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<mods:title id="6F2F9180779969115843022709F4314E">Taxonomic notes on the genus Tineovertex Moriuti (Insecta, Lepidoptera, Tineidae) with description of a new species</mods:title>
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<treatment id="03EB87E6CE1B3629FF15F9CE0E4EFB8A" ID-DOI="http://doi.org/10.5281/zenodo.6244679" ID-GBIF-Taxon="119378625" ID-Zenodo-Dep="6244679" LSID="urn:lsid:plazi:treatment:03EB87E6CE1B3629FF15F9CE0E4EFB8A" httpUri="http://treatment.plazi.org/id/03EB87E6CE1B3629FF15F9CE0E4EFB8A" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
<subSubSection id="C358657BCE1B362AFF15F9CE0EA2F995" box="[151,471,1595,1621]" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="8BFD36F0CE1B362AFF15F9CE0EA2F995" blockId="1.[151,471,1595,1621]" box="[151,471,1595,1621]" pageId="1" pageNumber="38">
<heading id="D0B5819CCE1B362AFF15F9CE0EA2F995" bold="true" box="[151,471,1595,1621]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<taxonomicName id="4C424D73CE1B362AFF15F9CE0EA2F995" ID-CoL="7X3V" ID-ENA="1178372" authority="Moriuti, 1982" authorityName="Moriuti" authorityYear="1982" box="[151,471,1595,1621]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE1B362AFF15F9CE0EA2F995" bold="true" box="[151,471,1595,1621]" pageId="1" pageNumber="38">
<emphasis id="B936EAE2CE1B362AFF15F9CE0E51F995" bold="true" box="[151,292,1595,1621]" italics="true" pageId="1" pageNumber="38">Tineovertex</emphasis>
<bibRefCitation id="EFD34B01CE1B362AFEAEF9CE0EA2F995" author="Moriuti" box="[300,471,1595,1621]" pageId="1" pageNumber="46" refString="Moriuti, S. (1982) Tineidae. In: Inoue, H. et al. (Ed.). Moths of Japan. Two Volumes. 1: 1 - 966, 2: 1 - 552. Kodansha, Tokyo, pp. 162 - 171. (In Japanese)" type="journal article" year="1982">Moriuti, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C358657BCE1B362AFF15F9700B72F960" box="[151,1031,1669,1696]" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="8BFD36F0CE1B362AFF15F9700B72F960" blockId="1.[151,1436,1669,2016]" box="[151,1031,1669,1696]" pageId="1" pageNumber="38">
<emphasis id="B936EAE2CE1B362AFF15F9700E4EF95F" bold="true" box="[151,315,1669,1695]" pageId="1" pageNumber="38">
<typeStatus id="54F98852CE1B362AFF15F9700FA1F95F" box="[151,212,1669,1695]" pageId="1" pageNumber="38">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C424D73CE1B362AFEC1F9730D90F960" authority="Meyrick, 1911" authorityName="Meyrick" authorityYear="1911" box="[323,741,1670,1696]" class="Insecta" family="Tineidae" genus="Tinea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="melanochrysa">
<emphasis id="B936EAE2CE1B362AFEC1F9730D5AF95F" box="[323,559,1670,1695]" italics="true" pageId="1" pageNumber="38">Tinea melanochrysa</emphasis>
<bibRefCitation id="EFD34B01CE1B362AFDB5F9730D90F960" author="Meyrick" box="[567,741,1670,1696]" pageId="1" pageNumber="46" refString="Meyrick, E. (1911) Descriptions of Indian Micro-Lepidoptera. Journal of the Bombay natural history Society, 21, 104 - 131." type="journal article" year="1911">Meyrick, 1911</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C358657BCE1B362AFF45F9580CF3F880" pageId="1" pageNumber="38" type="diagnosis">
<paragraph id="8BFD36F0CE1B362AFF45F9580CF3F880" blockId="1.[151,1436,1669,2016]" pageId="1" pageNumber="38">
<emphasis id="B936EAE2CE1B362AFF45F9580E32F907" bold="true" box="[199,327,1709,1735]" pageId="1" pageNumber="38">Diagnosis.</emphasis>
Medium-sized species with lateral bristles on the labial palpi; wings elongate without upturned forewing tip. Male genitalia simple, with extremely narrow ring and separated bilobed uncus; gnathos absent; saccus long rod-shaped; valva simple with apex rounded. Female genitalia with corethrogyne absent; papilla analis strongly sclerotized to form piercing apex.
