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<mods:title id="085A7075911B1062DE551C21E6CF8C6F">Decapoda: Anomura: Paguridae) from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest)</mods:title>
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<paragraph id="8BC0AD18FF95F96C350461EA19D46996" blockId="1.[264,736,1633,1760]" box="[264,736,1734,1760]" pageId="1" pageNumber="2">
<heading id="D0881A74FF95F96C350461EA19D46996" bold="true" box="[264,736,1734,1760]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B90B710AFF95F96C350461EA19D46996" bold="true" box="[264,736,1734,1760]" pageId="1" pageNumber="2">
<taxonomicName id="4C7FD69BFF95F96C350461EA1A8F6996" ID-CoL="31ff653c-22c9-4ed6-a697-b4a02a832f6e" box="[264,443,1734,1760]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF95F96C350461EA1A8F6996" bold="true" box="[264,443,1734,1760]" italics="true" pageId="1" pageNumber="2">Trichopagurus</emphasis>
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de
<bibRefCitation id="EFEED0E9FF95F96C35E461EA19D46996" author="Saint" box="[488,736,1734,1760]" pageId="1" pageNumber="20" refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 39, 923 - 954. [Dated 1967, published 1968.]" type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
</emphasis>
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<taxonomicName id="4C7FD69BFF95F96C3504602B1A916868" ID-CoL="31ff653c-22c9-4ed6-a697-b4a02a832f6e" box="[264,421,1799,1822]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF95F96C3504602B1A916868" box="[264,421,1799,1822]" italics="true" pageId="1" pageNumber="2">Trichopagurus</emphasis>
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de
<treatmentCitation id="0ADE8B09FF95F96C35C0602A19EC686B" author="Saint" box="[460,728,1798,1821]" page="927" pageId="1" pageNumber="2" year="1968">
<bibRefCitation id="EFEED0E9FF95F96C35C0602A1996686B" author="Saint" box="[460,674,1798,1821]" pageId="1" pageNumber="20" refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 39, 923 - 954. [Dated 1967, published 1968.]" type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
: 927
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; 1970: 210;
<treatmentCitation id="0ADE8B09FF95F96C3750602A1F5C686B" author="McLaughlin" box="[860,1128,1798,1821]" page="128" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="EFEED0E9FF95F96C3750602A1F06686B" author="McLaughlin" box="[860,1074,1798,1821]" pageId="1" pageNumber="19" refString="McLaughlin, P. A. (2003 a) Illustrated keys to families and genera of the superfamily Paguroidea (Crustacea: Decapoda: Anomura), with diagnoses of genera of Paguridae. Memoirs of Museum Victoria, 60, 111 - 144." type="journal article" year="2003" yearSuffix="a">McLaughlin, 2003a</bibRefCitation>
: 128
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.
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</paragraph>
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<typeStatus id="54C413BAFF95F96C350460681A08682B" box="[264,316,1860,1885]" pageId="1" pageNumber="2">Type</typeStatus>
species
</emphasis>
.
<emphasis id="B90B710AFF95F96C35BD6068194F682B" box="[433,635,1860,1885]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="4C7FD69BFF95F96C35BD60681944682B" box="[433,624,1860,1885]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Catapaguroides</taxonomicName>
?
</emphasis>
<taxonomicName id="4C7FD69BFF95F96C368F606818D46828" authority="Forest, 1954" authorityName="Forest" authorityYear="1954" box="[643,992,1860,1886]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF95F96C368F60681877682B" box="[643,835,1860,1885]" italics="true" pageId="1" pageNumber="2">trichophthalmus</emphasis>
<bibRefCitation id="EFEED0E9FF95F96C3746606818D46828" author="Forest" box="[842,992,1860,1886]" pageId="1" pageNumber="19" refString="Forest, J. (1954) Crustaces Decapodes Marcheurs des Iles de Tahiti et des Tuamotu. I. Paguridea (suite). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 26, 71 - 79." type="journal article" year="1954">Forest, 1954</bibRefCitation>
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. Gender: Masculine.
