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<document id="18630038E69A7F309F2FE812D249803A" ID-CLB-Dataset="2588" ID-DOI="10.11646/zootaxa.5091.3.2" ID-GBIF-Dataset="f6fc2fe0-6f9c-48cf-b252-e49db45b4a98" ID-ISSN="1175-5326" ID-Zenodo-Dep="5848281" ID-ZooBank="57DE1453-41B1-43B2-9B9E-B72BDA973208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642146879451" checkinUser="plazi" docAuthor="Durante, Antonio &amp; Apinda-Legnouo, Emelie Arlette" docDate="2022" docId="03ACBD4BFFA1FFFE0D8FFEC33292F815" docLanguage="en" docName="zootaxa.5091.3.2.pdf" docOrigin="Zootaxa 5091 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Novilema bifurcum Durante &amp; Apinda-Legnouo 2022, new species" docType="treatment" docVersion="6" lastPageNumber="424" masterDocId="FF95C533FFA9FFF60D18FFAD337BFF81" masterDocTitle="Fifth contribution to the study of the Lithosiini of Gabon: two genera new for Gabon, with a description of two new species (Lepidoptera: Erebidae: Arctiinae)" masterLastPageNumber="428" masterPageNumber="416" pageNumber="424" updateTime="1699302283932" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FC2CA3D377718DF4952E313FD57D24E9">Fifth contribution to the study of the Lithosiini of Gabon: two genera new for Gabon, with a description of two new species (Lepidoptera: Erebidae: Arctiinae)</mods:title>
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<mods:namePart id="56A5C79BC899951EBC7872AC158DC981">Durante, Antonio</mods:namePart>
<mods:affiliation id="C5778C2228D34987528ABD1A68CCEE44">Museo di Storia naturale del Salento. S. P. Calimera-Borgagne, Km 1, 73021 Calimera, Italy.</mods:affiliation>
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<mods:namePart id="B89C4B8B38DA5352FBD5DDA4E5533CE4">Apinda-Legnouo, Emelie Arlette</mods:namePart>
<mods:affiliation id="60087719F316F731AA35F22AD0432836">Institut de Recherches sur lÉcologie Tropicale, Libreville, Gabon.</mods:affiliation>
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<mods:date id="9B7C6382DC45250E73F5DF713663CE81">2022</mods:date>
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<treatment id="03ACBD4BFFA1FFFE0D8FFEC33292F815" ID-DOI="http://doi.org/10.5281/zenodo.5848267" ID-GBIF-Taxon="192131087" ID-Zenodo-Dep="5848267" LSID="urn:lsid:plazi:treatment:03ACBD4BFFA1FFFE0D8FFEC33292F815" httpUri="http://treatment.plazi.org/id/03ACBD4BFFA1FFFE0D8FFEC33292F815" lastPageNumber="424" pageId="8" pageNumber="424">
<subSubSection id="C31F5FD6FFA1FFFE0D8FFEC331F2FE09" box="[151,649,366,393]" pageId="8" pageNumber="424" type="nomenclature">
<paragraph id="8BBA0C5DFFA1FFFE0D8FFEC331F2FE09" blockId="8.[151,649,366,428]" box="[151,649,366,393]" pageId="8" pageNumber="424">
<heading id="D0F2BB31FFA1FFFE0D8FFEC331F2FE09" bold="true" box="[151,649,366,393]" fontSize="11" level="1" pageId="8" pageNumber="424" reason="1">
<emphasis id="B971D04FFFA1FFFE0D8FFEC331F2FE09" bold="true" box="[151,649,366,393]" pageId="8" pageNumber="424">
<taxonomicName id="4C0577DEFFA1FFFE0D8FFEC33289FE09" ID-CoL="9BDFL" authority="Durante" authorityName="Durante &amp; Apinda-Legnouo" authorityYear="2022" box="[151,498,366,393]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="bifurcum" status="sp. nov.">
<emphasis id="B971D04FFFA1FFFE0D8FFEC332F9FE08" bold="true" box="[151,386,366,393]" italics="true" pageId="8" pageNumber="424">Novilema bifurcum</emphasis>
Durante
</taxonomicName>
<taxonomicNameLabel id="A2426D34FFA1FFFE0CE2FEC331F2FE09" box="[506,649,366,392]" pageId="8" pageNumber="424" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31F5FD6FFA1FFFE0D8FFE3F3263FE2D" box="[151,280,402,428]" pageId="8" pageNumber="424" type="description">
