treatments-xml/data/03/CD/B6/03CDB626CCDF4E8CFEE87E158ADA85D5.xml
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<mods:title id="D04D401DA5A714B999EC8334AC334CC5">Revision of Autolytinae (Syllidae: Polychaeta).</mods:title>
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<emphasis id="B910DB22CCDF4E88FEE87E158BDB8355" bold="true" box="[264,965,804,830]" pageId="168" pageNumber="169">
<taxonomicName id="4C647CB3CCDF4E88FEE87E158AB28355" ID-CoL="6FTH4" authority="Gravier, 1906" authorityName="Gravier" authorityYear="1906" box="[264,684,804,830]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="species" species="charcoti" status="comb. nov.">
<emphasis id="B910DB22CCDF4E88FEE87E1589F98355" bold="true" box="[264,487,804,830]" italics="true" pageId="168" pageNumber="169">Epigamia charcoti</emphasis>
(
<bibRefCitation id="EFF57AC1CCDF4E88FE177E158ABA8355" author="Gravier" box="[503,676,804,830]" pageId="168" pageNumber="169" refString="Gravier, C. (1906) Sur les annelides polychetes recueillies par l'expedition Antarctique francaise (Syllidiens). Bulletin du Museum d'histoire naturelle, 12, 283 - 290." type="journal article" year="1906">Gravier, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2236659CCDF4E88FD547E158B018355" box="[692,799,804,830]" pageId="168" pageNumber="169">comb. n.</taxonomicNameLabel>
(Fig. 85AE)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37E54BBCCDF4E88FEE87E5E8CBA842E" pageId="168" pageNumber="169" type="reference_group">
<paragraph id="8BDB0730CCDF4E88FEE87E5E8CB483AE" blockId="168.[264,1323,878,1094]" pageId="168" pageNumber="169">
<treatmentCitationGroup id="AB74201ECCDF4E88FEE87E5E8CB483AE" pageId="168" pageNumber="169">
<taxonomicName id="4C647CB3CCDF4E88FEE87E5E8ABE83EE" ID-CoL="JZ36" authority="Gravier, 1906: 283" authorityName="Gravier" authorityPageNumber="283" authorityYear="1906" box="[264,672,878,902]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="species" species="charcoti">
<emphasis id="B910DB22CCDF4E88FEE87E5E89D283ED" box="[264,460,879,902]" italics="true" pageId="168" pageNumber="169">Autolytus charcoti</emphasis>
<treatmentCitation id="0AC52121CCDF4E88FE347E5F8ABE83EE" author="Gravier" box="[468,672,878,901]" page="283" pageId="168" pageNumber="169" year="1906">
<bibRefCitation id="EFF57AC1CCDF4E88FE347E5F8A7583EE" author="Gravier" box="[468,619,878,901]" pageId="168" pageNumber="169" refString="Gravier, C. (1906) Sur les annelides polychetes recueillies par l'expedition Antarctique francaise (Syllidiens). Bulletin du Museum d'histoire naturelle, 12, 283 - 290." type="journal article" year="1906">Gravier, 1906</bibRefCitation>
: 283
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285; 1907: 78, pl. 1, fig. 12, textfigs 12;
<treatmentCitation id="0AC52121CCDF4E88FB6D7E5F894A83CE" author="Benham" page="27" pageId="168" pageNumber="169" year="1921">
<bibRefCitation id="EFF57AC1CCDF4E88FB6D7E5F8D3883EE" author="Benham" box="[1165,1318,878,901]" pageId="168" pageNumber="169" refString="Benham, W. B. (1921) Polychaeta. Scientific Reports of the Australasian Antarctic Expedition, 1911 - 1914, 6, 1 - 128." type="journal article" year="1921">Benham 1921</bibRefCitation>
: 27
</treatmentCitation>
31, pl 5, figs 710; 1927: 60.
<treatmentCitation id="0AC52121CCDF4E88FD7E7EBF8B5183CE" author="Monro" box="[670,847,910,933]" page="97" pageId="168" pageNumber="169" year="1930">
<bibRefCitation id="EFF57AC1CCDF4E88FD7E7EBF8B3983CE" author="Monro" box="[670,807,910,933]" pageId="168" pageNumber="169" refString="Monro, C. C. A. (1930) Polychaete worms. Discovery reports, 2, 1 - 222." type="journal article" year="1930">Monro 1930</bibRefCitation>
: 97
</treatmentCitation>
; 1936: 131132, fig. 22;
<treatmentCitation id="0AC52121CCDF4E88FBBE7EBF8D3A83CE" author="Hartman" box="[1118,1316,910,933]" page="27" pageId="168" pageNumber="169" year="1953">
<bibRefCitation id="EFF57AC1CCDF4E88FBBE7EBF8CE283CE" author="Hartman" box="[1118,1276,910,933]" pageId="168" pageNumber="169" refString="Hartman, O. (1953) Non-pelagic polychaeta of the Swedish Antarctic Expedition 1901 - 1903. Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, 4, 1 - 83." type="journal article" year="1953">Hartman 1953</bibRefCitation>
: 27
</treatmentCitation>
; 1964: 77, pl. 24, fig. 1; Hartmann­Schröder &amp; Rosenfeldt 1988: 4546; 1992: 102.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDF4E88FEE87EFE8944846E" blockId="168.[264,1323,878,1094]" pageId="168" pageNumber="169">
<treatmentCitationGroup id="AB74201ECCDF4E88FEE87EFE8944846E" pageId="168" pageNumber="169">
<emphasis id="B910DB22CCDF4E88FEE87EFE890B838D" box="[264,277,975,998]" italics="true" pageId="168" pageNumber="169">?</emphasis>
<taxonomicName id="4C647CB3CCDF4E88FEFE7EFE8AB3838E" ID-CoL="JZ36" authority="Fauvel 1950: 758" authorityName="Fauvel" authorityPageNumber="758" authorityYear="1950" box="[286,685,974,998]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="species" species="charcoti">
<emphasis id="B910DB22CCDF4E88FEFE7EFE89FA838D" box="[286,484,975,998]" italics="true" pageId="168" pageNumber="169">Autolytus charcoti</emphasis>
<treatmentCitation id="0AC52121CCDF4E88FE0D7EFF8AB3838E" author="Fauvel" box="[493,685,974,997]" page="758" pageId="168" pageNumber="169" year="1950">
<bibRefCitation id="EFF57AC1CCDF4E88FE0D7EFF8A69838E" author="Fauvel" box="[493,631,974,997]" pageId="168" pageNumber="169" refString="Fauvel, P. (1950) Missions du batiment polaire &quot; Commandant-Charcot &quot;. Recoltes faites en Terre Adelie (1950) par Paul Tchernia. Annelides polychetes. Bulletin du Museum national d´histoire naturelle, 22, 753 - 773." type="journal article" year="1950">Fauvel 1950</bibRefCitation>
: 758
</treatmentCitation>
</taxonomicName>
759;
<treatmentCitation id="0AC52121CCDF4E88FD117EFF8C0D838E" author="Wesenberg-Lund" box="[753,1043,974,997]" page="97" pageId="168" pageNumber="169" year="1961">
<bibRefCitation id="EFF57AC1CCDF4E88FD117EFF8BF7838E" author="Wesenberg-Lund" box="[753,1001,974,997]" pageId="168" pageNumber="169" refString="Wesenberg-Lund, E. (1961) Reports of the Lund University Chile Expedition 1948 - 49. 43. Polychaeta Errantia. Lunds universitets arsskrift, 57, 1 - 137." type="journal article" year="1961">Wesenberg­Lund 1961</bibRefCitation>
: 97
</treatmentCitation>
;
<treatmentCitation id="0AC52121CCDF4E88FBC17EFF8948846E" author="Knox" page="79" pageId="168" pageNumber="169" year="1970">
<bibRefCitation id="EFF57AC1CCDF4E88FBC17EFF8D38838E" author="Knox" box="[1057,1318,974,997]" pageId="168" pageNumber="169" refString="Knox, G. A. &amp; Cameron, D. B. (1970) Polychaeta from the Snares Islands, New Zealand. Transactions of the Royal Society of New Zealand, 12, 73 - 85." type="journal article" year="1970">
Knox &amp;
<collectingCountry id="F37347A0CCDF4E88FB637EFF8CFB838E" box="[1155,1253,974,997]" name="Cameroon" pageId="168" pageNumber="169">Cameron</collectingCountry>
1970
</bibRefCitation>
: 79
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDF4E88FEE8793E8B59844E" blockId="168.[264,1323,878,1094]" box="[264,839,1038,1062]" pageId="168" pageNumber="169">
<treatmentCitationGroup id="AB74201ECCDF4E88FEE8793E8B59844E" box="[264,839,1038,1062]" pageId="168" pageNumber="169">
<taxonomicName id="4C647CB3CCDF4E88FEE8793E8B06844E" ID-CoL="JZ36" authority="Hartman 1967: 53" authorityName="Hartman" authorityPageNumber="53" authorityYear="1967" box="[264,792,1038,1062]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="species" species="charcoti" subGenus="Regulatus">
<emphasis id="B910DB22CCDF4E88FEE8793E8A50844D" box="[264,590,1039,1062]" italics="true" pageId="168" pageNumber="169">Autolytus (Regulatus) charcoti</emphasis>
<treatmentCitation id="0AC52121CCDF4E88FDB5793F8B06844E" author="Hartman" box="[597,792,1038,1061]" page="53" pageId="168" pageNumber="169" year="1967">
<bibRefCitation id="EFF57AC1CCDF4E88FDB5793F8AEC844E" author="Hartman" box="[597,754,1038,1061]" pageId="168" pageNumber="169" refString="Hartman, O. (1967) Polychaetous annelids collected by the USNS Eltanin and Staten Island cruises, chiefly from Antarctic seas. Allan Hancock Monographs in Marine Biology, 2, 1 - 387." type="journal article" year="1967">Hartman 1967</bibRefCitation>
: 53
</treatmentCitation>
</taxonomicName>
54.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDF4E88FEE8791F8CBA842E" blockId="168.[264,1323,878,1094]" box="[264,1188,1070,1094]" pageId="168" pageNumber="169">
<treatmentCitationGroup id="AB74201ECCDF4E88FEE8791F8CBA842E" box="[264,1188,1070,1094]" pageId="168" pageNumber="169">
Not
<taxonomicName id="4C647CB3CCDF4E88FED7791E8ABE842E" ID-CoL="JZ36" authority="Day 1960: 317" authorityName="Day" authorityPageNumber="317" authorityYear="1960" box="[311,672,1070,1094]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="species" species="charcoti">
<emphasis id="B910DB22CCDF4E88FED7791E89E4842D" box="[311,506,1071,1094]" italics="true" pageId="168" pageNumber="169">Autolytus charcoti</emphasis>
<treatmentCitation id="0AC52121CCDF4E88FDE1791F8ABE842E" author="Day" box="[513,672,1070,1093]" page="317" pageId="168" pageNumber="169" year="1960">
<bibRefCitation id="EFF57AC1CCDF4E88FDE1791F8A73842E" author="Day" box="[513,621,1070,1093]" pageId="168" pageNumber="169" refString="Day, J. H. (1960) The polychaete fauna of South Africa. Part 5. Errant species dredged off Cape coasts. Annals of the South African Museum, 45, 261 - 373." type="journal article" year="1960">Day 1960</bibRefCitation>
: 317
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</taxonomicName>
318; 1967: 283, fig. 12.12KN (=
<taxonomicName id="4C647CB3CCDF4E88FBF1791F8C68842E" ID-CoL="85GQD" box="[1041,1142,1070,1093]" class="Polychaeta" family="Syllidae" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="tribe" tribe="Procerini">Procerini</taxonomicName>
sp).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDF4E88FEE879478D3A84FA" box="[264,1316,1142,1169]" pageId="168" pageNumber="169" type="materials_examined">
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<emphasis id="B910DB22CCDF4E88FEE8794789EA84FB" bold="true" box="[264,500,1142,1168]" pageId="168" pageNumber="169">Material examined</emphasis>
.
