treatments-xml/data/03/DC/6C/03DC6C76CC284C3EF7DFFE81FCB6F821.xml
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<document id="06451CE2B246B81302EE787D59DBFAEC" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC284C3EF7DFFE81FCB6F821" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana chapmani Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="A6EC50CC-1FF8-4FB7-9E01-497C083A4A61" docUuidSource="ZooBank" docVersion="3" lastPageNumber="27" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="26" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="1F5AA0E12AB980757221FEBE2C564F94">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:namePart id="C03471033615300B3C011F3AFBFD4333">Lehmann, Ingo</mods:namePart>
<mods:nameIdentifier id="736147F1F805D39475413391865E5EA4" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:namePart id="457E86652C969BD1CADC20F640A384C0">Zahiri, Reza</mods:namePart>
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<mods:namePart id="9A4B717A1D6AB164F443953386F2C58B">Husemann, Martin</mods:namePart>
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<taxonomicName id="4C75A6E3CC284C3DF7DFFE81FCB6FE3D" ID-CoL="B8R83" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,808,366,393]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chapmani" status="sp. nov.">
<emphasis id="B9010172CC284C3DF7DFFE81FE02FE3C" bold="true" box="[151,412,366,393]" italics="true" pageId="25" pageNumber="26">Lukeniana chapmani</emphasis>
Lehmann, Zahiri &amp; Husemann
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<paragraph id="8BCADD60CC284C3DF7DFFE7DFEBEFE19" blockId="25.[151,970,366,465]" box="[151,288,402,429]" pageId="25" pageNumber="26">
<figureCitation id="134EC1E5CC284C3DF7DFFE7DFF75FE18" box="[151,235,402,429]" captionStart="FIGURE 5" captionStartId="26.[151,250,1484,1508]" captionTargetBox="[151,1436,181,1460]" captionTargetId="figure-28@26.[151,1436,181,1460]" captionTargetPageId="26" captionText="FIGURE 5. a. Lukeniana chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; b. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; c. L. michaelgrzimeki, paratype, female, Tanzania, Arusha Region, Mount Meru, Momella; d. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave; e. L. mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; f. L. mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkhorongo." figureDoi="http://doi.org/10.5281/zenodo.7840798" httpUri="https://zenodo.org/record/7840798/files/figure.png" pageId="25" pageNumber="26">Figs 5a</figureCitation>
,
<figureCitation id="134EC1E5CC284C3DF7BFFE7DFEBEFE19" box="[247,288,402,428]" captionStart="FIGURE 16" captionStartId="67.[151,250,1673,1697]" captionTargetBox="[159,1428,391,1636]" captionTargetId="figure-109@67.[151,1436,381,1649]" captionTargetPageId="67" captionText="FIGURE 16. a. Lukeniana madrandelei, paratype, male, DRC, Katanga Province, Lubumbashi; b. L. friederikebauerae, paratype, male, Zambia, Lusaka Province, Lusaka; c. L. chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; d. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; e. L. kammeri, holotype, male, Kenya, County Kiambu, Thika; f. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave." figureDoi="http://doi.org/10.5281/zenodo.7840829" httpUri="https://zenodo.org/record/7840829/files/figure.png" pageId="25" pageNumber="26">16c</figureCitation>
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urn:lsid:zoobank.org:act:
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<materialsCitation id="3B1DD73DCC284C3DF7DFFE10FD43FD89" collectionCode="ZMHU, ZMHB" country="Malawi" county="Berlin" location="Natural History Museum" municipality="Biodiversity Research" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC284C3DF7DFFE10FF51FDAC" box="[151,207,511,537]" pageId="25" pageNumber="26">Type</typeStatus>
