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<document id="1EF9558CE890D87232FAB23D6F65300A" ID-CLB-Dataset="20050" ID-DOI="10.11646/zootaxa.5120.4.5" ID-GBIF-Dataset="28a09134-c557-4b45-b524-02fde7feb24e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6392851" ID-ZooBank="A9208B48-E265-4E87-BE79-F0CD49B36D6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648537908093" checkinUser="plazi" docAuthor="Wu, San-An, Xu, Han &amp; Zheng, Xinyi" docDate="2022" docId="03EB87D4FFC7FFCEA9A11C1D866FF993" docLanguage="en" docName="zootaxa.5120.4.5.pdf" docOrigin="Zootaxa 5120 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Qinococcidae Wu" docType="treatment" docVersion="7" lastPageNumber="545" masterDocId="FFD2FFACFFC6FFCCA9361B038218FF9A" masterDocTitle="A new coccoid family (Hemiptera: Coccomorpha) for an unusual species of scale insect on Podocarpus macrophyllus (Podocarpaceae) from southern China" masterLastPageNumber="558" masterPageNumber="543" pageNumber="544" updateTime="1699318806928" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="422861D3275258E23C0D0065D9CF4E2D">A new coccoid family (Hemiptera: Coccomorpha) for an unusual species of scale insect on Podocarpus macrophyllus (Podocarpaceae) from southern China</mods:title>
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<mods:namePart id="F29A4443AB54B22C523A79AE2912EA68">Wu, San-An</mods:namePart>
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<mods:namePart id="9EA857F245A1F38DBA3BDA26DD64E7AB">Xu, Han</mods:namePart>
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<mods:affiliation id="A58C59D044A31544A11561A3992894C0">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China sananwu @ bjfu. edu. cn; https: // orcid. org / 0000 - 0002 - 9671 - 9401 &amp; xuhan @ bjfu. edu. cn; https: // orcid. org / 0000 - 0001 - 7226 - 1742</mods:affiliation>
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<mods:namePart id="F3C92E3873712DCCF00BB382DC105E0E">Zheng, Xinyi</mods:namePart>
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<mods:affiliation id="0246B03CEB0BD3A39C60D6A3F873E88F">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China sananwu @ bjfu. edu. cn; https: // orcid. org / 0000 - 0002 - 9671 - 9401 &amp; zxy 780346931 @ 126. com; https: // orcid. org / 0000 - 0002 - 7396 - 7488</mods:affiliation>
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<subSubSection id="C3586549FFC7FFCDA9A11C1D8007F8A2" box="[151,543,1822,1848]" pageId="1" pageNumber="544" type="nomenclature">
<paragraph id="8BFD36C2FFC7FFCDA9A11C1D8007F8A2" blockId="1.[151,543,1822,1848]" box="[151,543,1822,1848]" pageId="1" pageNumber="544">
<heading id="D0B581AEFFC7FFCDA9A11C1D8007F8A2" bold="true" box="[151,543,1822,1848]" fontSize="11" level="1" pageId="1" pageNumber="544" reason="1">
<emphasis id="B936EAD0FFC7FFCDA9A11C1D8007F8A2" bold="true" box="[151,543,1822,1848]" pageId="1" pageNumber="544">
<taxonomicName id="4C424D41FFC7FFCDA9A11C1D83A3F8A2" authority="Wu &amp; Xu &amp; Zheng, 2022" box="[151,443,1822,1848]" family="Qinococcidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="545" rank="family">QINOCOCCIDAE Wu</taxonomicName>
, fam. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586549FFC7FFCDA9A11C608015F8E1" box="[151,525,1891,1915]" pageId="1" pageNumber="544" type="materials_examined">
