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<document ID-DOI="http://doi.org/10.5281/zenodo.3792206" ID-GBIF-Dataset="6bed073a-175c-42e6-bb99-645c99d55b4f" ID-GBIF-Taxon="163971058" ID-ISSN="13132970" ID-Zenodo-Dep="3792206" ID-ZooBank="C631A347-306E-4773-84A4-E4712329186B" LSID="urn:lsid:zoobank.org:act:04A2A54B-80C1-4672-BEEA-4C9886BBF3FF" checkinTime="1587984202003" checkinUser="plazi" docAuthor="Miller, Jeremy, Griswold, Charles &amp; Yin, Chang" docDate="2009" docId="03ACB915FF8E7406FF4E463BFE9FFEC9" docLanguage="en" docName="ZK_article_1993.pdf" docOrigin="ZooKeys 11 (11)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Theridiosoma shuangbi Miller, Griswold &amp; Yin 2009, sp. n." docType="treatment" docUuid="04A2A54B-80C1-4672-BEEA-4C9886BBF3FF" docUuidSource="ZooBank" docVersion="5" lastPageId="18" lastPageNumber="27" masterDocId="FF95C16DFF9F7414FFC2455DFFF1FFB8" masterDocTitle="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers" masterLastPageNumber="195" masterPageNumber="9" pageId="17" pageNumber="26" updateTime="1635491590422" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Griswold, Charles</mods:namePart>
<mods:affiliation>California Academy of Sciences ,, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Chang</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University ,, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-05-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>11</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>9</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.11.160</mods:identifier>
<mods:identifier type="GBIF-Dataset">6bed073a-175c-42e6-bb99-645c99d55b4f</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576459</mods:identifier>
<mods:identifier type="ZooBank">C631A347-306E-4773-84A4-E4712329186B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792206" ID-GBIF-Taxon="163971058" ID-Zenodo-Dep="3792206" LSID="urn:lsid:zoobank.org:act:04A2A54B-80C1-4672-BEEA-4C9886BBF3FF" httpUri="http://treatment.plazi.org/id/03ACB915FF8E7406FF4E463BFE9FFEC9" lastPageId="18" lastPageNumber="27" pageId="17" pageNumber="26">
<subSubSection box="[140,778,870,897]" pageId="17" pageNumber="26" type="nomenclature">
<paragraph blockId="17.[140,778,870,897]" box="[140,778,870,897]" pageId="17" pageNumber="26">
<heading box="[140,778,870,897]" level="2" pageId="17" pageNumber="26">
<emphasis bold="true" box="[140,778,870,897]" pageId="17" pageNumber="26">
<taxonomicName authority="Miller, Griswold &amp; Yin" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[140,701,870,897]" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="26" phylum="Arthropoda" rank="species" species="shuangbi" status="sp. n.">
<emphasis bold="true" box="[140,416,870,897]" italics="true" pageId="17" pageNumber="26">
<emphasis box="[140,302,870,897]" inLineHeading="true" pageId="17" pageNumber="26" reason="1">Theridiosoma</emphasis>
shuangbi
</emphasis>
Miller, Griswold &amp; Yin
</taxonomicName>
,
<taxonomicNameLabel box="[713,778,871,897]" pageId="17" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="26" type="description">
<paragraph blockId="17.[140,939,906,968]" box="[140,939,906,933]" pageId="17" pageNumber="26">
<uri box="[140,939,906,933]" pageId="17" pageNumber="26">
urn:lsid:zoobank.org:act:
<uuid box="[418,939,906,933]" pageId="17" pageNumber="26">624239B5-E929-4492-872E-25575391B796</uuid>
</uri>
</paragraph>
<paragraph blockId="17.[140,939,906,968]" box="[140,293,941,968]" pageId="17" pageNumber="26">
<figureCitation box="[140,206,941,967]" captionStart="Figure 3" captionStartId="75.[140,212,957,979]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@75.[140,1108,164,937]" captionTargetPageId="75" captionText="Figure 3. A, B, Epeirotypus dalong sp. n. from Fugong Co.; C, D, Ogulnius barbandrewsi sp. n.; C from Guocai He at Fucai; D from No. 12 Bridge Camp area; E, Baalzebub nemesis sp. n. from No. 12 Bridge Camp area; F, G, Theridiosoma diwang sp. n. from Xiao Hei Shan Nature Reserve; H, I, Theridiosoma shuangbi sp. n. from Xiao Hei Shan Nature Reserve. A, C, E, F, H, epigynum, ventral view; B, epigynum, posterior view; D, G, I, cleared vulva, dorsal view. CD: copulatory duct; S: spermatheca; arrow in A indicates transverse submarginal groove; in G and I indicate lateral pits, in H indicates base of one of two opposing lateral apophyses." figureDoi="http://doi.org/10.5281/zenodo.3768781" httpUri="https://zenodo.org/record/3768781/files/figure.png" pageId="17" pageNumber="26">Figs 3</figureCitation>
