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<document id="2118BF2A3F4834DB537CCD518845C9B7" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. &amp; Gordon, Dennis P." docDate="2018" docId="03EB8789FFE54479A1DA7062FDE8FDCB" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Cheethamia Shaw 1967" docType="treatment" docVersion="4" lastPageNumber="1670" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of onychocellid cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1669" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="57CDAF3DEAF139A70CB82A5FD618D7F6">Synopsis of onychocellid cheilostome bryozoan genera</mods:title>
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<mods:namePart id="4886AD99754BF7EA2BA9299552D0C86F">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="26AF732A18D5C7A4570632DD92E67A52">Martha, Silviu O.</mods:namePart>
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<mods:title id="5CA075A1FF6C7071CBFEDF4255D985B1">Journal of Natural History</mods:title>
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<mods:date id="D0DE7BF902DD9C3D835E915ACF195080">2018</mods:date>
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<mods:number id="6898BB3CB2CF88A5D5D34992E92D9E1F">2018-07-03</mods:number>
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<mods:number id="D4EEDA5B196F0D2A6E40E6D809520DE1">25 - 26</mods:number>
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<subSubSection id="C3586514FFE5447AA1DA7062FCDFFB80" box="[470,845,1061,1086]" pageId="13" pageNumber="1669" type="nomenclature">
<paragraph id="8BFD369FFFE5447AA1DA7062FCDFFB80" blockId="13.[470,845,1061,1120]" box="[470,845,1061,1086]" pageId="13" pageNumber="1669">
<heading id="D0B581F3FFE5447AA1DA7062FCDFFB80" box="[470,845,1061,1086]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="1669" reason="2">
Genus
<taxonomicName id="4C424D1CFFE5447AA2257062FCDFFB80" ID-CoL="859Y7" authority="Shaw, 1967" authorityName="Shaw" authorityYear="1967" box="[553,845,1061,1086]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="1669" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE5447AA2257062FD2EFB80" bold="true" box="[553,700,1061,1086]" italics="true" pageId="13" pageNumber="1669">Cheethamia</emphasis>
<bibRefCitation id="EFD34B6EFFE5447AA2CA7062FCDFFB80" author="Shaw NG" box="[710,845,1061,1086]" pageId="13" pageNumber="1669" pagination="1393 - 1432" refId="ref28072" refString="Shaw NG. 1967. Cheilostomata from Gulfian (Upper Cretaceous) rocks of southwestern Arkansas. J Paleontol. 41: 1393 - 1432." type="journal article" year="1967">Shaw, 1967</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFE5447AA2557000FD59FBDE" box="[601,715,1095,1120]" pageId="13" pageNumber="1669" type="description">
<paragraph id="8BFD369FFFE5447AA2557000FD59FBDE" blockId="13.[470,845,1061,1120]" box="[601,715,1095,1120]" pageId="13" pageNumber="1669">
(
<figureCitation id="13792A1AFFE5447AA26D7000FD50FBDE" box="[609,706,1095,1120]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFE5447AA0AC70D2FDC6FB4A" pageId="13" pageNumber="1669" type="type_taxon">
<paragraph id="8BFD369FFFE5447AA0AC70D2FEA6FB10" blockId="13.[160,1155,1173,1268]" box="[160,308,1173,1198]" pageId="13" pageNumber="1669">
<emphasis id="B936EA8DFFE5447AA0AC70D2FEA6FB10" bold="true" box="[160,308,1173,1198]" italics="true" pageId="13" pageNumber="1669">
<typeStatus id="54F9883DFFE5447AA0AC70D2FF4AFB10" box="[160,216,1173,1198]" pageId="13" pageNumber="1669">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE5447AA0AC70FEFDC6FB4A" blockId="13.[160,1155,1173,1268]" pageId="13" pageNumber="1669">
<taxonomicName id="4C424D1CFFE5447AA0AC70FEFE53FB6F" ID-CoL="86FQZ" box="[160,449,1209,1233]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="1669" phylum="Bryozoa" rank="species" species="howei">
<emphasis id="B936EA8DFFE5447AA0AC70FEFEE1FB6F" box="[160,371,1209,1233]" italics="true" pageId="13" pageNumber="1669">Cheethamia howei</emphasis>
, 1967
</taxonomicName>
, by original designation.