<emphasis id="B910DB22CCDF4E88FDE479478AB384FB" bold="true" box="[516,685,1142,1168]" pageId="168" pageNumber="169">
<collectingCountry id="F37347A0CCDF4E88FDE479478A6484FB" box="[516,634,1142,1168]" name="Antarctica" pageId="168" pageNumber="169">Antarctic</collectingCountry>
Sea
</emphasis>
: 2
<typeStatus id="54DFB992CCDF4E88FD3379468B2784FA" box="[723,825,1143,1169]" pageId="168" pageNumber="169" type="syntype">syntypes</typeStatus>
<collectionCode id="ED759FF5CCDF4E88FCA179468B8584FA" LSID="urn:lsid:biocol.org:col:34988" box="[833,923,1143,1169]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="168" pageNumber="169">MNHN</collectionCode>
A72,
<collectingCountry id="F37347A0CCDF4E88FC0379468C4E84FA" box="[995,1104,1143,1169]" name="Antarctica" pageId="168" pageNumber="169">Antarctic</collectingCountry>
expedition, 1905.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDF4E88FED879AF8B54848A" pageId="168" pageNumber="169" type="diagnosis">
<paragraph id="8BDB0730CCDF4E88FED879AF8B54848A" blockId="168.[264,1324,1142,1889]" pageId="168" pageNumber="169">
<emphasis id="B910DB22CCDF4E88FED879AF89AE84D3" bold="true" box="[312,432,1182,1208]" pageId="168" pageNumber="169">Diagnosis</emphasis>
.
<taxonomicName id="4C647CB3CCDF4E88FE2179AE8A2D84D3" box="[449,563,1183,1208]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="168" pageNumber="169" phylum="Annelida" rank="genus">
<emphasis id="B910DB22CCDF4E88FE2179AE8A2D84D3" box="[449,563,1183,1208]" italics="true" pageId="168" pageNumber="169">Epigamia</emphasis>
</taxonomicName>
with long alternating dorsal cirri and a trepan consisting of 27 teeth, 1 large alternating with 2 smaller in 2 rings
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDF4E89FED879DF8A3C8415" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169" type="description">
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<emphasis id="B910DB22CCDF4E88FED879DF89D78563" bold="true" box="[312,457,1262,1288]" pageId="168" pageNumber="169">Description</emphasis>
. Two anterior fragments, length
<quantity id="4C9CAAD5CCDF4E88FCB679DE8BC98562" box="[854,983,1263,1289]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.8" metricValueMin="3.5" pageId="168" pageNumber="169" unit="mm" value="41.5" valueMax="48.0" valueMin="35.0">3548 mm</quantity>
for 2644 chaetigers, width
<quantity id="4C9CAAD5CCDF4E88FEE8782689A4855A" box="[264,442,1303,1329]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="168" pageNumber="169" unit="mm" value="0.5" valueMax="0.55" valueMin="0.45">0.450.55 mm</quantity>
. Preserved material yellowish brown, no colour markings. Ciliation as 2 trochs per segment, seen in posterior part of larger fragment.
</paragraph>
<paragraph id="8BDB0730CCDF4E88FED878568C8685C2" blockId="168.[264,1324,1142,1889]" pageId="168" pageNumber="169">Eyes separated; eye spots absent. Palps in dorsal view projecting 1/51/4 of prostomial length, fused. Extension of nuchal epaulettes to end of chaetiger 3 (Fig. 85A).</paragraph>
<paragraph id="8BDB0730CCDF4E88FED878868C21877A" blockId="168.[264,1324,1142,1889]" pageId="168" pageNumber="169">
Median antenna reaching chaetiger 1014. Lateral antennae and dorsal tentacular cirri, length 2/3 of median antenna. Ventral tentacular cirri 1/3 as long as dorsal pair. First dorsal cirri as long as median antenna. Second dorsal cirri as long as ventral tentacular cirri. From chaetiger 127 cirri with usual alternation in direction, more posterior difficult to assess due to lost cirri. Dorsal cirri from chaetiger 3, alternate in length; short cirri equals 1/24/5 of body width, long cirri 3/4 to slightly longer than body width. Cirrophores present on tentacular cirri, and all dorsal cirri. Cirrophores equal, cirrostyles unequal (Fig. 85A, B); short cirrostyles 1/22/
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length of long cirrostyles; cirrophores shorter than parapodial lobes; cirrophores shorter than cirrostyles. All appendages cylindrical.
</paragraph>
<paragraph id="8BDB0730CCDF4E89FED87A2E8B978135" blockId="168.[264,1324,1142,1889]" lastBlockId="169.[264,1324,284,1350]" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169">Parapodial lobes large, rounded, slightly swollen dorsal to chaetal fascicle (Fig. 85B). Chaetiger 24 with 2 aciculae. Chaetal fascicle with c. 10 compounds in all present chaetigers. Compound chaetae with large distal tooth (Fig. 85D); serration present. Single thin bayonet chaetae (Fig. 85E), beginning between 2628.</paragraph>
<paragraph id="8BDB0730CCDE4E89FED87C5D89AA824D" blockId="169.[264,1324,284,1350]" pageId="169" pageNumber="170">Pharynx with 1 sinuation anterior to proventricle. Trepan in tentacular segment or chaetiger 2, with 27 unequal teeth, 9 large teeth and 18 smaller; 1 large alternating with 2 small, in 2 rings (Fig. 85C). Basal ring present; infradental spines present. Proventricle equal in length to 45 segments in chaetiger 611 with c. 40 rows of muscle cells (n=2). Pygidium lost.</paragraph>
<paragraph id="8BDB0730CCDE4E89FED87F0289AE82F5" blockId="169.[264,1324,284,1350]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED87F0289FA8226" bold="true" box="[312,484,563,589]" pageId="169" pageNumber="170">Reproduction</emphasis>
. Epigamy recorded by
<bibRefCitation id="EFF57AC1CCDE4E89FCE17F058BAA8225" author="Benham" box="[769,948,564,590]" pageId="169" pageNumber="170" refString="Benham, W. B. (1921) Polychaeta. Scientific Reports of the Australasian Antarctic Expedition, 1911 - 1914, 6, 1 - 128." type="journal article" year="1921">Benham (1921</bibRefCitation>
;
<bibRefCitation id="EFF57AC1CCDE4E89FC237F058BE18225" author="Benham" box="[963,1023,564,590]" pageId="169" pageNumber="170" refString="Benham, W. B. (1927) Polychaeta. British Antarctic ' Terra Nova' Expedition, 1910. Natural History Report, Zoology, 7, 47 - 182." type="journal article" year="1927">1927</bibRefCitation>
) and
<bibRefCitation id="EFF57AC1CCDE4E89FBA57F058CEF8225" author="Monro" box="[1093,1265,564,590]" pageId="169" pageNumber="170" refString="Monro, C. C. A. (1936) Polychaete worms. 2. Discovery reports, 12, 59 - 198." type="journal article" year="1936">Monro (1936)</bibRefCitation>
. The descriptions are sufficiently good for a reliable connection of these epitokes to the atokes of
<taxonomicName id="4C647CB3CCDE4E89FEC87FB589B682F6" box="[296,424,644,669]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="charcoti">
<emphasis id="B910DB22CCDE4E89FEC87FB589B682F6" box="[296,424,644,669]" italics="true" pageId="169" pageNumber="170">E. charcoti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB0730CCDE4E89FED87F9D8CFE82AD" blockId="169.[264,1324,284,1350]" box="[312,1248,684,710]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED87F9D8ADA82AD" bold="true" box="[312,708,684,710]" pageId="169" pageNumber="170">Morphology of epitokous stages</emphasis>
. Descriptions extracted from
<bibRefCitation id="EFF57AC1CCDE4E89FBC17F9D8CC282AD" author="Benham" box="[1057,1244,684,710]" pageId="169" pageNumber="170" refString="Benham, W. B. (1921) Polychaeta. Scientific Reports of the Australasian Antarctic Expedition, 1911 - 1914, 6, 1 - 128." type="journal article" year="1921">Benham (1921)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDB0730CCDE4E89FED87FE58C8583B5" blockId="169.[264,1324,284,1350]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED87FE589698285" bold="true" box="[312,375,724,750]" pageId="169" pageNumber="170">
<collectingCountry id="F37347A0CCDE4E89FED87FE589698285" box="[312,375,724,750]" name="Malta" pageId="169" pageNumber="170">Male</collectingCountry>
</emphasis>
. Length
<quantity id="4C9CAAD5CCDE4E89FE057FE58A248285" box="[485,570,724,750]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="169" pageNumber="170" unit="mm" value="32.0">32 mm</quantity>
for 14+45+20 chaetigers. All appendages more or less brown. Palps small, do not project in front of prostomium. Nuchal epaulettes reaching end of chaetiger 3. Median antenna c.