<paragraph id="8BFD36C2FFC7FFCDA9A11C608015F8E1" blockId="1.[151,525,1891,1915]" box="[151,525,1891,1915]" pageId="1" pageNumber="544">
<typeStatus id="54F98860FFC7FFCDA9A11C6082D1F8E1" box="[151,201,1891,1915]" pageId="1" pageNumber="544">Type</typeStatus>
genus:
<taxonomicName id="4C424D41FFC7FFCDA82E1C6083A4F8E0" authority="Wu" authorityName="Wu" authorityYear="2022" box="[280,444,1891,1914]" class="Insecta" family="Margarodidae" genus="Qinococcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="544" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936EAD0FFC7FFCDA82E1C608388F8E0" box="[280,400,1891,1914]" italics="true" pageId="1" pageNumber="544">Qinococcus</emphasis>
Wu
</taxonomicName>
,
<emphasis id="B936EAD0FFC7FFCDA8F01C608015F8E1" bold="true" box="[454,525,1891,1915]" pageId="1" pageNumber="544">
<taxonomicNameLabel id="A20557ABFFC7FFCDA8F01C608015F8E1" box="[454,525,1891,1915]" pageId="1" pageNumber="544" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3586549FFC7FFCEA9A11CAA8627FE4B" lastPageId="2" lastPageNumber="545" pageId="1" pageNumber="544" type="diagnosis">
<paragraph id="8BFD36C2FFC7FFCEA9A11CAA80A8FF63" blockId="1.[151,1436,1961,2023]" lastBlockId="2.[151,1437,150,1545]" lastPageId="2" lastPageNumber="545" pageId="1" pageNumber="544">
<emphasis id="B936EAD0FFC7FFCDA9A11CAA8378F859" bold="true" box="[151,352,1961,1987]" pageId="1" pageNumber="544">Family diagnosis</emphasis>
.
<emphasis id="B936EAD0FFC7FFCDA85B1CAA801FF859" bold="true" box="[365,519,1961,1987]" pageId="1" pageNumber="544">Adult female</emphasis>
. Body tadpole-shaped, with head and thorax wide and abdomen narrow. Antennae much reduced, plate-like. Mouthparts developed. Legs much reduced. Thoracic spiracles with pores in atrium; abdominal spiracles numbering 5 pairs, also each with pores in atrium. Anal ring simple. Venter of abdominal segment I developed. Vulva opening longitudinally. Pores of several
<typeStatus id="54F98860FFC4FFCEAAA51BB881D6FF4F" box="[915,974,187,213]" pageId="2" pageNumber="545">types</typeStatus>
present, including large simple pores and multilocular pores, each with 26 central loculi.
</paragraph>
<paragraph id="8BFD36C2FFC4FFCEA9F11A018627FE4B" blockId="2.[151,1437,150,1545]" pageId="2" pageNumber="545">
<emphasis id="B936EAD0FFC4FFCEA9F11A018356FE87" bold="true" box="[199,334,258,285]" pageId="2" pageNumber="545">Adult male</emphasis>
. Xylococcid-like; antennae each filiform, long, and 10-segmented. A pair of compound eyes and two dorsal ocelli present. Legs well developed, with tarsi each 2-segmented; each claw with 2 denticles. Abdomen with large tubular ducts on tergites VI and VII. Head with a box-like arrangement of ventral ridges. Prothorax dorsally with strong pronotal ridge extending uninterrupted across hind part of neck, and an inverted “
<emphasis id="B936EAD0FFC4FFCEAC381A6C8705FE13" bold="true" box="[1294,1309,367,393]" pageId="2" pageNumber="545">π</emphasis>
<emphasis id="B936EAD0FFC4FFCEAC1C1A6C872BFE13" bold="true" box="[1322,1331,367,393]" pageId="2" pageNumber="545">-</emphasis>
like posttergite. Fore wing large, with a pterostigma, anterior flexing patch branched, and a clear line present posterior to cubital ridge. Hind wing reduced, lobe-like, with apex bilobate. Endophalus absent.
