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<document id="3FF3A808462CD2FC18475CE0D633C532" ID-DOI="10.5281/zenodo.5331014" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331014" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168815873" checkinUser="diego" docAuthor="Megna, Yoandri S. &amp; Epler, John H." docDate="2012" docId="03CA7F7DFFE14C57FE219A594A57FE06" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.383-410.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Copelatus cubaensis Schaeffer 1908" docType="treatment" docVersion="3" lastPageNumber="393" masterDocId="FFF30705FFE84C5DFFAE99134914FFB2" masterDocTitle="A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="410" masterPageNumber="383" pageNumber="392" updateTime="1699171915230" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FE3A13DB2958C8A566D83C7CCAEF74A9">A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="12C2D3A691980BF7C1937A749591CDFF">Megna, Yoandri S.</mods:namePart>
<mods:affiliation id="B4E4928A64C0EFB6C224E9AF61ACA4A9">Departamento de Biología, Universidad de Oriente. Patricio Lumumba s / n, Santiago, Santiago de Cuba, Cuba; e-mail: ysuarez 1976 @ yahoo. es</mods:affiliation>
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<mods:namePart id="0AAF6EC3B54C752B025797B312FAE956">Epler, John H.</mods:namePart>
<mods:affiliation id="371E21D0733FB7F9C5DBD5AA48FC1DD2">461 Tiger Hammock Road, Crawfordville, Florida, U. S. A. 32327; e-mail: johnepler 3 @ comcast. net</mods:affiliation>
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<mods:title id="78D2BCCCE9092505E7CD7E10FD7F5CAB">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="7E9D840C908762FF94BC9B636A3B67FF">2012</mods:date>
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<treatment id="03CA7F7DFFE14C57FE219A594A57FE06" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE14C57FE219A594A57FE06" httpUri="http://treatment.plazi.org/id/03CA7F7DFFE14C57FE219A594A57FE06" lastPageId="10" lastPageNumber="393" pageId="9" pageNumber="392">
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<taxonomicName id="4C63B5E8FFE14C54FE219A594A36FCD0" ID-CoL="Y36G" authority="Schaeffer, 1908" authorityName="Schaeffer" authorityYear="1908" box="[399,802,842,867]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE14C54FE219A594A36FCD0" bold="true" box="[399,802,842,867]" pageId="9" pageNumber="392">
<emphasis id="B9171279FFE14C54FE219A594B64FCD1" bold="true" box="[399,624,842,867]" italics="true" pageId="9" pageNumber="392">Copelatus cubaensis</emphasis>
<bibRefCitation id="EFF2B39AFFE14C54FDD89A594A36FCD0" author="SCHAEFFER C." box="[630,802,842,866]" pageId="9" pageNumber="392" pagination="16 - 18" refId="ref13877" refString="SCHAEFFER C. 1908: On North American and some Cuban Copelatus. Journal of the New York Entomological Society 16: 16 - 18." type="journal article" year="1908">Schaeffer, 1908</bibRefCitation>
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<paragraph id="8BDCCE6BFFE14C54FDBB9A7A4B89FCCE" blockId="9.[113,1090,873,980]" box="[533,669,873,892]" pageId="9" pageNumber="392">
(
<figureCitation id="1358D2EEFFE14C54FDB59A7A4B43FCCE" box="[539,599,873,892]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="9" pageNumber="392">Figs. 4</figureCitation>
,
<figureCitation id="1358D2EEFFE14C54FDCF9A7A4B6CFCCE" box="[609,632,873,892]" captionStart="Figs" captionStartId="14.[151,190,1544,1563]" captionTargetBox="[209,1062,729,1508]" captionTargetId="figure-11@14.[179,1099,723,1525]" captionTargetPageId="14" captionText="Figs. 20–23. Male genitalia of Cuban species of Copelatus. 20 – C. caelatipennis angustatus author, year; 21 – C. cordovai sp. nov.; 22 – C. cubaensis Schaeffer, 1908; 23 – C. posticatus (Fabricius, 1801). Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="9" pageNumber="392">22</figureCitation>
,