<quantity id="4C9CAAD5CCDE4E89FD837E158AB48355" box="[611,682,804,830]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="169" pageNumber="170" unit="mm" value="5.0">5 mm</quantity>
in length. Lateral bifid antennae, 1/3 as long as the median antenna; basal part 1/3 of total length, outer ventral rami longer than inner dorsal rami. Frontal processes, equal to 1/2 of prostomium width. First dorsal cirri, longer than median antenna. Alternation in direction of cirri not described. Cirri length not described. Large cirrophores on first dorsal cirri present, other cirrophores not described.
</paragraph>
<paragraph id="8BDB0730CCDE4E89FED87EDD8A7D843D" blockId="169.[264,1324,284,1350]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED87EDD898D846D" bold="true" box="[312,403,1004,1030]" pageId="169" pageNumber="170">Female</emphasis>
. Length
<quantity id="4C9CAAD5CCDE4E89FE1C7EDD8A50846D" box="[508,590,1004,1030]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="169" pageNumber="170" unit="mm" value="38.0">38 mm</quantity>
for 14+40+30 chaetigers. All appendages more or less brown. Dorsal surface of body with intersegmental brown bands. Distinct nuchal epaulettes. Egg sac from behind chaetiger 15.
</paragraph>
<paragraph id="8BDB0730CCDE4E89FED879558A3C8415" blockId="169.[264,1324,284,1350]" box="[312,546,1124,1150]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED8795589868415" bold="true" box="[312,408,1124,1150]" pageId="169" pageNumber="170">Habitat</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDE4E89FED879BD8A4A84CD" box="[312,596,1164,1190]" pageId="169" pageNumber="170" type="distribution">
<paragraph id="8BDB0730CCDE4E89FED879BD8A4A84CD" blockId="169.[264,1324,284,1350]" box="[312,596,1164,1190]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED879BD89CF84CD" bold="true" box="[312,465,1164,1190]" pageId="169" pageNumber="170">Distribution</emphasis>
.
<collectingCountry id="F37347A0CCDE4E89FE3F79BD8A4E84CD" box="[479,592,1164,1190]" name="Antarctica" pageId="169" pageNumber="170">Antarctic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDE4E89FED879858C7385D5" pageId="169" pageNumber="170" type="discussion">
<paragraph id="8BDB0730CCDE4E89FED879858B69852E" blockId="169.[264,1324,284,1350]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED8798589B484A5" bold="true" box="[312,426,1204,1230]" pageId="169" pageNumber="170">Remarks</emphasis>
.
<taxonomicName id="4C647CB3CCDE4E89FE5B79858A8684A6" box="[443,664,1204,1229]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="charcoti">
<emphasis id="B910DB22CCDE4E89FE5B79858A8684A6" box="[443,664,1204,1229]" italics="true" pageId="169" pageNumber="170">Epigamia charcoti</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C647CB3CCDE4E89FC6079858C1184A6" box="[896,1039,1204,1229]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="alternata">
<emphasis id="B910DB22CCDE4E89FC6079858C1184A6" box="[896,1039,1204,1229]" italics="true" pageId="169" pageNumber="170">E. alternata</emphasis>
</taxonomicName>
, both taxa have nuchal epaulettes reaching end of chaetiger 3, and long alternating dorsal cirri. The taxa may be separated on the detailed structure of the trepan, as
<taxonomicName id="4C647CB3CCDE4E89FC8A78358BF28576" box="[874,1004,1284,1309]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="charcoti">
<emphasis id="B910DB22CCDE4E89FC8A78358BF28576" box="[874,1004,1284,1309]" italics="true" pageId="169" pageNumber="170">E. charcoti</emphasis>
</taxonomicName>
lacks the additional spines at the base of the larger teeth, present in
<emphasis id="B910DB22CCDE4E89FD03781D8B69852E" box="[739,887,1324,1349]" italics="true" pageId="169" pageNumber="170">
<taxonomicName id="4C647CB3CCDE4E89FD03781D8B6C852E" box="[739,882,1324,1349]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="alternata">E. alternata</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDB0730CCDE4E89FEE878958C7385D5" blockId="169.[264,1133,1444,1470]" box="[264,1133,1444,1470]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FEE878958C7385D5" bold="true" box="[264,1133,1444,1470]" pageId="169" pageNumber="170">
<taxonomicName id="4C647CB3CCDE4E89FEE8789589F785D5" box="[264,489,1444,1470]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDE4E89FEE8789589F785D5" bold="true" box="[264,489,1444,1470]" italics="true" pageId="169" pageNumber="170">Epigamia labordai</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF57AC1CCDE4E89FE1978958B5785D5" author="San" box="[505,841,1444,1470]" pageId="169" pageNumber="170" refString="San Martin, G. &amp; Lopez, E. (2002) New species of Autolytus Grube, 1850, Paraprocerastea San Martin and Alos, 1989, and Sphaerosyllis Claparede, 1863 (Syllidae, Polychaeta) from the Iberian Peninsula. Sarsia, 87, 135 - 143." type="journal article" year="2002">San Martín &amp; López, 2002</bibRefCitation>
) comb. n. (Fig. 86AG)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDE4E89FEE878DE8A49864E" pageId="169" pageNumber="170" type="reference_group">
<paragraph id="8BDB0730CCDE4E89FEE878DE8A49864E" blockId="169.[264,1323,1518,1573]" pageId="169" pageNumber="170">
<treatmentCitationGroup id="AB74201ECCDE4E89FEE878DE8A49864E" pageId="169" pageNumber="170">
<taxonomicName id="4C647CB3CCDE4E89FEE878DE89D1866D" ID-CoL="JZ4V" box="[264,463,1519,1542]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDE4E89FEE878DE89D1866D" box="[264,463,1519,1542]" italics="true" pageId="169" pageNumber="170">Autolytus labordai</emphasis>
</taxonomicName>
<treatmentCitation id="0AC52121CCDE4E89FE3678DF8B31866E" author="San" box="[470,815,1518,1541]" page="136" pageId="169" pageNumber="170" year="2002">
<bibRefCitation id="EFF57AC1CCDE4E89FE3678DF8AE2866E" author="San" box="[470,764,1518,1541]" pageId="169" pageNumber="170" refString="San Martin, G. &amp; Lopez, E. (2002) New species of Autolytus Grube, 1850, Paraprocerastea San Martin and Alos, 1989, and Sphaerosyllis Claparede, 1863 (Syllidae, Polychaeta) from the Iberian Peninsula. Sarsia, 87, 135 - 143." type="journal article" year="2002">San Martín &amp; López, 2002</bibRefCitation>
: 136
</treatmentCitation>
139, figs 1AI, 2AF;
<treatmentCitation id="0AC52121CCDE4E89FBD278DF8D03866E" author="San" box="[1074,1309,1518,1541]" page="502" pageId="169" pageNumber="170" year="2003">
<bibRefCitation id="EFF57AC1CCDE4E89FBD278DF8CF4866E" author="San" box="[1074,1258,1518,1541]" pageId="169" pageNumber="170" refString="San Martin, G. (2003) Annelida, Polychaeta 2: Syllidae. In: Ramos, M. A. et al. (Eds) Fauna Iberica, Museo Nacional de Ciencias Naturales, Madrid, vol 21, 1 - 554." type="journal article" year="2003">San Martín 2003</bibRefCitation>
: 502
</treatmentCitation>
505, figs 278AI, 279AF.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDE4E89FEE87B678BBB86F2" pageId="169" pageNumber="170" type="materials_examined">
<paragraph id="8BDB0730CCDE4E89FEE87B678BBB86F2" blockId="169.[264,1323,1622,1889]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FEE87B6789E2861B" bold="true" box="[264,508,1622,1648]" pageId="169" pageNumber="170">Material examined</emphasis>
.
