; G–I—S. permutata; J–L—S. rubiginata; M–O—S. impersonata. A, E, G, K, N—genital segment; B, D, I, J, O—sternite A8; C, F, H, L, M—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965206" httpUri="https://zenodo.org/record/7965206/files/figure.png" pageId="21" pageNumber="22">Figs. 45D–45F</figureCitation>
) fibula rounded apically, left ceras shortened. In female genitalia (
<figureCitation id="13602A0A6904FFB0FBCB45BDFB3F0CB9" box="[1128,1217,151,173]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="21" pageNumber="22">Fig. 53C</figureCitation>
) lamella antevaginalis without V-shaped central fold, signum not developed, corpus bursae medially only with granulated surface.............................................................................. 
<taxonomicName id="4C5B4D0C6904FFB0FC7E45F8FBB50CFC" ID-CoL="4VVXV" authorityName=": Vasilyeva &amp; Epova" authorityYear="1987" box="[989,1099,210,232]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="decorata">
<emphasis id="B92FEA9D6904FFB0FC7E45F8FBB50CFC" box="[989,1099,210,232]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FC7E45F8FC140CF3" bold="true" box="[989,1002,210,231]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FC5545F8FBB50CFC" bold="true" box="[1014,1099,210,232]" pageId="21" pageNumber="22">decorata</emphasis>
</emphasis>
</taxonomicName>
([Denis &amp; Schiffermüller], 1775)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF3445C5FA620D36" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF3445C5FA620D11" blockId="21.[151,1437,151,2021]" box="[151,1436,239,261]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3445C5FA620D11" box="[151,1436,239,261]" pageId="21" pageNumber="22">6. Males.............................................................................................. 7</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344427FA620D36" blockId="21.[151,1437,151,2021]" box="[151,1436,269,291]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344427FA620D36" box="[151,1436,269,291]" pageId="21" pageNumber="22">- Females............................................................................................ 29</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF344401FA620D49" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF344401FA620D2B" blockId="21.[151,1437,151,2021]" box="[151,1436,298,320]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344401FA620D2B" box="[151,1436,298,320]" pageId="21" pageNumber="22">
7. Hindtibia with well-developed apical spurs (
<figureCitation id="13602A0A6904FFB0FD2F4400FD2B0D54" box="[652,725,298,320]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Fig. 6E</figureCitation>
)......................................................... 8
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344462FA620D49" blockId="21.[151,1437,151,2021]" box="[151,1436,327,349]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344462FA620D49" box="[151,1436,327,349]" pageId="21" pageNumber="22">- Hindtibia without apical spurs or they reduced............................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF34444FFA620DFB" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF34444FFA620DA1" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34444FFA620DA1" pageId="21" pageNumber="22">
8. Forewing length does not exceed 
<quantity id="4CA39B6A6904FFB0FD94444FFD8B0D6E" box="[567,629,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="22" unit="mm" value="9.0">9 mm</quantity>
, its terminal area conspicuously darker than rest part of wing (
<figureCitation id="13602A0A6904FFB0FB50444FFAB10D6F" box="[1267,1359,357,379]" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="21" pageNumber="22">Fig. 26C</figureCitation>
); fibula sclerotized not stronger than valvula, both cerata normally developed, near same length (
<figureCitation id="13602A0A6904FFB0FB9544A8FB330D8C" box="[1078,1229,386,408]" captionStart="FIGURE 45" captionStartId="90.[151,250,1859,1883]" captionTargetBox="[160,1424,191,1829]" captionTargetId="figure-25@90.[153,1434,181,1835]" captionTargetPageId="90" captionText="FIGURE 45. Male genitalia of Siberian Sterrhinae. A–C—Scopula ornata; D–F—S. decorata; G–I—S. permutata; J–L—S. rubiginata; M–O—S. impersonata. A, E, G, K, N—genital segment; B, D, I, J, O—sternite A8; C, F, H, L, M—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965206" httpUri="https://zenodo.org/record/7965206/files/figure.png" pageId="21" pageNumber="22">Figs. 45N, 45O</figureCitation>
)........................................................................................... 
<taxonomicName id="4C5B4D0C6904FFB0FBCF44B5FA620DA1" ID-CoL="4VW3P" authority="(Walker, 1861)" baseAuthorityName="Walker" baseAuthorityYear="1861" box="[1132,1436,415,437]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="impersonata">
<emphasis id="B92FEA9D6904FFB0FBCF44B5FAFF0DA0" box="[1132,1281,415,437]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBCF44B5FB870DA0" bold="true" box="[1132,1145,415,436]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB2544B5FAFF0DA0" bold="true" box="[1158,1281,415,436]" pageId="21" pageNumber="22">impersonata</emphasis>
</emphasis>
(Walker, 1861)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344497FA620DFB" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344497FA620DFB" pageId="21" pageNumber="22">
- Forewing length more than 
<quantity id="4CA39B6A6904FFB0FE4F4497FDCC0DC6" box="[492,562,445,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="mm" value="10.0">10 mm</quantity>
; fibula sclerotized clearly stronger than valvula, one or both cerata commonly shortened.. ................................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF3444D2FA620E8B" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF3444D2FA630E5C" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3444D2FA630E5C" pageId="21" pageNumber="22">
9. Antennal cilia comparatively long: 1.5–1.65 times width of flagellum, frons, vertex and collar light brown mixed with blackish scales; hindwing with distinct discal spot (
<figureCitation id="13602A0A6904FFB0FDDF473FFD2B0E3E" box="[636,725,533,555]" captionStart="FIGURE 29" captionStartId="74.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@74.[151,1435,181,1836]" captionTargetPageId="74" captionText="FIGURE 29. Adults of Siberian Sterrhinae. A—Scopula aequifasciata; B—S. cajanderi; C—S. albiceraria; D—Cyclophora albipunctata." figureDoi="http://doi.org/10.5281/zenodo.7965161" httpUri="https://zenodo.org/record/7965161/files/figure.png" pageId="21" pageNumber="22">Fig. 29B</figureCitation>
); both cerata very short and barely sclerotized, anterior margin of sternite A8 concave; vesica with elongate sclerite (
<figureCitation id="13602A0A6904FFB0FDDE4718FCEE0E53" box="[637,784,562,584]" captionStart="FIGURE 48" captionStartId="93.[151,250,1797,1821]" captionTargetBox="[152,1430,182,1767]" captionTargetId="figure-15@93.[151,1436,181,1773]" captionTargetPageId="93" captionText="FIGURE 48. Male genitalia of Siberian Sterrhinae. A–C—Scopula cajanderi; D–G—S. albiceraria; H, I—Cyclophora albipunctata; J, K—Timandra rectistrigaria; L–N—T. griseata. A, F—genital segment; H, J, L, N—genital segment; B, D, E—sternite A8; C, G, I, K, M—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965216" httpUri="https://zenodo.org/record/7965216/files/figure.png" pageId="21" pageNumber="22">Figs. 48B, 48C</figureCitation>
)................................ 
<taxonomicName id="4C5B4D0C6904FFB0FB024718FA630E5C" ID-CoL="4VVVY" authority="(Herz, 1904)" baseAuthorityName="Herz" baseAuthorityYear="1904" box="[1185,1437,562,584]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cajanderi">
<emphasis id="B92FEA9D6904FFB0FB024718FAE90E5C" box="[1185,1303,562,584]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FB024718FB500E53" bold="true" box="[1185,1198,562,583]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB194718FAE90E5C" bold="true" box="[1210,1303,562,584]" pageId="21" pageNumber="22">cajanderi</emphasis>
</emphasis>
(Herz, 1904)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF34477AFA620E8B" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34477AFA620E8B" pageId="21" pageNumber="22">
- Antennal cilia comparatively short: 1–1.3 times width of flagellum, frons black, vertex and collar sand coloured; hindwing without discal spot (
<figureCitation id="13602A0A6904FFB0FE064747FE010E96" box="[421,511,621,643]" captionStart="FIGURE 27" captionStartId="72.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@72.[151,1435,181,1836]" captionTargetPageId="72" captionText="FIGURE 27. Adults of Siberian Sterrhinae. A—Scopula incanata; B—S. immutata; C—S. frigidaria; D—S. ternata." figureDoi="http://doi.org/10.5281/zenodo.7965155" httpUri="https://zenodo.org/record/7965155/files/figure.png" pageId="21" pageNumber="22">Fig. 27D</figureCitation>
); sternite A8 usually with long right and short left cerata, anterior margin of sternite A8 convex; vesica without sclerite (
<figureCitation id="13602A0A6904FFB0FE6547A0FD6F0EB4" box="[454,657,650,672]" captionStart="FIGURE 46" captionStartId="91.[151,250,1867,1891]" captionTargetBox="[163,1425,198,1844]" captionTargetId="figure-15@91.[157,1431,181,1844]" captionTargetPageId="91" captionText="FIGURE 46. Male genitalia of Siberian Sterrhinae.A–C—Scopula beckeraria; D–F—S. incanata; G–I—S. immutata; J–L—S. frigidaria; M–P—S. ternata. A, D, G, K, O—genital segment; B, E, H, J, M, P—sternite A8; C, F, I, L, N—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965208" httpUri="https://zenodo.org/record/7965208/files/figure.png" pageId="21" pageNumber="22">Figs. 46M, 46N, 46P</figureCitation>
)........................................... 
<taxonomicName id="4C5B4D0C6904FFB0FB0747A0FA620E8B" ID-CoL="6XZ9G" authority="Schrank, 1802" authorityName="Schrank" authorityYear="1802" box="[1188,1436,650,672]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ternata">
<emphasis id="B92FEA9D6904FFB0FB0747A0FAFA0E8B" box="[1188,1284,650,672]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FB0747A0FB4F0E8B" bold="true" box="[1188,1201,650,671]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB1E47A0FAFA0E8B" bold="true" box="[1213,1284,650,671]" pageId="21" pageNumber="22">ternata</emphasis>
</emphasis>
Schrank, 1802
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF344782FA620ECE" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF344782FA630EA9" blockId="21.[151,1437,151,2021]" box="[151,1437,679,701]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344782FA630EA9" box="[151,1437,679,701]" pageId="21" pageNumber="22">
10. Hindleg: tarsus longer than tibia (
<figureCitation id="13602A0A6904FFB0FD87478DFD920EA9" box="[548,620,679,701]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Fig. 6F</figureCitation>
).......................................... 
<taxonomicName id="4C5B4D0C6904FFB0FBC9478DFA630EA9" ID-CoL="4VWBQ" authority="(Hufnagel, 1767)" baseAuthorityName="Hufnagel" baseAuthorityYear="1767" box="[1130,1437,679,701]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="rubiginata">
<emphasis id="B92FEA9D6904FFB0FBC9478DFB120EA9" box="[1130,1260,679,701]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBC9478DFB890EA8" bold="true" box="[1130,1143,679,700]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB27478DFB120EA9" bold="true" box="[1156,1260,679,701]" pageId="21" pageNumber="22">rubiginata</emphasis>
</emphasis>
(Hufnagel, 1767)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF3447EFFA620ECE" blockId="21.[151,1437,151,2021]" box="[151,1436,709,731]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3447EFFA620ECE" box="[151,1436,709,731]" pageId="21" pageNumber="22">
- Hindleg: tarsus shorter than tibia or nearly as long (
<figureCitation id="13602A0A6904FFB0FD7347EFFCC30ECF" box="[720,829,709,731]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Fig. 6G–6J</figureCitation>
)............................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF3447C8FA620F01" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF3447C8FA620EE3" blockId="21.[151,1437,151,2021]" box="[151,1436,738,760]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3447C8FA620EE3" box="[151,1436,738,760]" pageId="21" pageNumber="22">
11. Hind tarsus reduced, 3–4 times shorter than hindtibia (
<figureCitation id="13602A0A6904FFB0FD4F47C8FC950EEC" box="[748,875,738,760]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Figs. 6G, 6H</figureCitation>
)............................................ 12
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF34462AFA620F01" blockId="21.[151,1437,151,2021]" box="[151,1436,766,789]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34462AFA620F01" box="[151,1436,766,789]" pageId="21" pageNumber="22">
- Hind tarsus ½ length of hindtibia or more than ½ length of hindtibia (
<figureCitation id="13602A0A6904FFB0FCDF47D5FC170F01" box="[892,1001,767,789]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Figs. 6I, 6J</figureCitation>
)................................. 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF344637FA620FB3" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF344637FA630F79" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344637FA630F79" pageId="21" pageNumber="22">
12. Genital segment elongate: saccus elliptic, tegumen between socii sclerotized, prominent, reminiscent of uncus; aedeagus without ventrally protruding sclerite at tip (
<figureCitation id="13602A0A6904FFB0FD224610FCEC0F5B" box="[641,786,826,848]" captionStart="FIGURE 47" captionStartId="92.[151,250,1858,1882]" captionTargetBox="[155,1432,212,1827]" captionTargetId="figure-25@92.[151,1436,181,1834]" captionTargetPageId="92" captionText="FIGURE 47. Male genitalia of Siberian Sterrhinae. A–C—Scopula subpunctaria; D–F—S. prouti; G–I—S. flaccidaria; J–L— S. floslactata; M–P—S. aequifasciata. A, D, G, J, P—genital segment; B, E, H, L, O—sternite A8; C, F, I, K, M, N—phallos (M—lateral view, N—ventral view). Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965212" httpUri="https://zenodo.org/record/7965212/files/figure.png" pageId="21" pageNumber="22">Figs. 47J, 47K</figureCitation>
). Ground colour of wings yellowish white (
<figureCitation id="13602A0A6904FFB0FB174610FAF50F5B" box="[1204,1291,826,848]" captionStart="FIGURE 28" captionStartId="73.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@73.[151,1435,181,1836]" captionTargetPageId="73" captionText="FIGURE 28. Adults of Siberian Sterrhinae. A—Scopula subpunctaria; B—S. prouti; C—S. flaccidaria; D—S. floslactata." figureDoi="http://doi.org/10.5281/zenodo.7965159" httpUri="https://zenodo.org/record/7965159/files/figure.png" pageId="21" pageNumber="22">Fig. 28D</figureCitation>
)...................................................................................... 