<figureCitation id="1358D2EEFFE14C54FD2F9A7A4B83FCCE" box="[641,663,873,892]" captionStart="Figs" captionStartId="20.[151,190,1516,1535]" captionTargetBox="[165,1104,223,1503]" captionTargetId="figure-11@20.[161,1117,219,1504]" captionTargetPageId="20" captionText="Figs. 28–31. Known distribution of Copelatus species in Cuba. 28 – C. cubaensis Schaeffer, 1908; 29 – C. danyi sp. nov.; 30 – C. darlingtoni Young, 1942; 31 – C. insolitus Chevrolat, 1863. See Appendix 1 for additional information on the localities reported in each map." figureDoi="http://doi.org/10.5281/zenodo.5331032" httpUri="https://zenodo.org/record/5331032/files/figure.png" pageId="9" pageNumber="392">28</figureCitation>
)
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<treatmentCitationGroup id="AB73E945FFE14C54FFDF9A9F48C7FC2C" box="[113,467,907,926]" pageId="9" pageNumber="392">
<treatmentCitation id="0AC2E87AFFE14C54FFDF9A9F48C4FC2C" author="SCHAEFFER C." box="[113,464,907,926]" page="18" pageId="9" pageNumber="392" year="1908">
<taxonomicName id="4C63B5E8FFE14C54FFDF9A9F48C4FC2C" ID-CoL="Y36G" authority="Schaeffer, 1908: 18" authorityName="Schaeffer" authorityPageNumber="18" authorityYear="1908" box="[113,464,907,926]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE14C54FFDF9A9F4835FC2C" box="[113,289,908,926]" italics="true" pageId="9" pageNumber="392">Copelatus cubaensis</emphasis>
<bibRefCitation id="EFF2B39AFFE14C54FE899A9848C4FC2C" author="SCHAEFFER C." box="[295,464,907,926]" pageId="9" pageNumber="392" pagination="16 - 18" refId="ref13877" refString="SCHAEFFER C. 1908: On North American and some Cuban Copelatus. Journal of the New York Entomological Society 16: 16 - 18." type="journal article" year="1908">Schaeffer, 1908: 18</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BDCCE6BFFE14C54FFDF9AB448E7FC66" blockId="9.[113,1090,873,980]" pageId="9" pageNumber="392">
<treatmentCitationGroup id="AB73E945FFE14C54FFDF9AB448E7FC66" pageId="9" pageNumber="392">
<treatmentCitation id="0AC2E87AFFE14C54FFDF9AB448D5FC0B" author="YOUNG F. N." box="[113,449,934,953]" page=")" pageId="9" pageNumber="392" year="1963">
<taxonomicName id="4C63B5E8FFE14C54FFDF9AB448D5FC0B" ID-CoL="Y36G" authority=": YOUNG (1963: 60)" authorityName=": YOUNG" authorityPageNumber="60" authorityYear="1963" box="[113,449,934,953]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE14C54FFDF9AB44834FC0B" box="[113,288,935,953]" italics="true" pageId="9" pageNumber="392">Copelatus cubaensis</emphasis>
:
<bibRefCitation id="EFF2B39AFFE14C54FE849AB548D5FC0B" author="YOUNG F. N." box="[298,449,934,953]" pageId="9" pageNumber="392" pagination="56 - 77" refId="ref14144" refString="YOUNG F. N. 1963: The Nearctic species of Copelatus Erichson (Coleoptera: Dytiscidae). Quarterly Journal of the Florida Academy of Sciences 26: 56 - 77." type="journal article" year="1963">YOUNG (1963: 60)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<treatmentCitation id="0AC2E87AFFE14C54FE649AB54B97FC0B" author="SPANGLER P. J." box="[458,643,934,953]" page="167" pageId="9" pageNumber="392" year="1981">
<bibRefCitation id="EFF2B39AFFE14C54FE649AB54B97FC0B" author="SPANGLER P. J." box="[458,643,934,953]" pageId="9" pageNumber="392" pagination="145 - 171" refId="ref13928" refString="SPANGLER P. J. 1981: Supplement to the aquatic and semiaquatic Coleoptera of Cuba collected by the biospeleological expeditions to Cuba by the Academies of Science of Cuba and Romania. Resultats des Expeditions Biospeologiques Cubano Roumaines a Cuba 3: 145 - 171." type="journal article" year="1981">SPANGLER (1981: 167)</bibRefCitation>
</treatmentCitation>
,
<bibRefCitation id="EFF2B39AFFE14C54FD259AB54BE2FC0B" author="EPLER J. H." box="[651,758,934,953]" pageId="9" pageNumber="392" refId="ref12945" refString="EPLER J. H. 1996: Identification manual for the water beetles of Florida (Coleoptera: Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Hydraenidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae, Scirtidae). Florida Department of Environmental Protection, Tallahassee, iv + 253 pp." type="book" year="1996">EPLER (1996</bibRefCitation>