</paragraph>
</subSubSection>
<subSubSection id="C358657BCE1B3629FF45F8B80E5BFE0A" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="description">
<paragraph id="8BFD36F0CE1B362AFF45F8B80DC3F878" blockId="1.[151,1436,1669,2016]" pageId="1" pageNumber="38">
<emphasis id="B936EAE2CE1B362AFF45F8B80E2AF8A7" bold="true" box="[199,351,1869,1895]" pageId="1" pageNumber="38">Description.</emphasis>
Head: Vertex roughly clothed with black or white scales. Antenna filiform, about 1.0 × length of forewing; frons covered with dense scales; maxillary palpus moderately developed. Labial palpus with strong lateral bristles on second segment.
</paragraph>
<paragraph id="8BFD36F0CE1B3629FF45F8330BCCFF1A" blockId="1.[151,1436,1669,2016]" lastBlockId="2.[151,1437,152,1098]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
Thorax: Dorsum and tegula covered with black or white scales. Wings similarly patterned, moderately elongate, apices moderately rounded and not upturned. Wingspan
<quantity id="4CBA9B15CE183629FC1AFF6D0B6FFF72" box="[920,1050,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="39" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">1020 mm</quantity>
. Forewing with all veins free; R4 and R5 stalked or approximated basally; hindwing with all veins free and separate basally.
</paragraph>
<paragraph id="8BFD36F0CE183629FF45FF1D0E5BFE0A" blockId="2.[151,1437,152,1098]" pageId="2" pageNumber="39">Abdomen: Male genitalia with extremely narrow ring and a pair of separated uncus lobes; gnathos absent, juxta indistinct; saccus long, rod-shaped; valva simple and curved dorsally; aedeagus cylindrical, stout, nearly straight, vesica usually with cornuti. Female genitalia without corethrogyne; eighth abdominal tergum with a median keel; apophyses posteriors and anteriores long and stout; papilla analis strongly sclerotized, flattened, pointed to form piercing apex with minute serrations laterally; corpus bursae with or without a pair of symmetric signa.</paragraph>
</subSubSection>
<subSubSection id="C358657BCE183629FF45FE220E4EFB8A" pageId="2" pageNumber="39" type="discussion">
<paragraph id="8BFD36F0CE183629FF45FE220EECFD52" blockId="2.[151,1437,152,1098]" pageId="2" pageNumber="39">
<emphasis id="B936EAE2CE183629FF45FE220E35FE31" bold="true" box="[199,320,471,497]" pageId="2" pageNumber="39">Remarks.</emphasis>
All species of
<taxonomicName id="4C424D73CE183629FE70FE2D0D0CFE31" box="[498,633,472,497]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FE70FE2D0D0CFE31" box="[498,633,472,497]" italics="true" pageId="2" pageNumber="39">Tineovertex</emphasis>
</taxonomicName>
have a similar wing pattern. They can be separated into two groups by the color of the head and thorax. In
<taxonomicName id="4C424D73CE183629FDDEFDF50C68FDD9" box="[604,797,512,537]" class="Insecta" family="Tineidae" genus="Tinea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="melanochrysa">
<emphasis id="B936EAE2CE183629FDDEFDF50C68FDD9" box="[604,797,512,537]" italics="true" pageId="2" pageNumber="39">T. melanochrysa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C424D73CE183629FCA9FDF50CD1FDD9" box="[811,932,512,537]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="sartoria">
