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<document id="9853B7FFE0ABE35E8F3D853B0A872320" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. &amp; Gordon, Dennis P." docDate="2018" docId="03EB8789FFC84456A194759CFD1DFCFE" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Nudicella Schmidt and Bone 2004" docType="treatment" docVersion="4" lastPageNumber="1689" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of onychocellid cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1688" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5267DDD8BADD848B04DB0DEC48DA5DAD">Synopsis of onychocellid cheilostome bryozoan genera</mods:title>
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<mods:namePart id="139EC40FE5D4C0340FF409D49F0D2D04">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="4712424CB1B632FB47A9DFD3174139A5">Martha, Silviu O.</mods:namePart>
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<mods:date id="7C9CD2CC11998822472835B7F25BFAD4">2018</mods:date>
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<mods:number id="5DD44088372C21EF54F0BB0E747160B6">2018-07-03</mods:number>
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<subSubSection id="C3586514FFC84457A194759CFC19FE4A" box="[408,907,475,500]" pageId="32" pageNumber="1688" type="nomenclature">
<paragraph id="8BFD369FFFC84457A194759CFC19FE4A" blockId="32.[408,907,475,535]" box="[408,907,475,500]" pageId="32" pageNumber="1688">
<heading id="D0B581F3FFC84457A194759CFC19FE4A" box="[408,907,475,500]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="1688" reason="2">
Genus
<taxonomicName id="4C424D1CFFC84457A1E7759CFC19FE4A" ID-CoL="85GYB" authority="Schmidt and Bone, 2004" authorityName="Schmidt and Bone" authorityYear="2004" box="[491,907,475,500]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="1688" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFC84457A1E7759CFDCFFE4A" bold="true" box="[491,605,475,500]" italics="true" pageId="32" pageNumber="1688">Nudicella</emphasis>
<bibRefCitation id="EFD34B6EFFC84457A26B759CFC19FE4A" author="Schmidt R &amp; Bone Y" box="[615,907,475,500]" pageId="32" pageNumber="1688" pagination="185 - 203" refId="ref28003" refString="Schmidt R, Bone Y. 2004. Australian Cainozoic Bryozoa, 1: Nudicella gen. nov. (Onychocellidae, Cheilostomata): taxonomy, palaeoenvironments and biogeography. Alcheringa. 28: 185 - 203." type="journal article" year="2004">Schmidt and Bone, 2004</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFC84457A25D75B9FD40FDA9" box="[593,722,510,535]" pageId="32" pageNumber="1688" type="description">
<paragraph id="8BFD369FFFC84457A25D75B9FD40FDA9" blockId="32.[408,907,475,535]" box="[593,722,510,535]" pageId="32" pageNumber="1688">
(
<figureCitation id="13792A1AFFC84457A25575B9FD58FDA9" box="[601,714,510,535]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="32" pageNumber="1688">Figure 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFC84457A0AC7600FDC4FD18" pageId="32" pageNumber="1688" type="type_taxon">
<paragraph id="8BFD369FFFC84457A0AC7600FEA6FDDE" blockId="32.[160,1156,583,678]" box="[160,308,583,608]" pageId="32" pageNumber="1688">
<emphasis id="B936EA8DFFC84457A0AC7600FEA6FDDE" bold="true" box="[160,308,583,608]" italics="true" pageId="32" pageNumber="1688">
<typeStatus id="54F9883DFFC84457A0AC7600FF4AFDDE" box="[160,216,583,608]" pageId="32" pageNumber="1688">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="8BFD369FFFC84457A0AC762CFDC4FD18" blockId="32.[160,1156,583,678]" pageId="32" pageNumber="1688">
<taxonomicName id="4C424D1CFFC84457A0AC762CFE5FFD3D" ID-CoL="86NJ4" box="[160,461,619,643]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="1688" phylum="Bryozoa" rank="species" species="latiramosa">
<emphasis id="B936EA8DFFC84457A0AC762CFE11FD3D" box="[160,387,619,643]" italics="true" pageId="32" pageNumber="1688">Nudicella latiramosa</emphasis>
, 2004
</taxonomicName>
, by original designation. Middle Eocene to Early Oligocene,