<taxonomicName id="4C5B4D0C6904FFB0FBCD467DFA630F79" ID-CoL="4VW25" authority="(Haworth, 1809)" baseAuthorityName="Haworth" baseAuthorityYear="1809" box="[1134,1437,855,877]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="floslactata">
<emphasis id="B92FEA9D6904FFB0FBCD467DFB110F79" box="[1134,1263,855,877]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBCD467DFB850F78" bold="true" box="[1134,1147,855,876]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB2B467DFB110F79" bold="true" box="[1160,1263,855,877]" pageId="21" pageNumber="22">floslactata</emphasis>
</emphasis>
(Haworth, 1809)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF34465FFA620FB3" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34465FFA620FB3" pageId="21" pageNumber="22">- Genital segment not elongate: saccus broad and round, tegumen between socii not prominent; aedeagus with ventrally protruding sclerite at tip. Ground colour of wings glossy white......................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF34469AFA620843" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF34469AFA630814" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34469AFA630814" pageId="21" pageNumber="22">
13. Socii very long, terminally curved and dilated; sternite A8 with both cerata long and symmetrical, protruding beyond posterior margin of mappa, or left ceras shortened and strongly curved (
<figureCitation id="13602A0A6904FFB0FCE146E7FC250FF6" box="[834,987,973,995]" captionStart="FIGURE 47" captionStartId="92.[151,250,1858,1882]" captionTargetBox="[155,1432,212,1827]" captionTargetId="figure-25@92.[151,1436,181,1834]" captionTargetPageId="92" captionText="FIGURE 47. Male genitalia of Siberian Sterrhinae. A–C—Scopula subpunctaria; D–F—S. prouti; G–I—S. flaccidaria; J–L— S. floslactata; M–P—S. aequifasciata. A, D, G, J, P—genital segment; B, E, H, L, O—sternite A8; C, F, I, K, M, N—phallos (M—lateral view, N—ventral view). Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965212" httpUri="https://zenodo.org/record/7965212/files/figure.png" pageId="21" pageNumber="22">Figs. 47A, 47B</figureCitation>
). Wings with terminal dots, at least at anterior third of forewing (
<figureCitation id="13602A0A6904FFB0FE3046C0FE140FEB" box="[403,490,1002,1024]" captionStart="FIGURE 28" captionStartId="73.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@73.[151,1435,181,1836]" captionTargetPageId="73" captionText="FIGURE 28. Adults of Siberian Sterrhinae. A—Scopula subpunctaria; B—S. prouti; C—S. flaccidaria; D—S. floslactata." figureDoi="http://doi.org/10.5281/zenodo.7965159" httpUri="https://zenodo.org/record/7965159/files/figure.png" pageId="21" pageNumber="22">Fig. 28A</figureCitation>
)............................................ 
<taxonomicName id="4C5B4D0C6904FFB0FBA346C0FA630814" ID-CoL="4VWDS" authority="(Herrich-Schaffer, 1847)" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1847" box="[1024,1437,1002,1024]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subpunctaria">
<emphasis id="B92FEA9D6904FFB0FBA346C0FB630814" box="[1024,1181,1002,1024]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBA346C0FBF30FEB" bold="true" box="[1024,1037,1002,1023]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FBB946C0FB630814" bold="true" box="[1050,1181,1002,1024]" pageId="21" pageNumber="22">subpunctaria</emphasis>
</emphasis>
(Herrich-Schäffer, 1847)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344122FA620843" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344122FA620843" pageId="21" pageNumber="22">
- Socii shorter than in 
<taxonomicName id="4C5B4D0C6904FFB0FE054122FDC20808" ID-CoL="4VWDS" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1847" box="[422,572,1031,1053]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subpunctaria">
<emphasis id="B92FEA9D6904FFB0FE054122FDC20808" box="[422,572,1031,1053]" italics="true" pageId="21" pageNumber="22">S. subpunctaria</emphasis>
</taxonomicName>
, terminally straight and not dilated; sternite A8 with both cerata of almost equal length, not curved, cerata not protruding beyond posterior margin of mappa (
<figureCitation id="13602A0A6904FFB0FCF4410FFC13082E" box="[855,1005,1061,1083]" captionStart="FIGURE 47" captionStartId="92.[151,250,1858,1882]" captionTargetBox="[155,1432,212,1827]" captionTargetId="figure-25@92.[151,1436,181,1834]" captionTargetPageId="92" captionText="FIGURE 47. Male genitalia of Siberian Sterrhinae. A–C—Scopula subpunctaria; D–F—S. prouti; G–I—S. flaccidaria; J–L— S. floslactata; M–P—S. aequifasciata. A, D, G, J, P—genital segment; B, E, H, L, O—sternite A8; C, F, I, K, M, N—phallos (M—lateral view, N—ventral view). Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965212" httpUri="https://zenodo.org/record/7965212/files/figure.png" pageId="21" pageNumber="22">Figs. 47D, 47E</figureCitation>
). Wings without terminal dots (
<figureCitation id="13602A0A6904FFB0FAB8410FFA8F082E" box="[1307,1393,1061,1083]" captionStart="FIGURE 28" captionStartId="73.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@73.[151,1435,181,1836]" captionTargetPageId="73" captionText="FIGURE 28. Adults of Siberian Sterrhinae. A—Scopula subpunctaria; B—S. prouti; C—S. flaccidaria; D—S. floslactata." figureDoi="http://doi.org/10.5281/zenodo.7965159" httpUri="https://zenodo.org/record/7965159/files/figure.png" pageId="21" pageNumber="22">Fig. 28B</figureCitation>
)................................................................................. 
<taxonomicName id="4C5B4D0C6904FFB0FB014168FA620843" ID-CoL="6XZ6R" authority="Djakonov, 1935" authorityName="Djakonov" authorityYear="1935" box="[1186,1436,1090,1112]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="prouti">
<emphasis id="B92FEA9D6904FFB0FB014168FB060843" box="[1186,1272,1090,1112]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FB014168FB510843" bold="true" box="[1186,1199,1090,1111]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB1F4168FB060843" bold="true" box="[1212,1272,1090,1111]" pageId="21" pageNumber="22">prouti</emphasis>
</emphasis>
Djakonov, 1935
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF34414AFA6208BB" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF34414AFA620887" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34414AFA620887" pageId="21" pageNumber="22">14. Valvula and fibula sclerotized almost equally or fibula just slightly darker than valvula, fibula commonly with rounded tip; genitalia often asymmetric............................................................................. 15</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF3441B1FA6208BB" blockId="21.[151,1437,151,2021]" box="[151,1436,1178,1200]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3441B1FA6208BB" box="[151,1436,1178,1200]" pageId="21" pageNumber="22">- Fibula strongly sclerotized, significantly darker than valvula, often pointed; genitalia commonly symmetric............ 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF34419DFA620913" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF34419DFA6208FF" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34419DFA6208FF" pageId="21" pageNumber="22">
15. Hindleg: tarsus longer than 1/2 length of tibia, reaches about 2/3–3/4 length of tibia, occasionally almost equal to tibia (
<figureCitation id="13602A0A6904FFB0FAD6419DFF0808FF" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Fig. 6I</figureCitation>
)................................................................................................ 16
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF3441D9FA620913" blockId="21.[151,1437,151,2021]" box="[151,1436,1266,1288]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3441D9FA620913" box="[151,1436,1266,1288]" pageId="21" pageNumber="22">
- Hindleg: length of tarsus near 1/2 length of tibia (
<figureCitation id="13602A0A6904FFB0FD6741D8FCF9091C" box="[708,775,1266,1288]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="21" pageNumber="22">Fig. 6J</figureCitation>
).................................................... 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF344025FA620956" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF344025FA620931" blockId="21.[151,1437,151,2021]" box="[151,1436,1295,1317]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344025FA620931" box="[151,1436,1295,1317]" pageId="21" pageNumber="22">16. Both fibulae bifid.................................................................................... 17</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344007FA620956" blockId="21.[151,1437,151,2021]" box="[151,1436,1325,1347]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344007FA620956" box="[151,1436,1325,1347]" pageId="21" pageNumber="22">- If fibula bifid, than only on the one side.................................................................. 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF344060FA6209C0" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF344060FA63098E" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344060FA63098E" pageId="21" pageNumber="22">
17. Fibula with two separate curved projections, aedeagus curved, vesica with two elongate cornuti; right ceras long and strongly curved, left ceras short, broad, strongly spinose (
<figureCitation id="13602A0A6904FFB0FD16404DFCB00969" box="[693,846,1383,1405]" captionStart="FIGURE 44" captionStartId="89.[151,250,1867,1891]" captionTargetBox="[176,1433,206,1832]" captionTargetId="figure-15@89.[153,1434,181,1844]" captionTargetPageId="89" captionText="FIGURE 44. Male genitalia of Siberian Sterrhinae.A–D—Scopula umbelaria; E–G—S. nigropunctata; H–K—S. virgulata. A, E, H, K—genital segment; B, G, J—phallos; C, D, F, I—sternite A8. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965204" httpUri="https://zenodo.org/record/7965204/files/figure.png" pageId="21" pageNumber="22">Figs. 44H–44K</figureCitation>
). Forewing without discal spot (
<figureCitation id="13602A0A6904FFB0FB21404DFB260969" box="[1154,1240,1383,1405]" captionStart="FIGURE 25" captionStartId="70.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@70.[151,1435,181,1836]" captionTargetPageId="70" captionText="FIGURE 25. Adults of Siberian Sterrhinae. A—Scopula nigropunctata; B—S. virgulata; C—S. ornata; D—S. decorata." figureDoi="http://doi.org/10.5281/zenodo.7965151" httpUri="https://zenodo.org/record/7965151/files/figure.png" pageId="21" pageNumber="22">Fig. 25B</figureCitation>
).............................................................................. 