: 3.33, 2010: 5.45),
<treatmentCitation id="0AC2E87AFFE14C54FC369AB549B9FC66" author="LARSON D. J. &amp; ALARIE Y. &amp; ROUGHLEY R. E." page="47" pageId="9" pageNumber="392" year="2000">
<bibRefCitation id="EFF2B39AFFE14C54FC369AB549B9FC66" author="LARSON D. J. &amp; ALARIE Y. &amp; ROUGHLEY R. E." pageId="9" pageNumber="392" refId="ref13318" refString="LARSON D. J., ALARIE Y. &amp; ROUGHLEY R. E. 2000: Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, x + 971 pp." type="book" year="2000">LARSON et al. (2000: 47)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AC2E87AFFE14C54FF199AD2484FFC61" author="NILSSON A. N." box="[183,347,960,980]" page="68" pageId="9" pageNumber="392" year="2001">
<bibRefCitation id="EFF2B39AFFE14C54FF199AD2484FFC61" author="NILSSON A. N." box="[183,347,960,980]" pageId="9" pageNumber="392" pagination="1 - 395" refId="ref13702" refString="NILSSON A. N. 2001: Dytiscidae (Coleoptera). World Catalogue of Insects 3: 1 - 395." type="journal article" year="2001">NILSSON (2001: 68)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AC2E87AFFE14C54FECA9AD248FBFC66" author="PECK S. B." box="[356,495,960,980]" page="44" pageId="9" pageNumber="392" year="2005">
<bibRefCitation id="EFF2B39AFFE14C54FECA9AD248FBFC66" author="PECK S. B." box="[356,495,960,980]" pageId="9" pageNumber="392" pagination="1 - 241" refId="ref13788" refString="PECK S. B. 2005: A checklist of the beetles of Cuba with data on distribution and bionomics (Insecta: Coleoptera). Arthropods of Florida and Neighboring Land Areas 18: 1 - 241." type="journal article" year="2005">PECK (2005: 44)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C3799DE0FFE14C54FFDF9AF94BA9FBFC" pageId="9" pageNumber="392" type="materials_examined">
<paragraph id="8BDCCE6BFFE14C54FFDF9AF94B2DFC4F" blockId="9.[113,1090,1001,1102]" box="[113,569,1001,1021]" pageId="9" pageNumber="392">
<emphasis id="B9171279FFE14C54FFDF9AF949FEFC4F" bold="true" box="[113,234,1001,1021]" pageId="9" pageNumber="392">
<typeStatus id="54D870C9FFE14C54FFDF9AF9498AFC4F" box="[113,158,1002,1021]" pageId="9" pageNumber="392">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3748EFBFFE14C54FF419AF9480AFC4F" box="[239,286,1002,1021]" name="Cuba" pageId="9" pageNumber="392">Cuba</collectingCountry>
,
<collectingRegion id="49A70089FFE14C54FE899AF948CFFC4F" box="[295,475,1002,1021]" country="Cuba" name="Cienfuegos" pageId="9" pageNumber="392">Cienfuegos Province</collectingRegion>
, Cayamas.
</paragraph>
<paragraph id="8BDCCE6BFFE14C54FFDF9D174BA9FBFC" blockId="9.[113,1090,1001,1102]" pageId="9" pageNumber="392">
<emphasis id="B9171279FFE14C54FFDF9D17480AFBAA" bold="true" box="[113,286,1028,1048]" pageId="9" pageNumber="392">Material examined</emphasis>
:
<materialsCitation id="3B0BC436FFE14C54FE849D1748E6FB81" collectingDate="2005-08-06" collectingDateMax="2006-05-28" collectingDateMin="2005-08-06" collectorName="Y. S. Megna" country="Cuba" location="Nueva Gerona" pageId="9" pageNumber="392" specimenCount="2" stateProvince="Isla de la Juventud">
<emphasis id="B9171279FFE14C54FE849D17488CFBA5" bold="true" box="[298,408,1028,1048]" pageId="9" pageNumber="392">
<collectingCountry id="F3748EFBFFE14C54FE849D17487EFBAA" box="[298,362,1028,1048]" name="Cuba" pageId="9" pageNumber="392">CUBA</collectingCountry>
:
<collectingRegion id="49A70089FFE14C54FEDD9D16488CFBA5" box="[371,408,1029,1048]" country="Cuba" name="Isla de la Juventud" pageId="9" pageNumber="392">ISLA</collectingRegion>
</emphasis>
DE LA
<emphasis id="B9171279FFE14C54FE7D9D164B3BFBAA" bold="true" box="[467,559,1029,1048]" pageId="9" pageNumber="392">JUVENTUD:</emphasis>
<location id="8EBC98B0FFE14C54FD9A9D164BA7FBAA" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE14C57FE219A594A57FE06:8EBC98B0FFE14C54FD9A9D164BA7FBAA" box="[564,691,1029,1048]" country="Cuba" name="Nueva Gerona" pageId="9" pageNumber="392" stateProvince="Isla de la Juventud">Nueva Gerona</location>
