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<document id="AF1375442F16A1F6843D0BDAC53E4CC5" ID-CLB-Dataset="30511" ID-DOI="10.11646/zootaxa.4394.3.7" ID-GBIF-Dataset="cfaab9c0-29f0-4f25-90d3-0b810c656bc1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1199826" ID-ZooBank="29891032-F46F-45F5-B9CE-290271FBDC11" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521176689468" checkinUser="plazi" docAuthor="Novák, János &amp; Harvey, Mark S." docDate="2018" docId="03F2878BA930FFE2FF12FC0AFAEC3FC1" docLanguage="en" docName="zootaxa.4394.3.7.pdf" docOrigin="Zootaxa 4394 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geogarypus muchmorei Novák &amp; Harvey 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="421" masterDocId="FFCBFFF3A931FFE6FF85FFF8FFCA3C09" masterDocTitle="New species and records of the pseudoscorpion genus Geogarypus (Pseudoscorpiones: Geogarypidae) from India, Sri Lanka and New Guinea" masterLastPageNumber="427" masterPageNumber="417" pageNumber="418" updateTime="1698531126992" updateUser="ExternalLinkService">
<mods:mods id="02DF13D465B219D382AA83A286050C86" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="90D30EA103C2BC9DF633431B25D199B7">New species and records of the pseudoscorpion genus Geogarypus (Pseudoscorpiones: Geogarypidae) from India, Sri Lanka and New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="E6BCB0CF98A1BF928B0414803771FC17" type="personal">
<mods:role id="4C936111992F1FFF0B4F534D45EA9AB1">
<mods:roleTerm id="C6CE02DF89E1885BC0FFE00F2A385BFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E67B5E2815A22083075BEAD03503995">Novák, János</mods:namePart>
</mods:name>
<mods:name id="7B5DA04252E0BFA341CA22CA6F4FAFE6" type="personal">
<mods:role id="18F2B47DF695A72DBDF2BAED70ECF3A6">
<mods:roleTerm id="6DD362B05EB733C7C2A1064EAA09ADB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D927F67D6620AEC9EEB3FAC1574869B">Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0C200C6C5B8B6D6A01EFAD90D33A1AEF">text</mods:typeOfResource>
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<mods:title id="208D7743D0D8AC49753137994550CEDB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="481E060701DEDD075968524BAC34B00B">
<mods:date id="AAF5B065A8660006D609FD1523BDDB29">2018</mods:date>
<mods:detail id="4E636BE7DE79B4B2065EC50A4EE30894" type="pubDate">
<mods:number id="3A7627D950C6A0FC34680994F3D91FD9">2018-03-15</mods:number>
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<mods:detail id="1B464160769854EBDC4908DD7B7C70CB" type="volume">
<mods:number id="8C7F563CC8D062F67392A77CB9C5BC98">4394</mods:number>
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<mods:number id="EDF9443BDFF99FF12A81B8DA1334EDE4">3</mods:number>
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<mods:classification id="18FF2B90AEBB33575E82004BF518CC65">journal article</mods:classification>
<mods:identifier id="1F857510789AD29D1768448D6FA17356" type="CLB-Dataset">30511</mods:identifier>
<mods:identifier id="4CE5B038069A3C8706165DB86315D565" type="DOI">10.11646/zootaxa.4394.3.7</mods:identifier>
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<mods:identifier id="AD4EC5747590F0D269D652EA4E27B730" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F2878BA930FFE2FF12FC0AFAEC3FC1" ID-DOI="http://doi.org/10.5281/zenodo.5954798" ID-GBIF-Taxon="142289384" ID-Zenodo-Dep="5954798" LSID="urn:lsid:plazi:treatment:03F2878BA930FFE2FF12FC0AFAEC3FC1" httpUri="http://treatment.plazi.org/id/03F2878BA930FFE2FF12FC0AFAEC3FC1" lastPageId="4" lastPageNumber="421" pageId="1" pageNumber="418">
<subSubSection id="C3416516A930FFE7FF12FC0AFEE33827" pageId="1" pageNumber="418" type="nomenclature">
<paragraph id="8BE4369DA930FFE7FF12FC0AFDD53805" blockId="1.[151,543,1010,1070]" box="[151,543,1010,1036]" pageId="1" pageNumber="418">
<heading id="D0AC81F1A930FFE7FF12FC0AFDD53805" bold="true" box="[151,543,1010,1036]" fontSize="11" level="1" pageId="1" pageNumber="418" reason="1">
<emphasis id="B92FEA8FA930FFE7FF12FC0AFDD53805" bold="true" box="[151,543,1010,1036]" pageId="1" pageNumber="418">
<taxonomicName id="4C5B4D1EA930FFE7FF12FC0AFE733805" ID-CoL="3FPSV" authority="Novák &amp; Harvey, 2018" authorityName="Novák &amp; Harvey" authorityYear="2018" box="[151,441,1010,1036]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="418" phylum="Arthropoda" rank="species" species="muchmorei" status="sp. nov.">
<emphasis id="B92FEA8FA930FFE7FF12FC0AFE733805" bold="true" box="[151,441,1010,1036]" italics="true" pageId="1" pageNumber="418">Geogarypus muchmorei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57F4A930FFE7FE44FC0AFDD53805" box="[449,543,1010,1036]" pageId="1" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE4369DA930FFE7FF12FBEDFEE33827" blockId="1.[151,543,1010,1070]" box="[151,297,1045,1070]" pageId="1" pageNumber="418">
(
<figureCitation id="13602A18A930FFE7FF1AFBEDFEEB3827" box="[159,289,1045,1070]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="1" pageNumber="418">Figs 1AM</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3416516A930FFE7FF12FBA5FE723923" pageId="1" pageNumber="418" type="materials_examined">
<paragraph id="8BE4369DA930FFE7FF12FBA5FE723923" blockId="1.[151,1436,1117,2006]" pageId="1" pageNumber="418">
<materialsCitation id="3B333CC0A930FFE7FF12FBA5FEC138B7" ID-GBIF-Occurrence="1835647440" collectingDate="1967-09-23" collectionCode="HNHM" collectorName="Gyorgy Topal" country="India" location="Daitari" municipality="Jajpur-Keonjahr District" pageId="1" pageNumber="418" specimenCount="1" specimenCount-male="1" stateProvince="Orissa" typeStatus="holotype">
<emphasis id="B92FEA8FA930FFE7FF12FBA5FE8F387F" bold="true" box="[151,325,1117,1142]" pageId="1" pageNumber="418">
<typeStatus id="54E0883FA930FFE7FF12FBA5FF18387F" box="[151,210,1117,1142]" pageId="1" pageNumber="418">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54E0883FA930FFE7FEC9FBA5FE73387F" box="[332,441,1117,1142]" pageId="1" pageNumber="418" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D5DFD14A930FFE7FE41FBA6FE13387C" box="[452,473,1118,1141]" pageId="1" pageNumber="418" type="male"></specimenCount>
(
<collectionCode id="ED4AAE58A930FFE7FE6DFBA6FD8A387F" box="[488,576,1118,1142]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="418">HNHM</collectionCode>
Pseud-1988;
<figureCitation id="13602A18A930FFE7FD5FFBA5FCF8387F" box="[730,818,1117,1142]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="1" pageNumber="418">Fig. 1A</figureCitation>
),
<location id="8E846046A930FFE7FCC2FBA5FC5F387F" LSID="urn:lsid:plazi:treatment:03F2878BA930FFE2FF12FC0AFAEC3FC1:8E846046A930FFE7FCC2FBA5FC5F387F" box="[839,917,1117,1142]" country="India" municipality="Jajpur-Keonjahr District" name="Daitari" pageId="1" pageNumber="418" stateProvince="Orissa">Daitari</location>
[21°07ʹN, 85°50ʹE],
<collectingMunicipality id="6B80ACE7A930FFE7FB01FBA5FA52387F" box="[1156,1432,1117,1142]" pageId="1" pageNumber="418">Jajpur-Keonjahr District</collectingMunicipality>
,
<collectingRegion id="499FF87FA930FFE7FF12FB79FF283893" box="[151,226,1153,1178]" country="India" name="Odisha" pageId="1" pageNumber="418">Orissa</collectingRegion>
,
<collectingCountry id="F34C760DA930FFE7FF74FB79FEE73893" box="[241,301,1153,1178]" name="India" pageId="1" pageNumber="418">India</collectingCountry>
,
<collectingDate id="EFA1E9B5A930FFE7FEB8FB7AFE2E3893" box="[317,484,1153,1178]" pageId="1" pageNumber="418" value="1967-09-23">
<date id="FFE5105DA930FFE7FEB8FB7AFE2E3893" box="[317,484,1153,1178]" pageId="1" pageNumber="418" value="1967-09-23">23 Sept. 1967</date>
</collectingDate>
, beaten from bushes in forest, leg.