<paragraph id="8BBA0C5DFFA1FFFE0D8FFE3F3263FE2D" blockId="8.[151,649,366,428]" box="[151,280,402,428]" pageId="8" pageNumber="424">
(
<figureCitation id="133E10D8FFA1FFFE0D87FE3F3393FE2D" box="[159,232,402,428]" captionStart="FIGURES 18" captionStartId="4.[151,264,1866,1890]" captionTargetBox="[170,1414,194,1826]" captionTargetId="figure-21@4.[160,1427,179,1841]" captionTargetPageId="4" captionText="FIGURES 18. Lithosiini spp.: adults. 1, Pusiola unicolor n. sp.—holotype male. Gabon, Ivindo National Park. 2, ditto—paratype, male. Gabon, Ivindo National Park. 3, ditto—paratype, female. Gabon, Ivindo National Park. 4, ditto—paratype, female. Gabon, Ivindo National Park. 5, Pusiola celida—male. Gabon, Mondah forest. 6, Novilema bifurcum n. sp.—holotype, male. Gabon, Ivindo National Park. 7, Novilema fluminale—male. Gabon, Ivindo National Park. 8, ditto—male. Gabon, Ivindo National Park." figureDoi="http://doi.org/10.5281/zenodo.5848283" httpUri="https://zenodo.org/record/5848283/files/figure.png" pageId="8" pageNumber="424">Figs 6</figureCitation>
,
<figureCitation id="133E10D8FFA1FFFE0DECFE3F3274FE2D" box="[244,271,402,428]" captionStart="FIGURE 11" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[211,1391,194,1860]" captionTargetId="figure-27@7.[196,1391,181,1867]" captionTargetPageId="7" captionText="FIGURE 11. Novilema bifurcum n. sp.—AD: male genitalia. A: genitalia, aedeagus in situ; B: Uncus, socius and terminal portion of the juxta, enlarged; C: aedeagus, vesica everted, enlarged; D: granicula of the vesica, enlarged. ae: aedeagus; av: ala valvae; cr: cornuti; gr: granicula; ju: juxta; pav: distal process of the ala valvae; so: socius; sv: supravalva; te: tegumen; un: uncus; tr: transtilla; ve: vesica; vi: vinculum; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848289" httpUri="https://zenodo.org/record/5848289/files/figure.png" pageId="8" pageNumber="424">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31F5FD6FFA1FFFE0D8FFE7732BBFD99" pageId="8" pageNumber="424" type="materials_examined">
<paragraph id="8BBA0C5DFFA1FFFE0D8FFE7732BBFD99" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<materialsCitation id="3B6D0600FFA1FFFE0D8FFE7732BBFD99" ID-GBIF-Occurrence="3436732319" collectingDate="2017-11-24" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="8" pageNumber="424" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B971D04FFFA1FFFE0D8FFE77327FFE75" bold="true" box="[151,260,474,500]" pageId="8" pageNumber="424">
<typeStatus id="54BEB2FFFFA1FFFE0D8FFE77327FFE75" box="[151,260,474,500]" pageId="8" pageNumber="424" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D03C7D4FFA1FFFE0C0AFE763252FE74" box="[274,297,475,501]" pageId="8" pageNumber="424" type="male"></specimenCount>
,
<collectingCountry id="F3124CCDFFA1FFFE0C2DFE7732E0FE75" box="[309,411,474,500]" name="Gabon" pageId="8" pageNumber="424">GABON</collectingCountry>
,
<location id="8EDA5A86FFA1FFFE0CBEFE763297FE74" LSID="urn:lsid:plazi:treatment:03ACBD4BFFA1FFFE0D8FFEC33292F815:8EDA5A86FFA1FFFE0CBEFE763297FE74" box="[422,492,475,501]" country="Gabon" latitude="0.5119445" longLatPrecision="21" longitude="12.803612" name="Ipassa" pageId="8" pageNumber="424">Ipassa</location>
(Makokou),
<quantity id="4CFDA1B8FFA1FFFE0F66FE7731BDFE74" box="[638,710,474,501]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="424" unit="m" value="500.0">
