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<document id="CBB5E0549C5008C5E3E4506D024DF192" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC204C36F7DFFF78FB86FCA0" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana rajaeii Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="318C7422-38B6-47CD-ACA9-57EA7B67CB32" docUuidSource="ZooBank" docVersion="3" lastPageNumber="19" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="18" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="EE6B0B1B9619433D9DD25097A6CFD43D">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="A85CDE25F8C93F3057FE54AE0AA1FEF3">Author</mods:roleTerm>
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<mods:namePart id="836523BA9338936161980EC2AA1FAFD0">Lehmann, Ingo</mods:namePart>
<mods:nameIdentifier id="110FF39BC32A5B89483F30E7A4BCBA20" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:namePart id="D8820FD409C801932BAC135BE758DB46">Zahiri, Reza</mods:namePart>
<mods:nameIdentifier id="1CE5E68A1AA2B99AAE8A6347F55ED8B4" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3E70060CDFB751A2C66E59A40B9806A5">Husemann, Martin</mods:namePart>
<mods:nameIdentifier id="22EAB7E27CC636046351220A6E886B69" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="6E4A09C8B76E38F499406DB8568F734F">Zootaxa</mods:title>
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<mods:date id="3929E3974215A9D78561F7BAB6D27976">2023</mods:date>
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<mods:number id="D3CC5D17EAF86B7DF71F29F2497DC6BA">2023-04-18</mods:number>
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<paragraph id="8BCADD60CC204C35F7DFFF78FCFDFF04" blockId="17.[151,977,151,250]" box="[151,867,151,178]" pageId="17" pageNumber="18">
<heading id="D0826A0CCC204C35F7DFFF78FCFDFF04" bold="true" box="[151,867,151,178]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<emphasis id="B9010172CC204C35F7DFFF78FCFDFF04" bold="true" box="[151,867,151,178]" pageId="17" pageNumber="18">
<taxonomicName id="4C75A6E3CC204C35F7DFFF78FD63FF04" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,765,151,178]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rajaeii" status="sp. nov.">
<emphasis id="B9010172CC204C35F7DFFF78FEEFFF04" bold="true" box="[151,369,151,178]" italics="true" pageId="17" pageNumber="18">Lukeniana rajaeii</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC204C35F44DFF78FCFDFF04" box="[773,867,151,177]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC204C35F7DFFF54FAA9FEF4" pageId="17" pageNumber="18" type="description">
<paragraph id="8BCADD60CC204C35F7DFFF54FEC7FF60" blockId="17.[151,977,151,250]" box="[151,345,187,213]" pageId="17" pageNumber="18">
<figureCitation id="134EC1E5CC204C35F7DFFF54FF74FF60" box="[151,234,187,213]" captionStart="FIGURE 3" captionStartId="19.[151,250,1753,1777]" captionTargetBox="[151,1436,381,1729]" captionTargetId="figure-103@19.[151,1436,381,1729]" captionTargetPageId="19" captionText="FIGURE 3. a. Lukeniana enaiposha, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha; b. L. obliqualinea, male, Kenya, County Nairobi, Nairobi; c. L. obliqualinea, female, Kenya, County Nairobi, Loresho Ridge; d. L. raymondrevellii, holotype, male, Namibia, Otjozondjupa Region, Ozombanda; e. L. rajaeii, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole; f. L. tubiraensis, holotype, male, Tanzania, Kigoma Region, Tubira Forest." figureDoi="http://doi.org/10.5281/zenodo.7840792" httpUri="https://zenodo.org/record/7840792/files/figure.png" pageId="17" pageNumber="18">Figs 3e</figureCitation>
,