locality and repository:
<collectingCountry id="F3629DF0CC284C3DF6ACFE11FDA4FDAD" box="[484,570,510,536]" name="Malawi" pageId="25" pageNumber="26">Malawi</collectingCountry>
, the
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and
<location id="8EAA8BBBCC284C3DF48CFE11FBE0FDAD" LSID="urn:lsid:plazi:treatment:03DC6C76CC284C3EF7DFFE81FCB6F821:8EAA8BBBCC284C3DF48CFE11FBE0FDAD" box="[964,1150,510,536]" country="Malawi" county="Berlin" municipality="Biodiversity Research" name="Leibniz Institute" pageId="25" pageNumber="26">Leibniz Institute</location>
for
<location id="8EAA8BBBCC284C3DF3E6FE11FA82FDAD" LSID="urn:lsid:plazi:treatment:03DC6C76CC284C3EF7DFFE81FCB6F821:8EAA8BBBCC284C3DF3E6FE11FA82FDAD" box="[1198,1308,510,536]" country="Malawi" county="Berlin" municipality="Biodiversity Research" name="Evolution" pageId="25" pageNumber="26">Evolution</location>
and
<collectingMunicipality id="6BAE471ACC284C3DF21DFE11FECAFD89" pageId="25" pageNumber="26">Biodiversity Research</collectingMunicipality>
,
<collectingCounty id="62ABA5ECCC284C3DF617FDCDFE3BFD89" box="[351,421,546,572]" pageId="25" pageNumber="26">Berlin</collectingCounty>
(
<collectionCode id="ED6445A5CC284C3DF6FCFDCDFD95FD89" box="[436,523,546,572]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="25" pageNumber="26">ZMHU</collectionCode>
, formerly
<collectionCode id="ED6445A5CC284C3DF537FDCDFD4BFD89" box="[639,725,546,572]" pageId="25" pageNumber="26">ZMHB</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C36F8EEBCC284C3DF7DFFD85FE9CFD79" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BCADD60CC284C3DF7DFFD85FE9CFD79" blockId="25.[151,1437,618,2013]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF7DFFD85FE1DFD31" bold="true" box="[151,387,618,644]" pageId="25" pageNumber="26">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC284C3DF6C4FD85FE9CFD79" collectingDate="1996-10-19" collectionCode="ZMHB" collectorName="M. Nu &amp; W. Mey &amp; I. Lehmann &amp; D. Stuning" country="Malawi" elevation="800" location="Likabula" municipality="Southern Region" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Mulanje" typeStatus="holotype">
<typeStatus id="54CE63C2CC284C3DF6C4FD85FE6AFD31" box="[396,500,618,644]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC284C3DF50CFD85FD02FD31" box="[580,668,618,644]" name="Malawi" pageId="25" pageNumber="26">Malawi</collectingCountry>
,
<collectingMunicipality id="6BAE471ACC284C3DF5E2FD85FCF3FD31" box="[682,877,618,644]" pageId="25" pageNumber="26">Southern Region</collectingMunicipality>
,
<collectingRegion id="49B11382CC284C3DF432FD85FBA9FD31" box="[890,1079,618,644]" country="Malawi" name="Mulanje" pageId="25" pageNumber="26">Mulanje District</collectingRegion>
,
<location id="8EAA8BBBCC284C3DF30DFD85FB35FD31" LSID="urn:lsid:plazi:treatment:03DC6C76CC284C3EF7DFFE81FCB6F821:8EAA8BBBCC284C3DF30DFD85FB35FD31" box="[1093,1195,618,644]" country="Malawi" municipality="Southern Region" name="Likabula" pageId="25" pageNumber="26" stateProvince="Mulanje">Likabula</location>
,
<taxonomicName id="4C75A6E3CC284C3DF3F0FD85FAD2FD31" box="[1208,1356,618,644]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B9010172CC284C3DF3F0FD85FAD2FD31" box="[1208,1356,618,644]" italics="true" pageId="25" pageNumber="26">Brachystegia</emphasis>
</taxonomicName>
forest,
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<elevation id="00583A53CC284C3DF7DFFD61FF7AFD1C" box="[151,228,654,681]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FFCBFBA0CC284C3DF7BBFD61FE24FD1D" box="[243,442,654,680]" pageId="25" pageNumber="26" value="1996-10-19">