</paragraph>
</subSubSection>
<subSubSection id="C3586549FFC4FFCEA9F11AD981D6FE6F" box="[199,974,474,501]" pageId="2" pageNumber="545" type="description">
<paragraph id="8BFD36C2FFC4FFCEA9F11AD981D6FE6F" blockId="2.[151,1437,150,1545]" box="[199,974,474,501]" pageId="2" pageNumber="545">
<emphasis id="B936EAD0FFC4FFCEA9F11AD983B0FE6F" bold="true" box="[199,424,474,501]" pageId="2" pageNumber="545">First instar nymph</emphasis>
. Antennae each 7-segmented. Cicatrices present.
</paragraph>
</subSubSection>
<subSubSection id="C3586549FFC4FFCEA9F11AFD8746FD83" box="[199,1374,510,537]" pageId="2" pageNumber="545" type="biology_ecology">
<paragraph id="8BFD36C2FFC4FFCEA9F11AFD8746FD83" blockId="2.[151,1437,150,1545]" box="[199,1374,510,537]" pageId="2" pageNumber="545">
<emphasis id="B936EAD0FFC4FFCEA9F11AFD8339FD82" bold="true" box="[199,289,510,536]" pageId="2" pageNumber="545">Biology</emphasis>
. Living under the bark of the trunk and large branches. The life history includes cyst-like stages.
</paragraph>
</subSubSection>
<subSubSection id="C3586549FFC4FFCEA9F11920866FF993" pageId="2" pageNumber="545" type="discussion">
<paragraph id="8BFD36C2FFC4FFCEA9F1192083C4FD33" blockId="2.[151,1437,150,1545]" pageId="2" pageNumber="545">
<emphasis id="B936EAD0FFC4FFCEA9F119208098FDA6" bold="true" box="[199,640,546,573]" pageId="2" pageNumber="545">Remarks on taxonomic relationships</emphasis>
. This new family was established mainly based on the structure of the adult male of
<taxonomicName id="4C424D41FFC4FFCEA8191944801DFDFB" authority="Wu" authorityName="Wu &amp; Xu &amp; Zheng" authorityYear="2022" box="[303,517,583,609]" class="Insecta" family="Margarodidae" genus="Qinococcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="podocarpus" status="sp. nov.">
<emphasis id="B936EAD0FFC4FFCEA819194483CDFDFB" box="[303,469,583,609]" italics="true" pageId="2" pageNumber="545">Q. podocarpus</emphasis>
Wu
</taxonomicName>
,
<taxonomicNameLabel id="A20557ABFFC4FFCEAB2619448054FDFB" box="[528,588,583,609]" pageId="2" pageNumber="545" rank="species">sp. n.</taxonomicNameLabel>
, which possesses: 1) prothorax with an inverted “
<emphasis id="B936EAD0FFC4FFCEAD4519458682FDFA" bold="true" box="[1139,1178,582,608]" pageId="2" pageNumber="545">π”-</emphasis>
like post-tergite, and 2) fore wing with pterostigma and unusual venation, with the anterior flexing patch branched and a clear line present posterior to the cubital ridge.