<figureCitation id="134EC1E5CC204C35F7BFFF54FEBDFF60" box="[247,291,187,213]" captionStart="FIGURE 10" captionStartId="60.[151,250,1754,1778]" captionTargetBox="[151,1436,677,1731]" captionTargetId="figure-255@60.[151,1436,669,1731]" captionTargetPageId="60" captionText="FIGURE 10. An intact CuP vein on the forewing is a plesiomorphy (cf. Fig. 9) and occurs only in basal Metarbelidae in northeastern parts of the Republic of South Africa, mainly on the Southern African Plateau, and on the plateau area of Mount Mulanje, only 90 km to the North of the Zambezi River (Lehmann 2019b). Figured are examples of apomorphies in regard to the CuP on forewing (red arrows): a. CuP absent, CuP fold is weak but not continuous in Lukeniana michaelgrzimeki, female, paratype; b. The CuP fold is almost entirely absent in L. rajaeii, male, paratype; c. Zambezia darrelplowesi, female, holotype and d. Z. madambae, male, paratype, both have a strong CuP fold, continuous and partly sclerotized. Images were taken by Dr. Hossein Rajaei (SMNS) in November 2015." figureDoi="http://doi.org/10.5281/zenodo.7840810" httpUri="https://zenodo.org/record/7840810/files/figure.png" pageId="17" pageNumber="18">10b</figureCitation>
,
<figureCitation id="134EC1E5CC204C35F667FF54FEC7FF60" box="[303,345,187,213]" captionStart="FIGURE 13" captionStartId="64.[151,250,1283,1307]" captionTargetBox="[163,1424,181,1249]" captionTargetId="figure-28@64.[151,1436,166,1259]" captionTargetPageId="64" captionText="FIGURE 13. a. Lukeniana enaiposha, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha; b. L. obliqualinea, male, Kenya, County Nairobi, Nairobi; c. L. raymondrevellii, holotype, male, Namibia, Otjozondjupa Region, Ozombanda; d. L. rajaeii, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole; e. L. tubiraensis, holotype, male, Tanzania, Kigoma Region, Tubira Forest; f. L. lutztoepferi, holotype, male, Kenya, County Nairobi, Ololua Forest." figureDoi="http://doi.org/10.5281/zenodo.7840820" httpUri="https://zenodo.org/record/7840820/files/figure.png" pageId="17" pageNumber="18">13d</figureCitation>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="8BCADD60CC204C35F7DFFEC7FAA9FEF4" blockId="17.[151,1335,295,322]" box="[151,1335,295,322]" pageId="17" pageNumber="18">
<materialsCitation id="3B1DD73DCC204C35F7DFFEC7FAADFEF4" box="[151,1331,295,322]" collectionCode="SMNS" country="Ethiopia" location="State Museum of Natural History Stuttgart" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC204C35F7DFFEC7FF51FEF7" box="[151,207,296,322]" pageId="17" pageNumber="18">Type</typeStatus>
locality and repository:
<collectingCountry id="F3629DF0CC204C35F6A8FEC8FDDCFEF4" box="[480,578,295,321]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
, the
<location id="8EAA8BBBCC204C35F53FFEC8FBC8FEF4" LSID="urn:lsid:plazi:treatment:03DC6C76CC204C36F7DFFF78FB86FCA0:8EAA8BBBCC204C35F53FFEC8FBC8FEF4" box="[631,1110,295,322]" country="Ethiopia" name="State Museum of Natural History Stuttgart" pageId="17" pageNumber="18">State Museum of Natural History Stuttgart</location>
,
<collectingCountry id="F3629DF0CC204C35F329FEC8FB57FEF4" box="[1121,1225,295,321]" name="Germany" pageId="17" pageNumber="18">Germany</collectingCountry>
(
<collectionCode id="ED6445A5CC204C35F391FEC8FAB5FEF4" box="[1241,1323,295,321]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="18">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C36F8EEBCC204C35F7DFFE80FA1DFD88" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BCADD60CC204C35F7DFFE80FA1DFD88" blockId="17.[151,1437,367,2014]" pageId="17" pageNumber="18">
<emphasis id="B9010172CC204C35F7DFFE80FE1DFE3C" bold="true" box="[151,387,367,393]" pageId="17" pageNumber="18">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC204C35F6C4FE80FCA8FE40" collectingDate="1960-02-23" collectingDateMax="1960-03-05" collectingDateMin="1960-02-23" collectionCode="SMNS" collectorName="W. Richter" country="Ethiopia" county="Peoples' Region" elevation="2200" location="Gamo Gofa" municipality="Gamu Gofa Province" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch" typeStatus="holotype">
<typeStatus id="54CE63C2CC204C35F6C4FE80FE6AFE3C" box="[396,500,367,393]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC204C35F50DFE80FD39FE3C" box="[581,679,367,393]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
,
<collectingRegion id="49B11382CC204C35F5FCFE80FCE3FE3C" box="[692,893,367,393]" country="Ethiopia" name="Ye Debub Biheroch Bihereseboch na Hizboch" pageId="17" pageNumber="18">Southern Nations</collectingRegion>
,
<collectingRegion id="49B11382CC204C35F4C2FE80FB82FE3C" box="[906,1052,367,393]" country="Ethiopia" name="Nationalities" pageId="17" pageNumber="18">Nationalities</collectingRegion>