<collectorName id="26AE534BA930FFE7FC0FFB7AFBE03893" box="[906,1066,1153,1178]" pageId="1" pageNumber="418">György Topál</collectorName>
(in good condition, right chela dissected)
</materialsCitation>
.
<materialsCitation id="3B333CC0A930FFE7FE90FB5EFC6638EB" ID-GBIF-Occurrence="1835647298" collectingDate="1967-05-07" collectionCode="HNHM" collectorName="Gyorgy Topal" country="India" location="Barkalikapur" pageId="1" pageNumber="418" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="54E0883FA930FFE7FE90FB5EFE4C38B7" box="[277,390,1190,1214]" pageId="1" pageNumber="418" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5DFD14A930FFE7FE14FB5EFE7138B4" box="[401,443,1190,1214]" pageId="1" pageNumber="418" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4AAE58A930FFE7FE4FFB5EFDE838B7" box="[458,546,1190,1214]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="418">HNHM</collectionCode>
Pseud-1989;
<figureCitation id="13602A18A930FFE7FD38FB5DFCDB38B7" box="[701,785,1189,1214]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="1" pageNumber="418">Fig. 1B</figureCitation>
),
<location id="8E846046A930FFE7FCACFB5DFC7138B7" LSID="urn:lsid:plazi:treatment:03F2878BA930FFE2FF12FC0AFAEC3FC1:8E846046A930FFE7FCACFB5DFC7138B7" box="[809,955,1189,1214]" country="India" name="Barkalikapur" pageId="1" pageNumber="418" stateProvince="West Bengal">Barkalikapur</location>
[22°23ʹN, 88°14ʹE],
<collectingRegion id="499FF87FA930FFE7FB2FFB5EFAF038B7" box="[1194,1338,1189,1214]" country="India" name="West Bengal" pageId="1" pageNumber="418">West Bengal</collectingRegion>
,
<collectingCountry id="F34C760DA930FFE7FAC3FB5DFA4838B7" box="[1350,1410,1189,1214]" name="India" pageId="1" pageNumber="418">India</collectingCountry>
,
<collectingDate id="EFA1E9B5A930FFE7FA0BFB5EFEC438EB" pageId="1" pageNumber="418" value="1967-05-07">
<date id="FFE5105DA930FFE7FA0BFB5EFEC438EB" pageId="1" pageNumber="418" value="1967-05-07">7 May 1967</date>
</collectingDate>
, associated with bat (
<taxonomicName id="4C5B4D1EA930FFE7FD81FB31FD0B38E8" authorityName="E. Geoffroy" authorityYear="1810" box="[516,705,1225,1249]" class="Mammalia" family="Megadermatidae" genus="Megaderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="418" phylum="Chordata" rank="species" species="lyra">
<emphasis id="B92FEA8FA930FFE7FD81FB31FD0B38E8" box="[516,705,1225,1249]" italics="true" pageId="1" pageNumber="418">Megaderma lyra</emphasis>
</taxonomicName>
), leg.
<collectorName id="26AE534BA930FFE7FC89FB32FC6638EB" box="[780,940,1225,1250]" pageId="1" pageNumber="418">György Topál</collectorName>
</materialsCitation>
;
<materialsCitation id="3B333CC0A930FFE7FC3CFB32FE723923" ID-GBIF-Occurrence="1835647296" collectingDate="1967-01-01" collectionCode="HNHM" collectorName="Gyorgy Topal" country="India" location="Daitari" municipality="Keonjahr District" pageId="1" pageNumber="418" specimenCount="1" specimenCount-female="1" stateProvince="Orissa" typeStatus="paratype">
<specimenCount id="9D5DFD14A930FFE7FC3CFB32FC2E38E8" box="[953,996,1226,1250]" pageId="1" pageNumber="418" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4AAE58A930FFE7FC70FB32FB8638EB" box="[1013,1100,1226,1250]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="418">HNHM</collectionCode>
Pseud-1990),
<location id="8E846046A930FFE7FB74FB31FA8838EB" LSID="urn:lsid:plazi:treatment:03F2878BA930FFE2FF12FC0AFAEC3FC1:8E846046A930FFE7FB74FB31FA8838EB" box="[1265,1346,1225,1250]" country="India" municipality="Keonjahr District" name="Daitari" pageId="1" pageNumber="418" stateProvince="Orissa">Daitari</location>
, Jajpur-
<collectingMunicipality id="6B80ACE7A930FFE7FF12FB15FE96390F" box="[151,348,1261,1286]" pageId="1" pageNumber="418">Keonjahr District</collectingMunicipality>
,
<collectingRegion id="499FF87FA930FFE7FEE2FB15FE78390F" box="[359,434,1261,1286]" country="India" name="Odisha" pageId="1" pageNumber="418">Orissa</collectingRegion>
,
<collectingCountry id="F34C760DA930FFE7FE39FB15FE32390F" box="[444,504,1261,1286]" name="India" pageId="1" pageNumber="418">India</collectingCountry>
,
<collectingDate id="EFA1E9B5A930FFE7FD86FB16FD4E390F" box="[515,644,1261,1286]" pageId="1" pageNumber="418" value="1967-01-01">
<date id="FFE5105DA930FFE7FD86FB16FD4E390F" box="[515,644,1261,1286]" pageId="1" pageNumber="418" value="1967-01-01">1 Jan. 1967</date>
</collectingDate>
, netting in grasses from undergrowth in forest, near brook, leg.
<collectorName id="26AE534BA930FFE7FAC2FB16FF1C3923" pageId="1" pageNumber="418">György Topál</collectorName>
(in poor condition).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3416516A930FFE7FF42FACDFEBD397B" pageId="1" pageNumber="418" type="etymology">
<paragraph id="8BE4369DA930FFE7FF42FACDFEBD397B" blockId="1.[151,1436,1117,2006]" pageId="1" pageNumber="418">
<emphasis id="B92FEA8FA930FFE7FF42FACDFE843947" bold="true" box="[199,334,1333,1358]" pageId="1" pageNumber="418">Etymology.</emphasis>
This species is named after the late William B. Muchmore, for his contributions to our knowledge of pseudoscorpions.