</paragraph>
<paragraph id="8BFD36C2FFC4FFCEA9F119B0815EFC3F" blockId="2.[151,1437,150,1545]" pageId="2" pageNumber="545">
Among the adult males studied in the
<taxonomicName id="4C424D41FFC4FFCEAB4719B08109FD57" box="[625,785,691,717]" class="Insecta" infraOrder="Coccomorpha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="558" phylum="Arthropoda" rank="infraOrder">Coccomorpha</taxonomicName>
, the post-tergites are present in all groups (except perhaps the
<taxonomicName id="4C424D41FFC4FFCEA9F619D4836BFD68" box="[192,371,727,754]" class="Insecta" family="Stigmacoccidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">Stigmacoccidae</taxonomicName>
) in two morphological states: either (i) lying more-or-less longitudinally along the prothorax in the families
<taxonomicName id="4C424D41FFC4FFCEA81519F883BEFC8F" box="[291,422,763,789]" class="Insecta" family="Ortheziidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">Ortheziidae</taxonomicName>
and
<taxonomicName id="4C424D41FFC4FFCEA8E819F8812BFC8F" authority="sensu Morrison" authorityName="Morrison" box="[478,819,763,789]" class="Insecta" family="Margarodidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">
Margarodidae
<emphasis id="B936EAD0FFC4FFCEABB419F880D8FC8E" box="[642,704,763,788]" italics="true" pageId="2" pageNumber="545">sensu</emphasis>
Morrison
</taxonomicName>
, or (ii) transversely just anterior to the anterior margin of the scutum and parallel with each prealare of the mesothorax in the families
<taxonomicName id="4C424D41FFC4FFCEAD16181C86D8FCA3" authorityName="McKenzie" authorityYear="1942" box="[1056,1216,799,825]" class="Insecta" family="Pityococcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">Pityococcidae</taxonomicName>
, Phenacoleachidae, Steingellidae,
<taxonomicName id="4C424D41FFC4FFCEA80C18408384FCC7" authorityName="Beardsley" authorityYear="1969" box="[314,412,835,861]" class="Insecta" family="Putoidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">Putoidae</taxonomicName>
and most neococcoids (
<bibRefCitation id="EFD34B33FFC4FFCEAB85184081DCFCC7" author="Hodgson, C. J. &amp; Foldi, I." box="[691,964,835,861]" pageId="2" pageNumber="545" pagination="1 - 250" refId="ref11069" refString="Hodgson, C. J. &amp; Foldi, I. (2006) A review of the Margarodidae sensu Morrison (Hemiptera: Coccoidea) and some related taxa based on the morphology of adult males. Zootaxa, 1263 (1), 1 - 250. https: // doi. org / 10.11646 / zootaxa. 1263.1.1" type="journal article" year="2006">Hodgson &amp; Foldi 2006</bibRefCitation>
;
<bibRefCitation id="EFD34B33FFC4FFCEAAE418408665FCC7" author="Hodgson, C. J." box="[978,1149,835,861]" pageId="2" pageNumber="545" pagination="1 - 264" refId="ref11130" refString="Hodgson, C. J. (2020) A review of neococcid scale insects (Hemiptera: Sternorryncha: Coccomorpha) based on the morphology of the adult males. Zootaxa, 4765 (1), 1 - 264. https: // doi. org / 10.11646 / zootaxa. 4765.1.1" type="journal article" year="2020">Hodgson 2020</bibRefCitation>
). The post-tergites in the new family are well developed and complicated, showing an inverted “
<emphasis id="B936EAD0FFC4FFCEAA80186581C6FC1A" bold="true" box="[950,990,870,896]" pageId="2" pageNumber="545">π”-</emphasis>
like shape. Possibly this is the primitive condition, with the above two states being reductions from it.
</paragraph>
<paragraph id="8BFD36C2FFC4FFCEA9F118AC83C8FBC3" blockId="2.[151,1437,150,1545]" pageId="2" pageNumber="545">
The venation of the fore wing in adult male
<taxonomicName id="4C424D41FFC4FFCEAB8918AC815BFC53" authorityName="Wu" authorityYear="2022" box="[703,835,943,969]" class="Insecta" family="Margarodidae" genus="Qinococcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAD0FFC4FFCEAB8918AC815BFC53" box="[703,835,943,969]" italics="true" pageId="2" pageNumber="545">Qinococcus</emphasis>
</taxonomicName>
is the most complicated found in the described scale insects, with a pterostigma, the anterior flexing patch or light line between the subcostal ridge and cubital ridge forked, and the posterior flexing patches between the cubital ridge and posterior wing margin with two branches. The wings of adult males in other scale families have the anterior flexing patch unbranched and the posterior flexing patch with only one branch.