, and
<collectingCounty id="62ABA5ECCC204C35F314FE80FA8BFE3C" box="[1116,1301,367,393]" pageId="17" pageNumber="18">Peoples Region</collectingCounty>
(during the 1960s until 1995
<collectingMunicipality id="6BAE471ACC204C35F629FE7CFDC9FE18" box="[353,599,403,429]" pageId="17" pageNumber="18">Gamu Gofa Province</collectingMunicipality>
), today
<location id="8EAA8BBBCC204C35F5FBFE7CFCA7FE18" LSID="urn:lsid:plazi:treatment:03DC6C76CC204C36F7DFFF78FB86FCA0:8EAA8BBBCC204C35F5FBFE7CFCA7FE18" box="[691,825,403,429]" country="Ethiopia" county="Peoples' Region" municipality="Gamu Gofa Province" name="Gamo Gofa" pageId="17" pageNumber="18" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch">Gamo Gofa</location>
administrative zone, formerly
<location id="8EAA8BBBCC204C35F3D3FE7CFAAEFE18" LSID="urn:lsid:plazi:treatment:03DC6C76CC204C36F7DFFF78FB86FCA0:8EAA8BBBCC204C35F3D3FE7CFAAEFE18" box="[1179,1328,403,429]" country="Ethiopia" county="Peoples' Region" municipality="Gamu Gofa Province" name="Semien Omo" pageId="17" pageNumber="18" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch">Semien Omo</location>
(=
<location id="8EAA8BBBCC204C35F213FE7CFF4FFE64" LSID="urn:lsid:plazi:treatment:03DC6C76CC204C36F7DFFF78FB86FCA0:8EAA8BBBCC204C35F213FE7CFF4FFE64" country="Ethiopia" county="Peoples' Region" municipality="Gamu Gofa Province" name="North Omo" pageId="17" pageNumber="18" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch">North Omo</location>
) zone,
<location id="8EAA8BBBCC204C35F66AFE58FEEFFE64" LSID="urn:lsid:plazi:treatment:03DC6C76CC204C36F7DFFF78FB86FCA0:8EAA8BBBCC204C35F66AFE58FEEFFE64" box="[290,369,439,465]" country="Ethiopia" county="Peoples' Region" municipality="Gamu Gofa Province" name="Gidole" pageId="17" pageNumber="18" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch">Gidole</location>
, 0534 N (correct: 05380539 N), 3726 E (correct: 3721 E),
<quantity id="4C8D7085CC204C35F378FE58FB10FE67" box="[1072,1166,439,466]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="18" unit="m" value="2200.0">
<elevation id="00583A53CC204C35F378FE58FB10FE67" box="[1072,1166,439,466]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="18" unit="m" value="2200.0">2,200 m</elevation>
</quantity>
,
<date id="FFCBFBA0CC204C35F3D2FE58FF4CFE40" pageId="17" pageNumber="18" value="1960-02-23" valueMax="1960-03-05" valueMin="1960-02-23">
<collectingDate id="EF8F0248CC204C35F3D2FE58FF4CFE40" pageId="17" pageNumber="18" value="1960-02-23" valueMax="1960-03-05" valueMin="1960-02-23">23 February05 March 1960</collectingDate>
</date>
,
<collectorName id="2680B8B6CC204C35F795FE33FECBFE40" box="[221,341,475,502]" pageId="17" pageNumber="18">W. Richter</collectorName>
leg., no genitalia dissection done (
<collectionCode id="ED6445A5CC204C35F594FE34FCB0FE40" box="[732,814,475,501]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="18">SMNS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD73DCC204C35F409FE33FC8EFDAC" collectingDate="1960-02-23" collectingDateMax="1960-03-05" collectingDateMin="1960-02-23" collectionCode="SMNS" collectorName="I. Lehmann" country="Ethiopia" county="Peoples' Region" elevation="2200" location="Gamo Gofa" municipality="Gamu Gofa Province" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch" typeStatus="paratype">
<typeStatus id="54CE63C2CC204C35F409FE33FC2CFE43" box="[833,946,476,502]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7316E9CC204C35F4F5FE33FBBBFE40" box="[957,1061,475,502]" pageId="17" pageNumber="18" type="male">one male</specimenCount>
,
<collectingCountry id="F3629DF0CC204C35F379FE34FB0DFE40" box="[1073,1171,475,501]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
, same data as holotype, genitalia slide number 23/052015
<collectorName id="2680B8B6CC204C35F568FDEFFD3AFDAC" box="[544,676,511,538]" pageId="17" pageNumber="18">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC204C35F5FEFE10FC96FDAC" box="[694,776,511,537]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="18">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC204C35F456FDEFFAE1FD88" collectingDate="1960-02-23" collectingDateMax="1960-03-05" collectingDateMin="1960-02-23" collectionCode="SMNS" collectorName="I. Lehmann" country="Ethiopia" county="Peoples' Region" elevation="2200" location="Gamo Gofa" municipality="Gamu Gofa Province" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Ye Debub Biheroch Bihereseboch na Hizboch" typeStatus="paratype">