<taxonomicName id="4C5B4D0C6904FFB0FC7A40AFFBB5098E" ID-CoL="4VWFN" baseAuthorityName="Denis et Schiffermuller" baseAuthorityYear="1775" box="[985,1099,1412,1434]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="virgulata">
<emphasis id="B92FEA9D6904FFB0FC7A40AFFBB5098E" box="[985,1099,1412,1434]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FC7A40AFFC18098E" bold="true" box="[985,998,1413,1434]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FC5140AEFBB5098E" bold="true" box="[1010,1099,1412,1434]" pageId="21" pageNumber="22">virgulata</emphasis>
</emphasis>
</taxonomicName>
([Denis &amp; Schiffermüller], 1775)
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344089FA6209C0" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344089FA6209C0" pageId="21" pageNumber="22">
- Fibulae asymmetrically bifurcated only in apical parts, vesica with one strongly curved dentate cornutus; cerata near same length (
<figureCitation id="13602A0A6904FFB0FE884095FE4709C1" box="[299,441,1471,1493]" captionStart="FIGURE 42" captionStartId="87.[151,250,1867,1891]" captionTargetBox="[166,1418,193,1841]" captionTargetId="figure-15@87.[158,1430,181,1844]" captionTargetPageId="87" captionText="FIGURE 42. Male genitalia of Siberian Sterrhinae. A–C—Idaea descitaria; D–F—Holarctias rufinaria; G–I—H. rufinularia; J–L—Scopula agutsaensis; M–O—S. immorata. A, E, I, L, N—genital segment, ventral view; B—ditto, valvae shifted; C, D, H, K, O—phallos; G, F, J, M—sternite A8. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965198" httpUri="https://zenodo.org/record/7965198/files/figure.png" pageId="21" pageNumber="22">Figs. 42J–42L</figureCitation>
). Forewing with discal spot (
<figureCitation id="13602A0A6904FFB0FD6C4095FCDB09C1" box="[719,805,1471,1493]" captionStart="FIGURE 23" captionStartId="68.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@68.[151,1435,181,1836]" captionTargetPageId="68" captionText="FIGURE 23. Adults of Siberian Sterrhinae. A—Holarctias rufinularia; B—Scopula agutsaensis; C—S. immorata; D—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965143" httpUri="https://zenodo.org/record/7965143/files/figure.png" pageId="21" pageNumber="22">Fig. 23B</figureCitation>
).......................... 
<taxonomicName id="4C5B4D0C6904FFB0FBC84095FA6209C0" authority="Vasilenko, 1997" authorityName="Vasilenko" authorityYear="1997" box="[1131,1436,1471,1493]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="agutsaensis">
<emphasis id="B92FEA9D6904FFB0FBC84095FB0909C0" box="[1131,1271,1471,1493]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBC84095FB8609C0" bold="true" box="[1131,1144,1471,1492]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB274095FB0909C0" bold="true" box="[1156,1271,1471,1492]" pageId="21" pageNumber="22">agutsaensis</emphasis>
</emphasis>
Vasilenko, 1997
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF3440F7FA630A7C" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF3440F7FA630A04" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3440F7FA630A04" pageId="21" pageNumber="22">
18. Length of socii near equal to distance between outer margins of their bases, right ceras longer than left one (
<figureCitation id="13602A0A6904FFB0FB5B40F7FA6F09E6" box="[1272,1425,1501,1523]" captionStart="FIGURE 43" captionStartId="88.[151,250,1862,1886]" captionTargetBox="[159,1430,186,1825]" captionTargetId="figure-25@88.[153,1434,181,1837]" captionTargetPageId="88" captionText="FIGURE 43. Male genitalia of Siberian Sterrhinae.A–C—Scopula corrivalaria; D–F—S. virginalis; G–J—S. dignata; K–M— S. nemoraria. A, E, G, K—genital segment; B, D, H, M—sternite A8; C, F, I, L—phallos; J—ditto, vesica everted. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965202" httpUri="https://zenodo.org/record/7965202/files/figure.png" pageId="21" pageNumber="22">Figs. 43G, 43H</figureCitation>
); forewing with discal and terminal dots (
<figureCitation id="13602A0A6904FFB0FDC240D0FD490A1B" box="[609,695,1530,1552]" captionStart="FIGURE 24" captionStartId="69.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@69.[151,1435,181,1836]" captionTargetPageId="69" captionText="FIGURE 24. Adults of Siberian Sterrhinae. A—Scopula virginalis; B—S. dignata; C—S. nemoraria; D—S. umbelaria." figureDoi="http://doi.org/10.5281/zenodo.7965145" httpUri="https://zenodo.org/record/7965145/files/figure.png" pageId="21" pageNumber="22">Fig. 24B</figureCitation>
)....................................... 
<taxonomicName id="4C5B4D0C6904FFB0FB3A40D0FA630A04" ID-CoL="79YW9" authority="(Guenee, 1858)" baseAuthorityName="Guenee" baseAuthorityYear="1858" box="[1177,1437,1529,1552]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="dignata">
<emphasis id="B92FEA9D6904FFB0FB3A40D0FB030A04" box="[1177,1277,1530,1552]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FB3A40D0FB580A1B" bold="true" box="[1177,1190,1530,1551]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB1140D0FB030A04" bold="true" box="[1202,1277,1530,1552]" pageId="21" pageNumber="22">dignata</emphasis>
</emphasis>
(Guenée, 1858)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344332FA630A7C" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344332FA630A7C" pageId="21" pageNumber="22">
- Length of socii does not exceed ½ distance between outer margins of their bases, right ceras no longer than left one (
<figureCitation id="13602A0A6904FFB0FA94433DFEF20A5E" captionStart="FIGURE 43" captionStartId="88.[151,250,1862,1886]" captionTargetBox="[159,1430,186,1825]" captionTargetId="figure-25@88.[153,1434,181,1837]" captionTargetPageId="88" captionText="FIGURE 43. Male genitalia of Siberian Sterrhinae.A–C—Scopula corrivalaria; D–F—S. virginalis; G–J—S. dignata; K–M— S. nemoraria. A, E, G, K—genital segment; B, D, H, M—sternite A8; C, F, I, L—phallos; J—ditto, vesica everted. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965202" httpUri="https://zenodo.org/record/7965202/files/figure.png" pageId="21" pageNumber="22">Figs. 43D, 43E</figureCitation>
); forewing without discal and terminal dots (rarely forewing with weak discal dot) (
<figureCitation id="13602A0A6904FFB0FB92431FFB770A5E" box="[1073,1161,1589,1611]" captionStart="FIGURE 24" captionStartId="69.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@69.[151,1435,181,1836]" captionTargetPageId="69" captionText="FIGURE 24. Adults of Siberian Sterrhinae. A—Scopula virginalis; B—S. dignata; C—S. nemoraria; D—S. umbelaria." figureDoi="http://doi.org/10.5281/zenodo.7965145" httpUri="https://zenodo.org/record/7965145/files/figure.png" pageId="21" pageNumber="22">Fig. 24A</figureCitation>
).................................................................................................. 
<taxonomicName id="4C5B4D0C6904FFB0FBD54378FA630A7C" ID-CoL="94DMS" authority="(Fourcroy, 1785)" baseAuthorityName="Fourcroy" baseAuthorityYear="1785" box="[1142,1437,1618,1640]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="virginalis">
<emphasis id="B92FEA9D6904FFB0FBD54378FB120A7C" box="[1142,1260,1618,1640]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBD54378FB7D0A73" bold="true" box="[1142,1155,1618,1639]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB2C4378FB120A7C" bold="true" box="[1167,1260,1618,1640]" pageId="21" pageNumber="22">virginalis</emphasis>
</emphasis>
(Fourcroy, 1785)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF34435AFA620AC9" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF34435AFA630AB6" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34435AFA630AB6" pageId="21" pageNumber="22">
19. Genital segment symmetric, fibula and valvula tapering; both cerata short, strongly spinose (
<figureCitation id="13602A0A6904FFB0FBF94345FB110A91" box="[1114,1263,1647,1669]" captionStart="FIGURE 43" captionStartId="88.[151,250,1862,1886]" captionTargetBox="[159,1430,186,1825]" captionTargetId="figure-25@88.[153,1434,181,1837]" captionTargetPageId="88" captionText="FIGURE 43. Male genitalia of Siberian Sterrhinae.A–C—Scopula corrivalaria; D–F—S. virginalis; G–J—S. dignata; K–M— S. nemoraria. A, E, G, K—genital segment; B, D, H, M—sternite A8; C, F, I, L—phallos; J—ditto, vesica everted. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965202" httpUri="https://zenodo.org/record/7965202/files/figure.png" pageId="21" pageNumber="22">Figs. 43A, 43B</figureCitation>
).................................................................................... 
<taxonomicName id="4C5B4D0C6904FFB0FBE143A7FA630AB6" ID-CoL="6XZ7X" authority="(Kretschmar, 1862)" baseAuthorityName="Kretschmar" baseAuthorityYear="1862" box="[1090,1437,1676,1698]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="corrivalaria">
<emphasis id="B92FEA9D6904FFB0FBE143A7FB2F0AB6" box="[1090,1233,1676,1698]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBE143A7FBB10AB6" bold="true" box="[1090,1103,1677,1698]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FBFF43A6FB2F0AB6" bold="true" box="[1116,1233,1676,1698]" pageId="21" pageNumber="22">corrivalaria</emphasis>
</emphasis>
(Kretschmar, 1862)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344381FA620AC9" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344381FA620AC9" pageId="21" pageNumber="22">- Genital segment clearly asymmetric: right and left fibulae of different structure; cerata of different lengths, at least one ceras rather long: protruding beyond posterior margin of mappa.................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF3443CFFA630B46" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF3443CFFA630B0C" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF3443CFFA630B0C" pageId="21" pageNumber="22">
20. Right fibula flat, bilobed, both valvulae strongly curved at tips; vesica with large terminally rounded sclerites; left ceras very short, right ceras long and thin (
<figureCitation id="13602A0A6904FFB0FDB74228FD560B03" box="[532,680,1794,1816]" captionStart="FIGURE 44" captionStartId="89.[151,250,1867,1891]" captionTargetBox="[176,1433,206,1832]" captionTargetId="figure-15@89.[153,1434,181,1844]" captionTargetPageId="89" captionText="FIGURE 44. Male genitalia of Siberian Sterrhinae.A–D—Scopula umbelaria; E–G—S. nigropunctata; H–K—S. virgulata. A, E, H, K—genital segment; B, G, J—phallos; C, D, F, I—sternite A8. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965204" httpUri="https://zenodo.org/record/7965204/files/figure.png" pageId="21" pageNumber="22">Figs. 44E–44G</figureCitation>
).................................. 
<taxonomicName id="4C5B4D0C6904FFB0FBE54228FA630B0C" ID-CoL="4VW7T" authority="(Hufnagel, 1767)" baseAuthorityName="Hufnagel" baseAuthorityYear="1767" box="[1094,1437,1794,1816]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="nigropunctata">
<emphasis id="B92FEA9D6904FFB0FBE54228FB120B03" box="[1094,1260,1794,1816]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBE54228FBAD0B03" bold="true" box="[1094,1107,1794,1815]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FBC34228FB120B03" bold="true" box="[1120,1260,1794,1815]" pageId="21" pageNumber="22">nigropunctata</emphasis>
</emphasis>
(Hufnagel, 1767)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF34420AFA630B46" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF34420AFA630B46" pageId="21" pageNumber="22">
- Right fibula furcate, pincer-shaped; vesica with 3–4 terminal sclerites, one of them claw-shaped: long, curved and tapered; sternite A8 with both cerata long and narrow (
<figureCitation id="13602A0A6904FFB0FD3F4217FCCC0B46" box="[668,818,1853,1875]" captionStart="FIGURE 44" captionStartId="89.[151,250,1867,1891]" captionTargetBox="[176,1433,206,1832]" captionTargetId="figure-15@89.[153,1434,181,1844]" captionTargetPageId="89" captionText="FIGURE 44. Male genitalia of Siberian Sterrhinae.A–D—Scopula umbelaria; E–G—S. nigropunctata; H–K—S. virgulata. A, E, H, K—genital segment; B, G, J—phallos; C, D, F, I—sternite A8. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965204" httpUri="https://zenodo.org/record/7965204/files/figure.png" pageId="21" pageNumber="22">Figs. 44A–44D</figureCitation>
).......................... 
<taxonomicName id="4C5B4D0C6904FFB0FBDD4217FA630B46" ID-CoL="93MFZ" authority="(Hubner, 1813)" baseAuthorityName="Hubner" baseAuthorityYear="1813" box="[1150,1437,1852,1874]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="umbelaria">
<emphasis id="B92FEA9D6904FFB0FBDD4217FB020B46" box="[1150,1276,1852,1874]" italics="true" pageId="21" pageNumber="22">
<emphasis id="B92FEA9D6904FFB0FBDD4217FB750B46" bold="true" box="[1150,1163,1853,1874]" italics="true" pageId="21" pageNumber="22">S</emphasis>
. 