<elevation id="0028EB6EFFA1FFFE0F66FE7731BDFE74" box="[638,710,474,501]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="424" unit="m" value="500.0">500 m</elevation>
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,
<geoCoordinate id="EE316A9AFFA1FFFE0FCAFE773036FE75" box="[722,845,474,500]" degrees="0" direction="north" minutes="30" orientation="latitude" pageId="8" pageNumber="424" precision="15" seconds="43" value="0.5119445">0°3043”N</geoCoordinate>
<geoCoordinate id="EE316A9AFFA1FFFE0E4BFE7730A0FE75" box="[851,987,474,500]" degrees="12" direction="east" minutes="48" orientation="longitude" pageId="8" pageNumber="424" precision="15" seconds="13" value="12.803612">12°4813”E</geoCoordinate>
, 19/
<date id="FFBB2A9DFFA1FFFE0913FE7737F4FE75" box="[1035,1167,474,500]" pageId="8" pageNumber="424" value="2017-11-24">
<collectingDate id="EFFFD375FFA1FFFE0913FE7737F4FE75" box="[1035,1167,474,500]" pageId="8" pageNumber="424" value="2017-11-24">24-11-2017</collectingDate>
</date>
,
<collectorName id="26F0698BFFA1FFFE0982FE76378FFE74" box="[1178,1268,475,501]" pageId="8" pageNumber="424">Durante</collectorName>
leg., gen. sl. n. 997
<collectionCode id="ED149498FFA1FFFE0DD0FE533277FD99" box="[200,268,510,536]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="8" pageNumber="424">MAD</collectionCode>
, in coll.
<collectionCode id="ED149498FFA1FFFE0C74FE5332C7FD99" box="[364,444,510,536]" pageId="8" pageNumber="424">MSNS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F5FD6FFA1FFFE0DDFFD8F3009FC94" pageId="8" pageNumber="424" type="diagnosis">
<paragraph id="8BBA0C5DFFA1FFFE0DDFFD8F3613FDE1" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFFD8F3241FDBD" bold="true" box="[199,314,546,572]" pageId="8" pageNumber="424">Diagnosis</emphasis>
. The differences between
<taxonomicName id="4C0577DEFFA1FFFE0F79FD8F304CFDBD" authority="Durante &amp; Apinda-Legnouo, 2022" authorityName="Durante &amp; Apinda-Legnouo" authorityYear="2022" box="[609,823,546,572]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="bifurcum" status="sp. nov.">
<emphasis id="B971D04FFFA1FFFE0F79FD8F304CFDBD" box="[609,823,546,572]" italics="true" pageId="8" pageNumber="424">Novilema bifurcum</emphasis>
</taxonomicName>
<emphasis id="B971D04FFFA1FFFE0E26FD8F3006FDBD" bold="true" box="[830,893,546,572]" pageId="8" pageNumber="424">
<taxonomicNameLabel id="A2426D34FFA1FFFE0E26FD8F3006FDBD" box="[830,893,546,572]" pageId="8" pageNumber="424" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and its congeneric
<taxonomicName id="4C0577DEFFA1FFFE094EFD8E379AFDBD" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[1110,1249,546,572]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="fluminale">
<emphasis id="B971D04FFFA1FFFE094EFD8E379AFDBD" box="[1110,1249,546,572]" italics="true" pageId="8" pageNumber="424">N. fluminale</emphasis>
</taxonomicName>
in the shape and colour of the habitus are practically null, so the diagnosis is entirely based on the morphology of the genitalia.
</paragraph>
<paragraph id="8BBA0C5DFFA1FFFE0DDFFDC732F0FD28" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
The generic features are unmistakeably maintained, and it is thus very easy to assign this species as a
<taxonomicName id="4C0577DEFFA1FFFE0829FDC736E7FD05" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[1329,1436,618,644]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B971D04FFFA1FFFE0829FDC736E7FD05" box="[1329,1436,618,644]" italics="true" pageId="8" pageNumber="424">Novilema</emphasis>
</taxonomicName>
(see Remarks below).