<specimenCount id="9D7316E9CC204C35F456FDEFFC18FDAC" box="[798,902,511,538]" pageId="17" pageNumber="18" type="male">one male</specimenCount>
(the largest and most intact specimen and described/figured herein),
<collectingCountry id="F3629DF0CC204C35F6EDFDCCFD99FD88" box="[421,519,547,573]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
, same data as holotype, genitalia slide number 26/082015
<collectorName id="2680B8B6CC204C35F3DDFDCBFA88FD88" box="[1173,1302,547,574]" pageId="17" pageNumber="18">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC204C35F26DFDCCFAE9FD88" box="[1317,1399,547,573]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="18">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC204C35F78FFDA8FE3CF908" pageId="17" pageNumber="18" type="description">
<paragraph id="8BCADD60CC204C35F78FFDA8FB5BFD78" blockId="17.[151,1437,367,2014]" pageId="17" pageNumber="18">
<emphasis id="B9010172CC204C35F78FFDA8FECFFDD4" bold="true" box="[199,337,583,609]" pageId="17" pageNumber="18">Description</emphasis>
(based on largest
<typeStatus id="54CE63C2CC204C35F56CFDA8FD14FDD4" box="[548,650,583,609]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
). Male.
<emphasis id="B9010172CC204C35F5AFFDA8FCBAFDD4" box="[743,804,583,609]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
: Honey-yellow, shiny, with long hair-like scales on the lower part of the frons; eyes vinaceous brown with small black spots; antenna unusually long (0.50-0.53 length of forewing), bipectinate, with branches 4.5 width of shaft, branches and shaft covered with cream-coloured scales dorsally; antennal tips with slightly longer scales, bent towards apex; labial palpi honey yellow.
</paragraph>
<paragraph id="8BCADD60CC204C35F78FFD38FC05FBC8" blockId="17.[151,1437,367,2014]" pageId="17" pageNumber="18">
<emphasis id="B9010172CC204C35F78FFD38FE86FD44" box="[199,280,727,753]" italics="true" pageId="17" pageNumber="18">Thorax</emphasis>
: Patagia and tegulae with shiny, long hair-like scales of honey-yellow mixed with ochre and cream with few small patches of sepia. Metathorax with small crest of cream-coloured scales. Hindlegs yellow ochre mixed with sepia, with fine shiny hair-like scales; one pair of narrow tibial spurs present, outer spur 1.0 mm, inner spur
<quantity id="4C8D7085CC204C35F231FCF0FF58FCEB" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="mm" value="0.9">0.9 mm</quantity>
. Forewing length of
<typeStatus id="54CE63C2CC204C35F6FCFCACFD88FCE8" box="[436,534,835,861]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
15.0 mm (wingspan
<quantity id="4C8D7085CC204C35F442FCACFCEDFCE8" box="[778,883,835,862]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="17" pageNumber="18" unit="mm" value="31.5">31.5 mm</quantity>
), of
<typeStatus id="54CE63C2CC204C35F4E0FCABFB8DFCEB" box="[936,1043,836,862]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
<quantity id="4C8D7085CC204C35F353FCACFB1FFCEB" box="[1051,1153,835,862]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="17" pageNumber="18" unit="mm" value="15.5">15.5 mm</quantity>
and 14.0 mm (wingspan 32.5 and
<quantity id="4C8D7085CC204C35F648FC88FEF7FC34" box="[256,361,871,898]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="17" pageNumber="18" unit="mm" value="29.5">29.5 mm</quantity>
). Forewing upperside light honey yellow, not glossy; a broad white band below CuA
<subScript id="17F1DF25CC204C35F261FC95FAAFFC3D" attach="left" box="[1321,1329,890,904]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
, twice as broad as sepia band above, extending from near base of forewing to CuA
<subScript id="17F1DF25CC204C35F4ABFC71FC75FC19" attach="left" box="[995,1003,926,940]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