,
<date id="FFDDE8ABFFE14C54FD129D164A06FBAA" box="[700,786,1029,1048]" pageId="9" pageNumber="392" value="2006-05-28">
<collectingDate id="EF991143FFE14C54FD129D164A06FBAA" box="[700,786,1029,1048]" pageId="9" pageNumber="392" value="2006-05-28">28.v.2006</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE14C54FCB49D164A90FBAA" box="[794,900,1029,1048]" pageId="9" pageNumber="392">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE14C54FC1A9D164ACBFBAA" box="[948,991,1029,1048]" pageId="9" pageNumber="392" type="generic">1 ex.</specimenCount>
<collectingRegion id="49A70089FFE14C54FC4A9D164D2AFBAA" box="[996,1086,1029,1048]" country="Cuba" name="Las Tunas" pageId="9" pageNumber="392">Las Tunas</collectingRegion>
. Sábalo,
<date id="FFDDE8ABFFE14C54FF199D334834FB81" box="[183,288,1056,1075]" pageId="9" pageNumber="392" value="2005-08-06">
<collectingDate id="EF991143FFE14C54FF199D334834FB81" box="[183,288,1056,1075]" pageId="9" pageNumber="392" value="2005-08-06">06.viii.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE14C54FE879D334887FB81" box="[297,403,1056,1075]" pageId="9" pageNumber="392">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE14C54FE6B9D3348E6FB81" box="[453,498,1056,1075]" pageId="9" pageNumber="392" type="generic">1 ex.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE14C54FE559D334A85FB81" box="[507,913,1056,1075]" collectingDate="2009-01-12" collectorName="Y. S. Megna" country="Cuba" location="Comunales" pageId="9" pageNumber="392" specimenCount="2" stateProvince="Isla de la Juventud">
<location id="8EBC98B0FFE14C54FE559D334B49FB81" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE14C57FE219A594A57FE06:8EBC98B0FFE14C54FE559D334B49FB81" box="[507,605,1056,1075]" country="Cuba" name="Comunales" pageId="9" pageNumber="392" stateProvince="Isla de la Juventud">Comunales</location>
,
<date id="FFDDE8ABFFE14C54FDC89D334BADFB81" box="[614,697,1056,1075]" pageId="9" pageNumber="392" value="2009-01-12">
<collectingDate id="EF991143FFE14C54FDC89D334BADFB81" box="[614,697,1056,1075]" pageId="9" pageNumber="392" value="2009-01-12">12.i.2009</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE14C54FD6C9D334A38FB81" box="[706,812,1056,1075]" pageId="9" pageNumber="392">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE14C54FCF39D334A85FB81" box="[861,913,1056,1075]" pageId="9" pageNumber="392" type="generic">2 exs.</specimenCount>
</materialsCitation>
<materialsCitation id="3B0BC436FFE14C54FC389D0C4BA9FBFC" collectingDate="2006-09-03" collectorName="Y. S. Megna &amp; Deler-Hernandez" country="Cuba" location="San Rafael" pageId="9" pageNumber="392" specimenCount="17" stateProvince="Santiago de Cuba">
<emphasis id="B9171279FFE14C54FC389D0C4D54FB81" bold="true" box="[918,1088,1055,1075]" pageId="9" pageNumber="392">
<collectingRegion id="49A70089FFE14C54FC389D0C4D2EFB83" box="[918,1082,1055,1075]" country="Cuba" name="Santiago de Cuba" pageId="9" pageNumber="392">SANTIAGO DE CUBA</collectingRegion>
:
</emphasis>
<location id="8EBC98B0FFE14C54FFDF9D2949C5FBFF" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE14C57FE219A594A57FE06:8EBC98B0FFE14C54FFDF9D2949C5FBFF" box="[113,209,1082,1101]" country="Cuba" name="San Rafael" pageId="9" pageNumber="392" stateProvince="Santiago de Cuba">San Rafael</location>
,
<date id="FFDDE8ABFFE14C54FF779D294822FBFF" box="[217,310,1082,1101]" pageId="9" pageNumber="392" value="2006-09-03">
<collectingDate id="EF991143FFE14C54FF779D294822FBFF" box="[217,310,1082,1101]" pageId="9" pageNumber="392" value="2006-09-03">03.ix.2006</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE14C54FE909D2948B6FBFC" box="[318,418,1082,1102]" pageId="9" pageNumber="392">Y.S. Megna</collectorName>
&amp;
<collectorName id="2696ABBDFFE14C54FE139D294B5BFBFF" box="[445,591,1082,1101]" pageId="9" pageNumber="392">Deler-Hernández</collectorName>