H-I,
<figureCitation box="[264,293,941,968]" captionStart="Figure 95" captionStartId="166.[140,212,1410,1432]" captionTargetBox="[140,1108,165,1390]" captionTargetId="figure@166.[140,1109,164,1390]" captionTargetPageId="166" captionText="Figure 95. Distribution map, inset map shows study area. Blue: Baalzebub nemesis sp. n.; red: Theridiosoma shuangbi sp. n.; purple: Mysmena jinlong sp. n.; yellow: Crassignatha longtou sp. n." figureDoi="http://doi.org/10.5281/zenodo.3768922" httpUri="https://zenodo.org/record/3768922/files/figure.png" pageId="17" pageNumber="26">95</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="26" type="materials_examined">
<paragraph blockId="17.[140,1108,1011,1636]" pageId="17" pageNumber="26">
<emphasis bold="true" box="[140,379,1011,1037]" pageId="17" pageNumber="26">
<emphasis box="[140,241,1011,1037]" inLineHeading="true" pageId="17" pageNumber="26" reason="1">Material</emphasis>
Examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2610636303" collectingDate="2005-05-26" collectionCode="HNU" country="China" county="Long" elevation="2010" latitude="24.82886" location="Xiao" longLatPrecision="1" longitude="98.75917" municipality="Longjiang Township" pageId="17" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus box="[388,499,1012,1038]" pageId="17" pageNumber="26">Holotype</typeStatus>
:
<collectingCountry box="[513,612,1012,1038]" name="China" pageId="17" pageNumber="26">CHINA</collectingCountry>
:
<collectingRegion box="[625,717,1012,1038]" country="China" name="Yunnan" pageId="17" pageNumber="26">Yunnan</collectingRegion>
:
<collectingCounty box="[730,789,1012,1038]" pageId="17" pageNumber="26">Long</collectingCounty>
ling Co.,
<collectingMunicipality pageId="17" pageNumber="26">Longjiang Township</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03ACB915FF8E7406FF4E463BFE9FFEC9:8EDA5ED8FF8E7405FF0A414AFF0CFB89" box="[200,253,1047,1073]" country="China" county="Long" latitude="24.82886" longLatPrecision="1" longitude="98.75917" municipality="Longjiang Township" name="Xiao" pageId="17" pageNumber="26" stateProvince="Yunnan">Xiao</location>
hei shan
<location LSID="urn:lsid:plazi:treatment:03ACB915FF8E7406FF4E463BFE9FFEC9:8EDA5ED8FF8E7405FEA6414AFDFEFB89" box="[356,527,1047,1073]" country="China" county="Long" latitude="24.82886" longLatPrecision="1" longitude="98.75917" municipality="Longjiang Township" name="Nature Reserve" pageId="17" pageNumber="26" stateProvince="Yunnan">Nature Reserve</location>
(Gu Cheng Shan Mtn.),
<geoCoordinate box="[809,954,1047,1073]" degrees="24.82886" direction="north" orientation="latitude" pageId="17" pageNumber="26" precision="1" value="24.82886">24.82886°N</geoCoordinate>
,
<geoCoordinate box="[964,1103,1047,1074]" degrees="98.75917" direction="east" orientation="longitude" pageId="17" pageNumber="26" precision="1" value="98.75917">98.75917°E</geoCoordinate>
,
<quantity box="[140,234,1082,1109]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="17" pageNumber="26" unit="m" value="2010.0">
<elevation box="[140,234,1082,1109]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="17" pageNumber="26" unit="m" value="2010.0">2010 m</elevation>
</quantity>
,
<date box="[248,409,1082,1109]" pageId="17" pageNumber="26" value="2005-05-26">
<collectingDate box="[248,409,1082,1109]" pageId="17" pageNumber="26" value="2005-05-26">26 May 2005</collectingDate>
</date>
, in the forest, Yan Heng-mei, GKJ026 (CASENT 9022033,
<collectionCode box="[140,212,1118,1144]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="17" pageNumber="26">HNU</collectionCode>
),
<specimenCount box="[232,269,1117,1144]" pageId="17" pageNumber="26" type="female">1 ♀</specimenCount>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="26" type="etymology">
<paragraph blockId="17.[140,1108,1011,1636]" pageId="17" pageNumber="26">
<emphasis bold="true" box="[185,320,1152,1178]" pageId="17" pageNumber="26">
<emphasis box="[185,315,1152,1178]" inLineHeading="true" pageId="17" pageNumber="26" reason="1">Etymology</emphasis>
.