<emphasis id="B936EAE2CE183629FCA9FDF50CD1FDD9" box="[811,932,512,537]" italics="true" pageId="2" pageNumber="39">T. sartoria</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C424D73CE183629FC49FDF50B3FFDD9" box="[971,1098,512,537]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="gladiata">
<emphasis id="B936EAE2CE183629FC49FDF50B3FFDD9" box="[971,1098,512,537]" italics="true" pageId="2" pageNumber="39">T. gladiata</emphasis>
</taxonomicName>
the head and thorax are covered with blackish scales; in
<emphasis id="B936EAE2CE183629FE6DFDDD0C60FD81" box="[495,789,552,577]" italics="true" pageId="2" pageNumber="39">
<taxonomicName id="4C424D73CE183629FE6DFDDD0DF7FD81" box="[495,642,552,577]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="canicoma">T. canicoma</taxonomicName>
,
<taxonomicName id="4C424D73CE183629FD0DFDDD0C60FD81" box="[655,789,552,577]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="melliflua">T. melliflua</taxonomicName>
</emphasis>
&amp;
<taxonomicName id="4C424D73CE183629FCBFFDDD0CA1FD81" box="[829,980,552,577]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="antidroma">
<emphasis id="B936EAE2CE183629FCBFFDDD0CA1FD81" box="[829,980,552,577]" italics="true" pageId="2" pageNumber="39">T. antidroma</emphasis>
</taxonomicName>
they are covered with white or ochreous-yellow scales. However, within each species group, it is difficult to distinguish the species on the basis of superficial characters.
</paragraph>
<paragraph id="8BFD36F0CE183629FF45FD550E4EFB8A" blockId="2.[151,1437,152,1098]" pageId="2" pageNumber="39">
Female genitalia of the genus
<taxonomicName id="4C424D73CE183629FDA5FD550DC5FD79" box="[551,688,672,697]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FDA5FD550DC5FD79" box="[551,688,672,697]" italics="true" pageId="2" pageNumber="39">Tineovertex</emphasis>
</taxonomicName>
are characteristic in having long apophyses, the ovipositor with a serrate piercing apex, and the eighth abdominal tergum with a median keel. These features are similar to those of the tineid genus
<taxonomicName id="4C424D73CE183629FE40FD050B57FCCA" authority="Sauber, 1902 (Davis &amp; Heppner, 1987)" authorityName="Sauber, 1902 (Davis &amp; Heppner" authorityYear="1987" box="[450,1058,752,778]" class="Insecta" family="Tineidae" genus="Ischnuridia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FE40FD050D3CFCC9" box="[450,585,752,777]" italics="true" pageId="2" pageNumber="39">Ischnuridia</emphasis>
Sauber, 1902 (
<bibRefCitation id="EFD34B01CE183629FD7FFD050B6FFCCA" author="Davis" box="[765,1050,752,778]" pageId="2" pageNumber="46" refString="Davis, D. R. &amp; Heppner, J. B. (1987) New discoveries concerning Ischnuridia, a remarkable genus of Indo-Australian Tineidae (Lepidoptera). Tinea, 12 (Supplement), 145 - 151." type="journal article" year="1987">Davis &amp; Heppner, 1987</bibRefCitation>
)
</taxonomicName>
and most incurvarioid families.