leg.,
<specimenCount id="9D6505E2FFE14C54FDD19D294BA9FBFC" box="[639,701,1082,1102]" pageId="9" pageNumber="392" type="generic">17 exs.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE14C54FFDF9D75489FFA0C" pageId="9" pageNumber="392" type="diagnosis">
<paragraph id="8BDCCE6BFFE14C54FFDF9D754AC3FB6C" blockId="9.[113,1089,1126,1566]" pageId="9" pageNumber="392">
<emphasis id="B9171279FFE14C54FFDF9D7549F0FBCC" bold="true" box="[113,228,1126,1150]" pageId="9" pageNumber="392">Diagnosis.</emphasis>
TL
<quantity id="4C9B638EFFE14C54FEA29D754899FBCF" box="[268,397,1126,1150]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.4" metricValueMin="4.5" pageId="9" pageNumber="392" unit="mm" value="4.95" valueMax="5.4" valueMin="4.5">4.5–5.4 mm</quantity>
; EW
<quantity id="4C9B638EFFE14C54FE6D9D754B57FBCF" box="[451,579,1126,1150]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="9" pageNumber="392" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
; see
<tableCitation id="C6E1FBD0FFE14C54FDDF9D754BA9FBCC" box="[625,701,1126,1150]" captionStart="Table 1" captionStartId="16.[151,198,216,235]" captionText="Table 1. Measurements in mm (minimum, maximum, mean) of morphometric characters for Cuban species of Copelatus Erichson. HL: Head length; HW: Head width; DBE: Distance between eyes; PL: Pronotal length; PW: Pronotal width; EL: Elytral length; EW: Elytral width; TL: Total body length." pageId="9" pageNumber="392">Table 1</tableCitation>
for other body measurements.Adults of this species are recognized by the following combination of characters: elytron with one posterolateral submarginal stria and six discal striae (
<figureCitation id="1358D2EEFFE14C54FD1F9DB54BE0FB0C" box="[689,756,1190,1214]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="9" pageNumber="392">Fig. 4</figureCitation>
); elytra lighter than pronotum; genitalia with broad parameres and median lobe with broad lobate apex (
<figureCitation id="1358D2EEFFE14C54FCD59DD54ADFFB6C" box="[891,971,1222,1246]" captionStart="Figs" captionStartId="14.[151,190,1544,1563]" captionTargetBox="[209,1062,729,1508]" captionTargetId="figure-11@14.[179,1099,723,1525]" captionTargetPageId="14" captionText="Figs. 20–23. Male genitalia of Cuban species of Copelatus. 20 – C. caelatipennis angustatus author, year; 21 – C. cordovai sp. nov.; 22 – C. cubaensis Schaeffer, 1908; 23 – C. posticatus (Fabricius, 1801). Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="9" pageNumber="392">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8BDCCE6BFFE14C54FF3E9DF4489FFA0C" blockId="9.[113,1089,1126,1566]" pageId="9" pageNumber="392">
<taxonomicName id="4C63B5E8FFE14C54FF3E9DF44878FB4C" authorityName=": YOUNG" authorityYear="1963" box="[144,364,1255,1278]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE14C54FF3E9DF44878FB4C" box="[144,364,1255,1278]" italics="true" pageId="9" pageNumber="392">Copelatus cubaensis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C63B5E8FFE14C54FD819DF44BA3FB4C" authorityName="Young" authorityYear="1953" box="[559,695,1255,1278]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="blatchleyi">
<emphasis id="B9171279FFE14C54FD819DF44BA3FB4C" box="[559,695,1255,1278]" italics="true" pageId="9" pageNumber="392">C. blatchleyi</emphasis>
</taxonomicName>
, but differs from it by the characters given in the identification key, the slightly different coloration of elytra, which are usually lighter than pronotum, the male genitalia and the generally larger size (TL
<quantity id="4C9B638EFFE14C54FC299C354D13FA8C" box="[903,1031,1318,1342]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.4" metricValueMin="4.5" pageId="9" pageNumber="392" unit="mm" value="4.95" valueMax="5.4" valueMin="4.5">4.5-5.4 mm</quantity>
). The two species have similar genitalia with broad parameres, but the median lobe of