</emphasis>
Formed from the Chinese words for pair (shuāng) and arm (bì), referring to the paired processes on the epigynum.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="26" type="diagnosis">
<paragraph blockId="17.[140,1108,1011,1636]" pageId="17" pageNumber="26">
<emphasis bold="true" box="[185,312,1222,1248]" pageId="17" pageNumber="26">
<emphasis box="[185,307,1222,1248]" inLineHeading="true" pageId="17" pageNumber="26" reason="1">Diagnosis</emphasis>
.
</emphasis>
Distinguished from
<taxonomicName authorityName="Bosenberg &amp; Strand" authorityYear="1906" box="[557,693,1223,1249]" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="26" phylum="Arthropoda" rank="species" species="epeiroides">
<emphasis box="[557,693,1223,1249]" italics="true" pageId="17" pageNumber="26">T. epeiroides</emphasis>
</taxonomicName>
(the only other Asian
<taxonomicName box="[962,1107,1223,1249]" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis box="[962,1107,1223,1249]" italics="true" pageId="17" pageNumber="26">Theridiosoma</emphasis>
</taxonomicName>
species with lateral processes of the epigynum) by the shape and orientation of the apophyses, directly towards each other in
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[618,746,1293,1319]" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="26" phylum="Arthropoda" rank="species" species="shuangbi">
<emphasis box="[618,746,1293,1319]" italics="true" pageId="17" pageNumber="26">T. shuangbi</emphasis>
</taxonomicName>
(
<figureCitation box="[762,856,1293,1319]" captionStart="Figure 3" captionStartId="75.[140,212,957,979]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@75.[140,1108,164,937]" captionTargetPageId="75" captionText="Figure 3. A, B, Epeirotypus dalong sp. n. from Fugong Co.; C, D, Ogulnius barbandrewsi sp. n.; C from Guocai He at Fucai; D from No. 12 Bridge Camp area; E, Baalzebub nemesis sp. n. from No. 12 Bridge Camp area; F, G, Theridiosoma diwang sp. n. from Xiao Hei Shan Nature Reserve; H, I, Theridiosoma shuangbi sp. n. from Xiao Hei Shan Nature Reserve. A, C, E, F, H, epigynum, ventral view; B, epigynum, posterior view; D, G, I, cleared vulva, dorsal view. CD: copulatory duct; S: spermatheca; arrow in A indicates transverse submarginal groove; in G and I indicate lateral pits, in H indicates base of one of two opposing lateral apophyses." figureDoi="http://doi.org/10.5281/zenodo.3768781" httpUri="https://zenodo.org/record/3768781/files/figure.png" pageId="17" pageNumber="26">Fig. 3H</figureCitation>
), posteromesally in
<taxonomicName authorityName="Bosenberg &amp; Strand" authorityYear="1906" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="26" phylum="Arthropoda" rank="species" species="epeiroides">
<emphasis italics="true" pageId="17" pageNumber="26">T. epeiroides</emphasis>
</taxonomicName>
(
<bibRefCitation author="Brignoli PM" box="[258,429,1328,1355]" pageId="17" pageNumber="26" pagination="9 - 19" refId="ref45186" refString="Brignoli PM (1981) Spiders from the Philippines IV. A new Ogulnius and notes on some other oriental and Japanese Theridiosomatidae (Araneae). Acta arachnologica 30: 9 - 19." type="journal article" year="1981">Brignoli, 1981</bibRefCitation>
: figs 9, 10).
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="27" pageId="17" pageNumber="26" type="description">
<paragraph blockId="17.[140,1108,1011,1636]" pageId="17" pageNumber="26">
<emphasis bold="true" box="[185,335,1363,1389]" pageId="17" pageNumber="26">
<emphasis box="[185,330,1363,1389]" inLineHeading="true" pageId="17" pageNumber="26" reason="1">Description</emphasis>
.