and dorsum; costa from base of wing along half its length honey-yellow with few weak striae of sepia; a faded terminal line almost parallel along termen, subterminal line and postmedial line dark ochre, almost continuous from costa to CuA
<subScript id="17F1DF25CC204C35F326FC09FBE8FC41" attach="left" box="[1134,1142,998,1012]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
and slightly bent towards base of wing between R
<subScript id="17F1DF25CC204C35F6EEFBE6FE30FBA2" attach="left" box="[422,430,1033,1047]" fontSize="6" pageId="17" pageNumber="18">5</subScript>
and M
<subScript id="17F1DF25CC204C35F6B7FBE5FD99FBAD" attach="left" box="[511,519,1034,1048]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
; veins not distinctly coloured; cilia long,
<quantity id="4C8D7085CC204C35F495FC18FBABFBA7" box="[989,1077,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
, cream-coloured, shiny; lunules faded or absent along termen. Underside of forewing rough-scaled, cream-coloured, without striae, glossy; only costal margin darker, honey-yellow mixed with few sepia striae. Hindwing upperside and cilia light cream, glossy,
<quantity id="4C8D7085CC204C35F7DFFB8CFF73FBCB" box="[151,237,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
long. Underside light cream, costa light ochre without striae.
</paragraph>
<paragraph id="8BCADD60CC204C35F78FFB68FC5CF92C" blockId="17.[151,1437,367,2014]" pageId="17" pageNumber="18">
<emphasis id="B9010172CC204C35F78FFB68FEAFFB14" box="[199,305,1159,1185]" italics="true" pageId="17" pageNumber="18">Abdomen</emphasis>
: Mainly honey-yellow mixed with cream, shiny; abdominal tuft short and less than one-third of abdomen length. Genitalia (
<figureCitation id="134EC1E5CC204C35F6F2FB44FD85FB70" box="[442,539,1195,1221]" captionStart="FIGURE 13" captionStartId="64.[151,250,1283,1307]" captionTargetBox="[163,1424,181,1249]" captionTargetId="figure-28@64.[151,1436,166,1259]" captionTargetPageId="64" captionText="FIGURE 13. a. Lukeniana enaiposha, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha; b. L. obliqualinea, male, Kenya, County Nairobi, Nairobi; c. L. raymondrevellii, holotype, male, Namibia, Otjozondjupa Region, Ozombanda; d. L. rajaeii, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole; e. L. tubiraensis, holotype, male, Tanzania, Kigoma Region, Tubira Forest; f. L. lutztoepferi, holotype, male, Kenya, County Nairobi, Ololua Forest." figureDoi="http://doi.org/10.5281/zenodo.7840820" httpUri="https://zenodo.org/record/7840820/files/figure.png" pageId="17" pageNumber="18">Fig. 13d</figureCitation>
) one of the largest of any congener. Uncus subrectangular, outer edge slightly C-shaped near basal edge, inner edge rounded at base, lobes densely covered with short setae ventrally, basal edge of uncus bent at middle; gnathos arms slightly longer than width of valva, not touching coastal margin of valva, bent towards uncus and connected posteriorly with a thin membrane, probably not chitinized; valva broadly rectangular, costa nearly straight but bent upward at middle, semi-transtilla subrectangular, without setae; sacculus with few short setae; weakly-sclerotized projection with short setae, broad rounded tip slightly longer than short thorn-like process below; thorn-like process appearing reduced and slightly curved upward, hollow, tip rounded, no dots, without tiny setae, its base 0.7 as broad as weakly-sclerotized projection above, and without a second thorn-like process; median sector of valva with very few short setae on inner side; short, broad, ovoid emargination extending between weakly-sclerotized projection and thorn-like process,
<emphasis id="B9010172CC204C35F42AFA1FFC1DF9BC" box="[866,899,1520,1545]" italics="true" pageId="17" pageNumber="18">ca.</emphasis>
25% length of valva; ventral side of valva not bent at middle. Saccus half as large as juxta, finger-shaped with a rounded tip; vinculum very broad, plate-like, opposite saccus (plate-like structure 12 x larger than saccus). Juxta narrow, tall (1.5 as tall as width of juxta) with two acuminate tips, emargination between tips very deep, 90% length of juxta. Phallus short, as long as width of valva, slightly trumpet-like, bent near middle, bilobed with a cleft on each end.