</paragraph>
</subSubSection>
<subSubSection id="C3416516A930FFE7FF42FA85FD08399F" box="[199,706,1405,1430]" pageId="1" pageNumber="418" type="distribution">
<paragraph id="8BE4369DA930FFE7FF42FA85FD08399F" blockId="1.[151,1436,1117,2006]" box="[199,706,1405,1430]" pageId="1" pageNumber="418">
<emphasis id="B92FEA8FA930FFE7FF42FA85FEAA399F" bold="true" box="[199,352,1405,1430]" pageId="1" pageNumber="418">Distribution.</emphasis>
<collectingRegion id="499FF87FA930FFE7FEE2FA85FE64399F" box="[359,430,1405,1430]" country="India" name="Odisha" pageId="1" pageNumber="418">Orissa</collectingRegion>
and
<collectingRegion id="499FF87FA930FFE7FE60FA86FDBF399F" box="[485,629,1405,1430]" country="India" name="West Bengal" pageId="1" pageNumber="418">West Bengal</collectingRegion>
,
<collectingCountry id="F34C760DA930FFE7FD04FA85FD74399F" box="[641,702,1405,1430]" name="India" pageId="1" pageNumber="418">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3416516A930FFE7FF42FA59FEC53A9B" pageId="1" pageNumber="418" type="diagnosis">
<paragraph id="8BE4369DA930FFE7FF42FA59FEC53A9B" blockId="1.[151,1436,1117,2006]" pageId="1" pageNumber="418">
<emphasis id="B92FEA8FA930FFE7FF42FA59FE8B39B3" bold="true" box="[199,321,1441,1466]" pageId="1" pageNumber="418">Diagnosis.</emphasis>
Carapace dark brown, apart from 2 cone-shaped white areas in posterior third. Prolateral face of chelal hand swollen. Dorsal face of fixed chelal finger with dorsal constriction near middle. Without prolateral constriction of chelal hand at base of fingers. Trichobothrium
<emphasis id="B92FEA8FA930FFE7FCC8FA12FC963A08" box="[845,860,1514,1537]" italics="true" pageId="1" pageNumber="418">it</emphasis>
closer to
<emphasis id="B92FEA8FA930FFE7FC4EFA12FC153A08" box="[971,991,1514,1537]" italics="true" pageId="1" pageNumber="418">et</emphasis>
than to
<emphasis id="B92FEA8FA930FFE7FBBFFA12FB9F3A08" box="[1082,1109,1514,1537]" italics="true" pageId="1" pageNumber="418">ist</emphasis>
. Trichobothrium
<emphasis id="B92FEA8FA930FFE7FA9FFA12FAE83A08" box="[1306,1314,1514,1537]" italics="true" pageId="1" pageNumber="418">t</emphasis>
two-thirds from base of movable finger,
<taxonomicName id="4C5B4D1EA930FFE7FE60F9F6FD903A2F" box="[485,602,1549,1574]" pageId="1" pageNumber="418" rank="subSpecies" subSpecies="midway">
<emphasis id="B92FEA8FA930FFE7FE60F9F6FE323A2C" box="[485,504,1550,1573]" italics="true" pageId="1" pageNumber="418">st</emphasis>
midway
</taxonomicName>
between
<emphasis id="B92FEA8FA930FFE7FD43F9F6FD043A2C" box="[710,718,1550,1573]" italics="true" pageId="1" pageNumber="418">t</emphasis>
and
<emphasis id="B92FEA8FA930FFE7FC83F9F5FCDE3A2C" box="[774,788,1549,1573]" italics="true" pageId="1" pageNumber="418">b</emphasis>
. Distance between
<taxonomicName id="4C5B4D1EA930FFE7FC74F9F6FBF93A2F" box="[1009,1075,1549,1574]" pageId="1" pageNumber="418" rank="subSpecies" subSpecies="and">
<emphasis id="B92FEA8FA930FFE7FC74F9F6FBCE3A2C" box="[1009,1028,1550,1573]" italics="true" pageId="1" pageNumber="418">st</emphasis>
and
</taxonomicName>
<emphasis id="B92FEA8FA930FFE7FBBEF9F5FB9E3A2C" box="[1083,1108,1549,1573]" italics="true" pageId="1" pageNumber="418">sb</emphasis>
twice that between
<emphasis id="B92FEA8FA930FFE7FAB3F9F5FA853A2C" box="[1334,1359,1549,1573]" italics="true" pageId="1" pageNumber="418">sb</emphasis>
and
<emphasis id="B92FEA8FA930FFE7FA02F9F5FA5F3A2C" box="[1415,1429,1549,1573]" italics="true" pageId="1" pageNumber="418">b</emphasis>
. Teeth of fixed chelal finger homodentate, triangular and retrorse, 37 accessory teeth on prolateral side. Retrolateral face of fixed finger with 46 pit-like structures. See Remarks for comparisons with other species from the region.
</paragraph>
</subSubSection>
<subSubSection id="C3416516A930FFE4FF42F965FA453BB1" lastPageId="2" lastPageNumber="419" pageId="1" pageNumber="418" type="description">
<paragraph id="8BE4369DA930FFE7FF42F965FB923B84" blockId="1.[151,1436,1117,2006]" pageId="1" pageNumber="418">
<emphasis id="B92FEA8FA930FFE7FF42F965FE933ABF" bold="true" box="[199,345,1693,1718]" pageId="1" pageNumber="418">Description.</emphasis>
Carapace dark brown, with the exception of 2 cone-shaped white areas in posterior third which are smaller in male than in female. Chela dark brown, other pedipalpal segments somewhat lighter. Legs yellowishbrown, without darker stripes. Tergites generally yellowish brown, with dark brown pattern; all tergites with 2 lateral brown spots, each connected by a dark band running along anterior half of tergites (these are wider in female than in male), with the exception of tergite I, which lacks such a band, and tergite XI, which is entirely brown; tergites III with a larger and elongated medial dark spot; tergite III without medial spot. Sternites IVX with 2 dark medial spots. Sternites yellowish-brown, IVIX with 2 dark brown medial spots.
</paragraph>
<paragraph id="8BE4369DA930FFE4FF42F862FEB03CF1" blockId="1.[151,1436,1117,2006]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="419" pageId="1" pageNumber="418">
<emphasis id="B92FEA8FA930FFE7FF42F862FEFF3BB8" box="[199,309,1946,1969]" italics="true" pageId="1" pageNumber="418">Carapace</emphasis>
. Sub-triangular, with 4 corneate eyes and cucullus. Covered with star-shaped granules and small, mostly clavate and some acuminate setae (
<figureCitation id="13602A18A930FFE7FD01F845FCC33BDF" box="[644,777,1981,2006]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="1" pageNumber="418">Figs 1C, D</figureCitation>
), including 2 small and 2 longer clavate setae with three terminal denticles near anterior margin (
<figureCitation id="13602A18A933FFE4FDDFFF6FFD643CB9" box="[602,686,151,176]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">Fig. 1E</figureCitation>
) and 12 near posterior margin. Two pairs of slit-like lyrifissures in posterior third and one pair in ocular region. Anterior transverse furrow well-developed, posterior furrow weak and situated sub-basally.