</paragraph>
<paragraph id="8BBA0C5DFFA1FFFE0DDFFD1F3009FC94" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
The main differences from
<taxonomicName id="4C0577DEFFA1FFFE0CE3FD1E31FCFD4D" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[507,647,690,716]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="fluminale">
<emphasis id="B971D04FFFA1FFFE0CE3FD1E31FCFD4D" box="[507,647,690,716]" italics="true" pageId="8" pageNumber="424">N. fluminale</emphasis>
</taxonomicName>
are: the distal process of the ala valvae (sacculus) has a larger pincershaped bifurcation (slender in
<taxonomicName id="4C0577DEFFA1FFFE0CEBFD7A3104FD71" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[499,639,726,752]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="fluminale">
<emphasis id="B971D04FFFA1FFFE0CEBFD7A3104FD71" box="[499,639,726,752]" italics="true" pageId="8" pageNumber="424">N. fluminale</emphasis>
</taxonomicName>
); the apex of the supravalva is pointed (rounded in
<taxonomicName id="4C0577DEFFA1FFFE09CCFD7A361BFD71" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[1236,1376,726,752]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="fluminale">
<emphasis id="B971D04FFFA1FFFE09CCFD7A361BFD71" box="[1236,1376,726,752]" italics="true" pageId="8" pageNumber="424">N. fluminale</emphasis>
</taxonomicName>
); the juxta is Y-shaped at its distal end (not bifurcated in
<taxonomicName id="4C0577DEFFA1FFFE0FCFFD563019FC95" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[727,866,762,788]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="fluminale">
<emphasis id="B971D04FFFA1FFFE0FCFFD563019FC95" box="[727,866,762,788]" italics="true" pageId="8" pageNumber="424">N. fluminale</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F5FD6FFA1FFFE0DDFFCB3302EF9D1" pageId="8" pageNumber="424" type="description">
<paragraph id="8BBA0C5DFFA1FFFE0DDFFCB33056FCB9" blockId="8.[151,1437,474,1940]" box="[199,813,798,825]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFFCB3322AFCB9" bold="true" box="[199,337,798,824]" pageId="8" pageNumber="424">Description</emphasis>
. Wingspan
<specimenCount id="9D03C7D4FFA1FFFE0CCFFCB23297FCB8" box="[471,492,799,825]" pageId="8" pageNumber="424" type="male"></specimenCount>
<quantity id="4CFDA1B8FFA1FFFE0CEBFCB33121FCB8" box="[499,602,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="8" pageNumber="424" unit="mm" value="18.5">18.5 mm</quantity>
. Female unknown.
</paragraph>
<paragraph id="8BBA0C5DFFA1FFFE0DDFFCEF37D4FC01" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFFCEF327FFCDD" box="[199,260,834,860]" italics="true" pageId="8" pageNumber="424">Head</emphasis>
slightly paler than forewing upperside; antennae ciliated, dorsally same colour as the head, ventrally unpigmented; labial palpi porrect, not extending beyond the frons, slightly darker than the head.
</paragraph>
<paragraph id="8BBA0C5DFFA1FFFE0DDFFC2731D0FBB5" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFFC273263FC25" box="[199,280,906,932]" italics="true" pageId="8" pageNumber="424">Thorax</emphasis>
. Anterior margin of the patagia slightly paler than forewing upperside; posterior area of the patagia, thorax, tegulae, legs and anal tuft same colour as the forewing upperside; spurs formula 0-2-4. Forewing upperside ground colour ochreous yellow, paler towards the termen. Forewing underside slightly darker than upperside, costa and inner margin paler. Hindwing upperside straw yellow with costal margin and apex darker. Hindwing underside same colour as upperside. Fringes concolorous.
</paragraph>
<paragraph id="8BBA0C5DFFA1FFFE0DDFFB9332B2FBD9" blockId="8.[151,1437,474,1940]" box="[199,457,1086,1112]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFFB93324AFBD9" box="[199,305,1086,1112]" italics="true" pageId="8" pageNumber="424">Abdomen</emphasis>
slightly pale.