<materialsCitation id="3B2A3CC2FFE5447AA2F770FEFDC2FB4A" country="United States of America" county="Howard County" location="Howard County" pageId="13" pageNumber="1669" specimenCount="1" stateProvince="Arkansas" typeStatus="holotype">
Cretaceous, Campanian, Saratoga Chalk,
<collectingCounty id="629C4E13FFE5447AA0E0709BFE33FB4A" box="[236,417,1244,1268]" pageId="13" pageNumber="1669">Howard County</collectingCounty>
,
<collectingRegion id="4986F87DFFE5447AA1A1709BFD81FB4A" box="[429,531,1244,1268]" country="United States of America" name="Arkansas" pageId="13" pageNumber="1669">Arkansas</collectingRegion>
,
<collectingCountry id="F355760FFFE5447AA212709BFDC2FB4A" box="[542,592,1244,1268]" name="United States of America" pageId="13" pageNumber="1669">USA</collectingCountry>
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.
</paragraph>
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<subSubSection id="C3586514FFE54479A0AC716EFDE8FDCB" lastPageId="14" lastPageNumber="1670" pageId="13" pageNumber="1669" type="diagnosis">
<paragraph id="8BFD369FFFE5447AA0AC716EFE86FAFC" blockId="13.[160,1156,1321,1727]" box="[160,276,1321,1346]" pageId="13" pageNumber="1669">
<emphasis id="B936EA8DFFE5447AA0AC716EFE86FAFC" bold="true" box="[160,276,1321,1346]" italics="true" pageId="13" pageNumber="1669">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE54479A0AC710AFE94FF18" blockId="13.[160,1156,1321,1727]" lastBlockId="14.[160,1156,141,513]" lastPageId="14" lastPageNumber="1670" pageId="13" pageNumber="1669">
Colony encrusting. Autozooids hexagonal (
<figureCitation id="13792A1AFFE5447AA278710AFD5CFADB" box="[628,718,1357,1381]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
(b)); zooidal boundaries grooved or with a raised edge. Cryptocyst convex, extensive, granular. Gymnocyst lacking. Opesia terminal, occupying about one-third of frontal surface, semielliptical, arch-shaped with small opesiular indentations, proximal edge more or less straight, shelf-like occlusor lamina distally; a pair of closely spaced oral spine bases in non-ovicellate zooids (
<figureCitation id="13792A1AFFE5447AA32D7190FCECFA51" box="[801,894,1495,1519]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
(c)), four spine bases in zooids budded directly from the ancestrula. Ovicell hyperstomial (
<figureCitation id="13792A1AFFE5447AA37271BDFC48F9AC" box="[894,986,1530,1554]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
(d)), small, ooecium interior-walled, a convex transverse wall visible within the opesia separating the ovicell from the rest of the zooid. Avicularia vicarious, normally slightly asymmetrical (
<figureCitation id="13792A1AFFE5447AA40D7278FBCEF9E9" box="[1025,1116,1599,1623]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
(e)), sometimes symmetrical (
<figureCitation id="13792A1AFFE5447AA1A07225FD96F9C4" box="[428,516,1634,1658]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
(a)), dagger-like, about the same length as an autozooid but half the width; opesia longitudinally elliptical; rostrum long, acutely triangular, straight or slightly curved. Ancestrula similar in size and morphology to budded autozooids (
<figureCitation id="13792A1AFFE5447AA40572E0FBF3F901" box="[1033,1121,1703,1727]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="13" pageNumber="1669">Figure 4</figureCitation>
(f)).
<emphasis id="B936EA8DFFE64479A0AC74CAFE94FF18" bold="true" box="[160,262,141,166]" italics="true" pageId="14" pageNumber="1670">Remarks</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE64479A0AC74F6FCD6FDBF" blockId="14.[160,1156,141,513]" pageId="14" pageNumber="1670">
Unlike
<taxonomicName id="4C424D1CFFE64479A0FD74F6FEE4FF77" authorityName="Jullien" authorityYear="1882" box="[241,374,177,201]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE64479A0FD74F6FEE4FF77" box="[241,374,177,201]" italics="true" pageId="14" pageNumber="1670">Onychocella</emphasis>
</taxonomicName>
, the ovicell of
<taxonomicName id="4C424D1CFFE64479A22574F6FD3EFF77" authorityName="Shaw" authorityYear="1967" box="[553,684,177,201]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE64479A22574F6FD3EFF77" box="[553,684,177,201]" italics="true" pageId="14" pageNumber="1670">Cheethamia</emphasis>