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42FEFCFE953D81" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42FEFCFEFC3D15" box="[199,310,260,284]" italics="true" pageId="2" pageNumber="419">Chelicera</emphasis>
(
<figureCitation id="13602A18A933FFE4FECEFEFCFE6F3D15" box="[331,421,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">Fig. 1F</figureCitation>
). Hand with 5 acuminate setae and movable finger with 1 acuminate subdistal seta. Subterminal teeth of fingers poorly developed. Galea without rami in both sexes. Rallum composed of 1 smooth blade. Serrula exterior with 1314 blades, the distal one enlarged, straight and pointed, the basal 2 enlarged and hooked (
<figureCitation id="13602A18A933FFE4FF7CFE97FE983D81" box="[249,338,367,392]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42FE6CFD6B3FAD" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42FE6CFEE63DA5" box="[199,300,404,428]" italics="true" pageId="2" pageNumber="419">Pedipalp</emphasis>
(
<figureCitation id="13602A18A933FFE4FEB8FE6CFE533DA5" box="[317,409,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">Fig. 1H</figureCitation>
). All pedipalpal segments granulate, except the movable chelal finger and the distal twothirds of fixed finger. Vestitural setae short, most of them weakly clavate, some acuminate (
<figureCitation id="13602A18A933FFE4FB1EFE4FFACC3DD9" box="[1179,1286,439,464]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">
Figs
<date id="FFE5105DA933FFE4FB56FE40FB263DD9" box="[1235,1260,440,464]" pageId="2" pageNumber="419">1I</date>
, J
</figureCitation>
). Dorsal face of patella with 4 lyrifissures near pedicel and one at distal margin. Prolateral face of trochanter rounded. Fixed chelal finger (
<figureCitation id="13602A18A933FFE4FEBCFE07FE583E11" box="[313,402,511,536]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">Fig. 1K</figureCitation>
) with 26 slightly separated teeth in male and
<quantity id="4CA39B78A933FFE4FC2EFDF8FBDB3E11" box="[939,1041,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.096" metricValueMin="5.842" pageId="2" pageNumber="419" unit="in" value="23.5" valueMax="24.0" valueMin="23.0">2324 in</quantity>
females, which are triangular and retrorse, the basal ones flattened, accompanied by 6 accessory teeth in male and
<quantity id="4CA39B78A933FFE4FBA1FDDCFBA13E34" box="[1060,1131,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.778" metricValueMin="0.762" pageId="2" pageNumber="419" unit="in" value="5.0" valueMax="7.0" valueMin="3.0">37 in</quantity>
females on prolateral side, between
<emphasis id="B92FEA8FA933FFE4FF7EFDB0FEC03E56" box="[251,266,584,607]" italics="true" pageId="2" pageNumber="419">it</emphasis>
and
<emphasis id="B92FEA8FA933FFE4FEC5FDB0FE9C3E69" box="[320,342,584,608]" italics="true" pageId="2" pageNumber="419">ib</emphasis>
. In retrolateral view they are obscured by the regular teeth. Retrolateral face of fixed chelal finger with 46 pit-like structures between
<emphasis id="B92FEA8FA933FFE4FDB0FD95FD853E8D" box="[565,591,621,644]" italics="true" pageId="2" pageNumber="419">ist</emphasis>
and
<emphasis id="B92FEA8FA933FFE4FD0DFD94FD673E8D" box="[648,685,620,644]" italics="true" pageId="2" pageNumber="419">esb</emphasis>
. Movable chelal finger with 2023 slightly separated teeth, which are retrorse and pointed in distal half, those in proximal half progressively becoming rounded. Dental row on movable finger reaching to
<emphasis id="B92FEA8FA933FFE4FE4CFD4CFE283EC5" box="[457,482,692,716]" italics="true" pageId="2" pageNumber="419">sb</emphasis>
. Nodus ramosus not discernible. Prolateral face of hand swollen, without constriction and with 1 long, straight spine near base of fixed finger. Fixed finger as long as hand with pedicel. Dorsal face of fixed finger undulate in lateral view, with dorsal constriction near middle of finger. Fixed finger with 8 trichobothria, movable finger with 4 trichobothria;
<emphasis id="B92FEA8FA933FFE4FD56FCD8FD283F3E" box="[723,738,800,823]" italics="true" pageId="2" pageNumber="419">it</emphasis>
closer to
<emphasis id="B92FEA8FA933FFE4FCCAFCD9FCA93F31" box="[847,867,801,824]" italics="true" pageId="2" pageNumber="419">et</emphasis>
than to
<emphasis id="B92FEA8FA933FFE4FC38FCD8FC1D3F3E" box="[957,983,800,823]" italics="true" pageId="2" pageNumber="419">ist</emphasis>
,
<emphasis id="B92FEA8FA933FFE4FC63FCD8FBC13F31" box="[998,1035,800,824]" italics="true" pageId="2" pageNumber="419">esb</emphasis>
usually near base of finger, but in
<specimenCount id="9D5DFD14A933FFE4FA0BFCD8FF293F54" pageId="2" pageNumber="419" type="female">1 female</specimenCount>
<typeStatus id="54E0883FA933FFE4FF69FCBDFE863F54" box="[236,332,837,861]" pageId="2" pageNumber="419" type="paratype">paratype</typeStatus>
(HNHM Pseud-1990) it is situated halfway between
<emphasis id="B92FEA8FA933FFE4FC29FCBCFC0D3F55" box="[940,967,836,860]" italics="true" pageId="2" pageNumber="419">eb</emphasis>
and
<emphasis id="B92FEA8FA933FFE4FB85FCBCFBDC3F55" box="[1024,1046,836,860]" italics="true" pageId="2" pageNumber="419">ib</emphasis>
(the trichobothriotaxy of the right chela of this specimen is normal);
<emphasis id="B92FEA8FA933FFE4FDA5FC91FDE23F89" box="[544,552,873,896]" italics="true" pageId="2" pageNumber="419">t</emphasis>
two-thirds from base of movable finger;
<taxonomicName id="4C5B4D1EA933FFE4FC7AFC91FBBE3F89" box="[1023,1140,871,896]" pageId="2" pageNumber="419" rank="subSpecies" subSpecies="midway">
<emphasis id="B92FEA8FA933FFE4FC7AFC91FBD83F89" box="[1023,1042,873,896]" italics="true" pageId="2" pageNumber="419">st</emphasis>
midway
</taxonomicName>
between
<emphasis id="B92FEA8FA933FFE4FB67FC91FB203F89" box="[1250,1258,873,896]" italics="true" pageId="2" pageNumber="419">t</emphasis>
and
<emphasis id="B92FEA8FA933FFE4FAA7FC90FAFA3F89" box="[1314,1328,872,896]" italics="true" pageId="2" pageNumber="419">b</emphasis>
; distance between
<taxonomicName id="4C5B4D1EA933FFE4FF7EFC75FEF73FAC" box="[251,317,908,933]" pageId="2" pageNumber="419" rank="subSpecies" subSpecies="and">
<emphasis id="B92FEA8FA933FFE4FF7EFC75FEC43FAD" box="[251,270,909,932]" italics="true" pageId="2" pageNumber="419">st</emphasis>
and
</taxonomicName>
<emphasis id="B92FEA8FA933FFE4FEC1FC74FE973FAD" box="[324,349,908,932]" italics="true" pageId="2" pageNumber="419">sb</emphasis>
twice that between
<emphasis id="B92FEA8FA933FFE4FDB8FC74FD9C3FAD" box="[573,598,908,932]" italics="true" pageId="2" pageNumber="419">sb</emphasis>
and
<emphasis id="B92FEA8FA933FFE4FD09FC74FD503FAD" box="[652,666,908,932]" italics="true" pageId="2" pageNumber="419">b</emphasis>
.
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42FC48FF20395C" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42FC48FE913FC1" box="[199,347,944,968]" italics="true" pageId="2" pageNumber="419">Opisthosoma</emphasis>
. Pleural membrane irregularly striate, with sporadic vestitural setae. Chaetotaxy of tergites IXI (number of lyrifissures in parentheses): 1012 (4): 1014 (6): 1014 (6): 1214 (68): 1214 (68): 1214 (68): 12 (6): 1214 (46): 1214 (46): 1014 (4): 812. Chaetotaxy of sternites IVXI (number of lyrifissures in parentheses): 4 (2): 812 (2): 1014 (16): 1214 (46): 1214 (46): 1012 (4): 68 (4): 46. Tergite XII with 2 small setae, sternite XII without setae. Vestitural setae weakly clavate on tergites and acuminate on sternites. Median setae of sternites show no external differences from the other setae. No tactile setae on tergite and sternite XI. Surface of tergites granulated, sternites smooth. Male genital region: sternite II with 7 microsetae and two lyrifissures, sternite III with 5 microsetae around genital opening. Ejaculatory canal atrium, lateral apodeme and lateral rod generally similar to those illustrated by
<bibRefCitation id="EFCA4B6CA933FFE4FD5EFB29FC1438E1" box="[731,990,1231,1257]" pageId="2" pageNumber="419" refId="ref6979" refString="Harvey, M. S. (1986) The Australian Geogarypidae, new status, with a review of the generic classification (Arachnida: Pseudoscorpionida). Australian Journal of Zoology, 34, 753 - 778. https: // doi. org / 10.1071 / ZO 9860753" type="journal article">Harvey (1986: fig. 12)</bibRefCitation>
, but not clearly visible. Female genital region: sternite II with 4+4 microsetae and two lyrifissures, sternite III with 2 microsetae and 2 lyrifissures around genital opening. One pair of elongated lateral cribriform plates and a medial cribriform plate present, but weakly visible.
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42FA98FCC23994" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42FA98FE90397E" box="[199,346,1376,1400]" italics="true" pageId="2" pageNumber="419">Coxal region</emphasis>
. Pedipalpal coxa with 1 long and 1 short seta on anterior face, plus a further 1012 small setae. Coxae IIV with setal formula 57: 79: 1415: 1621.
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42FA51FE7539EC" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42FA51FF3639C9" box="[199,252,1449,1472]" italics="true" pageId="2" pageNumber="419">Legs</emphasis>
(
<figureCitation id="13602A18A933FFE4FE8AFA5FFE5039C9" box="[271,410,1447,1472]" captionStart="FIGURE 1" captionStartId="3.[151,250,1817,1839]" captionTargetBox="[230,1343,195,1772]" captionTargetId="figure@3.[230,1364,195,1796]" captionTargetPageId="3" captionText="FIGURE 1. Geogarypus muchmorei, sp. nov. A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur,male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989)." httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="2" pageNumber="419">Figs 1L, M</figureCitation>
). Diplotarsate and without tactile setae. Surface mostly granulate. Claws simple. Arolia slightly longer than claws.