<collectingDate id="EF8F0248CC284C3DF7BBFD61FE24FD1D" box="[243,442,654,680]" pageId="25" pageNumber="26" value="1996-10-19">19 October 1996</collectingDate>
</date>
,
<collectorName id="2680B8B6CC284C3DF680FD60FD8BFD1C" box="[456,533,655,681]" pageId="25" pageNumber="26">M. Nu</collectorName>
&amp;
<collectorName id="2680B8B6CC284C3DF508FD60FD02FD1C" box="[576,668,655,681]" pageId="25" pageNumber="26">W. Mey</collectorName>
leg., genitalia slide number 20/092010
<collectorName id="2680B8B6CC284C3DF327FD60FB6AFD1D" box="[1135,1268,654,681]" pageId="25" pageNumber="26">I. Lehmann</collectorName>
&amp;
<collectorName id="2680B8B6CC284C3DF257FD60FA02FD1D" box="[1311,1436,654,681]" pageId="25" pageNumber="26">D. Stüning</collectorName>
(
<collectionCode id="ED6445A5CC284C3DF7D7FD5DFF68FD79" box="[159,246,690,716]" pageId="25" pageNumber="26">ZMHB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC284C3DF78FFD39FE3CF999" pageId="25" pageNumber="26" type="description">
<paragraph id="8BCADD60CC284C3DF78FFD39FD89FCE9" blockId="25.[151,1437,618,2013]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF78FFD39FEC7FD45" bold="true" box="[199,345,726,752]" pageId="25" pageNumber="26">Description.</emphasis>
Male.
<emphasis id="B9010172CC284C3DF6E1FD39FE78FD45" box="[425,486,726,752]" italics="true" pageId="25" pageNumber="26">Head</emphasis>
: Cream-coloured, shiny, long, dense hair-like scales between eyes; eyes brown with small black patches; antenna 0.54 length of forewing, bipectinate, with branches 4.5 width of shaft, branches and shaft covered with cream-coloured scales dorsally; antennal tips spatulate with two long scales, bending towards apex; labial palpi cream-coloured.
</paragraph>
<paragraph id="8BCADD60CC284C3DF78FFC89FE98FB71" blockId="25.[151,1437,618,2013]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF78FFC89FE86FC35" box="[199,280,870,896]" italics="true" pageId="25" pageNumber="26">Thorax</emphasis>
: Patagia and tegulae shiny, with long hair-like cream-coloured scales. A small cream-coloured crest on metathorax. Hindlegs yellow-ochre with fine hair-like scales, shiny; one pair of small tibial spurs present,
<emphasis id="B9010172CC284C3DF233FC64FA02FC11" box="[1403,1436,907,932]" italics="true" pageId="25" pageNumber="26">ca.</emphasis>
<quantity id="4C8D7085CC284C3DF7DFFC41FF70FC7C" box="[151,238,942,969]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="mm" value="0.7">0.7 mm</quantity>
long. Forewing length 12.0 mm (wingspan
<quantity id="4C8D7085CC284C3DF597FC41FCD6FC7D" box="[735,840,942,969]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="25" pageNumber="26" unit="mm" value="26.5">26.5 mm</quantity>
). Forewing upperside cream, glossy towards termen; straight terminal line of sepia from near apex to dorsum, not strongly bent towards base of wing at M
<subScript id="17F1DF25CC284C3DF3B3FC0AFA9DFC46" attach="left" box="[1275,1283,997,1011]" fontSize="6" pageId="25" pageNumber="26">3</subScript>
; CuA
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broad, ivory-yellow, edged sepia above; subterminal line sepia, extending from costa towards CuA
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and reduced to several striae near latter; striae of sepia along costa and on entire upperside of wing; veins cream, not distinctly coloured except sepia 1A+2A; veinlet in cell sepia; cilia long,
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, cream-coloured, shiny; lunules of sepia along termen. Forewing underside roughly scaled, ivory-yellow, glossy; costal margin with few cream-coloured striae, terminal line of same colour as striae. Hindwing upperside and cilia ivory-yellow, glossy; underside as in forewing but without striae.