</paragraph>
<paragraph id="8BFD36C2FFC4FFCEA9F11F608674FAE3" blockId="2.[151,1437,150,1545]" pageId="2" pageNumber="545">
Based on the morphology of the adult male, the new family
<taxonomicName id="4C424D41FFC4FFCEAAA21F608636FBE7" box="[916,1070,1123,1149]" family="Qinococcidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" rank="family">Qinococcidae</taxonomicName>
is closest to the extinct family
<taxonomicName id="4C424D41FFC4FFCEA9A11F8483CCFB3B" authority="Koteja 2008" authorityName="Koteja" authorityYear="2008" box="[151,468,1159,1185]" class="Insecta" family="Weitschatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">
Weitschatidae
<bibRefCitation id="EFD34B33FFC4FFCEA8761F8483CCFB3B" author="Koteja, J." box="[320,468,1159,1185]" pageId="2" pageNumber="545" pagination="1 - 56" refId="ref11244" refString="Koteja, J. (2008) Xylococcidae and related groups (Hemiptera: Coccoidea) from Baltic amber. Prace Muzeum Ziemi, No. 49, 1 - 56." type="book chapter" year="2008">Koteja 2008</bibRefCitation>
</taxonomicName>
. They share the following characters: antennae each 10-segmented, with flagellar segments cylindrical and lacking capitate setae; compound eyes present; scutum without a membranous area; scutellum without membranous lateral areas; basisternum hexagonal; fore wings hyaline, broad at base, with a narrow alar fold for holding hamuli, subcostal ridge with distinct club-shaped pterostigma, cubital ridge situated beyond middle of wing; tarsus two-segmented; tail tufts present on abdominal tergites VI and VII (
<bibRefCitation id="EFD34B33FFC4FFCEADEC1E148770FAAB" author="Koteja, J." box="[1242,1384,1303,1329]" pageId="2" pageNumber="545" pagination="1 - 56" refId="ref11244" refString="Koteja, J. (2008) Xylococcidae and related groups (Hemiptera: Coccoidea) from Baltic amber. Prace Muzeum Ziemi, No. 49, 1 - 56." type="book chapter" year="2008">Koteja 2008</bibRefCitation>
;
<bibRefCitation id="EFD34B33FFC4FFCEAC441E148342FACF" author="Vea, I. M. &amp; Grimaldi, D. A." pageId="2" pageNumber="545" pagination="1 - 80" refId="ref11524" refString="Vea, I. M. &amp; Grimaldi, D. A. (2015) Diverse new scale insects (Hemiptera: Coccoidea) in amber from the Cretaceous and Eocene with a phylogenetic framework for fossil Coccoidea. American Museum Novitates, 3823, 1 - 80. https: // doi. org / 10.1206 / 3823.1" type="journal article" year="2015">Vea &amp; Grimaldi 2015</bibRefCitation>
;
<bibRefCitation id="EFD34B33FFC4FFCEA8531E388015FACF" author="Lin, S. &amp; Yao, Y. Z. &amp; Ren, D." box="[357,525,1339,1365]" pageId="2" pageNumber="545" pagination="427 - 434" refId="ref11276" refString="Lin, S., Yao, Y. Z. &amp; Ren, D. (2018) A new scale insect of the extinct family Weitschatidae (Insecta: Hemiptera: Coccomorpha) from mid-Cretaceous Burmese amber. Zootaxa, 4407 (3), 427 - 434. https: // doi. org / 10.11646 / zootaxa. 4407.3.9" type="journal article" year="2018">
Lin
<emphasis id="B936EAD0FFC4FFCEA8A41E3883D2FACF" box="[402,458,1339,1365]" italics="true" pageId="2" pageNumber="545">et al.</emphasis>
2018
</bibRefCitation>
); but
<taxonomicName id="4C424D41FFC4FFCEAB7C1E3880FCFACF" box="[586,740,1339,1365]" family="Qinococcidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" rank="family">Qinococcidae</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C424D41FFC4FFCEAD611E3886EBFACF" authorityName="Koteja" authorityYear="2008" box="[1111,1267,1339,1365]" class="Insecta" family="Weitschatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="family">Weitschatidae</taxonomicName>
by the anterior flexing patch being forked and in having two posterior flexing patches in the fore wing.