<taxonomicName id="4C63B5E8FFE14C54FC149C544D54FAEC" authorityName=": YOUNG" authorityYear="1963" box="[954,1088,1351,1374]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE14C54FC149C544D54FAEC" box="[954,1088,1351,1374]" italics="true" pageId="9" pageNumber="392">C. cubaensis</emphasis>
</taxonomicName>
is more truncate apically (
<figureCitation id="1358D2EEFFE14C54FE279C7548F7FACC" box="[393,483,1382,1406]" captionStart="Figs" captionStartId="14.[151,190,1544,1563]" captionTargetBox="[209,1062,729,1508]" captionTargetId="figure-11@14.[179,1099,723,1525]" captionTargetPageId="14" captionText="Figs. 20–23. Male genitalia of Cuban species of Copelatus. 20 – C. caelatipennis angustatus author, year; 21 – C. cordovai sp. nov.; 22 – C. cubaensis Schaeffer, 1908; 23 – C. posticatus (Fabricius, 1801). Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="9" pageNumber="392">Fig. 22a</figureCitation>
) while that of
<taxonomicName id="4C63B5E8FFE14C54FD2F9C744A1DFACC" authorityName="Young" authorityYear="1953" box="[641,777,1383,1406]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="blatchleyi">
<emphasis id="B9171279FFE14C54FD2F9C744A1DFACC" box="[641,777,1383,1406]" italics="true" pageId="9" pageNumber="392">C. blatchleyi</emphasis>
</taxonomicName>
is more rounded (see
<bibRefCitation id="EFF2B39AFFE14C54FC599C7549BEFA2C" author="YOUNG F. N." pageId="9" pageNumber="392" pagination="56 - 77" refId="ref14144" refString="YOUNG F. N. 1963: The Nearctic species of Copelatus Erichson (Coleoptera: Dytiscidae). Quarterly Journal of the Florida Academy of Sciences 26: 56 - 77." type="journal article" year="1963">YOUNG 1963</bibRefCitation>
:
<figureCitation id="1358D2EEFFE14C54FF189C9549E3FA2C" box="[182,247,1414,1438]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="9" pageNumber="392">Fig. 7</figureCitation>
).
<taxonomicName id="4C63B5E8FFE14C54FEA29C9448FCFA2C" authorityName="Young" authorityYear="1953" box="[268,488,1415,1438]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="392" phylum="Arthropoda" rank="species" species="blatchleyi">
<emphasis id="B9171279FFE14C54FEA29C9448FCFA2C" box="[268,488,1415,1438]" italics="true" pageId="9" pageNumber="392">Copelatus blatchleyi</emphasis>
</taxonomicName>
has the elytra usually darker than the pronotum and TL
<quantity id="4C9B638EFFE14C54FFDF9CB549E5FA0F" box="[113,241,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="9" pageNumber="392" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8 mm</quantity>
(
<bibRefCitation id="EFF2B39AFFE14C54FEAE9CB54894FA0C" author="EPLER J. H." box="[256,384,1446,1470]" pageId="9" pageNumber="392" refId="ref12999" refString="EPLER J. H. 2010: The water beetles of Florida - an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae. Florida Department of Environmental Protection, Tallahassee, 399 + iv pp." type="book" year="2010">EPLER 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE14C54FFDF9CD54D55F9AC" pageId="9" pageNumber="392" type="biology_ecology">
<paragraph id="8BDCCE6BFFE14C54FFDF9CD54D55F9AC" blockId="9.[113,1089,1126,1566]" pageId="9" pageNumber="392">
<emphasis id="B9171279FFE14C54FFDF9CD549C5FA6C" bold="true" box="[113,209,1478,1502]" pageId="9" pageNumber="392">Ecology.</emphasis>
<bibRefCitation id="EFF2B39AFFE14C54FF7B9CD5487FFA6C" author="YOUNG F. N." box="[213,363,1478,1502]" pageId="9" pageNumber="392" pagination="56 - 77" refId="ref14144" refString="YOUNG F. N. 1963: The Nearctic species of Copelatus Erichson (Coleoptera: Dytiscidae). Quarterly Journal of the Florida Academy of Sciences 26: 56 - 77." type="journal article" year="1963">YOUNG (1963)</bibRefCitation>
reported that this species was collected using a blacklight trap in Florida. In
<collectingCountry id="F3748EFBFFE14C54FF6D9CF549EAFA4C" box="[195,254,1510,1534]" name="Cuba" pageId="9" pageNumber="392">Cuba</collectingCountry>
, the species was collected in lentic (permanent and temporary) habitats located in lowlands, where it was predominantly associated with turbid waters with muddy bottom.