<bibRefCitation id="EFD34B01CE183629FF15FCED0EAAFCF2" author="Robinson" box="[151,479,792,818]" pageId="2" pageNumber="46" refString="Robinson, G. S. &amp; Nielsen E. S. (1993) Tineid Genera of Australia (Lepidoptera). Monographs on Australian Lepidoptera 2, i - xvi, 1 - 344. CSIRO Publications, Melborune, Australia." type="journal article" year="1993">Robinson &amp; Nielsen (1993)</bibRefCitation>
noted that the ovipositor of the genus
<taxonomicName id="4C424D73CE183629FC2BFCED0A7AFCF2" authority="Diakonoff, 1955" authorityName="Diakonoff" authorityYear="1955" box="[937,1295,792,818]" class="Insecta" family="Tineidae" genus="Ectropoceros" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FC2BFCED0B30FCF1" box="[937,1093,792,817]" italics="true" pageId="2" pageNumber="39">Ectropoceros</emphasis>
Diakonoff, 1955
</taxonomicName>
, which was considered a close-relative of
<emphasis id="B936EAE2CE183629FE79FCB50DFEFC99" box="[507,651,832,857]" italics="true" pageId="2" pageNumber="39">
<taxonomicName id="4C424D73CE183629FE79FCB50DF2FC99" box="[507,647,832,857]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">Tineovertex</taxonomicName>
,
</emphasis>
greatly resembles that of certain
<taxonomicName id="4C424D73CE183629FB98FCB50BCFFC9A" box="[1050,1210,832,858]" class="Insecta" family="Incurvariidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="family">Incurvariidae</taxonomicName>
. They inferred that
<taxonomicName id="4C424D73CE183629FF15FC9D0E46FC41" box="[151,307,872,897]" class="Insecta" family="Tineidae" genus="Ectropoceros" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FF15FC9D0E46FC41" box="[151,307,872,897]" italics="true" pageId="2" pageNumber="39">Ectropoceros</emphasis>
</taxonomicName>
represents the sister-group of
<emphasis id="B936EAE2CE183629FD1EFC9D0CADFC41" box="[668,984,872,897]" italics="true" pageId="2" pageNumber="39">
<taxonomicName id="4C424D73CE183629FD1EFC9D0C50FC41" box="[668,805,872,897]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">Tineovertex</taxonomicName>
+
<taxonomicName id="4C424D73CE183629FCC8FC9D0CA4FC41" box="[842,977,872,897]" class="Insecta" family="Tineidae" genus="Ischnuridia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">Ischnuridia</taxonomicName>
.
</emphasis>
According to T. Saito (personal communication) and
<bibRefCitation id="EFD34B01CE183629FEDDFC650E8CFC6A" author="Sakai" box="[351,505,912,938]" pageId="2" pageNumber="46" refString="Sakai, M. (2002) Systematic study of the family Tineidae of Japan (Insecta: Lepidoptera). Unpublished Ph. D. dissertation, Kyushu University, Japan." type="book" year="2002">Sakai (2002)</bibRefCitation>
, in
<collectingCountry id="F3557660CE183629FDA6FC650D12FC6A" box="[548,615,912,938]" name="Japan" pageId="2" pageNumber="39">Japan</collectingCountry>
the larvae of
<taxonomicName id="4C424D73CE183629FC89FC650CB8FC69" box="[779,973,912,937]" class="Insecta" family="Tineidae" genus="Tinea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="melanochrysa">
<emphasis id="B936EAE2CE183629FC89FC650CB8FC69" box="[779,973,912,937]" italics="true" pageId="2" pageNumber="39">T. melanochrysa</emphasis>
</taxonomicName>
feed on dead leaves on the ground and construct a flat-oblong portable case. Based on these observations, the first instar larvae of
<taxonomicName id="4C424D73CE183629FB47FC4D0A3BFC11" box="[1221,1358,952,977]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FB47FC4D0A3BFC11" box="[1221,1358,952,977]" italics="true" pageId="2" pageNumber="39">Tineovertex</emphasis>
</taxonomicName>
possibly may feed on or mine into living plant tissue. Later instars may feed on dead leaves and make a portable case. The larval feeding habits of
<taxonomicName id="4C424D73CE183629FDBAFBFD0DB4FBE1" box="[568,705,1032,1057]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAE2CE183629FDBAFBFD0DB4FBE1" box="[568,705,1032,1057]" italics="true" pageId="2" pageNumber="39">Tineovertex</emphasis>
</taxonomicName>
may be similar to those of
<taxonomicName id="4C424D73CE183629FB9AFBFD0BF7FBE2" box="[1048,1154,1032,1058]" class="Insecta" family="Adelidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="family">Adelidae</taxonomicName>
rather than to those of
<taxonomicName id="4C424D73CE183629FF15FBC50E42FB8A" box="[151,311,1072,1098]" class="Insecta" family="Incurvariidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="family">Incurvariidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>