<collectingRegion id="4986F87DFFC84457A0AC76C9FDC0FD18" box="[160,594,654,678]" isEnumeration="true" pageId="32" pageNumber="1688">South Australia and Western Australia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFC84456A0AC7691FB16FE8F" lastPageId="33" lastPageNumber="1689" pageId="32" pageNumber="1688" type="diagnosis">
<paragraph id="8BFD369FFFC84457A0AC7691FE86FD51" blockId="32.[160,1157,726,890]" box="[160,276,726,751]" pageId="32" pageNumber="1688">
<emphasis id="B936EA8DFFC84457A0AC7691FE86FD51" bold="true" box="[160,276,726,751]" italics="true" pageId="32" pageNumber="1688">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BFD369FFFC84456A0AC76BDFB16FE8F" blockId="32.[160,1157,726,890]" lastBlockId="33.[160,1156,142,305]" lastPageId="33" lastPageNumber="1689" pageId="32" pageNumber="1688">
Colony erect bifoliate, palmate with bifurcating branches (
<figureCitation id="13792A1AFFC84457A32C76BDFC1AFCAC" box="[800,904,762,786]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="32" pageNumber="1688">Figure 18</figureCitation>
(a,d)). Autozooids elongate hexagonal, distal edges rounded; zooidal boundaries grooved. Cryptocyst extensive, granular. Gymnocyst lacking. Opesia terminal, occupying less than one-third of frontal surface; semi-elliptical with opesiular indentations and an occlusor-lamina (
<figureCitation id="13792A1AFFC84457A3847725FC61FCC4" box="[904,1011,866,890]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="32" pageNumber="1688">Figure 18</figureCitation>
(b)), proximal edge straight or slightly concave. Ovicells immersed (
<figureCitation id="13792A1AFFC94456A30674C9FCE4FF18" box="[778,886,142,166]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="33" pageNumber="1689">Figure 18</figureCitation>
(e)); opesia in ovicellate autozooids enlarged, a transverse wall visible within the opesia separating the ovicell from the rest of the zooid. Avicularia vicarious, symmetrical (
<figureCitation id="13792A1AFFC94456A2EE7493FCD4FF52" box="[738,838,212,236]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="33" pageNumber="1689">Figure 18</figureCitation>
(c)), about half the width of an autozooid but of similar length; opesia transversely semi-elliptical with a small foramen; rostrum acutely triangular. Kenozooids present along branch edges and in axils (
<figureCitation id="13792A1AFFC94456A3F1755EFBF1FE8F" box="[1021,1123,281,305]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="33" pageNumber="1689">Figure 18</figureCitation>
(f)).
</paragraph>
</subSubSection>
<caption id="DF3D6617FFC84457A0AC7217FC7CF901" ID-DOI="http://doi.org/10.5281/zenodo.4747183" ID-Zenodo-Dep="4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="32" pageNumber="1688" startId="32.[160,225,1616,1639]" targetBox="[163,1153,948,1585]" targetPageId="32">
<paragraph id="8BFD369FFFC84457A0AC7217FC7CF901" blockId="32.[160,1156,1616,1727]" pageId="32" pageNumber="1688">
<emphasis id="B936EA8DFFC84457A0AC7217FE9AF9D9" bold="true" box="[160,264,1616,1639]" pageId="32" pageNumber="1688">Figure 18.</emphasis>
<taxonomicName id="4C424D1CFFC84457A11C7217FD05F9D9" authority="Schmidt and Bone" authorityName="Schmidt and Bone" box="[272,663,1616,1639]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="1688" phylum="Bryozoa" rank="species" species="latiramosa">
<emphasis id="B936EA8DFFC84457A11C7217FE44F9D9" box="[272,470,1616,1639]" italics="true" pageId="32" pageNumber="1688">Nudicella latiramosa</emphasis>
Schmidt and Bone
</taxonomicName>
; Early Oligocene, Port Willunga Formation, South Australia; (a
<emphasis id="B936EA8DFFC84457A114722AFEB7F93A" box="[280,293,1645,1668]" italics="true" pageId="32" pageNumber="1688"></emphasis>
c) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (d
<emphasis id="B936EA8DFFC84457A465722AFBE4F93A" box="[1129,1142,1645,1668]" italics="true" pageId="32" pageNumber="1688"></emphasis>
f) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm.