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE24C57FF3999C84A7AFEE6" pageId="10" pageNumber="393" type="distribution">
<paragraph id="8BDCCE6BFFE24C57FF3999C84A7AFEE6" blockId="10.[151,1127,219,436]" pageId="10" pageNumber="393">
<emphasis id="B9171279FFE24C57FF3999C8483EFF41" bold="true" box="[151,298,219,243]" pageId="10" pageNumber="393">Distribution.</emphasis>
In the
<collectingCountry id="F3748EFBFFE24C57FED499CF48BAFF46" box="[378,430,220,244]" name="United States of America" pageId="10" pageNumber="393">USA</collectingCountry>
known from southern
<collectingRegion id="49A70089FFE24C57FD0799CF4BE2FF46" box="[681,758,220,244]" country="United States of America" name="Florida" pageId="10" pageNumber="393">Florida</collectingRegion>
(
<bibRefCitation id="EFF2B39AFFE24C57FCA899CF4A92FF41" author="EPLER J. H." box="[774,902,219,244]" pageId="10" pageNumber="393" refId="ref12945" refString="EPLER J. H. 1996: Identification manual for the water beetles of Florida (Coleoptera: Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Hydraenidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae, Scirtidae). Florida Department of Environmental Protection, Tallahassee, iv + 253 pp." type="book" year="1996">EPLER 1996</bibRefCitation>
,
<bibRefCitation id="EFF2B39AFFE24C57FC3D99CF4AD8FF46" author="EPLER J. H." box="[915,972,220,244]" pageId="10" pageNumber="393" refId="ref12999" refString="EPLER J. H. 2010: The water beetles of Florida - an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae. Florida Department of Environmental Protection, Tallahassee, 399 + iv pp." type="book" year="2010">2010</bibRefCitation>
;
<bibRefCitation id="EFF2B39AFFE24C57FC7699CF49C5FEA6" author="LARSON D. J. &amp; ALARIE Y. &amp; ROUGHLEY R. E." pageId="10" pageNumber="393" refId="ref13318" refString="LARSON D. J., ALARIE Y. &amp; ROUGHLEY R. E. 2000: Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, x + 971 pp." type="book" year="2000">LARSON et al. 2000</bibRefCitation>
). Until now, all Cuban records of
<taxonomicName id="4C63B5E8FFE24C57FD8599EE4BA6FEA6" authorityName=": YOUNG" authorityYear="1963" box="[555,690,253,276]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE24C57FD8599EE4BA6FEA6" box="[555,690,253,276]" italics="true" pageId="10" pageNumber="393">C. cubaensis</emphasis>
</taxonomicName>
were from western
<collectingCountry id="F3748EFBFFE24C57FCD199EF4AA3FEA6" box="[895,951,252,276]" name="Cuba" pageId="10" pageNumber="393">Cuba</collectingCountry>
only (
<bibRefCitation id="EFF2B39AFFE24C57FC5699EF49DBFE86" author="SCHAEFFER C." pageId="10" pageNumber="393" pagination="16 - 18" refId="ref13877" refString="SCHAEFFER C. 1908: On North American and some Cuban Copelatus. Journal of the New York Entomological Society 16: 16 - 18." type="journal article" year="1908">SCHAEFFER 1908</bibRefCitation>
,
<bibRefCitation id="EFF2B39AFFE24C57FF77980F484AFE81" author="YOUNG F. N." box="[217,350,283,307]" pageId="10" pageNumber="393" pagination="56 - 77" refId="ref14144" refString="YOUNG F. N. 1963: The Nearctic species of Copelatus Erichson (Coleoptera: Dytiscidae). Quarterly Journal of the Florida Academy of Sciences 26: 56 - 77." type="journal article" year="1963">YOUNG 1963</bibRefCitation>
,
<bibRefCitation id="EFF2B39AFFE24C57FEC7980F48C9FE81" author="PECK S. B." box="[361,477,283,307]" pageId="10" pageNumber="393" pagination="1 - 241" refId="ref13788" refString="PECK S. B. 2005: A checklist of the beetles of Cuba with data on distribution and bionomics (Insecta: Coleoptera). Arthropods of Florida and Neighboring Land Areas 18: 1 - 241." type="journal article" year="2005">PECK 2005</bibRefCitation>