<emphasis id="B92FEA9D6904FFB0FB344216FB020B46" bold="true" box="[1175,1276,1852,1874]" pageId="21" pageNumber="22">umbelaria</emphasis>
</emphasis>
(Hübner, 1813)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB0FF344270FA620BBF" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB0FF344270FA620B7B" blockId="21.[151,1437,151,2021]" box="[151,1436,1882,1904]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344270FA620B7B" box="[151,1436,1882,1904]" pageId="21" pageNumber="22">21. Cerata almost equal in length, short, not reaching posterior margin of VIII sternite................................. 22</keyLead>
</paragraph>
<paragraph id="8BE4368F6904FFB0FF344252FA620BBF" blockId="21.[151,1437,151,2021]" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB0FF344252FA620BBF" pageId="21" pageNumber="22">- One or both cerata reaching posterior margin of sternite A8, or longer and protruding beyond posterior margin of sternite A8................................................................................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6904FFB3FF344298FA620CF3" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<paragraph id="8BE4368F6904FFB3FF344298FA630CB9" blockId="21.[151,1437,151,2021]" lastBlockId="22.[151,1437,151,2021]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<keyLead id="30AA95BA6904FFB3FF344298FA630CB9" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
22. Anterior margin of sternite A8 concave medially, tips of cerata not spinose (
<figureCitation id="13602A0A6904FFB0FC0B4298FBC50BD3" box="[936,1083,1970,1992]" captionStart="FIGURE 48" captionStartId="93.[151,250,1797,1821]" captionTargetBox="[152,1430,182,1767]" captionTargetId="figure-15@93.[151,1436,181,1773]" captionTargetPageId="93" captionText="FIGURE 48. Male genitalia of Siberian Sterrhinae. A–C—Scopula cajanderi; D–G—S. albiceraria; H, I—Cyclophora albipunctata; J, K—Timandra rectistrigaria; L–N—T. griseata. A, F—genital segment; H, J, L, N—genital segment; B, D, E—sternite A8; C, G, I, K, M—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965216" httpUri="https://zenodo.org/record/7965216/files/figure.png" pageId="21" pageNumber="22">Figs. 48D, 48E</figureCitation>
); wing pale sand coloured, transverse lines contrast, brown, fine, more or less straight, parallel to each other (
<figureCitation id="13602A0A6904FFB0FC2D42E5FC1A0BF1" box="[910,996,1999,2021]" captionStart="FIGURE 29" captionStartId="74.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@74.[151,1435,181,1836]" captionTargetPageId="74" captionText="FIGURE 29. Adults of Siberian Sterrhinae. A—Scopula aequifasciata; B—S. cajanderi; C—S. albiceraria; D—Cyclophora albipunctata." figureDoi="http://doi.org/10.5281/zenodo.7965161" httpUri="https://zenodo.org/record/7965161/files/figure.png" pageId="21" pageNumber="22">Fig. 29C</figureCitation>
)........................................................................................................ 
<taxonomicName id="4C5B4D0C6907FFB3FBB445BDFA630CB9" ID-CoL="93MDH" authority="(Herrich-Schaffer, 1844)" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1844" box="[1047,1437,151,173]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="albiceraria">
<emphasis id="B92FEA9D6907FFB3FBB445BDFB630CB9" box="[1047,1181,151,173]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FBB445BDFBDA0CB8" bold="true" box="[1047,1060,151,172]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FB9245BDFB630CB9" bold="true" box="[1073,1181,151,173]" pageId="22" pageNumber="23">albiceraria</emphasis>
</emphasis>
(Herrich-Schäffer, 1844)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF34459FFA620CF3" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34459FFA620CF3" pageId="22" pageNumber="23">- Anterior margin of sternite A8 convex medially, cerata with spinose tips (sometimes barely spinose); wing pattern different.. .................................................................................................. 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF3445DAFA620D49" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF3445DAFA630D36" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3445DAFA630D36" pageId="22" pageNumber="23">
23. Phallos almost straight, coecum of aedeagus not bent ventrally (
<figureCitation id="13602A0A6907FFB3FC9D45C5FC6A0D11" box="[830,916,239,261]" captionStart="FIGURE 46" captionStartId="91.[151,250,1867,1891]" captionTargetBox="[163,1425,198,1844]" captionTargetId="figure-15@91.[157,1431,181,1844]" captionTargetPageId="91" captionText="FIGURE 46. Male genitalia of Siberian Sterrhinae.A–C—Scopula beckeraria; D–F—S. incanata; G–I—S. immutata; J–L—S. frigidaria; M–P—S. ternata. A, D, G, K, O—genital segment; B, E, H, J, M, P—sternite A8; C, F, I, L, N—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965208" httpUri="https://zenodo.org/record/7965208/files/figure.png" pageId="22" pageNumber="23">Fig. 46L</figureCitation>
); ground colour of wings dark gray brown, discal spots and terminal dots absent (
<figureCitation id="13602A0A6907FFB3FE7A4427FDD10D36" box="[473,559,269,291]" captionStart="FIGURE 27" captionStartId="72.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@72.[151,1435,181,1836]" captionTargetPageId="72" captionText="FIGURE 27. Adults of Siberian Sterrhinae. A—Scopula incanata; B—S. immutata; C—S. frigidaria; D—S. ternata." figureDoi="http://doi.org/10.5281/zenodo.7965155" httpUri="https://zenodo.org/record/7965155/files/figure.png" pageId="22" pageNumber="23">Fig. 27C</figureCitation>
)............................................... 
<taxonomicName id="4C5B4D0C6907FFB3FBD74427FA630D36" ID-CoL="6XZ6B" authority="(Moschler, 1860)" baseAuthorityName="Moschler" baseAuthorityYear="1860" box="[1140,1437,268,290]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="frigidaria">
<emphasis id="B92FEA9D6907FFB3FBD74427FB130D36" box="[1140,1261,268,290]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FBD74427FB7F0D36" bold="true" box="[1140,1153,269,290]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FB2D4426FB130D36" bold="true" box="[1166,1261,268,290]" pageId="22" pageNumber="23">frigidaria</emphasis>
</emphasis>
(Möschler, 1860)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344401FA620D49" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344401FA620D49" pageId="22" pageNumber="23">- Phallos arched: apex and coecum of aedeagus slightly bent ventrally; ground colour of wings different, discal spots or/and terminal dots present.................................................................................. 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF34444FFA620E19" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF34444FFA630DA1" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34444FFA630DA1" pageId="22" pageNumber="23">
24. Sternite A8 with constriction in middle part, cerata very short, slightly curved (
<figureCitation id="13602A0A6907FFB3FC6C444FFBDA0D6F" box="[975,1060,357,379]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="22" pageNumber="23">Figs. 6A</figureCitation>
, 
<figureCitation id="13602A0A6907FFB3FB8E444FFBA60D6F" box="[1069,1112,357,379]" captionStart="FIGURE 46" captionStartId="91.[151,250,1867,1891]" captionTargetBox="[163,1425,198,1844]" captionTargetId="figure-15@91.[157,1431,181,1844]" captionTargetPageId="91" captionText="FIGURE 46. Male genitalia of Siberian Sterrhinae.A–C—Scopula beckeraria; D–F—S. incanata; G–I—S. immutata; J–L—S. frigidaria; M–P—S. ternata. A, D, G, K, O—genital segment; B, E, H, J, M, P—sternite A8; C, F, I, L, N—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965208" httpUri="https://zenodo.org/record/7965208/files/figure.png" pageId="22" pageNumber="23">46B</figureCitation>
); wings sand-coloured, with light brown tinge; terminal dots present, elongate; forewing apex pointed, forewing postmedial line dark brown, often discontinuous (
<figureCitation id="13602A0A6907FFB3FF4A44B5FEBF0DA1" box="[233,321,415,437]" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="22" pageNumber="23">Fig. 26D</figureCitation>
).................................................................... 
<taxonomicName id="4C5B4D0C6907FFB3FBDB44B5FA630DA1" ID-CoL="4VVVG" authority="(Lederer, 1853)" baseAuthorityName="Lederer" baseAuthorityYear="1853" box="[1144,1437,415,437]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="beckeraria">
<emphasis id="B92FEA9D6907FFB3FBDB44B5FB040DA1" box="[1144,1274,415,437]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FBDB44B5FB7B0DA0" bold="true" box="[1144,1157,415,436]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FB3244B5FB040DA1" bold="true" box="[1169,1274,415,437]" pageId="22" pageNumber="23">beckeraria</emphasis>
</emphasis>
(Lederer, 1853)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344497FA620E19" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344497FA620E19" pageId="22" pageNumber="23">
- Sternite A8 rounded, with cerata strongly curved (
<figureCitation id="13602A0A6907FFB3FD624497FCF10DC7" box="[705,783,445,467]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="22" pageNumber="23">Fig. 6D</figureCitation>
); ground colour of wings white with yellow suffusion, terminal dots absent or very small and pale; forewing apex not pointed, forewing postmedial line yellowish, usually continuous (
<figureCitation id="13602A0A6907FFB3FA9244F0FA790DFB" box="[1329,1415,474,496]" captionStart="FIGURE 27" captionStartId="72.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@72.[151,1435,181,1836]" captionTargetPageId="72" captionText="FIGURE 27. Adults of Siberian Sterrhinae. A—Scopula incanata; B—S. immutata; C—S. frigidaria; D—S. ternata." figureDoi="http://doi.org/10.5281/zenodo.7965155" httpUri="https://zenodo.org/record/7965155/files/figure.png" pageId="22" pageNumber="23">Fig. 27B</figureCitation>
)................................................. 
<taxonomicName id="4C5B4D0C6907FFB3FC8444DDFBAE0E19" ID-CoL="4VW3N" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[807,1104,503,525]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="immutata">
<emphasis id="B92FEA9D6907FFB3FC8444DDFC610E18" box="[807,927,503,525]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FC8444DDFCCA0E18" bold="true" box="[807,820,503,524]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FCE344DDFC610E18" bold="true" box="[832,927,503,524]" pageId="22" pageNumber="23">immutata</emphasis>
</emphasis>
(Linnaeus, 1758)
</taxonomicName>
(
<emphasis id="B92FEA9D6907FFB3FBFD44D2FA6A0E19" box="[1118,1428,503,525]" italics="true" pageId="22" pageNumber="23">rare form with both short cerata</emphasis>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF34473FFA620E71" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF34473FFA630E5C" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34473FFA630E5C" pageId="22" pageNumber="23">
25. Left fibula markedly smaller than right one (
<figureCitation id="13602A0A6907FFB3FD2D473FFD160E3E" box="[654,744,533,555]" captionStart="FIGURE 47" captionStartId="92.[151,250,1858,1882]" captionTargetBox="[155,1432,212,1827]" captionTargetId="figure-25@92.[151,1436,181,1834]" captionTargetPageId="92" captionText="FIGURE 47. Male genitalia of Siberian Sterrhinae. A–C—Scopula subpunctaria; D–F—S. prouti; G–I—S. flaccidaria; J–L— S. floslactata; M–P—S. aequifasciata. A, D, G, J, P—genital segment; B, E, H, L, O—sternite A8; C, F, I, K, M, N—phallos (M—lateral view, N—ventral view). Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965212" httpUri="https://zenodo.org/record/7965212/files/figure.png" pageId="22" pageNumber="23">Fig. 47G</figureCitation>
); hindwing termen clearly angled: wing shortly tailed at 
<emphasis id="B92FEA9D6907FFB3FAA4473FFAD90E3E" box="[1287,1319,533,554]" italics="true" pageId="22" pageNumber="23">M3</emphasis>
(
<figureCitation id="13602A0A6907FFB3FA97473FFA720E3E" box="[1332,1420,533,555]" captionStart="FIGURE 28" captionStartId="73.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@73.[151,1435,181,1836]" captionTargetPageId="73" captionText="FIGURE 28. Adults of Siberian Sterrhinae. A—Scopula subpunctaria; B—S. prouti; C—S. flaccidaria; D—S. floslactata." figureDoi="http://doi.org/10.5281/zenodo.7965159" httpUri="https://zenodo.org/record/7965159/files/figure.png" pageId="22" pageNumber="23">Fig. 28C</figureCitation>
).............................................................................. 