</paragraph>
<paragraph id="8BCADD60CC204C35F78FF94CFE3CF908" blockId="17.[151,1437,367,2014]" box="[199,418,1699,1725]" pageId="17" pageNumber="18">
<emphasis id="B9010172CC204C35F78FF94CFEBBF908" bold="true" box="[199,293,1699,1725]" pageId="17" pageNumber="18">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC204C36F78FF928FDA3FE64" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BCADD60CC204C36F78FF928FDA3FE64" blockId="17.[151,1437,367,2014]" lastBlockId="18.[151,1437,151,789]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B9010172CC204C35F78FF928FEDCF954" bold="true" box="[199,322,1735,1761]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC204C35F600F928FD91F954" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[328,527,1735,1761]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC204C35F600F928FD91F954" box="[328,527,1735,1761]" italics="true" pageId="17" pageNumber="18">Lukeniana rajaeii</emphasis>
</taxonomicName>
has two unique character states: each lobe of the uncus has a nearly equally broad basal edge and upper edge and hence, the lobe has an rectangular shape; and the upper width of the uncus lobe is the broadest, if compared to all congeners, with
<emphasis id="B9010172CC204C35F5C3F8FFFD32F89C" box="[651,684,1808,1833]" italics="true" pageId="17" pageNumber="18">ca.</emphasis>
80% of width of valva in ventral view. Other diagnostic characters include: the thorn-like process is thick and much reduced in its length, it is the shortest in the genus; and the thornlike process has a broader base, but strongly reduced since it is only 0.7 as broad as the weakly-sclerotized projection above, hence this base is the smallest among the genus.
<taxonomicName id="4C75A6E3CC204C35F466F893FC62F820" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[814,1020,1915,1941]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="raymondrevellii">
<emphasis id="B9010172CC204C35F466F893FC62F820" box="[814,1020,1915,1941]" italics="true" pageId="17" pageNumber="18">L. raymondrevellii</emphasis>
</taxonomicName>
shares character states with
<taxonomicName id="4C75A6E3CC204C35F27DF893FA03F820" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1333,1437,1916,1941]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC204C35F27DF893FA03F820" box="[1333,1437,1916,1941]" italics="true" pageId="17" pageNumber="18">L. rajaeii</emphasis>
</taxonomicName>
due to the rectangular shape of the uncus and valvae, and due to the thorn-like process that is less pronounced, representing a somewhat transitional character state. Hence, both species share a less developed, thorn-like process that is weakly sclerotized in
<taxonomicName id="4C75A6E3CC234C36F6EAFF78FDEFFF04" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[418,625,151,177]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="raymondrevellii">
<emphasis id="B9010172CC234C36F6EAFF78FDEFFF04" box="[418,625,151,177]" italics="true" pageId="18" pageNumber="19">L. raymondrevellii</emphasis>
</taxonomicName>
, but slightly stronger sclerotized in
<taxonomicName id="4C75A6E3CC234C36F4B6FF78FBF8FF05" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1022,1126,151,176]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F4B6FF78FBF8FF05" box="[1022,1126,151,176]" italics="true" pageId="18" pageNumber="19">L. rajaeii</emphasis>
</taxonomicName>
. Major differences between the species are the lower half of the vein of the anterior cell, that always occurs between M
<subScript id="17F1DF25CC234C36F3CBFF22FB15FF6E" attach="left" box="[1155,1163,205,219]" fontSize="6" pageId="18" pageNumber="19">1</subScript>
and M
<quantity id="4C8D7085CC234C36F391FF22FB63FF60" box="[1241,1277,187,219]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">
<subScript id="17F1DF25CC234C36F391FF22FB7FFF6E" attach="left" box="[1241,1249,205,219]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
in
</quantity>
the hindwing, is not obsolete in the
<typeStatus id="54CE63C2CC234C36F6CAFF30FE7AFF4C" box="[386,484,223,249]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
and in the
<typeStatus id="54CE63C2CC234C36F516FF30FD57FF4C" box="[606,713,223,249]" pageId="18" pageNumber="19" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C75A6E3CC234C36F5A5FF30FCCBFF4D" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[749,853,223,248]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F5A5FF30FCCBFF4D" box="[749,853,223,248]" italics="true" pageId="18" pageNumber="19">L. rajaeii</emphasis>