), but the distribution range of the species is now extended to the
<collectingRegion id="49A70089FFE24C57FF10982F4891FEE6" box="[190,389,316,340]" country="Cuba" name="Isla de la Juventud" pageId="10" pageNumber="393">Isla de la Juventud</collectingRegion>
and the eastern parts of the country (
<figureCitation id="1358D2EEFFE24C57FCBC982F4A76FEE6" box="[786,866,316,340]" captionStart="Figs" captionStartId="20.[151,190,1516,1535]" captionTargetBox="[165,1104,223,1503]" captionTargetId="figure-11@20.[161,1117,219,1504]" captionTargetPageId="20" captionText="Figs. 28–31. Known distribution of Copelatus species in Cuba. 28 – C. cubaensis Schaeffer, 1908; 29 – C. danyi sp. nov.; 30 – C. darlingtoni Young, 1942; 31 – C. insolitus Chevrolat, 1863. See Appendix 1 for additional information on the localities reported in each map." figureDoi="http://doi.org/10.5281/zenodo.5331032" httpUri="https://zenodo.org/record/5331032/files/figure.png" pageId="10" pageNumber="393">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE24C57FF39984F4A57FE06" pageId="10" pageNumber="393" type="discussion">
<paragraph id="8BDCCE6BFFE24C57FF39984F4A57FE06" blockId="10.[151,1127,219,436]" pageId="10" pageNumber="393">
<emphasis id="B9171279FFE24C57FF39984F4812FEC6" bold="true" box="[151,262,348,372]" pageId="10" pageNumber="393">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C63B5E8FFE24C57FDB3984E4A2DFEC1" authority="Young, 1953" authorityName="Young" authorityYear="1953" box="[541,825,347,372]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="blatchleyi">
<emphasis id="B9171279FFE24C57FDB3984E4BB2FEC6" box="[541,678,349,372]" italics="true" pageId="10" pageNumber="393">C. blatchleyi</emphasis>
Young, 1953
</taxonomicName>
, which is not recorded from
<collectingCountry id="F3748EFBFFE24C57FF39986F49C7FE26" box="[151,211,380,404]" name="Cuba" pageId="10" pageNumber="393">Cuba</collectingCountry>
. However, the presence of
<taxonomicName id="4C63B5E8FFE24C57FE5D986E4B6FFE26" authorityName="Young" authorityYear="1953" box="[499,635,381,404]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="393" phylum="Arthropoda" rank="species" species="blatchleyi">
<emphasis id="B9171279FFE24C57FE5D986E4B6FFE26" box="[499,635,381,404]" italics="true" pageId="10" pageNumber="393">C. blatchleyi</emphasis>
</taxonomicName>
in extreme southern Florida and
<collectingCountry id="F3748EFBFFE24C57FC73986F4D73FE26" box="[989,1127,380,404]" name="Bahamas" pageId="10" pageNumber="393">the Bahamas</collectingCountry>
(
<bibRefCitation id="EFF2B39AFFE24C57FF31988F4827FE01" author="YOUNG F. N." box="[159,307,411,436]" pageId="10" pageNumber="393" pagination="21 - 27" refId="ref14050" refString="YOUNG F. N. 1953 a: Two new species of water beetles from Florida (Coleoptera: Dytiscidae). Psyche 60: 21 - 27." type="journal article" year="1953">YOUNG 1953a</bibRefCitation>
,
<bibRefCitation id="EFF2B39AFFE24C57FE90988F48AEFE06" author="EPLER J. H." box="[318,442,412,436]" pageId="10" pageNumber="393" refId="ref12999" refString="EPLER J. H. 2010: The water beetles of Florida - an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae. Florida Department of Environmental Protection, Tallahassee, 399 + iv pp." type="book" year="2010">EPLER 2010</bibRefCitation>
) indicates that it may occur in
<collectingCountry id="F3748EFBFFE24C57FCAD988F4A2BFE06" box="[771,831,412,436]" name="Cuba" pageId="10" pageNumber="393">Cuba</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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