<taxonomicName id="4C5B4D0C6907FFB3FB284718FA630E5C" ID-CoL="4VVZY" authority="(Zeller, 1852)" baseAuthorityName="Zeller" baseAuthorityYear="1852" box="[1163,1437,562,584]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="flaccidaria">
<emphasis id="B92FEA9D6907FFB3FB284718FAF10E5C" box="[1163,1295,562,584]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FB284718FB660E53" bold="true" box="[1163,1176,562,583]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FB074718FAF10E5C" bold="true" box="[1188,1295,562,584]" pageId="22" pageNumber="23">flaccidaria</emphasis>
</emphasis>
(Zeller, 1852)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF34477AFA620E71" blockId="22.[151,1437,151,2021]" box="[151,1436,591,613]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34477AFA620E71" box="[151,1436,591,613]" pageId="22" pageNumber="23">
- Both fibulae about same size; hindwing termen rather smooth, wing not tailed at 
<emphasis id="B92FEA9D6907FFB3FC48477AFBF50E71" box="[1003,1035,592,613]" italics="true" pageId="22" pageNumber="23">M3</emphasis>
............................... 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344747FA620E8B" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344747FA630E96" blockId="22.[151,1437,151,2021]" box="[151,1437,620,643]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344747FA630E96" box="[151,1437,620,643]" pageId="22" pageNumber="23">
26. Both cerata of equal length: long, protruding beyond posterior margin of mappa (
<figureCitation id="13602A0A6907FFB3FC4E4747FBBC0E97" box="[1005,1090,621,643]" captionStart="FIGURE 46" captionStartId="91.[151,250,1867,1891]" captionTargetBox="[163,1425,198,1844]" captionTargetId="figure-15@91.[157,1431,181,1844]" captionTargetPageId="91" captionText="FIGURE 46. Male genitalia of Siberian Sterrhinae.A–C—Scopula beckeraria; D–F—S. incanata; G–I—S. immutata; J–L—S. frigidaria; M–P—S. ternata. A, D, G, K, O—genital segment; B, E, H, J, M, P—sternite A8; C, F, I, L, N—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965208" httpUri="https://zenodo.org/record/7965208/files/figure.png" pageId="22" pageNumber="23">Fig. 46E</figureCitation>
).... 
<taxonomicName id="4C5B4D0C6907FFB3FBD84747FA630E96" ID-CoL="4VW3W" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1147,1437,620,642]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="incanata">
<emphasis id="B92FEA9D6907FFB3FBD84747FB120E96" box="[1147,1260,621,642]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FBD84747FB760E96" bold="true" box="[1147,1160,621,642]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FB364747FB120E96" bold="true" box="[1173,1260,621,642]" pageId="22" pageNumber="23">incanata</emphasis>
</emphasis>
(Linnaeus, 1758)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF3447A1FA620E8B" blockId="22.[151,1437,151,2021]" box="[151,1436,650,672]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3447A1FA620E8B" box="[151,1436,650,672]" pageId="22" pageNumber="23">- Cerata of unequal length, only one of them protruding beyond posterior margin of mappa........................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344782FA620F26" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344782FA630F01" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344782FA630F01" pageId="22" pageNumber="23">
27. Postmedial line of forewing (often also of hindwing) distally bordered by wide, contrast, dark gray fascia (occasionally discontinuous); ground colour of wings reddish brown (in fresh samples) to grayish ochreous (
<figureCitation id="13602A0A6907FFB3FBD347EFFB350ECF" box="[1136,1227,709,731]" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="22" pageNumber="23">Fig. 26A</figureCitation>
); sternite A8 elongate (its length is about twice its width), left ceras very short: about 5 times shorter than right one (
<figureCitation id="13602A0A6907FFB3FBCB47C8FB480EEC" box="[1128,1206,738,760]" captionStart="FIGURE 45" captionStartId="90.[151,250,1859,1883]" captionTargetBox="[160,1424,191,1829]" captionTargetId="figure-25@90.[153,1434,181,1835]" captionTargetPageId="90" captionText="FIGURE 45. Male genitalia of Siberian Sterrhinae. A–C—Scopula ornata; D–F—S. decorata; G–I—S. permutata; J–L—S. rubiginata; M–O—S. impersonata. A, E, G, K, N—genital segment; B, D, I, J, O—sternite A8; C, F, H, L, M—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965206" httpUri="https://zenodo.org/record/7965206/files/figure.png" pageId="22" pageNumber="23">Fig. 45I</figureCitation>
)........................................................................................... 
<taxonomicName id="4C5B4D0C6907FFB3FBFF47D5FA630F01" ID-CoL="4VW9N" authority="(Staudinger, 1897)" baseAuthorityName="Staudinger" baseAuthorityYear="1897" box="[1116,1437,767,789]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="permutata">
<emphasis id="B92FEA9D6907FFB3FBFF47D5FB250F01" box="[1116,1243,767,789]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FBFF47D5FB970F00" bold="true" box="[1116,1129,767,788]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FBD5462AFB250F01" bold="true" box="[1142,1243,768,789]" pageId="22" pageNumber="23">permutata</emphasis>
</emphasis>
(Staudinger, 1897)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344637FA620F26" blockId="22.[151,1437,151,2021]" box="[151,1436,797,819]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344637FA620F26" box="[151,1436,797,819]" pageId="22" pageNumber="23">- Wings without wide, contrast, dark gray fascia distal of postmedial line; sternite A8 different........................ 28</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344610FA630FB3" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344610FA620F79" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344610FA620F79" pageId="22" pageNumber="23">
28. Sternite A8 with constriction in middle part; cerata slightly curved, right ceras slightly longer than left one (
<figureCitation id="13602A0A6907FFB3FABE4610FA950F44" box="[1309,1387,826,848]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="22" pageNumber="23">Fig. 6B</figureCitation>
); 
<emphasis id="B92FEA9D6907FFB3FA234610FDE70F79" italics="true" pageId="22" pageNumber="23">for external differences see point 24</emphasis>
.................................... 
<taxonomicName id="4C5B4D0C6907FFB3FC77467DFBA80F79" ID-CoL="4VVVG" baseAuthorityName="Lederer" baseAuthorityYear="1853" box="[980,1110,855,877]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="beckeraria">
<emphasis id="B92FEA9D6907FFB3FC77467DFBA80F79" box="[980,1110,855,877]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FC77467DFC1F0F78" bold="true" box="[980,993,855,876]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FC4E467DFBA80F79" bold="true" box="[1005,1110,855,877]" italics="true" pageId="22" pageNumber="23">beckeraria</emphasis>
</emphasis>
</taxonomicName>
(
<emphasis id="B92FEA9D6907FFB3FBC74672FA6A0F79" box="[1124,1428,855,877]" italics="true" pageId="22" pageNumber="23">rare form with both long cerata</emphasis>
)
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF34465FFA630FB3" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34465FFA630FB3" pageId="22" pageNumber="23">
- Sternite A8 without constriction in middle part, cerata strongly curved, right ceras 3–4 times longer than left one, semicircular (
<figureCitation id="13602A0A6907FFB3FF4A46B8FEC30FBC" box="[233,317,914,936]" captionStart="FIGURE 6" captionStartId="17.[151,250,1855,1879]" captionTargetBox="[163,1428,191,1826]" captionTargetId="figure-15@17.[153,1434,181,1832]" captionTargetPageId="17" captionText="FIGURE 6. Structure details of Scopula males. A–D—polymorphism of sternite A8 in some species; E–J—distal part of hindleg (tibia and tarsus).A, B—S. beckeraria; C, D—S. immutata; E—S. cajanderi; F—S. rubiginata; G—S. floslactata; H—S. subpunctaria; I—S. virgulata; J—S. corrivalaria." figureDoi="http://doi.org/10.5281/zenodo.7965083" httpUri="https://zenodo.org/record/7965083/files/figure.png" pageId="22" pageNumber="23">Figs. 6C</figureCitation>
, 
<figureCitation id="13602A0A6907FFB3FEE446B8FE8B0FBC" box="[327,373,914,936]" captionStart="FIGURE 46" captionStartId="91.[151,250,1867,1891]" captionTargetBox="[163,1425,198,1844]" captionTargetId="figure-15@91.[157,1431,181,1844]" captionTargetPageId="91" captionText="FIGURE 46. Male genitalia of Siberian Sterrhinae.A–C—Scopula beckeraria; D–F—S. incanata; G–I—S. immutata; J–L—S. frigidaria; M–P—S. ternata. A, D, G, K, O—genital segment; B, E, H, J, M, P—sternite A8; C, F, I, L, N—phallos. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965208" httpUri="https://zenodo.org/record/7965208/files/figure.png" pageId="22" pageNumber="23">46H</figureCitation>
); 
<emphasis id="B92FEA9D6907FFB3FE2646B8FD200FB3" box="[389,734,914,936]" italics="true" pageId="22" pageNumber="23">for external differences see point 24</emphasis>
................................................ 
<taxonomicName id="4C5B4D0C6907FFB3FA8746B8FA630FB3" ID-CoL="4VW3N" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1316,1437,914,936]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="immutata">
<emphasis id="B92FEA9D6907FFB3FA8746B8FA630FB3" box="[1316,1437,914,936]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FA8746B8FACF0FB3" bold="true" box="[1316,1329,914,935]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA9D46B8FA630FB3" bold="true" box="[1342,1437,914,935]" italics="true" pageId="22" pageNumber="23">immutata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF34469AFA620FEB" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF34469AFA620FF6" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34469AFA620FF6" pageId="22" pageNumber="23">
29. Corpus bursae drop-shaped, its posterior part (1/3 to 1/2) strongly sclerotized, ductus bursae extremely short, signum consists of few tiny sclerites (near 10) (
<figureCitation id="13602A0A6907FFB3FDA946E7FD9F0FF7" box="[522,609,973,995]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="22" pageNumber="23">Fig. 53A</figureCitation>
).......................................................... 
<taxonomicName id="4C5B4D0C6907FFB3FA8946E7FA620FF6" ID-CoL="4VWFN" baseAuthorityName="Denis et Schiffermuller" baseAuthorityYear="1775" box="[1322,1436,972,994]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="virgulata">
<emphasis id="B92FEA9D6907FFB3FA8946E7FA620FF6" box="[1322,1436,972,994]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FA8946E7FAC90FF6" bold="true" box="[1322,1335,973,994]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FAE046E6FA620FF6" bold="true" box="[1347,1436,972,994]" italics="true" pageId="22" pageNumber="23">virgulata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF3446C1FA620FEB" blockId="22.[151,1437,151,2021]" box="[151,1436,1002,1024]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3446C1FA620FEB" box="[151,1436,1002,1024]" pageId="22" pageNumber="23">- Genitalia different................................................................................... 30</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344122FA62082E" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344122FA620809" blockId="22.[151,1437,151,2021]" box="[151,1436,1031,1053]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344122FA620809" box="[151,1436,1031,1053]" pageId="22" pageNumber="23">
30. Ductus bursae broad, with lateral longitudinal sclerotization (
<figureCitation id="13602A0A6907FFB3FC8D412DFC780809" box="[814,902,1031,1053]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Fig. 52A</figureCitation>
)............................... 
<taxonomicName id="4C5B4D0C6907FFB3FAAD412DFA620809" ID-CoL="6XZ7X" baseAuthorityName="Kretschmar" baseAuthorityYear="1862" box="[1294,1436,1031,1053]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corrivalaria">
<emphasis id="B92FEA9D6907FFB3FAAD412DFA620809" box="[1294,1436,1031,1053]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FAAD412DFAE50808" bold="true" box="[1294,1307,1031,1052]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA84412DFA620809" bold="true" box="[1319,1436,1031,1053]" italics="true" pageId="22" pageNumber="23">corrivalaria</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF34410FFA62082E" blockId="22.[151,1437,151,2021]" box="[151,1436,1061,1083]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34410FFA62082E" box="[151,1436,1061,1083]" pageId="22" pageNumber="23">- Ductus bursae different................................................................................ 31</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344168FA6208BB" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344168FA630860" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344168FA630860" pageId="22" pageNumber="23">
31. Ductus bursae very short and broad, posterior parts of corpus bursae with granulated surface opposite signum patch (
<figureCitation id="13602A0A6907FFB3FAD64168FEF70861" captionStart="FIGURE 51" captionStartId="96.[151,250,1867,1891]" captionTargetBox="[166,1421,193,1840]" captionTargetId="figure-25@96.[158,1429,181,1844]" captionTargetPageId="96" captionText="FIGURE 51. Female genitalia of Siberian Sterrhinae. A—Idaea straminata; B—I. dimidiata; C—I. descitaria; D—Holarctias rufinaria; E—H. rufinularia; F—Scopula agutsaensis; G—S. immorata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965222" httpUri="https://zenodo.org/record/7965222/files/figure.png" pageId="22" pageNumber="23">Fig. 51F</figureCitation>
).................................................................................... 