<emphasis id="B910DB22CCDE4E89FDF37B678A45861B" bold="true" box="[531,603,1622,1648]" pageId="169" pageNumber="170">
<collectingCountry id="F37347A0CCDE4E89FDF37B678A45861B" box="[531,603,1622,1648]" name="Spain" pageId="169" pageNumber="170">Spain</collectingCountry>
</emphasis>
:
<typeStatus id="54DFB992CCDE4E89FD917B668AC6861A" box="[625,728,1623,1649]" pageId="169" pageNumber="170" type="holotype">holotype</typeStatus>
<collectionCode id="ED759FF5CCDE4E89FD067B668B20861A" LSID="urn:lsid:biocol.org:col:35275" box="[742,830,1623,1649]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="169" pageNumber="170">MNCN</collectionCode>
16.01/6096a and 56
<typeStatus id="54DFB992CCDE4E89FBB37B668CDD861A" box="[1107,1219,1623,1649]" pageId="169" pageNumber="170" type="paratype">paratypes</typeStatus>
<collectionCode id="ED759FF5CCDE4E89FB327B668D34861A" LSID="urn:lsid:biocol.org:col:35275" box="[1234,1322,1623,1649]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="169" pageNumber="170">MNCN</collectionCode>
16.01/6096b, Banco de Galicia,
<quantity id="4C9CAAD5CCDE4E89FD627B4E8B1C86F2" box="[642,770,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="7.745" metricValueMax="7.8" metricValueMin="7.69" pageId="169" pageNumber="170" unit="m" value="774.5" valueMax="780.0" valueMin="769.0">769780m</quantity>
,
<date id="FFDA21F0CCDE4E89FCEE7B4E8BBF86F2" box="[782,929,1663,1689]" pageId="169" pageNumber="170" value="1991-06-28">28 Jun 1991</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDE4E89FED87B97896E877A" pageId="169" pageNumber="170" type="diagnosis">
<paragraph id="8BDB0730CCDE4E89FED87B97896E877A" blockId="169.[264,1323,1622,1889]" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED87B9789AE86AB" bold="true" box="[312,432,1702,1728]" pageId="169" pageNumber="170">Diagnosis</emphasis>
.
<taxonomicName id="4C647CB3CCDE4E89FE5E7B968A2E86AB" box="[446,560,1703,1728]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="169" pageNumber="170" phylum="Annelida" rank="genus">
<emphasis id="B910DB22CCDE4E89FE5E7B968A2E86AB" box="[446,560,1703,1728]" italics="true" pageId="169" pageNumber="170">Epigamia</emphasis>
</taxonomicName>
with long nuchal epaulettes; compounds with long blades; trepan with teeth in 3 sizes, 1 large alternating with 2 medium or with 1 medium, 1 small and 1 medium.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDE4E8AFED87A2F8A2C8635" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170" type="description">
<paragraph id="8BDB0730CCDE4E8AFED87A2F89E081ED" blockId="169.[264,1323,1622,1889]" lastBlockId="170.[264,1324,284,1870]" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170">
<emphasis id="B910DB22CCDE4E89FED87A2F89D78753" bold="true" box="[312,457,1822,1848]" pageId="169" pageNumber="170">Description</emphasis>
. All material incomplete anterior fragments. Length of
<typeStatus id="54DFB992CCDE4E89FB807A2E8CD88752" box="[1120,1222,1823,1849]" pageId="169" pageNumber="170" type="holotype">holotype</typeStatus>
<quantity id="4C9CAAD5CCDE4E89FB307A2E8D358752" box="[1232,1323,1823,1849]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="169" pageNumber="170" unit="mm" value="5.6">5.6 mm</quantity>
for 50 chaetigers, width
<quantity id="4C9CAAD5CCDE4E89FDD37A768A8F870A" box="[563,657,1863,1889]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="169" pageNumber="170" unit="mm" value="0.6">0.6 mm</quantity>
(in
<typeStatus id="54DFB992CCDE4E89FD277A768B29870A" box="[711,823,1863,1889]" pageId="169" pageNumber="170" type="paratype">paratypes</typeStatus>
<quantity id="4C9CAAD5CCDE4E89FCA27A768BCD870A" box="[834,979,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.6" metricValueMin="1.7" pageId="169" pageNumber="170" unit="mm" value="2.15" valueMax="2.6" valueMin="1.7">1.72.6 mm</quantity>
for 1420 chaetigers, width
<quantity id="4C9CAAD5CCDD4E8AFEE87C2D89B0815D" box="[264,430,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="5.249999999999999" metricValueMax="7.0" metricValueMin="3.5" pageId="170" pageNumber="171" unit="mm" value="0.5249999999999999" valueMax="0.7" valueMin="0.35">0.350.7 mm</quantity>
). Preserved material yellowish, in some specimens the nuchal epaulettes faintly brown­greenish; eyes dark red. Ciliation not possible to assess (scarce according to original description).
</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED87CA58C508195" blockId="170.[264,1324,284,1870]" pageId="170" pageNumber="171">Eyes confluent; eye spots present. Palps in dorsal view projecting 1/41/3 of prostomial length, fused at base (Fig. 86B). Extension of nuchal epaulettes from beginning of chaetiger 4 to end of chaetiger 6 (Fig. 86A), divergent after chaetiger 1.</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED87F3D8A49830D" blockId="170.[264,1324,284,1870]" pageId="170" pageNumber="171">
All specimens have lost their median antenna as well as their first dorsal cirri. In a few some part of the lateral antennae left. Dorsal tentacular cirri equal in length to body width, ventral pair and second dorsal cirri 1/2 as long. Alternation in direction of cirri not assessed. Dorsal cirri from chaetiger 3, alternate in length in at least to chaetiger 10, short cirri equals 1/3 of body width, long cirri equals 1/2 of body width. Cirrophores present on tentacular cirri, and all dorsal cirri. Cirrophores equal, cirrostyles unequal; short cirrostyles 2/
<quantity id="4C9CAAD5CCDD4E8AFEC17FCD894E837D" box="[289,336,764,790]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="170" pageNumber="171" unit="in" value="3.0">3 in</quantity>
length of long cirrostyles; cirrophores shorter than parapodial lobes; cirrophores shorter than cirrostyles. All appendages cylindrical, including lost anterior appendages (
<bibRefCitation id="EFF57AC1CCDD4E8AFEF07E7D8A55830D" author="San" box="[272,587,844,870]" pageId="170" pageNumber="171" refString="San Martin, G. &amp; Lopez, E. (2002) New species of Autolytus Grube, 1850, Paraprocerastea San Martin and Alos, 1989, and Sphaerosyllis Claparede, 1863 (Syllidae, Polychaeta) from the Iberian Peninsula. Sarsia, 87, 135 - 143." type="journal article" year="2002">San Martín &amp; López 2002</bibRefCitation>
).
</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED87E458D1D846D" blockId="170.[264,1324,284,1870]" pageId="170" pageNumber="171">
Parapodial lobes large, rounded rectangular. Anterior chaetigers with 3 aciculae,
<quantity id="4C9CAAD5CCDD4E8AFB1F7E45890183DD" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="170" pageNumber="171" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
posterior part of fragments. Chaetal fascicle with 1215 compounds. Compound chaetae with small distal tooth (Fig. 86D); serration present; chaetal blades longer than usual in autolytines. Single thin bayonet chaetae (Fig. 86E), beginning between chaetiger 730.
</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED879258D3784A5" blockId="170.[264,1324,284,1870]" pageId="170" pageNumber="171">Pharynx with 1 sinuation, anterior and sometimes lateral to proventricle. Trepan in chaetiger 12 with 3034 unequal teeth; 1 large alternating with 2 medium or 1 medium1 small1 medium (Fig. 86G); arranged in 2 or 3 rings (uncertain observation). Basal ring present; infradental spines may be present (uncertain observation). Proventricle equal in length to 34 segments in chaetiger 815 with 3134 rows of muscle cells. Pygidium lost.</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED879ED89ED858D" blockId="170.[264,1324,284,1870]" pageId="170" pageNumber="171">
<emphasis id="B910DB22CCDD4E8AFED879ED89FA849D" bold="true" box="[312,484,1244,1270]" pageId="170" pageNumber="171">Reproduction</emphasis>
. Epigamy. Several of the specimens have eggs from chaetiger 89 and backwards. In some of the larger specimens the palps are smaller and directed downwards, the eyes are larger and the anterior pair has moved to a more ventral position (Fig. 86C). In one specimen notopodial aciculae (Fig. 86F) and developing swimming chaetae was found from chaetiger 17 and in the same specimen the proventricle had begun to degenerate. In addition, fig. 2C in the original description shows a male with developing bifurcation on the lateral antennae.