</paragraph>
</caption>
<subSubSection id="C3586514FFC94456A0AC7526FD1DFCFE" pageId="33" pageNumber="1689" type="discussion">
<paragraph id="8BFD369FFFC94456A0AC7526FE94FEC4" blockId="33.[160,1156,353,724]" box="[160,262,353,378]" pageId="33" pageNumber="1689">
<emphasis id="B936EA8DFFC94456A0AC7526FE94FEC4" bold="true" box="[160,262,353,378]" italics="true" pageId="33" pageNumber="1689">Remarks</emphasis>
</paragraph>
<paragraph id="8BFD369FFFC94456A0AC75C2FC67FDD3" blockId="33.[160,1156,353,724]" pageId="33" pageNumber="1689">
Five species were recognised by the authors of
<taxonomicName id="4C424D1CFFC94456A2EB75C2FCD9FE23" authorityName="Schmidt and Bone" authorityYear="2004" box="[743,843,389,413]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="1689" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFC94456A2EB75C2FCD9FE23" box="[743,843,389,413]" italics="true" pageId="33" pageNumber="1689">Nudicella</emphasis>
</taxonomicName>
, all from the Cenozoic of
<collectingCountry id="F355760FFFC94456A0AC75E0FE93FE01" box="[160,257,423,447]" name="Australia" pageId="33" pageNumber="1689">Australia</collectingCountry>
(
<bibRefCitation id="EFD34B6EFFC94456A11F75E0FDBCFE01" author="Schmidt R &amp; Bone Y" box="[275,558,423,447]" pageId="33" pageNumber="1689" pagination="185 - 203" refId="ref28003" refString="Schmidt R, Bone Y. 2004. Australian Cainozoic Bryozoa, 1: Nudicella gen. nov. (Onychocellidae, Cheilostomata): taxonomy, palaeoenvironments and biogeography. Alcheringa. 28: 185 - 203." type="journal article" year="2004">Schmidt and Bone 2004</bibRefCitation>
). The key characters of the genus are the straight avicularia and occurrence of a small foramen in the proximal cryptocyst of the avicularia (
<figureCitation id="13792A1AFFC94456A0AB75AAFE88FDBB" box="[167,282,493,517]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="33" pageNumber="1689">Figure 18</figureCitation>
(c)). Most species of
<taxonomicName id="4C424D1CFFC94456A21175AAFD13FDBB" authorityName="Schmidt and Bone" authorityYear="2004" box="[541,641,493,517]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="1689" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFC94456A21175AAFD13FDBB" box="[541,641,493,517]" italics="true" pageId="33" pageNumber="1689">Nudicella</emphasis>
</taxonomicName>
have palmate erect colonies with narrow bifoliate branches that bifurcate. However,
<taxonomicName id="4C424D1CFFC94456A2997648FBC1FD99" authority="Schmidt and Bone, 2004" authorityName="Schmidt and Bone" authorityYear="2004" box="[661,1107,527,551]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="1689" phylum="Bryozoa" rank="species" species="cribriforma">
<emphasis id="B936EA8DFFC94456A2997648FCBFFD99" box="[661,813,527,551]" italics="true" pageId="33" pageNumber="1689">N. cribriforma</emphasis>
<bibRefCitation id="EFD34B6EFFC94456A33B7648FBC1FD99" author="Schmidt R &amp; Bone Y" box="[823,1107,527,551]" pageId="33" pageNumber="1689" pagination="185 - 203" refId="ref28003" refString="Schmidt R, Bone Y. 2004. Australian Cainozoic Bryozoa, 1: Nudicella gen. nov. (Onychocellidae, Cheilostomata): taxonomy, palaeoenvironments and biogeography. Alcheringa. 28: 185 - 203." type="journal article" year="2004">Schmidt and Bone, 2004</bibRefCitation>
</taxonomicName>
has characteristically cribrate colonies (see
<bibRefCitation id="EFD34B6EFFC94456A26E7675FD7AFDF4" author="Taylor PD" box="[610,744,562,586]" pageId="33" pageNumber="1689" pagination="32 - 38" refId="ref28287" refString="Taylor PD. 2012. Identity of Mesosecos Faura &amp; Canu, 1917, a neglected early genus of free-living Cupuladriidae (Bryozoa, Cheilostomata) from the Spanish Eocene. Batalleria. 17: 32 - 38." type="journal article" year="2012">Taylor 2012</bibRefCitation>
), and an unnamed species with an encrusting colony form was depicted by
<bibRefCitation id="EFD34B6EFFC94456A27A7612FC1DFDD3" author="Schmidt R &amp; Bone Y" box="[630,911,597,621]" pageId="33" pageNumber="1689" pagination="185 - 203" refId="ref28003" refString="Schmidt R, Bone Y. 2004. Australian Cainozoic Bryozoa, 1: Nudicella gen. nov. (Onychocellidae, Cheilostomata): taxonomy, palaeoenvironments and biogeography. Alcheringa. 28: 185 - 203." type="journal article" year="2004">Schmidt and Bone (2004</bibRefCitation>
; fig. 9D).