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42FA08FBA33AB4" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42FA08FE813A0E" box="[199,331,1520,1543]" italics="true" pageId="2" pageNumber="419">Dimensions</emphasis>
(in mm, ratios in parentheses). Male
<typeStatus id="54E0883FA933FFE4FD7CFA17FC953A01" box="[761,863,1519,1544]" pageId="2" pageNumber="419" type="holotype">holotype</typeStatus>
. Body 1.35; carapace 0.43/0.55 (0.78×). Chelicera 0.12/0.07, movable finger 0.08. Pedipalpal trochanter 0.18/0.10 (1.8×); femur 0.40/0.12 (3.33×); patella 0.29/0.13 (2.23×). Chela (with pedicel) 0.64 (3.04×); pedicel 0.04; hand 0.27/0.21 (1.29×); fingers 0.32. Leg I trochanter 0.11/0.09 (1.22×); femur 0.18/0.06 (3.01×); patella 0.11/0.07 (1.57×); tibia 0.13/0.06 (2.16×); metatarsus 0.08/0.04 (2.03×); tarsus 0.10/0.03 (3.34×). Leg IV. trochanter 0.12/0.10 (1.21×); femur 0.09/0.08 (1.13×); patella 0.27/0.11 (2.45×); tibia 0.23/0.07 (3.29×); metatarsus 0.13/0.05 (2.62×); tarsus 0.11/0.03 (3.65×).
</paragraph>
<paragraph id="8BE4369DA933FFE4FF42F930FA453BB1" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="419">
<emphasis id="B92FEA8FA933FFE4FF42F930FE593AE9" box="[199,403,1736,1760]" italics="true" pageId="2" pageNumber="419">
Female
<typeStatus id="54E0883FA933FFE4FEA0F931FE593AE9" box="[293,403,1737,1760]" pageId="2" pageNumber="419" type="paratype">paratypes</typeStatus>
</emphasis>
. Body 1.571.63; carapace 0.450.49/0.640.68 (0.700.72×). Chelicera 0.130.15/0.07 0.08, movable finger 0.090.10. Pedipalpal trochanter 0.26/0.140.15 (1.731.86×); femur 0.500.53/0.140.15 (3.343.79×); patella 0.350.37/0.140.15 (2.472.51×). Chela (with pedicel) 0.770.83 (3.353.46×); pedicel 0.05; hand 0.370.39/0.230.24 (1.171.44×); fingers 0.420.43. Leg I trochanter 0.14/0.09 (1.55×); femur 0.19 0.21/0.070.08 (2.632.71×); patella 0.120.13/0.08 (1.481.51×); tibia 0.15/0.06 (2.51×); metatarsus 0.100.11/ 0.040.05 (2.042.74×); tarsus 0.110.12/0.03 (3.674.03×). Leg IV. trochanter 0.18/0.11 (1.64×); femur 0.11/0.08 (1.38×); patella 0.32/0.10 (3.22×); tibia 0.27/0.06 (5.87×); metatarsus 0.16/0.05 (3.23×); tarsus 0.12/0.04 (3.08×).
</paragraph>
</subSubSection>
<caption id="DF246615A932FFE5FF12F8E1FF133BEE" httpUri="https://zenodo.org/record/1199828/files/figure.png" pageId="3" pageNumber="420" startId="3.[151,250,1817,1839]" targetBox="[230,1343,195,1772]" targetPageId="3">
<paragraph id="8BE4369DA932FFE5FF12F8E1FF133BEE" blockId="3.[151,1436,1817,2023]" pageId="3" pageNumber="420">
<emphasis id="B92FEA8FA932FFE5FF12F8E1FEDF3B27" bold="true" box="[151,277,1817,1839]" pageId="3" pageNumber="420">FIGURE 1.</emphasis>
<taxonomicName id="4C5B4D1EA932FFE5FE9BF8E2FDD93B26" authority="Novák &amp; Harvey, 2018" authorityName="Novák &amp; Harvey" authorityYear="2018" box="[286,531,1817,1839]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="muchmorei" status="sp. nov.">
<emphasis id="B92FEA8FA932FFE5FE9BF8E2FDD93B26" box="[286,531,1817,1839]" italics="true" pageId="3" pageNumber="420">Geogarypus muchmorei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21C57F4A932FFE5FDA6F8E1FDBC3B27" box="[547,630,1817,1838]" pageId="3" pageNumber="420" rank="species">
<emphasis id="B92FEA8FA932FFE5FDA6F8E1FDBC3B27" bold="true" box="[547,630,1817,1838]" pageId="3" pageNumber="420">sp. nov.</emphasis>
</taxonomicNameLabel>
A, male holotype (HNHM Pseud-1988); B, female paratype (HNHM Pseud- 1989); C, clavate seta of carapace, male (HNHM Pseud-1988); D, acuminate seta of carapace, male (HNHM Pseud-1988); E, anterior margin of carapace, male (HNHM Pseud-1988); F, right chelicera of male (HNHM Pseud-1988); G, serrula exterior of male (HNHM Pseud-1988); H, right pedipalp of male (HNHM Pseud-1988); I, acuminate seta of pedipalpal femur, male (HNHM Pseud-1988); J, bifurcate seta of pedipalpal femur, male (HNHM Pseud-1988); K, left chela of female (HNHM Pseud- 1990); L, right leg I of male, prolateral side (HNHM Pseud-1988); M, left leg IV of female, prolateral side (HNHM Pseud- 1989).
</paragraph>
</caption>
<subSubSection id="C3416516A935FFE2FF42FF6FFAEC3FC1" pageId="4" pageNumber="421" type="discussion">
<paragraph id="8BE4369DA935FFE2FF42FF6FFAEC3FC1" blockId="4.[151,1436,151,968]" pageId="4" pageNumber="421">
<emphasis id="B92FEA8FA935FFE2FF42FF6FFEF13CB9" bold="true" box="[199,315,151,176]" pageId="4" pageNumber="421">Remarks.</emphasis>
The new species can be distinguished from
<taxonomicName id="4C5B4D1EA935FFE2FCA9FF60FC593CB9" authority="Novák &amp; Harvey, 2018" authorityName="Novák &amp; Harvey" authorityYear="2018" box="[812,915,152,176]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="klarae" status="sp. nov.">
<emphasis id="B92FEA8FA935FFE2FCA9FF60FC593CB9" box="[812,915,152,176]" italics="true" pageId="4" pageNumber="421">G. klarae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57F4A935FFE2FC1FFF6FFC3E3CB9" box="[922,1012,151,176]" pageId="4" pageNumber="421" rank="species">
<emphasis id="B92FEA8FA935FFE2FC1FFF6FFC3E3CB9" bold="true" box="[922,1012,151,176]" pageId="4" pageNumber="421">sp. nov.</emphasis>
</taxonomicNameLabel>
by the colour pattern of the carapace, the shape of the chela and the chelal dentition. It differs from
<taxonomicName id="4C5B4D1EA935FFE2FC55FF44FBAD3CDD" authorityName="Beier" authorityYear="1965" box="[976,1127,188,212]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="sagittatus">
<emphasis id="B92FEA8FA935FFE2FC55FF44FC2B3CDA" box="[976,993,188,211]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FC7FFF45FBAD3CDD" box="[1018,1127,189,212]" italics="true" pageId="4" pageNumber="421">sagittatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5B4D1EA935FFE2FB05FF44FE553CF1" authority="Murthy &amp; Ananthakrishnan, 1977" authorityName="Murthy &amp; Ananthakrishnan" authorityYear="1977" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="asiaticus">
<emphasis id="B92FEA8FA935FFE2FB05FF44FB5B3CDA" box="[1152,1169,188,211]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FB2FFF45FAC43CDD" box="[1194,1294,189,212]" italics="true" pageId="4" pageNumber="421">asiaticus</emphasis>
<bibRefCitation id="EFCA4B6CA935FFE2FAA5FF44FE553CF1" pageId="4" pageNumber="421" refId="ref7428" refString="Murthy, V. A. &amp; Ananthakrishnan, T. N. (1977) Indian Chelonethi. Oriental Insects Monograph, 4, i - iii, 1 - 210." type="journal article">Murthy &amp; Ananthakrishnan, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5B4D1EA935FFE2FE2EFF18FDA53CF1" authorityName="Murthy &amp; Ananthakrishnan" authorityYear="1977" box="[427,623,224,248]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="heterodentatus">
<emphasis id="B92FEA8FA935FFE2FE2EFF18FE763CFE" box="[427,444,224,247]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FE4FFF18FDA53CF1" box="[458,623,224,248]" italics="true" pageId="4" pageNumber="421">heterodentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5B4D1EA935FFE2FD23FF18FCF13CF1" authorityName="Beier" authorityYear="1974" box="[678,827,224,248]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="B92FEA8FA935FFE2FD23FF18FCF13CF1" box="[678,827,224,248]" italics="true" pageId="4" pageNumber="421">G. nepalensis</emphasis>
</taxonomicName>
in the shape of the chelal hand, as it lacks a prolateral constriction at the base of the fingers (cf.