</taxonomicName>
is hyperstomial, albeit small in the
<typeStatus id="54F9883DFFE64479A45D74F6FB11FF77" box="[1105,1155,177,201]" pageId="14" pageNumber="1670">type</typeStatus>
species
<taxonomicName id="4C424D1CFFE64479A0F47493FD04FF52" authority="(Taylor and McKinney 2006)" baseAuthorityName="Taylor and McKinney" baseAuthorityYear="2006" box="[248,662,212,236]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="howei">
<emphasis id="B936EA8DFFE64479A0F47493FEC7FF52" box="[248,341,212,236]" italics="true" pageId="14" pageNumber="1670">C. howei</emphasis>
(
<bibRefCitation id="EFD34B6EFFE64479A16E7493FD1FFF52" author="Taylor PD &amp; McKinney FK" box="[354,653,212,236]" pageId="14" pageNumber="1670" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney 2006</bibRefCitation>
)
</taxonomicName>
. The oral spine bases are another feature of
<taxonomicName id="4C424D1CFFE64479A0AC74B1FE92FEB0" baseAuthorityName="Taylor and McKinney" baseAuthorityYear="2006" box="[160,256,246,270]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="howei">
<emphasis id="B936EA8DFFE64479A0AC74B1FE92FEB0" box="[160,256,246,270]" italics="true" pageId="14" pageNumber="1670">C. howei</emphasis>
</taxonomicName>
that are lacking in the
<typeStatus id="54F9883DFFE64479A21674B1FDDEFEB0" box="[538,588,246,270]" pageId="14" pageNumber="1670">type</typeStatus>
species of
<taxonomicName id="4C424D1CFFE64479A2D874B1FCCBFEB0" authorityName="Jullien" authorityYear="1882" box="[724,857,246,270]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE64479A2D874B1FCCBFEB0" box="[724,857,246,270]" italics="true" pageId="14" pageNumber="1670">Onychocella</emphasis>
</taxonomicName>
, although oral spines are present in a few Cretaceous species that have been referred to
<taxonomicName id="4C424D1CFFE64479A37C755EFC67FE8F" authorityName="Jullien" authorityYear="1882" box="[880,1013,281,305]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE64479A37C755EFC67FE8F" box="[880,1013,281,305]" italics="true" pageId="14" pageNumber="1670">Onychocella</emphasis>
</taxonomicName>
(see above).
<bibRefCitation id="EFD34B6EFFE64479A0CC757BFED4FEEA" author="Shaw NG" box="[192,326,316,340]" pageId="14" pageNumber="1670" pagination="1393 - 1432" refId="ref28072" refString="Shaw NG. 1967. Cheilostomata from Gulfian (Upper Cretaceous) rocks of southwestern Arkansas. J Paleontol. 41: 1393 - 1432." type="journal article" year="1967">Shaw (1967)</bibRefCitation>
transferred
<taxonomicName id="4C424D1CFFE64479A1CB757BFC88FEEA" authority="Voigt, 1962" authorityName="Voigt" authorityYear="1962" box="[455,794,316,340]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="pinguis">
<emphasis id="B936EA8DFFE64479A1CB757BFD09FEEA" box="[455,667,316,340]" italics="true" pageId="14" pageNumber="1670">Onychocella pinguis</emphasis>
<bibRefCitation id="EFD34B6EFFE64479A2AC757BFC88FEEA" author="Voigt E" box="[672,794,316,340]" pageId="14" pageNumber="1670" refId="ref28832" refString="Voigt E. 1962. Viрkнiмiлovыi мsaнki ivрopiйсkoй caсti СССР i нikotoрыk сopрidiльныk obлaсtiй [Upper Cretaceous bryozoans of the European part of the USSR and neighbouring territories]. Мoсkva [Moskva]: Izdatiльсtvo Мoсkovсkogo uнiviрсitita [Moscow University Press]." type="book" year="1962">Voigt, 1962</bibRefCitation>
</taxonomicName>
from the Maastrichtian of
<collectingRegion id="4986F87DFFE64479A43A757BFB16FEEA" box="[1078,1156,316,340]" country="Ukraine" name="Crimea" pageId="14" pageNumber="1670">Crimea</collectingRegion>
to
<taxonomicName id="4C424D1CFFE64479A0B07519FEAFFEC8" authorityName="Shaw" authorityYear="1967" box="[188,317,350,374]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE64479A0B07519FEAFFEC8" box="[188,317,350,374]" italics="true" pageId="14" pageNumber="1670">Cheethamia</emphasis>
</taxonomicName>
, an action supported by
<bibRefCitation id="EFD34B6EFFE64479A2447519FCB2FEC8" author="Koromyslova AV" box="[584,800,350,374]" pageId="14" pageNumber="1670" pagination="275 - 286" refId="ref26776" refString="Koromyslova AV. 2014. Morphological features and systematic position of the bryozoans Onychocella rowei and O. mimosa (Cheilostomata) from a Campanian erratic block (Belarus). Paleontol Zh. 48: 275 - 286." type="journal article" year="2014">Koromyslova (2014)</bibRefCitation>