</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED878C5899C865D" blockId="170.[264,1324,284,1870]" pageId="170" pageNumber="171">
<emphasis id="B910DB22CCDD4E8AFED878C58ADA8665" bold="true" box="[312,708,1524,1550]" pageId="170" pageNumber="171">Morphology of epitokous stages</emphasis>
. No fully developed epitokous individual found, but see above.
</paragraph>
<paragraph id="8BDB0730CCDD4E8AFED87B758A2C8635" blockId="170.[264,1324,284,1870]" box="[312,562,1604,1630]" pageId="170" pageNumber="171">
<emphasis id="B910DB22CCDD4E8AFED87B7589868635" bold="true" box="[312,408,1604,1630]" pageId="170" pageNumber="171">Habitat</emphasis>
.
<quantity id="4C9CAAD5CCDD4E8AFE467B758A308635" box="[422,558,1604,1630]" metricMagnitude="2" metricUnit="m" metricValue="7.745" metricValueMax="7.8" metricValueMin="7.69" pageId="170" pageNumber="171" unit="m" value="774.5" valueMax="780.0" valueMin="769.0">
769
<specimenCount id="9D62CCB9CCDD4E8AFE017B758A308635" box="[481,558,1604,1630]" pageId="170" pageNumber="171" type="generic">780 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDD4E8AFED87B5D8C8886ED" box="[312,1174,1644,1670]" pageId="170" pageNumber="171" type="distribution">
<paragraph id="8BDB0730CCDD4E8AFED87B5D8C8886ED" blockId="170.[264,1324,284,1870]" box="[312,1174,1644,1670]" pageId="170" pageNumber="171">
<emphasis id="B910DB22CCDD4E8AFED87B5D89CF86ED" bold="true" box="[312,465,1644,1670]" pageId="170" pageNumber="171">Distribution</emphasis>
. North East Atlantic.
<collectingCountry id="F37347A0CCDD4E8AFD337B5D8B0486ED" box="[723,794,1644,1670]" name="Spain" pageId="170" pageNumber="171">Spain</collectingCountry>
. Only known from
<typeStatus id="54DFB992CCDD4E8AFC1E7B5D8C2F86ED" box="[1022,1073,1644,1670]" pageId="170" pageNumber="171">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDD4E8BFED87BA5893081BD" lastPageId="171" lastPageNumber="172" pageId="170" pageNumber="171" type="discussion">
<paragraph id="8BDB0730CCDD4E8BFED87BA5893081BD" blockId="170.[264,1324,284,1870]" lastBlockId="171.[264,1323,284,470]" lastPageId="171" lastPageNumber="172" pageId="170" pageNumber="171">
<emphasis id="B910DB22CCDD4E8AFED87BA589B486C5" bold="true" box="[312,426,1684,1710]" pageId="170" pageNumber="171">Remarks</emphasis>
.
<taxonomicName id="4C647CB3CCDD4E8AFE5B7BA58A8586C6" box="[443,667,1684,1709]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDD4E8AFE5B7BA58A8586C6" box="[443,667,1684,1709]" italics="true" pageId="170" pageNumber="171">Epigamia labordai</emphasis>
</taxonomicName>
, together with
<taxonomicName id="4C647CB3CCDD4E8AFCB67BA58C3D86C6" box="[854,1059,1684,1709]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDD4E8AFCB67BA58C3D86C6" box="[854,1059,1684,1709]" italics="true" pageId="170" pageNumber="171">E. macrophtalma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C647CB3CCDD4E8AFB827BA5899486BD" authority="Imajima, 1966" authorityName="Imajima" authorityYear="1966" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDD4E8AFB827BA58CF786C6" box="[1122,1257,1684,1709]" italics="true" pageId="170" pageNumber="171">E. usaensis</emphasis>
<bibRefCitation id="EFF57AC1CCDD4E8AFB137BA5899486BD" author="Imajima" pageId="170" pageNumber="171" refString="Imajima, M. (1966) The Syllidae (Polychaetous Annelids) from Japan (2) Autolytinae. Publications of the Seto Marine Biological Laboratory, 14, 27 - 83." type="journal article" year="1966">Imajima, 1966</bibRefCitation>
</taxonomicName>
, make up one subclade in
<taxonomicName id="4C647CB3CCDD4E8AFD317B8D8B5D86BE" box="[721,835,1724,1749]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="genus">
<emphasis id="B910DB22CCDD4E8AFD317B8D8B5D86BE" box="[721,835,1724,1749]" italics="true" pageId="170" pageNumber="171">Epigamia</emphasis>
</taxonomicName>
. The clade is supported by a number of apomorphies, e.g.
<emphasis id="B910DB22CCDD4E8AFE047BD58BA28696" box="[484,956,1764,1789]" italics="true" pageId="170" pageNumber="171">number of different teeth­sizes equals 3</emphasis>
(character 41), and
<emphasis id="B910DB22CCDD4E8AFB517BD589AB874E" italics="true" pageId="170" pageNumber="171">number of rings equals 3</emphasis>
(character 44) (uncertain if 2 or
<quantity id="4C9CAAD5CCDD4E8AFCAD7A3D8B60874D" box="[845,894,1804,1830]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="170" pageNumber="171" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C647CB3CCDD4E8AFC687A3D8C0E874E" box="[904,1040,1804,1829]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDD4E8AFC687A3D8C0E874E" box="[904,1040,1804,1829]" italics="true" pageId="170" pageNumber="171">E. labordai</emphasis>
</taxonomicName>
).
<taxonomicName id="4C647CB3CCDD4E8AFBCC7A3D8D12874E" box="[1068,1292,1804,1829]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDD4E8AFBCC7A3D8D12874E" box="[1068,1292,1804,1829]" italics="true" pageId="170" pageNumber="171">Epigamia labordai</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C647CB3CCDD4E8AFE277A058A508726" box="[455,590,1844,1869]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="170" pageNumber="171" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDD4E8AFE277A058A508726" box="[455,590,1844,1869]" italics="true" pageId="170" pageNumber="171">E. usaensis</emphasis>
</taxonomicName>
with which it shares the long bent nuchal epaulettes and the long chaetal blades; however, the trepans differ between the two, in
<taxonomicName id="4C647CB3CCDC4E8BFBD17C2D8CA8815E" box="[1073,1206,284,309]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDC4E8BFBD17C2D8CA8815E" box="[1073,1206,284,309]" italics="true" pageId="171" pageNumber="172">E. usaensis</emphasis>
</taxonomicName>
there is a regular alternation of 1 large, 1 medium, 1 small and 1 medium tooth, in
<taxonomicName id="4C647CB3CCDC4E8BFB847C758CF78136" box="[1124,1257,324,349]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDC4E8BFB847C758CF78136" box="[1124,1257,324,349]" italics="true" pageId="171" pageNumber="172">E. labordai</emphasis>
</taxonomicName>
there is not always a small tooth in between the middle­sized teeth. In addition the mediumsized trepan teeth is not fused by two third to the larger teeth in
<taxonomicName id="4C647CB3CCDC4E8BFC137CA58C6981C6" box="[1011,1143,404,429]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDC4E8BFC137CA58C6981C6" box="[1011,1143,404,429]" italics="true" pageId="171" pageNumber="172">E. labordai</emphasis>
</taxonomicName>
as in
<taxonomicName id="4C647CB3CCDC4E8BFB5C7CA5893981BE" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDC4E8BFB5C7CA5893981BE" italics="true" pageId="171" pageNumber="172">E. usaensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDC4E8BFEE87F058A0483DE" pageId="171" pageNumber="172" type="reference_group">
<paragraph id="8BDB0730CCDC4E8BFEE87F058C1782FE" blockId="171.[264,1095,563,590]" lastBlockId="171.[264,1323,638,949]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87F058C1782FE" pageId="171" pageNumber="172">
<emphasis id="B910DB22CCDC4E8BFEE87F058C598226" bold="true" box="[264,1095,563,590]" pageId="171" pageNumber="172">
<taxonomicName id="4C647CB3CCDC4E8BFEE87F058B308226" authority="Marenzeller, 1875" authorityName="Marenzeller" authorityYear="1875" box="[264,814,563,590]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalma" status="comb. nov.">
<emphasis id="B910DB22CCDC4E8BFEE87F058A2A8225" bold="true" box="[264,564,564,590]" italics="true" pageId="171" pageNumber="172">Epigamia macrophtalma</emphasis>
(
<bibRefCitation id="EFF57AC1CCDC4E8BFDA37F028B388226" author="Marenzeller" box="[579,806,563,589]" pageId="171" pageNumber="172" refString="Marenzeller, E. (1875) Zur Kenntniss der adriatischen Anneliden. Zweiter Beitrag. (Polynoinen, Hesioneen, Syllideen). Sitzungsberichte der Kaiserliche Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Klasse, 72, 1 - 43." type="journal article" year="1875">Marenzeller, 1875</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2236659CCDC4E8BFCD57F028BBE8226" box="[821,928,563,589]" pageId="171" pageNumber="172">comb. n.</taxonomicNameLabel>
(Fig. 87AE)
</emphasis>
<taxonomicName id="4C647CB3CCDC4E8BFEE87F4E8B1582FE" authority="Marenzeller, 1875: 37" authorityName="Marenzeller" authorityPageNumber="37" authorityYear="1875" box="[264,779,638,662]" class="Polychaeta" family="Syllidae" genus="Proceraea" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDC4E8BFEE87F4E8A0682FD" box="[264,536,639,662]" italics="true" pageId="171" pageNumber="172">Proceraea macrophtalma</emphasis>