<bibRefCitation id="EFCA4B6CA935FFE2FDFEFEFCFD363D14" box="[635,764,260,285]" pageId="4" pageNumber="421" refId="ref6571" refString="Beier, M. (1957) Pseudoscorpionida. Insects of Micronesia, 3, 1 - 64." type="journal article">Beier 1957</bibRefCitation>
,
<bibRefCitation id="EFCA4B6CA935FFE2FC8CFEFCFC8E3D14" box="[777,836,260,285]" pageId="4" pageNumber="421" refId="ref6675" refString="Beier, M. (1965) Die Pseudoscorpioniden Neu-Guineas und der benachbarten Inseln. Pacific Insects, 7, 749 - 796." type="journal article">1965</bibRefCitation>
,
<bibRefCitation id="EFCA4B6CA935FFE2FCD4FEFCFC473D15" box="[849,909,260,284]" pageId="4" pageNumber="421" refId="ref6699" refString="Beier, M. (1974) Pseudoscorpione aus Nepal. Senckenbergiana Biologica, 55, 261 - 80." type="journal article">1974</bibRefCitation>
). The shape of the chelal hand in dorsal view closely resembles that of the Australian species
<taxonomicName id="4C5B4D1EA935FFE2FD30FED0FC1B3D49" authority="Harvey, 1986" authorityName="Harvey" authorityYear="1986" box="[693,977,295,320]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="exochus">
<emphasis id="B92FEA8FA935FFE2FD30FED0FD0C3D36" box="[693,710,296,319]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FD51FED0FCE43D49" box="[724,814,296,320]" italics="true" pageId="4" pageNumber="421">exochus</emphasis>
<bibRefCitation id="EFCA4B6CA935FFE2FCB3FED0FC1B3D49" box="[822,977,295,320]" pageId="4" pageNumber="421" refId="ref6979" refString="Harvey, M. S. (1986) The Australian Geogarypidae, new status, with a review of the generic classification (Arachnida: Pseudoscorpionida). Australian Journal of Zoology, 34, 753 - 778. https: // doi. org / 10.1071 / ZO 9860753" type="journal article">Harvey, 1986</bibRefCitation>
</taxonomicName>
, but the dorsal face of the fixed finger is undulate in lateral view in the new species, unlike
<taxonomicName id="4C5B4D1EA935FFE2FD63FEB4FBDF3D6C" authority="(Harvey 1986)" baseAuthorityName="Harvey" baseAuthorityYear="1986" box="[742,1045,332,357]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="exochus">
<emphasis id="B92FEA8FA935FFE2FD63FEB4FCAB3D6D" box="[742,865,332,356]" italics="true" pageId="4" pageNumber="421">G. exochus</emphasis>
(
<bibRefCitation id="EFCA4B6CA935FFE2FCF6FEB4FBC73D6C" box="[883,1037,332,357]" pageId="4" pageNumber="421" refId="ref6979" refString="Harvey, M. S. (1986) The Australian Geogarypidae, new status, with a review of the generic classification (Arachnida: Pseudoscorpionida). Australian Journal of Zoology, 34, 753 - 778. https: // doi. org / 10.1071 / ZO 9860753" type="journal article">Harvey 1986</bibRefCitation>
)
</taxonomicName>
. Trichobothrium
<emphasis id="B92FEA8FA935FFE2FB58FEB5FB2F3D6D" box="[1245,1253,333,356]" italics="true" pageId="4" pageNumber="421">t</emphasis>
of the movable chelal finger is located subterminally in
<taxonomicName id="4C5B4D1EA935FFE2FDF7FE88FD253D81" baseAuthorityName="Harvey" baseAuthorityYear="1986" box="[626,751,368,392]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="exochus">
<emphasis id="B92FEA8FA935FFE2FDF7FE88FD253D81" box="[626,751,368,392]" italics="true" pageId="4" pageNumber="421">G. exochus</emphasis>
</taxonomicName>
, but it is situated two-thirds from base of the finger in
<taxonomicName id="4C5B4D1EA935FFE2FA01FE88FEDE3DA5" authority="Novák &amp; Harvey, 2018" authorityName="Novák &amp; Harvey" authorityYear="2018" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="muchmorei" status="sp. nov.">
<emphasis id="B92FEA8FA935FFE2FA01FE88FEDE3DA5" italics="true" pageId="4" pageNumber="421">G. muchmorei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57F4A935FFE2FE99FE6CFEBC3DA4" box="[284,374,404,429]" pageId="4" pageNumber="421" rank="species">
<emphasis id="B92FEA8FA935FFE2FE99FE6CFEBC3DA4" bold="true" box="[284,374,404,429]" pageId="4" pageNumber="421">sp. nov.</emphasis>
</taxonomicNameLabel>
Also,
<taxonomicName id="4C5B4D1EA935FFE2FE44FE6CFDF13DA5" baseAuthorityName="Harvey" baseAuthorityYear="1986" box="[449,571,404,428]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="exochus">
<emphasis id="B92FEA8FA935FFE2FE44FE6CFE183DA2" box="[449,466,404,427]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FE64FE6CFDF13DA5" box="[481,571,404,428]" italics="true" pageId="4" pageNumber="421">exochus</emphasis>
</taxonomicName>
has
<taxonomicName id="4C5B4D1EA935FFE2FDF5FE6DFD2B3DA4" box="[624,737,404,429]" pageId="4" pageNumber="421" rank="subSpecies" subSpecies="situated">
<emphasis id="B92FEA8FA935FFE2FDF5FE6DFD493DA5" box="[624,643,405,428]" italics="true" pageId="4" pageNumber="421">st</emphasis>
situated
</taxonomicName>
distinctly nearer to
<emphasis id="B92FEA8FA935FFE2FC41FE6DFC063DA5" box="[964,972,405,428]" italics="true" pageId="4" pageNumber="421">t</emphasis>
, while it is closer to
<emphasis id="B92FEA8FA935FFE2FB3FFE6CFB193DA5" box="[1210,1235,404,428]" italics="true" pageId="4" pageNumber="421">sb</emphasis>
in
<taxonomicName id="4C5B4D1EA935FFE2FB7DFE6CFA5F3DA5" authorityName="Novák &amp; Harvey" authorityYear="2018" box="[1272,1429,404,428]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="muchmorei">
<emphasis id="B92FEA8FA935FFE2FB7DFE6CFAC33DA2" box="[1272,1289,404,427]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FA9DFE6CFA5F3DA5" box="[1304,1429,404,428]" italics="true" pageId="4" pageNumber="421">muchmorei</emphasis>
</taxonomicName>
. The chela of
<taxonomicName id="4C5B4D1EA935FFE2FEB5FE40FE613DD9" baseAuthorityName="Harvey" baseAuthorityYear="1986" box="[304,427,440,464]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="exochus">
<emphasis id="B92FEA8FA935FFE2FEB5FE40FE613DD9" box="[304,427,440,464]" italics="true" pageId="4" pageNumber="421">G. exochus</emphasis>
</taxonomicName>
has 10 accessory teeth on the fixed finger (
<bibRefCitation id="EFCA4B6CA935FFE2FC20FE40FBF53DD9" box="[933,1087,439,464]" pageId="4" pageNumber="421" refId="ref6979" refString="Harvey, M. S. (1986) The Australian Geogarypidae, new status, with a review of the generic classification (Arachnida: Pseudoscorpionida). Australian Journal of Zoology, 34, 753 - 778. https: // doi. org / 10.1071 / ZO 9860753" type="journal article">Harvey 1986</bibRefCitation>
), whereas the new species has only 47 such teeth. The new species differs from
<taxonomicName id="4C5B4D1EA935FFE2FD51FE24FBCC3DFD" authority="(Simon, 1899)" baseAuthorityName="Simon" baseAuthorityYear="1899" box="[724,1030,476,501]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="irrugatus">
<emphasis id="B92FEA8FA935FFE2FD51FE24FD2F3DFA" box="[724,741,476,499]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FD76FE25FC913DFD" box="[755,859,477,500]" italics="true" pageId="4" pageNumber="421">irrugatus</emphasis>
(Simon, 1899)
</taxonomicName>
in its chelal dentition (teeth of both fingers more widely separated in
<taxonomicName id="4C5B4D1EA935FFE2FD8FFDF8FD5A3E1E" baseAuthorityName="Simon" baseAuthorityYear="1899" box="[522,656,512,535]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="irrugatus">
<emphasis id="B92FEA8FA935FFE2FD8FFDF8FDD13E1E" box="[522,539,512,535]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FDADFDF8FD5A3E1E" box="[552,656,512,535]" italics="true" pageId="4" pageNumber="421">irrugatus</emphasis>
</taxonomicName>
) and in having the prolateral face of the chelal hand more swollen (cf.