<taxonomicName id="4C5B4D0C6907FFB3FAB34175FA630860" authorityName="Vasilenko" authorityYear="1997" box="[1296,1437,1119,1141]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="agutsaensis">
<emphasis id="B92FEA9D6907FFB3FAB34175FA630860" box="[1296,1437,1119,1141]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FAB34175FAE30860" bold="true" box="[1296,1309,1119,1140]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA894175FA630860" bold="true" box="[1322,1437,1119,1140]" italics="true" pageId="22" pageNumber="23">agutsaensis</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344157FA6208BB" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344157FA6208BB" pageId="22" pageNumber="23">- Ductus bursae commonly narrow and moderately long or short and broad; posterior parts of corpus bursae without granulated surface opposite signum patch.......................................................................... 32</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344192FA6208FF" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344192FA6208D9" blockId="22.[151,1437,151,2021]" box="[151,1436,1206,1229]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344192FA6208D9" box="[151,1436,1206,1229]" pageId="22" pageNumber="23">
32. Ostium broad (as broad as ½ width of posterior margin of tergite A8 or broader), well sclerotized (
<figureCitation id="13602A0A6907FFB3FB31419DFB0C08D9" box="[1170,1266,1207,1229]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Figs. 52B</figureCitation>
, 
<figureCitation id="13602A0A6907FFB3FB5F419DFAAB08D9" box="[1276,1365,1207,1229]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="22" pageNumber="23">53E, 53F</figureCitation>
)... 33
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF3441FFFA6208FF" blockId="22.[151,1437,151,2021]" box="[151,1436,1237,1259]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3441FFFA6208FF" box="[151,1436,1237,1259]" pageId="22" pageNumber="23">- Genitalia different................................................................................... 35</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF3441D8FA620931" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF3441D8FA620913" blockId="22.[151,1437,151,2021]" box="[151,1436,1266,1288]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3441D8FA620913" box="[151,1436,1266,1288]" pageId="22" pageNumber="23">33. Lamella antevaginalis not developed; ostium opened, funnel-shaped (tapered anteriorly), attached to narrow antrum...... 34</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF34403AFA620931" blockId="22.[151,1437,151,2021]" box="[151,1436,1295,1317]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34403AFA620931" box="[151,1436,1295,1317]" pageId="22" pageNumber="23">
- Ostium not funnel-shaped, lamella antevaginalis well developed: wide, near oval, covers ostium (
<figureCitation id="13602A0A6907FFB3FB294025FB1E0931" box="[1162,1248,1295,1317]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Fig. 52B</figureCitation>
)..... 
<taxonomicName id="4C5B4D0C6907FFB3FA854025FA620931" ID-CoL="94DMS" baseAuthorityName="Fourcroy" baseAuthorityYear="1785" box="[1318,1436,1295,1317]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="virginalis">
<emphasis id="B92FEA9D6907FFB3FA854025FA620931" box="[1318,1436,1295,1317]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FA854025FACD0930" bold="true" box="[1318,1331,1295,1316]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA9C4025FA620931" bold="true" box="[1343,1436,1295,1317]" italics="true" pageId="22" pageNumber="23">virginalis</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344007FA6309C0" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344007FA620969" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344007FA620969" pageId="22" pageNumber="23">
34. Wings brown, light brown or rosy brown, discal spot of forewing very small, often indistinct (
<figureCitation id="13602A0A6907FFB3FBC84007FB3D0957" box="[1131,1219,1325,1347]" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="22" pageNumber="23">Fig. 26B</figureCitation>
); ductus bursae longer than in 
<taxonomicName id="4C5B4D0C6907FFB3FE894060FE440974" ID-CoL="4VW3P" baseAuthorityName="Walker" baseAuthorityYear="1861" box="[298,442,1354,1376]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="impersonata">
<emphasis id="B92FEA9D6907FFB3FE894060FE440974" box="[298,442,1354,1376]" italics="true" pageId="22" pageNumber="23">S. impersonata</emphasis>
</taxonomicName>
, signum well developed: numerous spinules form oval or drop-shaped patch (
<figureCitation id="13602A0A6907FFB3FB054060FB050974" box="[1190,1275,1354,1376]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="22" pageNumber="23">Fig. 53E</figureCitation>
)..................................................................................................... 
<taxonomicName id="4C5B4D0C6907FFB3FAB8404DFA620969" ID-CoL="4VWBQ" baseAuthorityName="Hufnagel" baseAuthorityYear="1767" box="[1307,1436,1383,1405]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rubiginata">
<emphasis id="B92FEA9D6907FFB3FAB8404DFA620969" box="[1307,1436,1383,1405]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FAB8404DFAD60968" bold="true" box="[1307,1320,1383,1404]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA97404DFA620969" bold="true" box="[1332,1436,1383,1405]" italics="true" pageId="22" pageNumber="23">rubiginata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF3440AFFA6309C0" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3440AFFA6309C0" pageId="22" pageNumber="23">
- Wings different shades of gray, discal spot of forewing well pronounced (
<figureCitation id="13602A0A6907FFB3FC3840AFFC0D098F" box="[923,1011,1413,1435]" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="22" pageNumber="23">Fig. 26C</figureCitation>
); ductus bursae shorter than in 
<taxonomicName id="4C5B4D0C6907FFB3FAB940AFFA68098E" ID-CoL="4VWBQ" baseAuthorityName="Hufnagel" baseAuthorityYear="1767" box="[1306,1430,1413,1434]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rubiginata">
<emphasis id="B92FEA9D6907FFB3FAB940AFFA68098E" box="[1306,1430,1413,1434]" italics="true" pageId="22" pageNumber="23">S. rubiginata</emphasis>
</taxonomicName>
, signum weakly developed: few spinules unevenly scattered around narrow (often broken) longitudinal sclerite (
<figureCitation id="13602A0A6907FFB3FAB34088FA9D09AC" box="[1296,1379,1442,1464]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="22" pageNumber="23">Fig. 53F</figureCitation>
)............................................................................................ 
<taxonomicName id="4C5B4D0C6907FFB3FAAB4095FA6309C0" ID-CoL="4VW3P" baseAuthorityName="Walker" baseAuthorityYear="1861" box="[1288,1437,1471,1493]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="impersonata">
<emphasis id="B92FEA9D6907FFB3FAAB4095FA6309C0" box="[1288,1437,1471,1493]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FAAB4095FAEB09C0" bold="true" box="[1288,1301,1471,1492]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA814095FA6309C0" bold="true" box="[1314,1437,1471,1492]" italics="true" pageId="22" pageNumber="23">impersonata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF3440F7FA620A1B" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF3440F7FA6209E7" blockId="22.[151,1437,151,2021]" box="[151,1436,1501,1523]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3440F7FA6209E7" box="[151,1436,1501,1523]" pageId="22" pageNumber="23">35. Signum absent...................................................................................... 36</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF3440D1FA620A1B" blockId="22.[151,1437,151,2021]" box="[151,1436,1530,1552]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3440D1FA620A1B" box="[151,1436,1530,1552]" pageId="22" pageNumber="23">- Signum present...................................................................................... 37</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF34433DFA620A91" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF34433DFA620A5E" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF34433DFA620A5E" pageId="22" pageNumber="23">
36. Wings pale yellow, transverse lines yellowish brown (
<figureCitation id="13602A0A6907FFB3FD48433DFCBA0A39" box="[747,836,1559,1581]" captionStart="FIGURE 29" captionStartId="74.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@74.[151,1435,181,1836]" captionTargetPageId="74" captionText="FIGURE 29. Adults of Siberian Sterrhinae. A—Scopula aequifasciata; B—S. cajanderi; C—S. albiceraria; D—Cyclophora albipunctata." figureDoi="http://doi.org/10.5281/zenodo.7965161" httpUri="https://zenodo.org/record/7965161/files/figure.png" pageId="22" pageNumber="23">Fig. 29C</figureCitation>
); lamella antevaginalis rather slender, crescent-shaped, ductus bursae comparatively narrow and long (
<figureCitation id="13602A0A6907FFB3FDC3431FFD480A5F" box="[608,694,1589,1611]" captionStart="FIGURE 55" captionStartId="100.[151,250,1867,1891]" captionTargetBox="[157,1426,188,1842]" captionTargetId="figure-25@100.[151,1436,181,1844]" captionTargetPageId="100" captionText="FIGURE 55. Female genitalia of Siberian Sterrhinae. A—Scopula flaccidaria; B—S. cajanderi; C—S. albiceraria; D— Cyclophora albipunctata; E—Timandra rectistrigaria; F—T. griseata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965230" httpUri="https://zenodo.org/record/7965230/files/figure.png" pageId="22" pageNumber="23">Fig. 55C</figureCitation>
).................................................. 
<taxonomicName id="4C5B4D0C6907FFB3FAB4431FFA620A5E" ID-CoL="93MDH" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1844" box="[1303,1436,1588,1610]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="albiceraria">
<emphasis id="B92FEA9D6907FFB3FAB4431FFA620A5E" box="[1303,1436,1588,1610]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FAB4431FFADA0A5E" bold="true" box="[1303,1316,1589,1610]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA93431EFA620A5E" bold="true" box="[1328,1436,1588,1610]" italics="true" pageId="22" pageNumber="23">albiceraria</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344379FA620A91" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344379FA620A91" pageId="22" pageNumber="23">
- Wings pale gray (rarely with yellowish tinge) with dark gray transverse lines (
<figureCitation id="13602A0A6907FFB3FC6B4378FBDE0A73" box="[968,1056,1618,1640]" captionStart="FIGURE 29" captionStartId="74.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@74.[151,1435,181,1836]" captionTargetPageId="74" captionText="FIGURE 29. Adults of Siberian Sterrhinae. A—Scopula aequifasciata; B—S. cajanderi; C—S. albiceraria; D—Cyclophora albipunctata." figureDoi="http://doi.org/10.5281/zenodo.7965161" httpUri="https://zenodo.org/record/7965161/files/figure.png" pageId="22" pageNumber="23">Fig. 29B</figureCitation>
); lamella antevaginalis larger and wider than in 
<taxonomicName id="4C5B4D0C6907FFB3FE89435AFE500A90" ID-CoL="93MDH" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1844" box="[298,430,1647,1669]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="albiceraria">
<emphasis id="B92FEA9D6907FFB3FE89435AFE500A90" box="[298,430,1647,1669]" italics="true" pageId="22" pageNumber="23">S. albiceraria</emphasis>
</taxonomicName>
, ductus bursae shorter and broader than in 
<taxonomicName id="4C5B4D0C6907FFB3FCEF435AFC2E0A90" ID-CoL="93MDH" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1844" box="[844,976,1647,1669]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="albiceraria">
<emphasis id="B92FEA9D6907FFB3FCEF435AFC2E0A90" box="[844,976,1647,1669]" italics="true" pageId="22" pageNumber="23">S. albiceraria</emphasis>
</taxonomicName>
(
<figureCitation id="13602A0A6907FFB3FC7E4345FBCD0A91" box="[989,1075,1647,1669]" captionStart="FIGURE 55" captionStartId="100.[151,250,1867,1891]" captionTargetBox="[157,1426,188,1842]" captionTargetId="figure-25@100.[151,1436,181,1844]" captionTargetPageId="100" captionText="FIGURE 55. Female genitalia of Siberian Sterrhinae. A—Scopula flaccidaria; B—S. cajanderi; C—S. albiceraria; D— Cyclophora albipunctata; E—Timandra rectistrigaria; F—T. griseata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965230" httpUri="https://zenodo.org/record/7965230/files/figure.png" pageId="22" pageNumber="23">Fig. 55B</figureCitation>
)................... 