<treatmentCitation id="0AC52121CCDC4E8BFDFF7F4F8B1582FE" author="Marenzeller" box="[543,779,638,661]" page="37" pageId="171" pageNumber="172" year="1875">
<bibRefCitation id="EFF57AC1CCDC4E8BFDFF7F4F8AFB82FE" author="Marenzeller" box="[543,741,638,661]" pageId="171" pageNumber="172" refString="Marenzeller, E. (1875) Zur Kenntniss der adriatischen Anneliden. Zweiter Beitrag. (Polynoinen, Hesioneen, Syllideen). Sitzungsberichte der Kaiserliche Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Klasse, 72, 1 - 43." type="journal article" year="1875">Marenzeller, 1875</bibRefCitation>
: 37
</treatmentCitation>
</taxonomicName>
39, pl. 4, fig. 2, 2AB.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87FAF8B8882DE" blockId="171.[264,1323,638,949]" box="[264,918,670,694]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87FAF8B8882DE" box="[264,918,670,694]" pageId="171" pageNumber="172">
Not
<taxonomicName id="4C647CB3CCDC4E8BFED77FAE8B2082DE" authority="Langerhans 1879: 579" authorityName="Langerhans" authorityPageNumber="579" authorityYear="1879" box="[311,830,670,694]" class="Polychaeta" family="Syllidae" genus="Proceraea" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDC4E8BFED77FAE8A5982DD" box="[311,583,671,694]" italics="true" pageId="171" pageNumber="172">Proceraea macrophtalma</emphasis>
<treatmentCitation id="0AC52121CCDC4E8BFDAE7FAF8B2082DE" author="Langerhans" box="[590,830,670,693]" page="579" pageId="171" pageNumber="172" year="1879">
<bibRefCitation id="EFF57AC1CCDC4E8BFDAE7FAF8B1482DE" author="Langerhans" box="[590,778,670,693]" pageId="171" pageNumber="172" refString="Langerhans, P. (1879) Die Wurmfauna von Madeira. Zeitschrift fur wissenschaftliche Zoologie, 32, 513 - 592." type="journal article" year="1879">Langerhans 1879</bibRefCitation>
: 579
</treatmentCitation>
</taxonomicName>
, fig. 29.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87F8F8C1C82BE" blockId="171.[264,1323,638,949]" box="[264,1026,702,726]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87F8F8C1C82BE" box="[264,1026,702,726]" pageId="171" pageNumber="172">
Not
<taxonomicName id="4C647CB3CCDC4E8BFED77F8E8BD982BE" authority="Saint-Joseph 1887: 226" authorityName="Saint-Joseph" authorityPageNumber="226" authorityYear="1887" box="[311,967,702,726]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalma" subGenus="Proceraea">
<emphasis id="B910DB22CCDC4E8BFED77F8E8ADB82BD" box="[311,709,703,726]" italics="true" pageId="171" pageNumber="172">Autolytus (Proceraea) macrophtalma</emphasis>
<treatmentCitation id="0AC52121CCDC4E8BFD2B7F8F8BD982BE" author="Saint-Joseph" box="[715,967,702,725]" page="226" pageId="171" pageNumber="172" year="1887">
<bibRefCitation id="EFF57AC1CCDC4E8BFD2B7F8F8B8D82BE" author="Saint-Joseph" box="[715,915,702,725]" pageId="171" pageNumber="172" refString="Saint-Joseph, A. (1887) Les Annelides polychetes des cotes de Dinard. Premiere partie. Annales des sciences naturelles (Zoologie) (ser. 7), 1, 127 - 270." type="journal article" year="1887">Saint­Joseph 1887</bibRefCitation>
: 226
</treatmentCitation>
</taxonomicName>
228.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87FEF8C1E829E" blockId="171.[264,1323,638,949]" box="[264,1024,734,758]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87FEF8C1E829E" box="[264,1024,734,758]" pageId="171" pageNumber="172">
Not
<taxonomicName id="4C647CB3CCDC4E8BFED77FEE8B14829E" authority="Southern 1914: 41" authorityName="Southern" authorityPageNumber="41" authorityYear="1914" box="[311,778,734,758]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDC4E8BFED77FEE8A22829D" box="[311,572,735,758]" italics="true" pageId="171" pageNumber="172">Autolytus macrophtalma</emphasis>
<treatmentCitation id="0AC52121CCDC4E8BFDA37FEF8B14829E" author="Southern" box="[579,778,734,757]" page="41" pageId="171" pageNumber="172" year="1914">
<bibRefCitation id="EFF57AC1CCDC4E8BFDA37FEF8AFD829E" author="Southern" box="[579,739,734,757]" pageId="171" pageNumber="172" refString="Southern, R. (1914) Clare Island Survey. Archiannelida and Polychaeta. Proceedings of the Royal Irish Academy, 31, 1 - 160." type="journal article" year="1914">Southern 1914</bibRefCitation>
: 41
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AC52121CCDC4E8BFCF67FEF8BDB829E" author="Allen" box="[790,965,734,757]" page="870" pageId="171" pageNumber="172" year="1927">
<bibRefCitation id="EFF57AC1CCDC4E8BFCF67FEF8B8F829E" author="Allen" box="[790,913,734,757]" pageId="171" pageNumber="172" refString="Allen, E. J. (1927) Fragmentation in the genus Autolytus and in other syllids. Journal of the Marine Biological Association of the United Kingdom, 14, 869 - 876." type="journal article" year="1927">Allen 1927</bibRefCitation>
: 870
</treatmentCitation>
871.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87FCF8BBC837E" blockId="171.[264,1323,638,949]" box="[264,930,766,790]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87FCF8BBC837E" box="[264,930,766,790]" pageId="171" pageNumber="172">
Not
<taxonomicName id="4C647CB3CCDC4E8BFED77FCE8B10837E" authority="Cognetti 1957: 65" authorityName="Cognetti" authorityPageNumber="65" authorityYear="1957" box="[311,782,766,790]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalmus">
<emphasis id="B910DB22CCDC4E8BFED77FCE8A59837D" box="[311,583,767,790]" italics="true" pageId="171" pageNumber="172">Autolytus macrophtalmus</emphasis>
<treatmentCitation id="0AC52121CCDC4E8BFDAE7FCF8B10837E" author="Cognetti" box="[590,782,766,789]" page="65" pageId="171" pageNumber="172" year="1957">
<bibRefCitation id="EFF57AC1CCDC4E8BFDAE7FCF8AF7837E" author="Cognetti" box="[590,745,766,789]" pageId="171" pageNumber="172" refString="Cognetti, G. (1957) I Sillidi del Golfo di Napoli. Pubblicazioni della Stazione Zoologica di Napoli, 30, 1 - 100." type="journal article" year="1957">Cognetti 1957</bibRefCitation>
: 65
</treatmentCitation>
</taxonomicName>
66, fig. 12B.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87E2E8AEE835E" blockId="171.[264,1323,638,949]" box="[264,752,798,822]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87E2E8AEE835E" box="[264,752,798,822]" pageId="171" pageNumber="172">
<taxonomicName id="4C647CB3CCDC4E8BFEE87E2E8972835D" ID-CoL="d713879c-ca64-40ea-a2b9-b9e1c408eed9" box="[264,364,799,822]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="genus">
<emphasis id="B910DB22CCDC4E8BFEE87E2E8972835D" box="[264,364,799,822]" italics="true" pageId="171" pageNumber="172">Autolytus</emphasis>
</taxonomicName>
sp
<treatmentCitation id="0AC52121CCDC4E8BFE727E2F8A63835E" author="Hauenschild" box="[402,637,798,821]" page="42" pageId="171" pageNumber="172" year="1953">
<bibRefCitation id="EFF57AC1CCDC4E8BFE727E2F8A49835E" author="Hauenschild" box="[402,599,798,821]" pageId="171" pageNumber="172" refString="Hauenschild, C. (1953) Die phanotypische Geschlechtsbestimmung bei Grubea clavata Claparede (Annel. Polych.) und vergleichende Beobachtungen an anderen Sylliden. Zoologische Jahrbucher, Abteilung fur Allgemeine Zoologie und Physiologie der Tiere, 64, 14 - 54." type="journal article" year="1953">Hauenschild 1953</bibRefCitation>
: 42
</treatmentCitation>
43, fig. 5.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87E0E8986831E" blockId="171.[264,1323,638,949]" pageId="171" pageNumber="172">
<treatmentCitationGroup id="AB74201ECCDC4E8BFEE87E0E8986831E" pageId="171" pageNumber="172">
<taxonomicName id="4C647CB3CCDC4E8BFEE87E0E8ACC833E" ID-CoL="JZ66" authority="Martin &amp; Alos, 1987: 579" authorityName="Martin &amp; Alos" authorityPageNumber="579" authorityYear="1987" box="[264,722,830,854]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="sardai">
<emphasis id="B910DB22CCDC4E8BFEE87E0E89A8833D" box="[264,438,831,854]" italics="true" pageId="171" pageNumber="172">Autolytus sardai</emphasis>
<treatmentCitation id="0AC52121CCDC4E8BFE5D7E0F8ACC833E" author="Martin" box="[445,722,830,853]" page="579" pageId="171" pageNumber="172" year="1987">
<bibRefCitation id="EFF57AC1CCDC4E8BFE5D7E0F8A81833E" author="Martin" box="[445,671,830,853]" pageId="171" pageNumber="172" refString="Martin, D. &amp; Alos, C. (1987) Autolytus sardai n. sp. Nueva especie de Autolytinae (Syllidae, Polychaeta) en el Mediterraneo espanol. Cahiers de Biologie Marine, 28, 579 - 584." type="journal article" year="1987">Martin &amp; Alós, 1987</bibRefCitation>
: 579
</treatmentCitation>
</taxonomicName>
584, figs 2AC, 3AC;
<treatmentCitation id="0AC52121CCDC4E8BFC3C7E0F8CDB833E" author="San" box="[988,1221,830,853]" page="498" pageId="171" pageNumber="172" year="2003">