</taxonomicName>
; the aedeagus is broad and short, not longer than the width of valva in
<taxonomicName id="4C75A6E3CC234C36F615FEECFE5BFEA9" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[349,453,259,284]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F615FEECFE5BFEA9" box="[349,453,259,284]" italics="true" pageId="18" pageNumber="19">L. rajaeii</emphasis>
</taxonomicName>
, but narrow and
<emphasis id="B9010172CC234C36F5CCFEECFD3BFEA9" box="[644,677,259,284]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
20% longer than the width of valva in
<taxonomicName id="4C75A6E3CC234C36F313FEECFAB5FEA8" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1115,1323,259,285]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="raymondrevellii">
<emphasis id="B9010172CC234C36F313FEECFAB5FEA8" box="[1115,1323,259,285]" italics="true" pageId="18" pageNumber="19">L. raymondrevellii</emphasis>
</taxonomicName>
; the basal edge of one uncus lobe is
<emphasis id="B9010172CC234C36F6FBFEC8FE4BFEF5" box="[435,469,295,320]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
1.9 longer than the upper edge of the same uncus lobe in the latter species but of almost equal length in
<taxonomicName id="4C75A6E3CC234C36F60FFEA4FE2FFED1" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[327,433,331,356]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F60FFEA4FE2FFED1" box="[327,433,331,356]" italics="true" pageId="18" pageNumber="19">L. rajaeii</emphasis>
</taxonomicName>
; the ventral end of the gnathos arms is much narrower if compared to the basal width of the gnathos arms in
<taxonomicName id="4C75A6E3CC234C36F63EFE80FDDBFE3C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[374,581,367,393]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="raymondrevellii">
<emphasis id="B9010172CC234C36F63EFE80FDDBFE3C" box="[374,581,367,393]" italics="true" pageId="18" pageNumber="19">L. raymondrevellii</emphasis>
</taxonomicName>
but the ventral end is broader in
<taxonomicName id="4C75A6E3CC234C36F4F9FE80FB87FE3D" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[945,1049,367,392]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F4F9FE80FB87FE3D" box="[945,1049,367,392]" italics="true" pageId="18" pageNumber="19">L. rajaeii</emphasis>
</taxonomicName>
; the inner side of the gnathos arms is cut in
<taxonomicName id="4C75A6E3CC234C36F7BEFE7CFE5BFE18" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[246,453,403,429]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="raymondrevellii">
<emphasis id="B9010172CC234C36F7BEFE7CFE5BFE18" box="[246,453,403,429]" italics="true" pageId="18" pageNumber="19">L. raymondrevellii</emphasis>
</taxonomicName>
; and the thorn-like process is narrow and has an acuminate tip in the latter species but is broad with a rounded tip in
<taxonomicName id="4C75A6E3CC234C36F686FE58FDA9FE65" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[462,567,439,464]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F686FE58FDA9FE65" box="[462,567,439,464]" italics="true" pageId="18" pageNumber="19">L. rajaeii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC234C36F78FFE35FE77FD1C" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BCADD60CC234C36F78FFE35FB4EFD30" blockId="18.[151,1437,151,789]" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F78FFE35FEFFFE41" bold="true" box="[199,353,474,500]" pageId="18" pageNumber="19">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC234C36F62FFE34FDB1FE41" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[359,559,475,501]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F62FFE34FDB1FE41" box="[359,559,475,501]" italics="true" pageId="18" pageNumber="19">Lukeniana rajaeii</emphasis>
</taxonomicName>
is known from Gidole on the southern Ethiopian Plateau,
<emphasis id="B9010172CC234C36F3FFFE34FB47FE41" box="[1207,1241,475,500]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
<quantity id="4C8D7085CC234C36F397FE34FAB8FE40" box="[1247,1318,475,501]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="10.0">10 km</quantity>
southwest of the southern tip of Lake Chamo and
<emphasis id="B9010172CC234C36F505FE10FDF1FDAD" box="[589,623,511,536]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