</paragraph>
<paragraph id="8BCADD60CC284C3DF78FFB21FD27F9BD" blockId="25.[151,1437,618,2013]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF78FFB21FEAFFB5D" box="[199,305,1230,1256]" italics="true" pageId="25" pageNumber="26">Abdomen</emphasis>
: Mainly cream-coloured, shiny; abdominal tuft one-third length of abdomen. Genitalia (
<figureCitation id="134EC1E5CC284C3DF248FB21FAC2FB5D" box="[1280,1372,1230,1256]" captionStart="FIGURE 16" captionStartId="67.[151,250,1673,1697]" captionTargetBox="[159,1428,391,1636]" captionTargetId="figure-109@67.[151,1436,381,1649]" captionTargetPageId="67" captionText="FIGURE 16. a. Lukeniana madrandelei, paratype, male, DRC, Katanga Province, Lubumbashi; b. L. friederikebauerae, paratype, male, Zambia, Lusaka Province, Lusaka; c. L. chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; d. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; e. L. kammeri, holotype, male, Kenya, County Kiambu, Thika; f. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave." figureDoi="http://doi.org/10.5281/zenodo.7840829" httpUri="https://zenodo.org/record/7840829/files/figure.png" pageId="25" pageNumber="26">Fig. 16c</figureCitation>
) with uncus bearing pointed lobes, short setae ventrally, base of uncus strongly bent at middle; gnathos arms long (slightly longer than basal width of valva); valva rectangular, costa without setae; sacculus without setae; weakly-sclerotized projection densely setose, with rounded end and slightly longer than single thorn-like process below; latter hollow, narrow, with acuminate tip without setae; median sector of valva without setae on inner side; a rounded emargination extending between weakly-sclerotized projection and thorn-like process, emargination 30% of length of valva. Saccus short, triangular. Juxta with two acuminate tips, neither bearing a short process, between tips a deep, rectangular shaped emargination 60% of length of juxta. Phallus shorter than width of valva, not trumpet-like, slightly bent at middle, bilobate with a cleft on each end.
</paragraph>
<paragraph id="8BCADD60CC284C3DF78FF9FDFE3CF999" blockId="25.[151,1437,618,2013]" box="[199,418,1554,1580]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF78FF9FDFEBBF999" bold="true" box="[199,293,1554,1580]" pageId="25" pageNumber="26">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC284C3DF78FF9D9FE5CF8C5" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8BCADD60CC284C3DF78FF9D9FE5CF8C5" blockId="25.[151,1437,618,2013]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF78FF9D9FEDCF9E5" bold="true" box="[199,322,1590,1616]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC284C3DF600F9D9FDACF9E5" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[328,562,1590,1616]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chapmani">
<emphasis id="B9010172CC284C3DF600F9D9FDACF9E5" box="[328,562,1590,1616]" italics="true" pageId="25" pageNumber="26">Lukeniana chapmani</emphasis>
</taxonomicName>
bears similarities to
<taxonomicName id="4C75A6E3CC284C3DF451F9D8FC5EF9E5" authority="Lehmann &amp; Zahiri &amp; Husemann, 2023" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[793,960,1590,1616]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="stevecollinsi" status="sp. nov.">
<emphasis id="B9010172CC284C3DF451F9D8FC5EF9E5" box="[793,960,1590,1616]" italics="true" pageId="25" pageNumber="26">L. stevecollinsi</emphasis>
</taxonomicName>
<emphasis id="B9010172CC284C3DF48EF9D9FB81F9E5" bold="true" box="[966,1055,1590,1616]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A232BC09CC284C3DF48EF9D9FB81F9E5" box="[966,1055,1590,1616]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, including the light cream appearance of the forewings, rectangular valvae, and a sharply bent edge of the uncus at its middle.