<taxonomicName id="4C5B4D0C6907FFB3FA854345FA620A91" ID-CoL="4VVVY" baseAuthorityName="Herz" baseAuthorityYear="1904" box="[1318,1436,1647,1669]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cajanderi">
<emphasis id="B92FEA9D6907FFB3FA854345FA620A91" box="[1318,1436,1647,1669]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FA854345FACD0A90" bold="true" box="[1318,1331,1647,1668]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA9C4345FA620A91" bold="true" box="[1343,1436,1647,1669]" italics="true" pageId="22" pageNumber="23">cajanderi</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF3443A7FA620AAB" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF3443A7FA620AB6" blockId="22.[151,1437,151,2021]" box="[151,1436,1677,1699]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3443A7FA620AB6" box="[151,1436,1677,1699]" pageId="22" pageNumber="23">
37. Ostium near symmetric, commonly with semicircular or circular structures (
<figureCitation id="13602A0A6907FFB3FC1E43A7FBE00AB7" box="[957,1054,1677,1699]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Figs. 52C</figureCitation>
, 
<figureCitation id="13602A0A6907FFB3FB8443A7FB400AB7" box="[1063,1214,1677,1699]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="22" pageNumber="23">53D, 53G, 53H</figureCitation>
, 
<figureCitation id="13602A0A6907FFB3FB6443A7FADC0AB7" box="[1223,1314,1677,1699]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="22" pageNumber="23">54A–54F</figureCitation>
, 
<figureCitation id="13602A0A6907FFB3FA8F43A7FAA80AB7" box="[1324,1366,1677,1699]" captionStart="FIGURE 55" captionStartId="100.[151,250,1867,1891]" captionTargetBox="[157,1426,188,1842]" captionTargetId="figure-25@100.[151,1436,181,1844]" captionTargetPageId="100" captionText="FIGURE 55. Female genitalia of Siberian Sterrhinae. A—Scopula flaccidaria; B—S. cajanderi; C—S. albiceraria; D— Cyclophora albipunctata; E—Timandra rectistrigaria; F—T. griseata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965230" httpUri="https://zenodo.org/record/7965230/files/figure.png" pageId="22" pageNumber="23">55A</figureCitation>
)... 39
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344381FA620AAB" blockId="22.[151,1437,151,2021]" box="[151,1436,1706,1728]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344381FA620AAB" box="[151,1436,1706,1728]" pageId="22" pageNumber="23">
- Ostium asymmetric, without semicircular or circular structures, consists of several irregular sclerites (
<figureCitation id="13602A0A6907FFB3FB104380FABD0AD4" box="[1203,1347,1706,1728]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Figs. 52E, 52F</figureCitation>
).... 38
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF3443E2FA620B21" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF3443E2FA620AEE" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3443E2FA620AEE" pageId="22" pageNumber="23">
38. Wings with sharp discal spots and usually with almost continuous terminal lines, fringe of forewing without dark dots (
<figureCitation id="13602A0A6907FFB3FAD643EDFEF00AEF" captionStart="FIGURE 25" captionStartId="70.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@70.[151,1435,181,1836]" captionTargetPageId="70" captionText="FIGURE 25. Adults of Siberian Sterrhinae. A—Scopula nigropunctata; B—S. virgulata; C—S. ornata; D—S. decorata." figureDoi="http://doi.org/10.5281/zenodo.7965151" httpUri="https://zenodo.org/record/7965151/files/figure.png" pageId="22" pageNumber="23">Fig. 25A</figureCitation>
); lamella antevaginalis with distinct, irregular folds (
<figureCitation id="13602A0A6907FFB3FCA943CFFCA00AEF" box="[778,862,1765,1787]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Fig. 52F</figureCitation>
)................................. 
<taxonomicName id="4C5B4D0C6907FFB3FB5443CFFA620AEE" ID-CoL="4VW7T" baseAuthorityName="Hufnagel" baseAuthorityYear="1767" box="[1271,1436,1765,1786]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigropunctata">
<emphasis id="B92FEA9D6907FFB3FB5443CFFA620AEE" box="[1271,1436,1765,1786]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FB5443CFFAFA0AEE" bold="true" box="[1271,1284,1765,1786]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FAB343CFFA620AEE" bold="true" box="[1296,1436,1765,1786]" italics="true" pageId="22" pageNumber="23">nigropunctata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF344229FA620B21" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344229FA620B21" pageId="22" pageNumber="23">
- Discal spots commonly absent, terminal lines fine, discontinuous, usually broken into short and weak dashes or dots, fringe of forewing with dark dots (
<figureCitation id="13602A0A6907FFB3FE764235FDD10B21" box="[469,559,1823,1845]" captionStart="FIGURE 24" captionStartId="69.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@69.[151,1435,181,1836]" captionTargetPageId="69" captionText="FIGURE 24. Adults of Siberian Sterrhinae. A—Scopula virginalis; B—S. dignata; C—S. nemoraria; D—S. umbelaria." figureDoi="http://doi.org/10.5281/zenodo.7965145" httpUri="https://zenodo.org/record/7965145/files/figure.png" pageId="22" pageNumber="23">Fig. 24D</figureCitation>
); lamella antevaginalis with large lateral (left) sclerite (
<figureCitation id="13602A0A6907FFB3FBE14235FB680B21" box="[1090,1174,1823,1845]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="22" pageNumber="23">Fig. 52E</figureCitation>
)........... 
<taxonomicName id="4C5B4D0C6907FFB3FABD4235FA620B21" ID-CoL="93MFZ" baseAuthorityName="Hubner" baseAuthorityYear="1813" box="[1310,1436,1823,1845]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="umbelaria">
<emphasis id="B92FEA9D6907FFB3FABD4235FA620B21" box="[1310,1436,1823,1845]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FABD4235FAD50B20" bold="true" box="[1310,1323,1823,1844]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA944235FA620B21" bold="true" box="[1335,1436,1823,1845]" italics="true" pageId="22" pageNumber="23">umbelaria</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB3FF344217FA620BBE" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344217FA620B99" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344217FA620B99" pageId="22" pageNumber="23">
39. Postmedial line of forewing distinct, zigzagging, distally bordered by wide, contrast, dark gray fascia (occasionally discontinuous); ground colour of wings reddish brown (in fresh samples) to grayish ochreous (
<figureCitation id="13602A0A6907FFB3FBCD4270FB390B64" box="[1134,1223,1882,1904]" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="22" pageNumber="23">Fig. 26A</figureCitation>
); genitalia: 
<figureCitation id="13602A0A6907FFB3FA9B4270FAD50B98" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="22" pageNumber="23">
Fig. 53D........................................................................................... 
<taxonomicName id="4C5B4D0C6907FFB3FABD425DFAD50B98" ID-CoL="4VW9N" baseAuthorityName="Staudinger" baseAuthorityYear="1897" box="[1310,1323,1911,1932]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="permutata">
<emphasis id="B92FEA9D6907FFB3FABD425DFAD50B98" bold="true" box="[1310,1323,1911,1932]" italics="true" pageId="22" pageNumber="23">S</emphasis>
</taxonomicName>
</figureCitation>
. 
<emphasis id="B92FEA9D6907FFB3FA944252FA620B99" bold="true" box="[1335,1436,1912,1933]" italics="true" pageId="22" pageNumber="23">permutata</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6907FFB3FF3442BFFA620BBE" blockId="22.[151,1437,151,2021]" box="[151,1436,1941,1963]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF3442BFFA620BBE" box="[151,1436,1941,1963]" pageId="22" pageNumber="23">- Wing pattern different................................................................................. 40</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6907FFB2FF344298FA620CDE" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<paragraph id="8BE4368F6907FFB3FF344298FA630BF1" blockId="22.[151,1437,151,2021]" pageId="22" pageNumber="23">
<keyLead id="30AA95BA6907FFB3FF344298FA630BF1" pageId="22" pageNumber="23">
40. Corpus bursae near round, barely elongated; signum near ductus bursae, very small, not elongated, of undefined shape, spinules very small (
<figureCitation id="13602A0A6907FFB3FEF742E5FE590BF1" box="[340,423,1999,2021]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="22" pageNumber="23">Fig. 54F</figureCitation>
)........................................................................ 
<taxonomicName id="4C5B4D0C6907FFB3FABF42E5FA630BF1" ID-CoL="4VW25" baseAuthorityName="Haworth" baseAuthorityYear="1809" box="[1308,1437,1999,2021]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="floslactata">
<emphasis id="B92FEA9D6907FFB3FABF42E5FA630BF1" box="[1308,1437,1999,2021]" italics="true" pageId="22" pageNumber="23">
<emphasis id="B92FEA9D6907FFB3FABF42E5FAD70BF0" bold="true" box="[1308,1321,1999,2020]" italics="true" pageId="22" pageNumber="23">S</emphasis>
. 
<emphasis id="B92FEA9D6907FFB3FA9542E5FA630BF1" bold="true" box="[1334,1437,1999,2021]" italics="true" pageId="22" pageNumber="23">floslactata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3445B2FA620CDE" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3445B2FA620CDE" pageId="23" pageNumber="24">- Corpus bursae more or less elongated, signum rather large, oval or elongated, usually located in central part of corpus bursae .................................................................................................. 41</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF3445F8FA620D36" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF3445F8FA630D11" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3445F8FA630D11" pageId="23" pageNumber="24">
41. Lamella postvaginalis well developed, lamella antevaginalis and lamella postvaginalis flat (without folds or bulging), forming transverse cleft, in center of which ostium is located (
<figureCitation id="13602A0A6906FFB2FD4745C5FCC40D11" box="[740,826,239,261]" captionStart="FIGURE 52" captionStartId="97.[151,250,1867,1891]" captionTargetBox="[159,1430,196,1841]" captionTargetId="figure-15@97.[153,1434,181,1844]" captionTargetPageId="97" captionText="FIGURE 52. Female genitalia of Siberian Sterrhinae. A—Scopula corrivalaria; B—S. virginalis; C—S. dignata; D—S. nemoraria; E—S. umbelaria; F—S. nigropunctata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965224" httpUri="https://zenodo.org/record/7965224/files/figure.png" pageId="23" pageNumber="24">Fig. 52C</figureCitation>
).......................................... 
<taxonomicName id="4C5B4D0C6906FFB2FA9B45C5FA630D11" ID-CoL="79YW9" baseAuthorityName="Guenee" baseAuthorityYear="1858" box="[1336,1437,239,261]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="dignata">
<emphasis id="B92FEA9D6906FFB2FA9B45C5FA630D11" box="[1336,1437,239,261]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FA9B45C5FABB0D10" bold="true" box="[1336,1349,239,260]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FAF145C5FA630D11" bold="true" box="[1362,1437,239,261]" italics="true" pageId="23" pageNumber="24">dignata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF344427FA620D36" blockId="23.[151,1437,151,1053]" box="[151,1436,269,291]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF344427FA620D36" box="[151,1436,269,291]" pageId="23" pageNumber="24">- Lamella postvaginalis not developed, ostium usually surrounded by semicircular or ring-shaped lamella antevaginalis.... 42</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF344400FA620DA1" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF344400FA620D6E" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF344400FA620D6E" pageId="23" pageNumber="24">
42. Ground colour glossy white, transverse lines yellowish gray; postmedial line of forewing sharply bent inwards near costa, forming right or acute angle (
<figureCitation id="13602A0A6906FFB2FDA0446DFD630D49" box="[515,669,327,349]" captionStart="FIGURE 28" captionStartId="73.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@73.[151,1435,181,1836]" captionTargetPageId="73" captionText="FIGURE 28. Adults of Siberian Sterrhinae. A—Scopula subpunctaria; B—S. prouti; C—S. flaccidaria; D—S. floslactata." figureDoi="http://doi.org/10.5281/zenodo.7965159" httpUri="https://zenodo.org/record/7965159/files/figure.png" pageId="23" pageNumber="24">Figs. 28A, 28B</figureCitation>
); lamella antevaginalis posteriorly and laterally strongly sclerotized, anteriorly weakly sclerotized, not swollen, without transverse fold (
<figureCitation id="13602A0A6906FFB2FCA4444FFC640D6F" box="[775,922,357,379]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="23" pageNumber="24">Figs. 54D, 54E</figureCitation>
)........................................ 43
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3444A9FA620DA1" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3444A9FA620DA1" pageId="23" pageNumber="24">- Ground colour different or postmedial line of forewing not bent inwards near costa; lamella antevaginalis commonly anteriorly strongly sclerotized, or swollen, or with transverse fold...................................................... 44</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF344497FA620E53" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF344497FA630DE4" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF344497FA630DE4" pageId="23" pageNumber="24">
43. Lamella antevaginalis rather round; signum with distinct longitudinal medial ridge of fused spinules (
<figureCitation id="13602A0A6906FFB2FB134497FAF60DC7" box="[1200,1288,445,467]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="23" pageNumber="24">Fig. 54D</figureCitation>
).................................................................................................. 