</paragraph>
<paragraph id="8BFD36C2FFC4FFCEA9F11E80866FF993" blockId="2.[151,1437,150,1545]" pageId="2" pageNumber="545">
Based on the published descriptions of the adult females of
<taxonomicName id="4C424D41FFC4FFCEAA581E80866BFA07" authority="Foldi" authorityName="Foldi" authorityYear="1997" box="[878,1139,1411,1437]" class="Insecta" family="Margarodidae" genus="Jansenus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="burgeri">
<emphasis id="B936EAD0FFC4FFCEAA581E808628FA07" box="[878,1072,1411,1437]" italics="true" pageId="2" pageNumber="545">Jansenus burgeri</emphasis>
Foldi
</taxonomicName>
and
<taxonomicName id="4C424D41FFC4FFCEAD9D1E80877DFA07" box="[1195,1381,1411,1437]" class="Insecta" family="Margarodidae" genus="Neogreenia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B936EAD0FFC4FFCEAD9D1E808735FA06" box="[1195,1325,1411,1436]" italics="true" pageId="2" pageNumber="545">Neogreenia</emphasis>
spp.
</taxonomicName>
, it is possible that these two genera may also fall within
<taxonomicName id="4C424D41FFC4FFCEABE91EA48164FA5B" box="[735,892,1447,1473]" family="Qinococcidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="545" rank="family">Qinococcidae</taxonomicName>
, but none of their males have been studied. The adult females of these three genera all share (i) vulva opening longitudinally, and (ii) multilocular pores each with 26 central loculi (
<bibRefCitation id="EFD34B33FFC4FFCEA85C1EEC802FF993" author="Tang F. D. &amp; Hao J. J." box="[362,567,1518,1545]" pageId="2" pageNumber="545" refId="ref11485" refString="Tang F. D. &amp; Hao J. J. (1995) The Margarodidae and others of China (Homoptera: Coccoidea of Insecta). Chinese Agricultural Science Technology Press, Beijing, 738 pp." type="book" year="1995">Tang &amp; Hao 1995</bibRefCitation>
;
<bibRefCitation id="EFD34B33FFC4FFCEAB751EEC80D8F993" author="Foldi, I." box="[579,704,1519,1545]" pageId="2" pageNumber="545" pagination="185 - 195" refId="ref10955" refString="Foldi, I. (1997) Les Xylococcinae (Hemiptera: Coccoidea: Margarodidae): caracteristiques des especes et description d'un nouveau genre. Annales de la Soci e t e entomologique de France, New Series, 33, 185 - 195." type="journal article" year="1997">Foldi 1997</bibRefCitation>
;
<bibRefCitation id="EFD34B33FFC4FFCEABFA1EEC81BBF993" author="Wu, S. A. &amp; Cheng G. F." box="[716,931,1519,1545]" pageId="2" pageNumber="545" pagination="62 - 64" refId="ref11677" refString="Wu, S. A. &amp; Cheng G. F. (2006) A description of a new species of genus Neogreenia MacGillivray from China (Homoptera: Coccoidea: Margarodidae). Scientia Silvae Sinicae, 42, 62 - 64." type="journal article" year="2006">Wu &amp; Cheng 2006</bibRefCitation>
;
<bibRefCitation id="EFD34B33FFC4FFCEAA9B1EEC8673F993" author="Wu, S. A. &amp; Nan, N." box="[941,1131,1519,1545]" pageId="2" pageNumber="545" pagination="43 - 54" refId="ref11722" refString="Wu, S. A. &amp; Nan, N. (2012) Neogreenia lonicera sp. nov., a new species of Margarodidae sensu lato (Hemiptera: Coccoidea) from China, with a key to species of Neogreenia MacGillivray and placement of the genus in the family Kuwaniidae. Zootaxa, 3274 (1), 43 - 54. https: // doi. org / 10.11646 / zootaxa. 3274.1.5" type="journal article" year="2012">Wu &amp; Nan 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>