</paragraph>
<paragraph id="8BBA0C5DFFA1FFFE0DDFFBCF302EF9D1" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFFBCF324EFBFD" bold="true" box="[199,309,1122,1148]" pageId="8" pageNumber="424">Genitalia</emphasis>
<specimenCount id="9D03C7D4FFA1FFFE0C23FBCE3228FBFC" box="[315,339,1123,1149]" pageId="8" pageNumber="424" type="male"></specimenCount>
. Uncus laterally compressed, ending in a point, pedunculated, rising from an unpaired semi-circular structure probably belonging to the cephalic margin of A10, possibly a socius (
<figureCitation id="133E10D8FFA1FFFE0919FB2B371DFB21" box="[1025,1126,1158,1184]" captionStart="FIGURE 11" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[211,1391,194,1860]" captionTargetId="figure-27@7.[196,1391,181,1867]" captionTargetPageId="7" captionText="FIGURE 11. Novilema bifurcum n. sp.—AD: male genitalia. A: genitalia, aedeagus in situ; B: Uncus, socius and terminal portion of the juxta, enlarged; C: aedeagus, vesica everted, enlarged; D: granicula of the vesica, enlarged. ae: aedeagus; av: ala valvae; cr: cornuti; gr: granicula; ju: juxta; pav: distal process of the ala valvae; so: socius; sv: supravalva; te: tegumen; un: uncus; tr: transtilla; ve: vesica; vi: vinculum; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848289" httpUri="https://zenodo.org/record/5848289/files/figure.png" pageId="8" pageNumber="424">Fig. 11B</figureCitation>
); tegumen delicate, roughly triangular in dorsal view, with a bisinuated dorso-caudal margin; vinculum also delicate, with its arms approaching each other at the distal end (i.e. ventrally), assuming a bifid shape; saccus absent; juxta swollen at the basal 2/5, then protruding in a sturdy long stem ending in a fork; fultura superior a rod-like arched transtilla joining the opposite plicae centripetalis (sensu
<bibRefCitation id="EF9471ACFFA1FFFE0CD1FABB31DAFAB0" author="Birket-Smith, J." box="[457,673,1302,1329]" pageId="8" pageNumber="424" pagination="1 - 161" refId="ref6365" refString="Birket-Smith, J. (1965) A revision of the West African Eilemic moths, based on the male genitalia (Lep., Arctiidae, Lithosinae; incl. gen. Crocosia, Eilema, Lithosia, Pelosia, Phryganopsis a. o.). Haile Sellassie I University. Papers from the Faculty of science, Series C (Zoology), 1, 1 - 161." type="journal article" year="1965">Birket-Smith 1965</bibRefCitation>
); manica penis ventrally sclerotized at the base, joining the juxta; valva overall quite slender; basis valvae curved ventrally, touching each other; supravalva membranous, distally gently tapering, with costa almost straight, producing a very smooth, slightly swollen processus distalis plicae in the middle; ala valvae fairly sclerotized, with sacculus slender and distal process curved over the supravalva and ending in a strong pincer; aedeagus half as long as the ala valvae, tubular, slightly bent, with coecum penis shallow, ductus seminalis inserted dorsally; vesica with a flat sclerotization (cornutus) on the left side and five sturdy thorn-like cornuti varying in size, three of which are aligned at the junction with the aedeagus on the right, the smallest on the dorsal lobe and the largest on the ventral lobe (
<figureCitation id="133E10D8FFA1FFFE0F83F9BF3184F9AD" box="[667,767,1554,1580]" captionStart="FIGURE 11" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[211,1391,194,1860]" captionTargetId="figure-27@7.[196,1391,181,1867]" captionTargetPageId="7" captionText="FIGURE 11. Novilema bifurcum n. sp.—AD: male genitalia. A: genitalia, aedeagus in situ; B: Uncus, socius and terminal portion of the juxta, enlarged; C: aedeagus, vesica everted, enlarged; D: granicula of the vesica, enlarged. ae: aedeagus; av: ala valvae; cr: cornuti; gr: granicula; ju: juxta; pav: distal process of the ala valvae; so: socius; sv: supravalva; te: tegumen; un: uncus; tr: transtilla; ve: vesica; vi: vinculum; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848289" httpUri="https://zenodo.org/record/5848289/files/figure.png" pageId="8" pageNumber="424">Fig. 11C</figureCitation>
); finally, a scobination of small irregularly-shaped granicula on the distal surface between these last two cornuti (
<figureCitation id="133E10D8FFA1FFFE0FF9F99B3033F9D1" box="[737,840,1590,1616]" captionStart="FIGURE 11" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[211,1391,194,1860]" captionTargetId="figure-27@7.[196,1391,181,1867]" captionTargetPageId="7" captionText="FIGURE 11. Novilema bifurcum n. sp.—AD: male genitalia. A: genitalia, aedeagus in situ; B: Uncus, socius and terminal portion of the juxta, enlarged; C: aedeagus, vesica everted, enlarged; D: granicula of the vesica, enlarged. ae: aedeagus; av: ala valvae; cr: cornuti; gr: granicula; ju: juxta; pav: distal process of the ala valvae; so: socius; sv: supravalva; te: tegumen; un: uncus; tr: transtilla; ve: vesica; vi: vinculum; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848289" httpUri="https://zenodo.org/record/5848289/files/figure.png" pageId="8" pageNumber="424">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F5FD6FFA1FFFE0DDFF9F73063F919" pageId="8" pageNumber="424" type="etymology">
<paragraph id="8BBA0C5DFFA1FFFE0DDFF9F73063F919" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFF9F73235F9F5" bold="true" box="[199,334,1626,1652]" pageId="8" pageNumber="424">Etymology.</emphasis>
The species name, from the Latin “
<emphasis id="B971D04FFFA1FFFE0FC2F9F7304EF9F5" box="[730,821,1626,1652]" italics="true" pageId="8" pageNumber="424">bifurcus</emphasis>
”, means “forked” or “bifid”, and refers to the juxta and the distal process of the ala valvae, both ending in a fork.