</emphasis>
Carapace dusky pale yellow, darker centrally and through head region. Sternum dusky pale yellow with darker margin. Legs orange. Abdomen off white with pair of medium gray stripes running posteriorly from anterodorsal part of abdomen, converging to a single stripe running to spinneret region.
</paragraph>
<paragraph blockId="17.[140,1108,1011,1636]" pageId="17" pageNumber="26">
<taxonomicName authority=": Epigynum" authorityName="Epigynum" box="[185,383,1504,1530]" class="Polychaeta" family="Polychaeta (awaiting allocation)" genus="Vulva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polychaeta (awaiting allocation)" pageId="17" pageNumber="26" phylum="Annelida" rank="genus">Vulva: Epigynum</taxonomicName>
a flat plate with small median process, with pair of oblong lateral pits, lateral processes arise from near outside margin, run towards each other on longitudinal axis, tips rounded (
<figureCitation box="[513,609,1575,1601]" captionStart="Figure 3" captionStartId="75.[140,212,957,979]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@75.[140,1108,164,937]" captionTargetPageId="75" captionText="Figure 3. A, B, Epeirotypus dalong sp. n. from Fugong Co.; C, D, Ogulnius barbandrewsi sp. n.; C from Guocai He at Fucai; D from No. 12 Bridge Camp area; E, Baalzebub nemesis sp. n. from No. 12 Bridge Camp area; F, G, Theridiosoma diwang sp. n. from Xiao Hei Shan Nature Reserve; H, I, Theridiosoma shuangbi sp. n. from Xiao Hei Shan Nature Reserve. A, C, E, F, H, epigynum, ventral view; B, epigynum, posterior view; D, G, I, cleared vulva, dorsal view. CD: copulatory duct; S: spermatheca; arrow in A indicates transverse submarginal groove; in G and I indicate lateral pits, in H indicates base of one of two opposing lateral apophyses." figureDoi="http://doi.org/10.5281/zenodo.3768781" httpUri="https://zenodo.org/record/3768781/files/figure.png" pageId="17" pageNumber="26">Fig. 3H</figureCitation>
). Spermathecae and copulatory ducts both touching medially (
<figureCitation box="[360,439,1610,1636]" captionStart="Figure 3" captionStartId="75.[140,212,957,979]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@75.[140,1108,164,937]" captionTargetPageId="75" captionText="Figure 3. A, B, Epeirotypus dalong sp. n. from Fugong Co.; C, D, Ogulnius barbandrewsi sp. n.; C from Guocai He at Fucai; D from No. 12 Bridge Camp area; E, Baalzebub nemesis sp. n. from No. 12 Bridge Camp area; F, G, Theridiosoma diwang sp. n. from Xiao Hei Shan Nature Reserve; H, I, Theridiosoma shuangbi sp. n. from Xiao Hei Shan Nature Reserve. A, C, E, F, H, epigynum, ventral view; B, epigynum, posterior view; D, G, I, cleared vulva, dorsal view. CD: copulatory duct; S: spermatheca; arrow in A indicates transverse submarginal groove; in G and I indicate lateral pits, in H indicates base of one of two opposing lateral apophyses." figureDoi="http://doi.org/10.5281/zenodo.3768781" httpUri="https://zenodo.org/record/3768781/files/figure.png" pageId="17" pageNumber="26">Fig. 3I</figureCitation>
).
</paragraph>
<paragraph blockId="18.[140,1108,167,369]" pageId="18" pageNumber="27">
Female (
<typeStatus box="[280,385,167,194]" pageId="18" pageNumber="27">holotype</typeStatus>
): Total length 2.00, carapace 0.72 long, 0.61 wide, clypeus 0.14, sternum 0.39 long, 0.38 wide, coxa IV separated by 1.41 times their width. Posterior median eyes nearly juxtaposed. Macrosetae: Leg I: patella d1, tibia d2, p1, r1; Leg II: patella d1, tibia d2, r1; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.22; TmII: 0.25; TmIII: 0.21. Leg measurements: see Appendix A.
</paragraph>
<paragraph blockId="18.[140,1108,167,369]" box="[185,366,343,369]" pageId="18" pageNumber="27">Male unknown.</paragraph>
</subSubSection>
</treatment>
</document>