<bibRefCitation id="EFCA4B6CA935FFE2FF12FDDDFEE53E35" box="[151,303,548,573]" pageId="4" pageNumber="421" refId="ref7169" refString="Harvey, M. S. (2010) Redescription of Geogarypus irrugatus from Sumatra (Pseudoscorpiones: Geogarypidae). Journal of Arachnology, 38, 383 - 386. https: // doi. org / 10.1636 / A 08 - 101 SC. 1" type="journal article">Harvey 2010</bibRefCitation>
). The darker coloration of the carapace and the shape of the chelal hand clearly distinguish the new species from
<taxonomicName id="4C5B4D1EA935FFE2FEB2FDB0FE3D3E69" baseAuthorityName="Rainbow" baseAuthorityYear="1897" box="[311,503,584,608]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="longidigitatus">
<emphasis id="B92FEA8FA935FFE2FEB2FDB0FE823E56" box="[311,328,584,607]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FEDEFDB0FE3D3E69" box="[347,503,584,608]" italics="true" pageId="4" pageNumber="421">longidigitatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5B4D1EA935FFE2FDB2FDB0FCE63E69" authority="Beier, 1963" authorityName="Beier" authorityYear="1963" box="[567,812,583,608]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="B92FEA8FA935FFE2FDB2FDB0FD823E56" box="[567,584,584,607]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FDD9FDB0FD533E69" box="[604,665,584,608]" italics="true" pageId="4" pageNumber="421">albus</emphasis>
Beier, 1963
</taxonomicName>
(
<bibRefCitation id="EFCA4B6CA935FFE2FCBAFDBFFC183E69" box="[831,978,583,608]" pageId="4" pageNumber="421" refId="ref6590" refString="Beier, M. (1963 a) Pseudoscorpione aus Vogelnestern von Malaya. Pacific Insects, 5, 507 - 11." type="journal article">Beier 1963a</bibRefCitation>
;
<bibRefCitation id="EFCA4B6CA935FFE2FC66FDB0FBB63E69" box="[995,1148,584,608]" pageId="4" pageNumber="421" refId="ref7030" refString="Harvey, M. S. (1988). Pseudoscorpions from the Krakatau Islands and adjacent regions, Indonesia (Chelicerata: Pseudoscorpionida). Memoirs of the Museum of Victoria, 49, 309 - 353. https: // doi. org / 10.24199 / j. mmv. 1988.49.13" type="journal article">Harvey 1988</bibRefCitation>
,
<bibRefCitation id="EFCA4B6CA935FFE2FB08FDB0FB003E69" box="[1165,1226,584,608]" pageId="4" pageNumber="421" refId="ref7125" refString="Harvey, M. S. (2000) From Siam to Rapa Nui - the identity and distribution of Geogarypus longidigitatus (Rainbow) (Pseudoscorpiones: Geogarypidae). Bulletin of the British Arachnological Society, 11, 377 - 384." type="journal article">2000</bibRefCitation>
). The chela of
<taxonomicName id="4C5B4D1EA935FFE2FA01FDB0FE1D3E8D" authority="Chamberlin, 1930" authorityName="Chamberlin" authorityYear="1930" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B92FEA8FA935FFE2FA01FDB0FA5F3E56" box="[1412,1429,584,607]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FF12FD94FECF3E8D" box="[151,261,620,644]" italics="true" pageId="4" pageNumber="421">angulatus</emphasis>
<bibRefCitation id="EFCA4B6CA935FFE2FE89FD94FE1D3E8D" box="[268,471,620,645]" pageId="4" pageNumber="421" refId="ref6753" refString="Chamberlin, J. C. (1930) A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part 11. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History, Series 10, 5, 1 - 48; 585 - 620." type="journal article">Chamberlin, 1930</bibRefCitation>
</taxonomicName>
has a totally different shape in lateral view (
<bibRefCitation id="EFCA4B6CA935FFE2FC49FD94FB5C3E8D" box="[972,1174,620,645]" pageId="4" pageNumber="421" refId="ref6753" refString="Chamberlin, J. C. (1930) A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part 11. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History, Series 10, 5, 1 - 48; 585 - 620." type="journal article">Chamberlin 1930</bibRefCitation>
). Trichobothrium
<emphasis id="B92FEA8FA935FFE2FAE4FD94FA483E8D" box="[1377,1410,620,644]" italics="true" pageId="4" pageNumber="421">isb</emphasis>
is situated nearer to the dental margin in
<taxonomicName id="4C5B4D1EA935FFE2FDCFFD68FBA13EA1" authority="Murthy &amp; Ananthakrishnan, 1977" authorityName="Murthy &amp; Ananthakrishnan" authorityYear="1977" box="[586,1131,655,680]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B92FEA8FA935FFE2FDCFFD68FD913EAE" box="[586,603,656,679]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FDECFD68FD283EA1" box="[617,738,656,680]" italics="true" pageId="4" pageNumber="421">granulatus</emphasis>
<bibRefCitation id="EFCA4B6CA935FFE2FD6FFD77FBA13EA1" box="[746,1131,655,680]" pageId="4" pageNumber="421" refId="ref7428" refString="Murthy, V. A. &amp; Ananthakrishnan, T. N. (1977) Indian Chelonethi. Oriental Insects Monograph, 4, i - iii, 1 - 210." type="journal article">Murthy &amp; Ananthakrishnan, 1977</bibRefCitation>
</taxonomicName>
, and the vestitural setae on the carapace and the pedipalps of the new species are mostly clavate, whereas they are acuminate in
<taxonomicName id="4C5B4D1EA935FFE2FA86FD4CFDEE3EF9" authority="(Murthy &amp; Ananthakrishnan 1977)" baseAuthorityName="Murthy &amp; Ananthakrishnan" baseAuthorityYear="1977" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B92FEA8FA935FFE2FA86FD4CFADE3EC2" box="[1283,1300,692,715]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FAA6FD4CFA563EC5" box="[1315,1436,692,716]" italics="true" pageId="4" pageNumber="421">granulatus</emphasis>
(
<bibRefCitation id="EFCA4B6CA935FFE2FF1BFD2FFDD63EF9" box="[158,540,727,752]" pageId="4" pageNumber="421" refId="ref7428" refString="Murthy, V. A. &amp; Ananthakrishnan, T. N. (1977) Indian Chelonethi. Oriental Insects Monograph, 4, i - iii, 1 - 210." type="journal article">Murthy &amp; Ananthakrishnan 1977</bibRefCitation>
)
</taxonomicName>
. In
<taxonomicName id="4C5B4D1EA935FFE2FDCAFD20FD213EF9" authority="Novák &amp; Harvey, 2018" authorityName="Novák &amp; Harvey" authorityYear="2018" box="[591,747,728,752]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="muchmorei" status="sp. nov.">
<emphasis id="B92FEA8FA935FFE2FDCAFD20FDAA3EE6" box="[591,608,728,751]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FDEBFD20FD213EF9" box="[622,747,728,752]" italics="true" pageId="4" pageNumber="421">muchmorei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57F4A935FFE2FD76FD2FFC873EF9" box="[755,845,727,752]" pageId="4" pageNumber="421" rank="species">