<taxonomicName id="4C75A6E3CC284C3DF3F8F9B5FA02F9C1" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1200,1436,1626,1652]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chapmani">
<emphasis id="B9010172CC284C3DF3F8F9B5FA02F9C1" box="[1200,1436,1626,1652]" italics="true" pageId="25" pageNumber="26">Lukeniana chapmani</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C75A6E3CC284C3DF68AF990FDF4F92D" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[450,618,1662,1688]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="stevecollinsi">
<emphasis id="B9010172CC284C3DF68AF990FDF4F92D" box="[450,618,1662,1688]" italics="true" pageId="25" pageNumber="26">L. stevecollinsi</emphasis>
</taxonomicName>
by the more delicate appearance, lines and striae on the forewing upperside and by four diagnostic characters in the male genitalia: (i) the uncus has pointed lobes, a unique character state; (ii) the phallus is the shortest among this genus, one-third shorter than the width of valva, particularly if compared to
<taxonomicName id="4C75A6E3CC284C3DF7FDF904FEC3F8B1" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[181,349,1770,1796]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="stevecollinsi">
<emphasis id="B9010172CC284C3DF7FDF904FEC3F8B1" box="[181,349,1770,1796]" italics="true" pageId="25" pageNumber="26">L. stevecollinsi</emphasis>
</taxonomicName>
, where the phallus is almost one-third longer than the width of valva; (iii) the thorn-like process is well developed, narrow and looks delicate as in
<taxonomicName id="4C75A6E3CC284C3DF584F8E0FCE9F89D" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[716,887,1806,1832]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mikerobertsi">
<emphasis id="B9010172CC284C3DF584F8E0FCE9F89D" box="[716,887,1806,1832]" italics="true" pageId="25" pageNumber="26">L. mikerobertsi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75A6E3CC284C3DF4E5F8E0FB1AF89D" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[941,1156,1806,1832]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="georgeadamsoni">
<emphasis id="B9010172CC284C3DF4E5F8E0FB1AF89D" box="[941,1156,1806,1832]" italics="true" pageId="25" pageNumber="26">L. georgeadamsoni</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C75A6E3CC284C3DF3BCF8E0FA02F89D" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1268,1436,1806,1832]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="stevecollinsi">
<emphasis id="B9010172CC284C3DF3BCF8E0FA02F89D" box="[1268,1436,1806,1832]" italics="true" pageId="25" pageNumber="26">L. stevecollinsi</emphasis>
</taxonomicName>
the thorn-like process is very thick; (iv) the sacculus is narrow in
<taxonomicName id="4C75A6E3CC284C3DF4CCF8DCFB8EF8F9" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[900,1040,1842,1868]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chapmani">
<emphasis id="B9010172CC284C3DF4CCF8DCFB8EF8F9" box="[900,1040,1842,1868]" italics="true" pageId="25" pageNumber="26">L. chapmani</emphasis>
</taxonomicName>
, but very broad in
<taxonomicName id="4C75A6E3CC284C3DF3A4F8DCFA0BF8F9" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1260,1429,1842,1868]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="stevecollinsi">
<emphasis id="B9010172CC284C3DF3A4F8DCFA0BF8F9" box="[1260,1429,1842,1868]" italics="true" pageId="25" pageNumber="26">L. stevecollinsi</emphasis>
</taxonomicName>
, 40% of the width of valva.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC284C3EF78FF895FE77F909" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BCADD60CC284C3DF78FF895FAABF869" blockId="25.[151,1437,618,2013]" pageId="25" pageNumber="26">