<taxonomicName id="4C5B4D0C6906FFB2FAA344F0FA630DE4" ID-CoL="4VWDS" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1847" box="[1280,1437,474,496]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="subpunctaria">
<emphasis id="B92FEA9D6906FFB2FAA344F0FA630DE4" box="[1280,1437,474,496]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FAA344F0FAF30DFB" bold="true" box="[1280,1293,474,495]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FAB944F0FA630DE4" bold="true" box="[1306,1437,474,496]" italics="true" pageId="23" pageNumber="24">subpunctaria</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3444D2FA620E53" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3444D2FA620E53" pageId="23" pageNumber="24">
- Lamella antevaginalis rather elongated, its anterior end markedly oblique relatively posterior end; fused spinules of signum does not form more or less continuous longitudinal medial ridge but usually forms few groups along medial axis (
<figureCitation id="13602A0A6906FFB2FA85473FFA850E3F" box="[1318,1403,533,555]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="23" pageNumber="24">Fig. 54E</figureCitation>
).............................................................................................. 
<taxonomicName id="4C5B4D0C6906FFB2FAE44718FA620E53" ID-CoL="6XZ6R" authorityName="Djakonov" authorityYear="1935" box="[1351,1436,562,584]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="prouti">
<emphasis id="B92FEA9D6906FFB2FAE44718FA620E53" box="[1351,1436,562,584]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FAE44718FAAA0E53" bold="true" box="[1351,1364,562,583]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FAC34718FA620E53" bold="true" box="[1376,1436,562,583]" italics="true" pageId="23" pageNumber="24">prouti</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF34477AFA620EB4" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF34477AFA630E96" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF34477AFA630E96" pageId="23" pageNumber="24">
44. Sterigma with two lateral longitudinal sclerotized folds, signum very slender, spinules often fused to medial ridge (
<figureCitation id="13602A0A6906FFB2FA924765FA770E71" box="[1329,1417,591,613]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="23" pageNumber="24">Fig. 54A</figureCitation>
).......................................................................................... 
<taxonomicName id="4C5B4D0C6906FFB2FA874747FA630E96" ID-CoL="4VW3N" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1316,1437,621,642]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="immutata">
<emphasis id="B92FEA9D6906FFB2FA874747FA630E96" box="[1316,1437,621,642]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FA874747FACF0E96" bold="true" box="[1316,1329,621,642]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FA9D4747FA630E96" bold="true" box="[1342,1437,621,642]" italics="true" pageId="23" pageNumber="24">immutata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3447A1FA620EB4" blockId="23.[151,1437,151,1053]" box="[151,1436,650,672]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3447A1FA620EB4" box="[151,1436,650,672]" pageId="23" pageNumber="24">- Sterigma different: without lateral longitudinal sclerotized folds, signum more oval................................ 45</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF34478DFA620EEC" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF34478DFA630ECE" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF34478DFA630ECE" pageId="23" pageNumber="24">
45. Hindwing termen clearly angled: wing shortly tailed at 
<emphasis id="B92FEA9D6906FFB2FD554782FCE80EA9" box="[758,790,680,701]" italics="true" pageId="23" pageNumber="24">M3</emphasis>
, tornus almost right angled (
<figureCitation id="13602A0A6906FFB2FB89478DFB7C0EA9" box="[1066,1154,679,701]" captionStart="FIGURE 28" captionStartId="73.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@73.[151,1435,181,1836]" captionTargetPageId="73" captionText="FIGURE 28. Adults of Siberian Sterrhinae. A—Scopula subpunctaria; B—S. prouti; C—S. flaccidaria; D—S. floslactata." figureDoi="http://doi.org/10.5281/zenodo.7965159" httpUri="https://zenodo.org/record/7965159/files/figure.png" pageId="23" pageNumber="24">Fig. 28C</figureCitation>
); genitalia: 
<figureCitation id="13602A0A6906FFB2FB50478DFADB0ECE" captionStart="FIGURE 55" captionStartId="100.[151,250,1867,1891]" captionTargetBox="[157,1426,188,1842]" captionTargetId="figure-25@100.[151,1436,181,1844]" captionTargetPageId="100" captionText="FIGURE 55. Female genitalia of Siberian Sterrhinae. A—Scopula flaccidaria; B—S. cajanderi; C—S. albiceraria; D— Cyclophora albipunctata; E—Timandra rectistrigaria; F—T. griseata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965230" httpUri="https://zenodo.org/record/7965230/files/figure.png" pageId="23" pageNumber="24">
Fig. 55A................................................................................................ 
<taxonomicName id="4C5B4D0C6906FFB2FABB47EFFADB0ECE" ID-CoL="4VVZY" baseAuthorityName="Zeller" baseAuthorityYear="1852" box="[1304,1317,709,730]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="flaccidaria">
<emphasis id="B92FEA9D6906FFB2FABB47EFFADB0ECE" bold="true" box="[1304,1317,709,730]" italics="true" pageId="23" pageNumber="24">S</emphasis>
</taxonomicName>
</figureCitation>
. 
<emphasis id="B92FEA9D6906FFB2FA9147EEFA630ECE" bold="true" box="[1330,1437,708,730]" italics="true" pageId="23" pageNumber="24">flaccidaria</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3447C9FA620EEC" blockId="23.[151,1437,151,1053]" box="[151,1436,738,760]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3447C9FA620EEC" box="[151,1436,738,760]" pageId="23" pageNumber="24">- Hindwing rounded, termen rather smooth................................................................. 46</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF3447D5FA620F79" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF3447D5FA620F26" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3447D5FA620F26" pageId="23" pageNumber="24">46. Wings without discal spots (in rare cases discal dot very weak, not darker than medial line) and without terminal dots, forewing apex not pointed..................................................................................... 47</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF344611FA620F79" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF344611FA620F79" pageId="23" pageNumber="24">- Wings with distinct discal spots (they always darker than medial line) and distinct terminal dots, forewing apex slightly pointed............................................................................................ 48</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF34465FFA630FBC" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF34465FFA630F9E" blockId="23.[151,1437,151,1053]" box="[151,1437,885,907]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF34465FFA630F9E" box="[151,1437,885,907]" pageId="23" pageNumber="24">
47. Lamella antevaginalis semicircular, slightly swollen anteriorly (
<figureCitation id="13602A0A6906FFB2FCE5465FFC620F9F" box="[838,924,885,907]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="23" pageNumber="24">Fig. 54C</figureCitation>
).................................. 
<taxonomicName id="4C5B4D0C6906FFB2FA9F465FFA630F9E" ID-CoL="6XZ9G" authorityName="Schrank" authorityYear="1802" box="[1340,1437,885,906]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ternata">
<emphasis id="B92FEA9D6906FFB2FA9F465FFA630F9E" box="[1340,1437,885,906]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FA9F465FFAB70F9E" bold="true" box="[1340,1353,885,906]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FAF5465FFA630F9E" bold="true" box="[1366,1437,885,906]" italics="true" pageId="23" pageNumber="24">ternata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3446B9FA630FBC" blockId="23.[151,1437,151,1053]" box="[151,1437,914,936]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3446B9FA630FBC" box="[151,1437,914,936]" pageId="23" pageNumber="24">
- Lamella antevaginalis oval, not swollen anteriorly (
<figureCitation id="13602A0A6906FFB2FD7346B8FCD90FBC" box="[720,807,914,936]" captionStart="FIGURE 54" captionStartId="99.[151,250,1864,1888]" captionTargetBox="[166,1432,188,1833]" captionTargetId="figure-15@99.[151,1436,181,1840]" captionTargetPageId="99" captionText="FIGURE 54. Female genitalia of Siberian Sterrhinae. A—Scopula immutata; B—S. frigidaria; C—S. ternata; D—S. subpunctaria; E—S. prouti; F—S. floslactata; G—S. aequifasciata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965228" httpUri="https://zenodo.org/record/7965228/files/figure.png" pageId="23" pageNumber="24">Fig. 54B</figureCitation>
)......................................... 
<taxonomicName id="4C5B4D0C6906FFB2FA8746B8FA630FBC" ID-CoL="6XZ6B" baseAuthorityName="Moschler" baseAuthorityYear="1860" box="[1316,1437,914,936]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="frigidaria">
<emphasis id="B92FEA9D6906FFB2FA8746B8FA630FBC" box="[1316,1437,914,936]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FA8746B8FACF0FB3" bold="true" box="[1316,1329,914,935]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FA9D46B8FA630FBC" bold="true" box="[1342,1437,914,936]" italics="true" pageId="23" pageNumber="24">frigidaria</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AF2E2A6906FFB2FF34469AFA620809" pageId="23" pageNumber="24">
<paragraph id="8BE4368F6906FFB2FF34469AFA630FF6" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF34469AFA630FF6" pageId="23" pageNumber="24">
48. Lamella antevaginalis with swollen C-shaped pad (
<figureCitation id="13602A0A6906FFB2FD6E4685FCD80FD1" box="[717,806,943,965]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="23" pageNumber="24">Fig. 53H</figureCitation>
); ground colour of wings gray to pale gray (
<figureCitation id="13602A0A6906FFB2FB6D4685FADB0FD1" box="[1230,1317,943,965]" captionStart="FIGURE 27" captionStartId="72.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-25@72.[151,1435,181,1836]" captionTargetPageId="72" captionText="FIGURE 27. Adults of Siberian Sterrhinae. A—Scopula incanata; B—S. immutata; C—S. frigidaria; D—S. ternata." figureDoi="http://doi.org/10.5281/zenodo.7965155" httpUri="https://zenodo.org/record/7965155/files/figure.png" pageId="23" pageNumber="24">Fig. 27A</figureCitation>
).................................................................................................... 
<taxonomicName id="4C5B4D0C6906FFB2FA8F46E7FA630FF6" ID-CoL="4VW3W" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1324,1437,973,994]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="incanata">
<emphasis id="B92FEA9D6906FFB2FA8F46E7FA630FF6" box="[1324,1437,973,994]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FA8F46E7FAC70FF6" bold="true" box="[1324,1337,973,994]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FAE546E7FA630FF6" bold="true" box="[1350,1437,973,994]" italics="true" pageId="23" pageNumber="24">incanata</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE4368F6906FFB2FF3446C1FA620809" blockId="23.[151,1437,151,1053]" pageId="23" pageNumber="24">
<keyLead id="30AA95BA6906FFB2FF3446C1FA620809" pageId="23" pageNumber="24">
- Lamella antevaginalis with strongly sclerotized transverse fold (
<figureCitation id="13602A0A6906FFB2FCEC46C0FC570814" box="[847,937,1002,1024]" captionStart="FIGURE 53" captionStartId="98.[151,250,1867,1891]" captionTargetBox="[159,1429,188,1838]" captionTargetId="figure-25@98.[153,1434,181,1844]" captionTargetPageId="98" captionText="FIGURE 53. Female genitalia of Siberian Sterrhinae. A—Scopula virgulata; B—S. ornata; C—S. decorata; D—S. permutata; E—S. rubiginata; F—S. impersonata; G—S. beckeraria; H—S. incanata. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.7965226" httpUri="https://zenodo.org/record/7965226/files/figure.png" pageId="23" pageNumber="24">Fig. 53G</figureCitation>
); wings sand coloured, with light brown tinge (
<figureCitation id="13602A0A6906FFB2FAD646C0FEF30809" captionStart="FIGURE 26" captionStartId="71.[151,250,1859,1883]" captionTargetBox="[152,1434,182,1835]" captionTargetId="figure-15@71.[151,1435,181,1836]" captionTargetPageId="71" captionText="FIGURE 26. Adults of Siberian Sterrhinae. A—Scopula permutata; B—S. rubiginata; C—S. impersonata; D—S. beckeraria." figureDoi="http://doi.org/10.5281/zenodo.7965153" httpUri="https://zenodo.org/record/7965153/files/figure.png" pageId="23" pageNumber="24">Fig. 26D</figureCitation>
)..................................................................................... 
<taxonomicName id="4C5B4D0C6906FFB2FAB9412DFA620809" ID-CoL="4VVVG" baseAuthorityName="Lederer" baseAuthorityYear="1853" box="[1306,1436,1031,1053]" class="Insecta" family="Geometridae" genus="Scopula" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="beckeraria">
<emphasis id="B92FEA9D6906FFB2FAB9412DFA620809" box="[1306,1436,1031,1053]" italics="true" pageId="23" pageNumber="24">
<emphasis id="B92FEA9D6906FFB2FAB9412DFAD90808" bold="true" box="[1306,1319,1031,1052]" italics="true" pageId="23" pageNumber="24">S</emphasis>
. 
<emphasis id="B92FEA9D6906FFB2FA90412DFA620809" bold="true" box="[1331,1436,1031,1053]" italics="true" pageId="23" pageNumber="24">beckeraria</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>