<emphasis id="B92FEA8FA935FFE2FD76FD2FFC873EF9" bold="true" box="[755,845,727,752]" pageId="4" pageNumber="421">sp. nov.</emphasis>
</taxonomicNameLabel>
, the distance between
<emphasis id="B92FEA8FA935FFE2FBC8FD20FB963EE6" box="[1101,1116,728,751]" italics="true" pageId="4" pageNumber="421">it</emphasis>
and
<emphasis id="B92FEA8FA935FFE2FB11FD21FB623EF9" box="[1172,1192,729,752]" italics="true" pageId="4" pageNumber="421">et</emphasis>
on the fixed finger is half that between
<emphasis id="B92FEA8FA935FFE2FED8FD05FEA73F1D" box="[349,365,765,788]" italics="true" pageId="4" pageNumber="421">it</emphasis>
and
<emphasis id="B92FEA8FA935FFE2FE26FD05FE083F1D" box="[419,450,765,788]" italics="true" pageId="4" pageNumber="421">est</emphasis>
, whereas in
<taxonomicName id="4C5B4D1EA935FFE2FDCBFD04FD2C3F1D" baseAuthorityName="Murthy &amp; Ananthakrishnan" baseAuthorityYear="1977" box="[590,742,764,788]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B92FEA8FA935FFE2FDCBFD04FD953F1A" box="[590,607,764,787]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FDE8FD04FD2C3F1D" box="[621,742,764,788]" italics="true" pageId="4" pageNumber="421">granulatus</emphasis>
</taxonomicName>
these distances are nearly equal. Furthermore, trichobothrium
<emphasis id="B92FEA8FA935FFE2FF12FCD8FF763F31" box="[151,188,800,824]" italics="true" pageId="4" pageNumber="421">esb</emphasis>
is nearly equidistant between
<emphasis id="B92FEA8FA935FFE2FD96FCD8FDE43F31" box="[531,558,800,824]" italics="true" pageId="4" pageNumber="421">eb</emphasis>
and
<emphasis id="B92FEA8FA935FFE2FDE0FCD8FD4C3F31" box="[613,646,800,824]" italics="true" pageId="4" pageNumber="421">isb</emphasis>
in
<taxonomicName id="4C5B4D1EA935FFE2FD2EFCD8FC893F31" baseAuthorityName="Murthy &amp; Ananthakrishnan" baseAuthorityYear="1977" box="[683,835,800,824]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B92FEA8FA935FFE2FD2EFCD8FD763F3E" box="[683,700,800,823]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FD4FFCD8FC893F31" box="[714,835,800,824]" italics="true" pageId="4" pageNumber="421">granulatus</emphasis>
</taxonomicName>
, but much closer to
<emphasis id="B92FEA8FA935FFE2FBAEFCD8FB8C3F31" box="[1067,1094,800,824]" italics="true" pageId="4" pageNumber="421">eb</emphasis>
in
<taxonomicName id="4C5B4D1EA935FFE2FBEEFCD8FACD3F31" authority="Novák &amp; Harvey, 2018" authorityName="Novák &amp; Harvey" authorityYear="2018" box="[1131,1287,800,824]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="muchmorei" status="sp. nov.">
<emphasis id="B92FEA8FA935FFE2FBEEFCD8FBB63F3E" box="[1131,1148,800,823]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FB0FFCD8FACD3F31" box="[1162,1287,800,824]" italics="true" pageId="4" pageNumber="421">muchmorei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57F4A935FFE2FA8AFCE7FAA33F31" box="[1295,1385,799,824]" pageId="4" pageNumber="421" rank="species">
<emphasis id="B92FEA8FA935FFE2FA8AFCE7FAA33F31" bold="true" box="[1295,1385,799,824]" pageId="4" pageNumber="421">sp. nov.</emphasis>
</taxonomicNameLabel>
The teeth of the movable chelal finger are more spaced in
<taxonomicName id="4C5B4D1EA935FFE2FD73FCBCFC443F55" baseAuthorityName="Murthy &amp; Ananthakrishnan" baseAuthorityYear="1977" box="[758,910,836,860]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B92FEA8FA935FFE2FD73FCBCFCCD3F52" box="[758,775,836,859]" italics="true" pageId="4" pageNumber="421">G</emphasis>
.
<emphasis id="B92FEA8FA935FFE2FC90FCBCFC443F55" box="[789,910,836,860]" italics="true" pageId="4" pageNumber="421">granulatus</emphasis>
</taxonomicName>
than in the new species.
<taxonomicName id="4C5B4D1EA935FFE2FB2EFCBCFE9F3F89" authority="Sivaraman, 1980" authorityName="Sivaraman" authorityYear="1980" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="globulus">
<emphasis id="B92FEA8FA935FFE2FB2EFCBCFA563F55" box="[1195,1436,836,860]" italics="true" pageId="4" pageNumber="421">Geogarypus globulus</emphasis>
<bibRefCitation id="EFCA4B6CA935FFE2FF12FC9FFE9F3F89" box="[151,341,871,896]" pageId="4" pageNumber="421" refId="ref7581" refString="Sivaraman, S. (1980) Pseudoscorpions from South India: a new genus and some new species of the super-family Garypoidea Chamberlin (Pseudoscorpionida: Diplosphyronida). Oriental Insects, 14, 325 - 343. https: // doi. org / 10.1080 / 00305316.1980.10434812" type="journal article">Sivaraman, 1980</bibRefCitation>
</taxonomicName>
differs from the new species in having
<emphasis id="B92FEA8FA935FFE2FC90FC90FCEE3F76" box="[789,804,872,895]" italics="true" pageId="4" pageNumber="421">it</emphasis>
more distally located on the fixed chelal finger, and the prolateral side of chelal hand not swollen (
<bibRefCitation id="EFCA4B6CA935FFE2FDF0FC74FCF83FAD" box="[629,818,908,933]" pageId="4" pageNumber="421" refId="ref7581" refString="Sivaraman, S. (1980) Pseudoscorpions from South India: a new genus and some new species of the super-family Garypoidea Chamberlin (Pseudoscorpionida: Diplosphyronida). Oriental Insects, 14, 325 - 343. https: // doi. org / 10.1080 / 00305316.1980.10434812" type="journal article">Sivaraman 1980</bibRefCitation>
).
<taxonomicName id="4C5B4D1EA935FFE2FCC0FC74FBE13FAD" baseAuthorityName="With" baseAuthorityYear="1906" box="[837,1067,908,932]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B92FEA8FA935FFE2FCC0FC74FBE13FAD" box="[837,1067,908,932]" italics="true" pageId="4" pageNumber="421">Geogarypus elegans</emphasis>
</taxonomicName>
has a totally brown carapace, 12 accessory teeth on the fixed chelal finger (
<quantity id="4CA39B78A935FFE2FDF5FC48FD733FC1" box="[624,697,943,968]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.778" metricValueMin="1.016" pageId="4" pageNumber="421" unit="in" value="5.5" valueMax="7.0" valueMin="4.0">47 in</quantity>
the new species) and more slender chelae (
<bibRefCitation id="EFCA4B6CA935FFE2FB18FC57FAD33FC1" box="[1181,1305,943,968]" pageId="4" pageNumber="421" refId="ref7630" refString="With, C. J. (1906) The Danish expedition to Siam 1899 - 1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter, Series 7, 3, 1 - 214." type="journal article">With 1906</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>