who rediagnosed the genus and added two further Maastrichtian species from eastern
<collectingCountry id="F355760FFFE64479A2E675C6FCACFE27" box="[746,830,385,409]" name="Ukraine" pageId="14" pageNumber="1670">Ukraine</collectingCountry>
and
<collectingCountry id="F355760FFFE64479A37F75C6FC7DFE27" box="[883,1007,385,409]" name="Kazakhstan" pageId="14" pageNumber="1670">Kazakhstan</collectingCountry>
to the genus:
<taxonomicName id="4C424D1CFFE64479A0AC75E3FDE7FE02" authority="Koromyslova, 2014" authorityName="Koromyslova" authorityYear="2014" box="[160,629,420,444]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="subpinguis">
<emphasis id="B936EA8DFFE64479A0AC75E3FE09FE02" box="[160,411,420,444]" italics="true" pageId="14" pageNumber="1670">Cheethamia subpinguis</emphasis>
<bibRefCitation id="EFD34B6EFFE64479A1AF75E3FDE7FE02" author="Koromyslova AV" box="[419,629,420,444]" pageId="14" pageNumber="1670" pagination="275 - 286" refId="ref26776" refString="Koromyslova AV. 2014. Morphological features and systematic position of the bryozoans Onychocella rowei and O. mimosa (Cheilostomata) from a Campanian erratic block (Belarus). Paleontol Zh. 48: 275 - 286." type="journal article" year="2014">Koromyslova, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C424D1CFFE64479A2A375E3FC65FE02" authority="Koromyslova, 2014" authorityName="Koromyslova" authorityYear="2014" box="[687,1015,420,444]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="arcuata">
<emphasis id="B936EA8DFFE64479A2A375E3FC8CFE02" box="[687,798,420,444]" italics="true" pageId="14" pageNumber="1670">C. arcuata</emphasis>
<bibRefCitation id="EFD34B6EFFE64479A32A75E3FC65FE02" author="Koromyslova AV" box="[806,1015,420,444]" pageId="14" pageNumber="1670" pagination="275 - 286" refId="ref26776" refString="Koromyslova AV. 2014. Morphological features and systematic position of the bryozoans Onychocella rowei and O. mimosa (Cheilostomata) from a Campanian erratic block (Belarus). Paleontol Zh. 48: 275 - 286." type="journal article" year="2014">Koromyslova, 2014</bibRefCitation>
</taxonomicName>
. All of these species share with
<taxonomicName id="4C424D1CFFE64479A1617581FE5AFE60" baseAuthorityName="Taylor and McKinney" baseAuthorityYear="2006" box="[365,456,454,478]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="howei">
<emphasis id="B936EA8DFFE64479A1617581FE5AFE60" box="[365,456,454,478]" italics="true" pageId="14" pageNumber="1670">C. howei</emphasis>
</taxonomicName>
one or a pair of oral spine bases but
<taxonomicName id="4C424D1CFFE64479A3567581FC53FE60" authorityName="Voigt" authorityYear="1962" box="[858,961,454,478]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="pinguis">
<emphasis id="B936EA8DFFE64479A3567581FC53FE60" box="[858,961,454,478]" italics="true" pageId="14" pageNumber="1670">C. pinguis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C424D1CFFE64479A3FB7581FB11FE60" authorityName="Koromyslova" authorityYear="2014" box="[1015,1155,454,478]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="1670" phylum="Bryozoa" rank="species" species="subpinguis">
<emphasis id="B936EA8DFFE64479A3FB7581FB11FE60" box="[1015,1155,454,478]" italics="true" pageId="14" pageNumber="1670">C. subpinguis</emphasis>
</taxonomicName>
differ from the
<typeStatus id="54F9883DFFE64479A14475AEFEEBFDBF" box="[328,377,489,513]" pageId="14" pageNumber="1670">type</typeStatus>
species in having erect bifoliate colonies.
</paragraph>
<paragraph id="8BFD369FFFE64479A0AC767EFF79FDEC" blockId="14.[160,634,569,629]" box="[160,235,569,594]" pageId="14" pageNumber="1670">
<emphasis id="B936EA8DFFE64479A0AC767EFF79FDEC" bold="true" box="[160,235,569,594]" italics="true" pageId="14" pageNumber="1670">Range</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE64479A0AC761AFDE8FDCB" blockId="14.[160,634,569,629]" box="[160,634,605,629]" pageId="14" pageNumber="1670">Cretaceous (Campanian to Maastrichtian).</paragraph>
</subSubSection>
</treatment>
</document>