</paragraph>
</subSubSection>
<subSubSection id="C31F5FD6FFA1FFFE0DDFF90F3292F815" pageId="8" pageNumber="424" type="discussion">
<paragraph id="8BBA0C5DFFA1FFFE0DDFF90F3292F815" blockId="8.[151,1437,474,1940]" pageId="8" pageNumber="424">
<emphasis id="B971D04FFFA1FFFE0DDFF90F324FF93D" bold="true" box="[199,308,1698,1724]" pageId="8" pageNumber="424">Remarks</emphasis>
. This Gabonese species is the second species included in the genus
<taxonomicName id="4C0577DEFFA1FFFE0920F90F37D8F93D" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[1080,1187,1698,1724]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B971D04FFFA1FFFE0920F90F37D8F93D" box="[1080,1187,1698,1724]" italics="true" pageId="8" pageNumber="424">Novilema</emphasis>
</taxonomicName>
, previously described as monotypic from
<collectingCountry id="F3124CCDFFA1FFFE0C6DF96B32B2F961" box="[373,457,1734,1760]" name="Nigeria" pageId="8" pageNumber="424">Nigeria</collectingCountry>
on the
<typeStatus id="54BEB2FFFFA1FFFE0F07F96A3134F960" box="[543,591,1735,1761]" pageId="8" pageNumber="424">type</typeStatus>
species
<taxonomicName id="4C0577DEFFA1FFFE0FA8F96B30F6F961" authorityName="Durante &amp; Panzera" authorityYear="2001" box="[688,909,1734,1760]" class="Insecta" family="Erebidae" genus="Novilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="424" phylum="Arthropoda" rank="species" species="fluminale">
<emphasis id="B971D04FFFA1FFFE0FA8F96B30F6F961" box="[688,909,1734,1760]" italics="true" pageId="8" pageNumber="424">Novilema fluminale</emphasis>
</taxonomicName>
. All the characters of the genus, as well as the supposed autapomorphies (i.e. the absence of valvellae and the long independent juxta; see
<bibRefCitation id="EF9471ACFFA1FFFE09D8F9463252F8A8" author="Durante, M. A. &amp; Panzera, S." pageId="8" pageNumber="424" pagination="465 - 472" refId="ref6987" refString="Durante, M. A. &amp; Panzera, S. (2001 b) I Lepidotteri del Delta del Niger (terzo contributo) (Lepidoptera, Arctiidae, Lithosiinae). Lambillionea, 101 (3), 465 - 472." type="journal article" year="2001">Durante &amp; Panzera (2001b: 466)</bibRefCitation>
and
<bibRefCitation id="EF9471ACFFA1FFFE0C46F8A2314EF8A8" author="Kruger, M." box="[350,565,1806,1833]" pageId="8" pageNumber="424" pagination="1 - 176" refId="ref7647" refString="Kruger, M. (2015) Generic classification of Afrotropical footman moths sensu stricto (Lepidoptera: Erebidae: Arctiinae: Lithosiini (partim )). Transvaal Museum Monograph, 15, 1 - 176." type="journal article" year="2015">Krüger (2015: 46))</bibRefCitation>
, are clearly maintained: the generally slender valva; the bifurcated apex of the ala valvae process; the large uncus in lateral view; the valvae firmly juxtaposed to each other at the base; branches of the vinculum ventrally approaching each other without forming a saccus; the tubular aedeagus and the vesica with sturdy thorn-like cornuti.
</paragraph>
</subSubSection>
</treatment>
</document>