<bibRefCitation id="EFF57AC1CCDC4E8BFC3C7E0F8C8C833E" author="San" box="[988,1170,830,853]" pageId="171" pageNumber="172" refString="San Martin, G. (2003) Annelida, Polychaeta 2: Syllidae. In: Ramos, M. A. et al. (Eds) Fauna Iberica, Museo Nacional de Ciencias Naturales, Madrid, vol 21, 1 - 554." type="journal article" year="2003">San Martín 2003</bibRefCitation>
: 498
</treatmentCitation>
500, fig. 276AE.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFEE87E4E8A0483DE" blockId="171.[264,1323,638,949]" pageId="171" pageNumber="172">
<taxonomicName id="4C647CB3CCDC4E8BFEE87E4E8B4483FE" authority="Nygren &amp; Sundberg 2003" authorityName="Nygren &amp; Sundberg" authorityYear="2003" box="[264,858,894,918]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="species" species="macrophtalmus">
<emphasis id="B910DB22CCDC4E8BFEE87E4E8A0183FD" box="[264,543,895,918]" italics="true" pageId="171" pageNumber="172">Autolytus macrophtalmus</emphasis>
<bibRefCitation id="EFF57AC1CCDC4E8BFDCD7E4F8B4483FE" author="Nygren" box="[557,858,894,917]" pageId="171" pageNumber="172" refString="Nygren, A. &amp; Sundberg, P. (2003) Phylogeny and evolution of reproductive modes in Autolytinae (Syllidae, Annelida). Molecular phylogenetics and evolution, () 29, 235 - 249." type="journal article" year="2003">Nygren &amp; Sundberg 2003</bibRefCitation>
</taxonomicName>
: GenBank sequences, 18S rDNA partial sequence
<accessionNumber id="94379AD3CCDC4E8BFE407EAF8A0883DE" box="[416,534,926,949]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF474293" pageId="171" pageNumber="172">AF474293</accessionNumber>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDC4E8BFEE87ED78B458412" pageId="171" pageNumber="172" type="materials_examined">
<paragraph id="8BDB0730CCDC4E8BFEE87ED78B458412" blockId="171.[264,1324,998,1905]" pageId="171" pageNumber="172">
<emphasis id="B910DB22CCDC4E8BFEE87ED789E6846B" bold="true" box="[264,504,998,1024]" pageId="171" pageNumber="172">Material examined</emphasis>
.
<emphasis id="B910DB22CCDC4E8BFDEA7ED78A5A846B" bold="true" box="[522,580,998,1024]" pageId="171" pageNumber="172">
<collectingCountry id="F37347A0CCDC4E8BFDEA7ED78A5A846B" box="[522,580,998,1024]" name="Italy" pageId="171" pageNumber="172">Italy</collectingCountry>
</emphasis>
: 2
<typeStatus id="54DFB992CCDC4E8BFD8F7ED68AC8846A" box="[623,726,999,1025]" pageId="171" pageNumber="172" type="syntype">syntypes</typeStatus>
on slides
<collectionCode id="ED759FF5CCDC4E8BFCB57ED68BAB846A" box="[853,949,999,1025]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="171" pageNumber="172">NHMW</collectionCode>
2507, Adriatic Sea.
<emphasis id="B910DB22CCDC4E8BFB527ED78D14846B" bold="true" box="[1202,1290,998,1024]" pageId="171" pageNumber="172">
<collectingCountry id="F37347A0CCDC4E8BFB527ED78D14846B" box="[1202,1290,998,1024]" name="France" pageId="171" pageNumber="172">France</collectingCountry>
</emphasis>
: 9 spms (2 spms in formalin, 6 spms on slides (5 rear ends in author's collection for
<collectionCode id="ED759FF5CCDC4E8BFB0B793E8D358442" LSID="urn:lsid:biocol.org:col:15536" box="[1259,1323,1039,1065]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="171" pageNumber="172">DNA</collectionCode>
analyses), 1 spm in author's collection for
<collectionCode id="ED759FF5CCDC4E8BFCF179068B4F843A" LSID="urn:lsid:biocol.org:col:15536" box="[785,849,1079,1105]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="171" pageNumber="172">DNA</collectionCode>
analyses), Banyuls­sur­Mer,
<geoCoordinate id="EE5061F7CCDC4E8BFB5479068D35843A" box="[1204,1323,1079,1105]" direction="north" orientation="latitude" pageId="171" pageNumber="172" precision="92" value="42.476665">42°28.6'N</geoCoordinate>
<geoCoordinate id="EE5061F7CCDC4E8BFEE8796E89618412" box="[264,383,1119,1145]" direction="east" orientation="longitude" pageId="171" pageNumber="172" precision="92" value="3.1616666">03°09.7'E</geoCoordinate>
, dive,
<quantity id="4C9CAAD5CCDC4E8BFE2C796E8A148412" box="[460,522,1119,1145]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="171" pageNumber="172" unit="m" value="30.0">30 m</quantity>
, &quot;coralligene&quot;,
<date id="FFDA21F0CCDC4E8BFD5F796E8B498412" box="[703,855,1119,1145]" pageId="171" pageNumber="172" value="2001-04-26">26 Apr 2001</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDC4E8BFED879B78C6B84A2" pageId="171" pageNumber="172" type="diagnosis">
<paragraph id="8BDB0730CCDC4E8BFED879B78C6B84A2" blockId="171.[264,1324,998,1905]" pageId="171" pageNumber="172">
<emphasis id="B910DB22CCDC4E8BFED879B789AE84CB" bold="true" box="[312,432,1158,1184]" pageId="171" pageNumber="172">Diagnosis</emphasis>
.
<taxonomicName id="4C647CB3CCDC4E8BFE2079B68A2C84CB" box="[448,562,1159,1184]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="171" pageNumber="172" phylum="Annelida" rank="genus">
<emphasis id="B910DB22CCDC4E8BFE2079B68A2C84CB" box="[448,562,1159,1184]" italics="true" pageId="171" pageNumber="172">Epigamia</emphasis>
</taxonomicName>
with very large eyes, dorsal cirri from chaetiger 3 flattened and filled with &quot;empty&quot; glands, and a trepan with 36 teeth in 3 sizes, in 3 rings.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDC4E8CFED879E78ABC83B5" lastPageId="172" lastPageNumber="173" pageId="171" pageNumber="172" type="description">
<paragraph id="8BDB0730CCDC4E8BFED879E78CE88502" blockId="171.[264,1324,998,1905]" pageId="171" pageNumber="172">
<emphasis id="B910DB22CCDC4E8BFED879E789D7849B" bold="true" box="[312,457,1238,1264]" pageId="171" pageNumber="172">Description</emphasis>
. Length
<quantity id="4C9CAAD5CCDC4E8BFDDB79E68AD2849A" box="[571,716,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.8" metricValueMin="1.8" pageId="171" pageNumber="172" unit="mm" value="3.3" valueMax="4.8" valueMin="1.8">1.84.8 mm</quantity>
for 2043 chaetigers, width
<quantity id="4C9CAAD5CCDC4E8BFBD379E68CFB849A" box="[1075,1253,1239,1265]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="171" pageNumber="172" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.200.30 mm</quantity>
. Live specimens yellow in pharyngeal region, intestinal region orange to brownish (Fig. 87A, B). Dorsal cirri from chaetiger 3 with, what looks like, large empty glands, which gives the cirri a checkered appearance (Fig. 87A, B); eyes red. Ciliation as 1 troch per segment.
</paragraph>
<paragraph id="8BDB0730CCDC4E8BFED878468AD6858A" blockId="171.[264,1324,998,1905]" pageId="171" pageNumber="172">Eyes strikingly large, confluent (Fig. 87A, B); eye spots present. Palps in dorsal view projecting 1/5 of prostomial length (Fig. 87A, B), fused at base. Extension of nuchal epaulettes to end of chaetiger 1 (Fig. 87A).</paragraph>
<paragraph id="8BDB0730CCDC4E8BFED878DE8C6A874A" blockId="171.[264,1324,998,1905]" pageId="171" pageNumber="172">Median antenna reaching c. chaetiger 1011 (n=5) in live specimens. Lateral antennae and dorsal tentacular cirri, length 1/22/3 of median antenna. Ventral tentacular cirri 1/2 as long as dorsal pair. First dorsal cirri as long as median antenna, second dorsal cirri as long as ventral tentacular cirri. Alternation in direction of cirri not assessed. Dorsal cirri from chaetiger 3, of equal length (Fig. 87A), c. 1/2 of body width. Cirrophores present on tentacular cirri, and all dorsal cirri. Cirrophores and cirrostyles equal; cirrophores shorter than parapodial lobes; cirrophores shorter than cirrostyles. Antennae, tentacular cirri, first and second dorsal cirri and anal cirri cylindrical, all other dorsal cirri flattened.</paragraph>
<paragraph id="8BDB0730CCDC4E8CFED87A1E8CA28135" blockId="171.[264,1324,998,1905]" lastBlockId="172.[264,1323,284,1470]" lastPageId="172" lastPageNumber="173" pageId="171" pageNumber="172">
Parapodial lobes of medium size, rounded. Anterior chaetigers with 23 aciculae,
<quantity id="4C9CAAD5CCDC4E8BFB1F7A1E8901871A" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="171" pageNumber="172" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
median and posterior. Chaetal fascicle with 611 compounds in anterior chaetigers,
<quantity id="4C9CAAD5CCDC4E8CFB1F7A668901815D" lastPageId="172" lastPageNumber="173" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="171" pageNumber="172" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">46 in</quantity>
median and posterior. Compound chaetae with small distal tooth; serration present (Fig. 87D). Single thin bayonet chaetae (Fig. 87E), beginning between chaetiger 211.