</paragraph>
<paragraph id="8BFD369FFFC94456A0CC7630FB8FFD6A" blockId="33.[160,1156,353,724]" pageId="33" pageNumber="1689">
The presence of a transverse wall between the main chamber of the maternal zooid and the ovicell (
<figureCitation id="13792A1AFFC94456A0FC76DDFEC6FD0C" box="[240,340,666,690]" captionStart="Figure 18" captionStartId="32.[160,225,1616,1639]" captionTargetBox="[163,1153,948,1585]" captionTargetId="figure-158@32.[163,1153,948,1585]" captionTargetPageId="32" captionText="Figure 18. Nudicella latiramosa Schmidt and Bone; Early Oligocene, Port Willunga Formation, South Australia; (ac) NHMUK BZ 8216; (a) palmate colony; (b) autozooid opesia; (c) avicularium; (df) NHMUK BZ 8217; (d) bifurcating branch; (e) ovicellate autozooids; (f) non-ovicellate autozooids with a kenozooid in branch axis. Scale bars: a, d = 1 mm; b = 100 µm; c, e, f = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747183" httpUri="https://zenodo.org/record/4747183/files/figure.png" pageId="33" pageNumber="1689">Figure 18</figureCitation>
(e)) is reminiscent of
<taxonomicName id="4C424D1CFFC94456A22076DDFD38FD0C" authorityName="Shaw" authorityYear="1967" box="[556,682,666,690]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="1689" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFC94456A22076DDFD38FD0C" box="[556,682,666,690]" italics="true" pageId="33" pageNumber="1689">Cheethamia</emphasis>
</taxonomicName>
(
<figureCitation id="13792A1AFFC94456A2BA76DDFC9CFD0C" box="[694,782,666,690]" captionStart="Figure 4" captionStartId="13.[160,225,810,833]" captionTargetBox="[162,1152,142,779]" captionTargetId="figure-176@13.[162,1153,142,779]" captionTargetPageId="13" captionText="Figure 4. Cheethamia howei Shaw. (a) holotype, LSUMG-I 8237; Cretaceous, Campanian, Saratoga Chalk, Howard County, Arkansas, USA. (bf) NHMUK BZ 4809; Cretaceous, Maastrichtian, Lake Ridge Apartments, Livingston, Alabama, USA; (b) ovicellate and non-ovicellate autozooids and an avicularium; (c) opesia of non-ovicellate autozooid showing occlusor lamina and a pair of closely spaced oral spine bases; (d) ovicellate autozooid with transverse wall visible in opesia; (e) avicularium; (f) ancestrula (lower centre) and first-budded autozooids. Scale bars: a, b, e, f = 500 µm; c = 100 µm; d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747149" httpUri="https://zenodo.org/record/4747149/files/figure.png" pageId="33" pageNumber="1689">Figure 4</figureCitation>
(d)). Both
<taxonomicName id="4C424D1CFFC94456A37E76DDFC40FD0C" authorityName="Schmidt and Bone" authorityYear="2004" box="[882,978,666,690]" class="Gymnolaemata" family="Onychocellidae" genus="Nudicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="1689" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFC94456A37E76DDFC40FD0C" box="[882,978,666,690]" italics="true" pageId="33" pageNumber="1689">Nudicella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C424D1CFFC94456A40976DDFB16FD0C" authorityName="Shaw" authorityYear="1967" box="[1029,1156,666,690]" class="Gymnolaemata" family="Onychocellidae" genus="Cheethamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="1689" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFC94456A40976DDFB16FD0C" box="[1029,1156,666,690]" italics="true" pageId="33" pageNumber="1689">Cheethamia</emphasis>
</taxonomicName>
also have occlusor laminae, suggesting that the two genera may be closely related.
</paragraph>
<paragraph id="8BFD369FFFC94456A0AC7742FF79FCA0" blockId="33.[160,655,773,832]" box="[160,235,773,798]" pageId="33" pageNumber="1689">
<emphasis id="B936EA8DFFC94456A0AC7742FF79FCA0" bold="true" box="[160,235,773,798]" italics="true" pageId="33" pageNumber="1689">Range</emphasis>
</paragraph>
<paragraph id="8BFD369FFFC94456A0AC776FFD1DFCFE" blockId="33.[160,655,773,832]" box="[160,655,808,832]" pageId="33" pageNumber="1689">Paleocene (Thanetian) to Miocene (Middle).</paragraph>
</subSubSection>
</treatment>
</document>