<emphasis id="B9010172CC284C3DF78FF895FEFFF821" bold="true" box="[199,353,1914,1940]" pageId="25" pageNumber="26">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC284C3DF62FF895FDCFF821" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[359,593,1914,1940]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chapmani">
<emphasis id="B9010172CC284C3DF62FF895FDCFF821" box="[359,593,1914,1940]" italics="true" pageId="25" pageNumber="26">Lukeniana chapmani</emphasis>
</taxonomicName>
is known from the western slopes of Mount
<collectingRegion id="49B11382CC284C3DF377F895FB02F821" box="[1087,1180,1914,1940]" country="Malawi" name="Mulanje" pageId="25" pageNumber="26">Mulanje</collectingRegion>
(also Mount Mlanje or Mulanji) located in the
<collectingCountry id="F3629DF0CC284C3DF6D1F871FE71F80D" box="[409,495,1950,1976]" name="Malawi" pageId="25" pageNumber="26">Malawi</collectingCountry>
Rift of the EARS in southeastern
<collectingCountry id="F3629DF0CC284C3DF42DF871FC25F80D" box="[869,955,1950,1976]" name="Malawi" pageId="25" pageNumber="26">Malawi</collectingCountry>
and
<emphasis id="B9010172CC284C3DF4A6F870FB8EF80D" box="[1006,1040,1951,1976]" italics="true" pageId="25" pageNumber="26">ca.</emphasis>
<quantity id="4C8D7085CC284C3DF35DF871FBF7F80D" box="[1045,1129,1950,1976]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="26" unit="km" value="600.0">600 km</quantity>
southeast of Thazima Gate, the
<typeStatus id="54CE63C2CC284C3DF788F82CFF6EF868" box="[192,240,1987,2013]" pageId="25" pageNumber="26">type</typeStatus>
locality of
<taxonomicName id="4C75A6E3CC284C3DF639F82CFD86F869" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[369,536,1986,2012]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="stevecollinsi">
<emphasis id="B9010172CC284C3DF639F82CFD86F869" box="[369,536,1986,2012]" italics="true" pageId="25" pageNumber="26">L. stevecollinsi</emphasis>
</taxonomicName>
. This species is defined here as an Afromontane near-endemic species.
</paragraph>
<caption id="DF0A8DE8CC2B4C3EF7DFFA23FCB2F9C1" ID-DOI="http://doi.org/10.5281/zenodo.7840798" ID-Zenodo-Dep="7840798" httpUri="https://zenodo.org/record/7840798/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1484,1508]" targetBox="[151,1436,181,1460]" targetPageId="26">
<paragraph id="8BCADD60CC2B4C3EF7DFFA23FCB2F9C1" blockId="26.[151,1437,1484,1653]" pageId="26" pageNumber="27">
<emphasis id="B9010172CC2B4C3EF7DFFA23FD88FA51" bold="true" box="[151,534,1484,1508]" pageId="26" pageNumber="27">
FIGURE 5. a.
<taxonomicName id="4C75A6E3CC2B4C3EF67DFA23FD88FA51" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[309,534,1484,1508]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chapmani">
<emphasis id="B9010172CC2B4C3EF67DFA23FD88FA51" bold="true" box="[309,534,1484,1508]" italics="true" pageId="26" pageNumber="27">Lukeniana chapmani</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula;
<emphasis id="B9010172CC2B4C3EF3BAFA23FF45F9BD" bold="true" pageId="26" pageNumber="27">
b.
<taxonomicName id="4C75A6E3CC2B4C3EF245FA23FF45F9BD" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2B4C3EF245FA23FF45F9BD" bold="true" italics="true" pageId="26" pageNumber="27">L. michaelgrzimeki</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella;
<emphasis id="B9010172CC2B4C3EF4DCFA1FFBE9F9BD" bold="true" box="[916,1143,1520,1544]" pageId="26" pageNumber="27">
c.
<taxonomicName id="4C75A6E3CC2B4C3EF4E5FA1FFBE9F9BD" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[941,1143,1520,1544]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2B4C3EF4E5FA1FFBE9F9BD" bold="true" box="[941,1143,1520,1544]" italics="true" pageId="26" pageNumber="27">L. michaelgrzimeki</emphasis>
</taxonomicName>
</emphasis>
, paratype, female, Tanzania, Arusha Region, Mount Meru, Momella;
<emphasis id="B9010172CC2B4C3EF565F9FBFD6DF999" bold="true" box="[557,755,1556,1580]" pageId="26" pageNumber="27">
d.