</paragraph>
<paragraph id="8BDB0730CCDB4E8CFED87C5D898A8225" blockId="172.[264,1323,284,1470]" pageId="172" pageNumber="173">Pharynx with 1, 2 or 3 sinuations anterior and lateral to anterior half of proventricle (Fig. 87A). Trepan in chaetiger 1 (Fig. 87A), with 36 unequal teeth in 3 rings, in outer view 9 large alternating with 18 median and 9 smaller (n=6) (Fig. 87C). Basal ring present; infradental spines absent. Proventricle equal in length to 1.52 segments in chaetiger 79 (Fig. 87A) with 1618 rows of muscle cells (n=4). Anal cirri equal in length to 11.5 times body width.</paragraph>
<paragraph id="8BDB0730CCDB4E8CFED87F6A8AB983DD" blockId="172.[264,1323,284,1470]" pageId="172" pageNumber="173">
<emphasis id="B910DB22CCDB4E8CFED87F6A89FA821E" bold="true" box="[312,484,603,629]" pageId="172" pageNumber="173">Reproduction</emphasis>
. Epigamy, concluded from circumstantial evidence. Several of the examined specimens are females with eggs from just behind (Fig. 87B) and at the sides of the proventricle, in two of these specimens one can also see that the parapodia behind chaetiger 14 are larger and are developing notopodial lobes. However, no swimming chaetae could be detected. One of the specimens is also found to have larger eyes and smaller palps, indications of a transition to an epigamic state. The strange &quot;chimaera&quot; described by
<bibRefCitation id="EFF57AC1CCDB4E8CFEE87E7D89EC830D" author="Hauenschild" box="[264,498,844,870]" pageId="172" pageNumber="173" refString="Hauenschild, C. (1953) Die phanotypische Geschlechtsbestimmung bei Grubea clavata Claparede (Annel. Polych.) und vergleichende Beobachtungen an anderen Sylliden. Zoologische Jahrbucher, Abteilung fur Allgemeine Zoologie und Physiologie der Tiere, 64, 14 - 54." type="journal article" year="1953">Hauenschild (1953)</bibRefCitation>
from a photograph taken by Hartmann, is most likely a female of
<taxonomicName id="4C647CB3CCDB4E8CFAF17E7D89AC83E6" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFAF17E7D89AC83E6" italics="true" pageId="172" pageNumber="173">E. macrophtalma</emphasis>
</taxonomicName>
entangled with a posterior end of another individual of male sex. Eggs are found as far forward as chaetiger 5.
</paragraph>
<paragraph id="8BDB0730CCDB4E8CFED87EF58ABC83B5" blockId="172.[264,1323,284,1470]" box="[312,674,964,990]" pageId="172" pageNumber="173">
<emphasis id="B910DB22CCDB4E8CFED87EF5898683B5" bold="true" box="[312,408,964,990]" pageId="172" pageNumber="173">Habitat</emphasis>
. Coralligene, subtidal.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDB4E8CFED87EDD8A8C846D" box="[312,658,1004,1030]" pageId="172" pageNumber="173" type="distribution">
<paragraph id="8BDB0730CCDB4E8CFED87EDD8A8C846D" blockId="172.[264,1323,284,1470]" box="[312,658,1004,1030]" pageId="172" pageNumber="173">
<emphasis id="B910DB22CCDB4E8CFED87EDD89CF846D" bold="true" box="[312,465,1004,1030]" pageId="172" pageNumber="173">Distribution</emphasis>
. Mediterranean.
</paragraph>
</subSubSection>
<subSubSection id="C37E54BBCCDB4E8CFED879258ADA85D5" pageId="172" pageNumber="173" type="discussion">
<paragraph id="8BDB0730CCDB4E8CFED879258ADA85D5" blockId="172.[264,1323,284,1470]" pageId="172" pageNumber="173">
<emphasis id="B910DB22CCDB4E8CFED8792589B48445" bold="true" box="[312,426,1044,1070]" pageId="172" pageNumber="173">Remarks</emphasis>
.
<taxonomicName id="4C647CB3CCDB4E8CFE5C79258AFD8446" box="[444,739,1044,1069]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFE5C79258AFD8446" box="[444,739,1044,1069]" italics="true" pageId="172" pageNumber="173">Epigamia macrophtalma</emphasis>
</taxonomicName>
is easily recognized on its large eyes, flattened cirri, and trepan structure. Its trepan is very similar to
<taxonomicName id="4C647CB3CCDB4E8CFC6C790D8C0E843E" box="[908,1040,1084,1109]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDB4E8CFC6C790D8C0E843E" box="[908,1040,1084,1109]" italics="true" pageId="172" pageNumber="173">E. usaensis</emphasis>
</taxonomicName>
', but in
<taxonomicName id="4C647CB3CCDB4E8CFB92790D8CE9843E" box="[1138,1271,1084,1109]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDB4E8CFB92790D8CE9843E" box="[1138,1271,1084,1109]" italics="true" pageId="172" pageNumber="173">E. usaensis</emphasis>
</taxonomicName>
, the middle­sized teeth are fused with the large tooth.
<taxonomicName id="4C647CB3CCDB4E8CFCBD79558C9C8416" box="[861,1154,1124,1149]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFCBD79558C9C8416" box="[861,1154,1124,1149]" italics="true" pageId="172" pageNumber="173">Epigamia macrophtalma</emphasis>
</taxonomicName>
may be separated from
<taxonomicName id="4C647CB3CCDB4E8CFE6D79BD8A0D84CE" box="[397,531,1164,1189]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="labordai">
<emphasis id="B910DB22CCDB4E8CFE6D79BD8A0D84CE" box="[397,531,1164,1189]" italics="true" pageId="172" pageNumber="173">E. labordai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C647CB3CCDB4E8CFDAF79BD8ACA84CE" box="[591,724,1164,1189]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="usaensis">
<emphasis id="B910DB22CCDB4E8CFDAF79BD8ACA84CE" box="[591,724,1164,1189]" italics="true" pageId="172" pageNumber="173">E. usaensis</emphasis>
</taxonomicName>
on its shorter nuchal epaulettes reaching only end of chaetiger 1, its flattened cirri, and its generally more convoluted pharynx. None of the previous records of
<taxonomicName id="4C647CB3CCDB4E8CFE1179ED8AA5849E" box="[497,699,1244,1269]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFE1179ED8AA5849E" box="[497,699,1244,1269]" italics="true" pageId="172" pageNumber="173">E. macrophtalma</emphasis>
</taxonomicName>
by earlier authors appear to be correct. Although it is not evident which species they refer to,
<taxonomicName id="4C647CB3CCDB4E8CFD3D78358BB68576" box="[733,936,1284,1309]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFD3D78358BB68576" box="[733,936,1284,1309]" italics="true" pageId="172" pageNumber="173">E. macrophtalma</emphasis>
</taxonomicName>
can be excluded. Their &quot;
<taxonomicName id="4C647CB3CCDB4E8CFB2F7835899F852E" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFB2F7835899F852E" italics="true" pageId="172" pageNumber="173">A. macrophtalma</emphasis>
</taxonomicName>
&quot; is a species with 20 equal teeth that have a schizogamous reproduction. The trepan structure is difficult to see, and even Marenzeller himself did not give an accurate description, which is a cause to the confusion. The description of
<taxonomicName id="4C647CB3CCDB4E8CFBF6784D8CC985FE" box="[1046,1239,1404,1429]" class="Polychaeta" family="Syllidae" genus="Autolytus" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="sardai">
<emphasis id="B910DB22CCDB4E8CFBF6784D8CC985FE" box="[1046,1239,1404,1429]" italics="true" pageId="172" pageNumber="173">Autolytus sardai</emphasis>
</taxonomicName>
agrees in every detail with
<taxonomicName id="4C647CB3CCDB4E8CFE1378958AA385D6" box="[499,701,1444,1469]" class="Polychaeta" family="Syllidae" genus="Epigamia" kingdom="Animalia" order="Phyllodocida" pageId="172" pageNumber="173" phylum="Annelida" rank="species" species="macrophtalma">
<emphasis id="B910DB22CCDB4E8CFE1378958AA385D6" box="[499,701,1444,1469]" italics="true" pageId="172" pageNumber="173">E. macrophtalma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>