<quantity id="4C8D7085CC234C36F53DFE11FD54FDAC" box="[629,714,510,537]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="km" value="150.0">150 km</quantity>
north from the frontier to
<collectingCountry id="F3629DF0CC234C36F4B8FE10FBA3FDAC" box="[1008,1085,511,537]" name="Kenya" pageId="18" pageNumber="19">Kenya</collectingCountry>
. Gidole (elevation
<quantity id="4C8D7085CC234C36F258FE11FF30FD88" metricMagnitude="3" metricUnit="m" metricValue="2.0555" metricValueMax="2.259" metricValueMin="1.8519999999999999" pageId="18" pageNumber="19" unit="m" value="2055.5" valueMax="2259.0" valueMin="1852.0">
<geoCoordinate id="EE41BBA7CC234C36F258FE11FAD0FDAC" box="[1296,1358,510,537]" degrees="1,852" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="55" value="1.852">1,852</geoCoordinate>
<geoCoordinate id="EE41BBA7CC234C36F206FE11FA03FDAC" box="[1358,1437,510,537]" degrees="2,259" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="55" value="-2.259">2,259</geoCoordinate>
m
</quantity>
) is located within the Southern Main Ethiopian Rift (SMER)
<emphasis id="B9010172CC234C36F410FDCCFC08FD89" box="[856,918,547,572]" italics="true" pageId="18" pageNumber="19">sensu</emphasis>
<bibRefCitation id="EFE4A091CC234C36F4D4FDCCFB7BFD88" author="Keranen, K. M. &amp; Klemperer, S. L." box="[924,1253,547,573]" pageId="18" pageNumber="19" pagination="96 - 111" refId="ref78607" refString="Keranen, K. M. &amp; Klemperer, S. L. (2008) Discontinuous and diachronous evolution of the Main Ethiopian Rift: Implications for development of continental rifts. Earth and Planetary Science Letters, 265, 96 - 111. https: // doi. org / 10.1016 / j. epsl. 2007.09.038" type="journal article" year="2008">Keranen &amp; Klemperer (2008)</bibRefCitation>
and in the “high rainfall interval” of
<quantity id="4C8D7085CC234C36F637FDA8FDACFDD4" box="[383,562,583,609]" metricMagnitude="0" metricUnit="m" metricValue="1.8315" metricValueMax="2.331" metricValueMin="1.3319999999999999" pageId="18" pageNumber="19" unit="mm" value="1831.5" valueMax="2331.0" valueMin="1332.0">13322331 mm</quantity>
of the physiographic unit Southwestern Plateau of the Ethiopian Highlands
<emphasis id="B9010172CC234C36F7DFFD84FF4BFD31" box="[151,213,619,644]" italics="true" pageId="18" pageNumber="19">sensu</emphasis>
<bibRefCitation id="EFE4A091CC234C36F794FD84FEFAFD30" author="Friis, I." box="[220,356,619,645]" pageId="18" pageNumber="19" pagination="1 - 396" refId="ref77574" refString="Friis, I. (1992) Forests and Forest Trees of Northeast Tropical Africa. Their natural habitats and distribution patterns in Ethiopia, Djibouti and Somalia. Royal Botanic Gardens, Kew. Kew Bulletin Additional, Series XV, pp. i - iv + 1 - 396." type="book chapter" year="1992">Friis (1992)</bibRefCitation>
.
<taxonomicName id="4C75A6E3CC234C36F627FD84FDA6FD31" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[367,568,619,645]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC234C36F627FD84FDA6FD31" box="[367,568,619,645]" italics="true" pageId="18" pageNumber="19">Lukeniana rajaeii</emphasis>
</taxonomicName>
can be classified as an Afromontane near-endemic species.
</paragraph>
<paragraph id="8BCADD60CC234C36F78FFD61FE77FD1C" blockId="18.[151,1437,151,789]" box="[199,489,654,681]" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F78FFD61FEB4FD1D" bold="true" box="[199,298,654,680]" pageId="18" pageNumber="19">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC234C36F78FFD5CFB86FCA0" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BCADD60CC234C36F78FFD5CFB86FCA0" blockId="18.[151,1437,151,789]" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F78FFD5CFED0FD78" bold="true" box="[199,334,691,717]" pageId="18" pageNumber="19">Etymology.</emphasis>
The new species is named in honor of Hossein Rajaei Shoorcheh,
<taxonomicName id="4C75A6E3CC234C36F328FD5CFB79FD78" box="[1120,1255,691,717]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
curator and researcher at the State Museum of Natural History, Stuttgart. The first author is grateful to Hossein for significant help during studies for his doctoral dissertation on
<taxonomicName id="4C75A6E3CC234C36F5D1FD14FCB3FCA0" baseAuthorityName="Lehmann" baseAuthorityYear="2019" box="[665,813,763,789]" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
at the ZFMK, Bonn.
</paragraph>
</subSubSection>
</treatment>
</document>