<taxonomicName id="4C75A6E3CC2B4C3EF50FF9FBFD6DF999" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[583,755,1556,1580]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="timdavenporti">
<emphasis id="B9010172CC2B4C3EF50FF9FBFD6DF999" bold="true" box="[583,755,1556,1580]" italics="true" pageId="26" pageNumber="27">L. timdavenporti</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave;
<emphasis id="B9010172CC2B4C3EF6ECF9D7FDCCF9E5" bold="true" box="[420,594,1592,1616]" pageId="26" pageNumber="27">
e.
<taxonomicName id="4C75A6E3CC2B4C3EF6F5F9D7FDCCF9E5" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[445,594,1592,1616]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2B4C3EF6F5F9D7FDCCF9E5" bold="true" box="[445,594,1592,1616]" italics="true" pageId="26" pageNumber="27">L. mzuzuensis</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo;
<emphasis id="B9010172CC2B4C3EF3A4F9D7FA08F9E5" bold="true" box="[1260,1430,1592,1616]" pageId="26" pageNumber="27">
f.
<taxonomicName id="4C75A6E3CC2B4C3EF249F9D7FA08F9E5" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1281,1430,1592,1616]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2B4C3EF249F9D7FA08F9E5" bold="true" box="[1281,1430,1592,1616]" italics="true" pageId="26" pageNumber="27">L. mzuzuensis</emphasis>
</taxonomicName>
</emphasis>
, paratype, female, Malawi, Northern Region, Mzuzu, Nkhorongo.
</paragraph>
</caption>
<paragraph id="8BCADD60CC2B4C3EF78FF94DFE77F909" blockId="26.[151,1436,1698,1941]" box="[199,489,1698,1724]" pageId="26" pageNumber="27">
<emphasis id="B9010172CC2B4C3EF78FF94DFEB4F909" bold="true" box="[199,298,1698,1724]" pageId="26" pageNumber="27">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2B4C3EF78FF929FCB6F821" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8BCADD60CC2B4C3EF78FF929FCB6F821" blockId="26.[151,1436,1698,1941]" pageId="26" pageNumber="27">
<emphasis id="B9010172CC2B4C3EF78FF929FED0F955" bold="true" box="[199,334,1734,1760]" pageId="26" pageNumber="27">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC2B4C3EF61EF929FDDDF955" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[342,579,1734,1760]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chapmani">
<emphasis id="B9010172CC2B4C3EF61EF929FDDDF955" box="[342,579,1734,1760]" italics="true" pageId="26" pageNumber="27">Lukeniana chapmani</emphasis>
</taxonomicName>
is named for the late forest officer James Douglas “Jim” Chapman (1918 2011) who joined the
<collectingCountry id="F3629DF0CC2B4C3EF6D9F905FD99F8B1" box="[401,519,1770,1796]" name="Malawi" pageId="26" pageNumber="27">Nyasaland</collectingCountry>
Department of Forestry in 1952 and served for 13 years. Chapman collected not only a large number of plant specimens in most of the evergreen forests of
<collectingCountry id="F3629DF0CC2B4C3EF4A8F8E1FBC8F89D" box="[992,1110,1806,1832]" name="Malawi" pageId="26" pageNumber="27">Nyasaland</collectingCountry>
(now
<collectingCountry id="F3629DF0CC2B4C3EF3D6F8E1FB6BF89D" box="[1182,1269,1806,1832]" name="Malawi" pageId="26" pageNumber="27">Malawi</collectingCountry>
) for more than three decades, but also described most of their constituent species. James Chapman and his wife Elizabeth Grey collected extensively on and around Mount
<collectingRegion id="49B11382CC2B4C3EF5CCF8B9FD7FF8C5" box="[644,737,1878,1904]" country="Malawi" name="Mulanje" pageId="26" pageNumber="27">Mulanje</collectingRegion>
in 19851988 and added much new knowledge to the flora of this mountain (Hazel Chapman pers. com. to I.L. in 2014).
</paragraph>
</subSubSection>
</treatment>
</document>