</paragraph>
<paragraph id="8BC0AD18FF96F96F3534663018DB6C48" blockId="2.[264,1324,284,1590]" pageId="2" pageNumber="3">
<emphasis id="B90B710AFF96F96F35346630191D6E43" box="[312,553,284,309]" italics="true" pageId="2" pageNumber="3">Emended diagnosis.</emphasis>
Gills biserial, 11 pairs, including 1 pleurobranch above third pereopod (sixth thoracic somite) and 2 arthrobranchs on each third maxilliped to fourth pereopods; most lamellae distinctly narrowed distally. Rostrum distinct, triangular. Ocular acicles simple. Third maxilliped with 1 accessory tooth. Chelipeds strongly unequal, with weak degree of sexual dimorphism. Fourth pereopod semichelate, propodal rasp consisting of 1 row of corneous scales; no preungual process. Coxa of male right fifth pereopod with moderately long sexual tube directed toward exterior, distal part of tube semitransparent, coiled; left with very short tube. Female with single left gonopore; coxae of fifth pereopods slightly to somewhat asymmetrical, left largest. Anterior lobe of sixth thoracic somite transversely oblong. Eighth thoracic sternite composed of 2 distinct, rounded lobes.
<collectingCountry id="F368ED88FF96F96F355165801AAD6DB0" box="[349,409,684,710]" name="Malta" pageId="2" pageNumber="3">Male</collectingCountry>
with unpaired left third to fifth pleopods; female with unpaired left second to fifth pleopods, no paired first pleopods modified as gonopods. Uropods markedly asymmetrical. Telson with faint or distinct transverse indentations, terminal margins oblique; posterior lobes each with fringe of long setae on lateral margin.
</paragraph>
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<subSubSection id="C365FE93FF96F96F35346460181A6940" pageId="2" pageNumber="3" type="discussion">
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<emphasis id="B90B710AFF96F96F353464601A926C13" box="[312,422,844,869]" italics="true" pageId="2" pageNumber="3">Remarks.</emphasis>
<collectingCountry id="F368ED88FF96F96F35BF64601AE06C10" box="[435,468,844,870]" name="American Samoa" pageId="2" pageNumber="3">As</collectingCountry>
noted before, the gills of
<taxonomicName id="4C7FD69BFF96F96F372B64601F956C13" box="[807,1185,844,869]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF96F96F372B64601F956C13" box="[807,1185,844,869]" italics="true" pageId="2" pageNumber="3">Trichopagurus trichophthalmus</emphasis>
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have been reported as intermediaire (de
<bibRefCitation id="EFEED0E9FF96F96F368B645818476CF8" author="Saint" box="[647,883,884,910]" pageId="2" pageNumber="20" refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 39, 923 - 954. [Dated 1967, published 1968.]" type="journal article" year="1968">Saint Laurent 1968</bibRefCitation>
,
<bibRefCitation id="EFEED0E9FF96F96F3789645818F56CF8" author="Laurent" box="[901,961,884,910]" pageId="2" pageNumber="20" refString="Saint Laurent, M. de (1970) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. V. Trichopagurus de Saint Laurent. (Crustaces Decapodes Paguridae). VI. Conclusion. Bulletin du Museum national d'Histoire naturelle, Paris, (2) 42, 210 - 222." type="journal article" year="1970">1970</bibRefCitation>
) or quadriserial (
<bibRefCitation id="EFEED0E9FF96F96F309464581A606CC0" author="McLaughlin" pageId="2" pageNumber="19" refString="McLaughlin, P. A. (2003 a) Illustrated keys to families and genera of the superfamily Paguroidea (Crustacea: Decapoda: Anomura), with diagnoses of genera of Paguridae. Memoirs of Museum Victoria, 60, 111 - 144." type="journal article" year="2003" yearSuffix="a">McLaughlin 2003a</bibRefCitation>
).
<collectingCountry id="F368ED88FF96F96F357864B01AA26CC0" box="[372,406,924,950]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="3">De</collectingCountry>
<bibRefCitation id="EFEED0E9FF96F96F35AD64B019AD6CC0" box="[417,665,924,950]" pageId="2" pageNumber="20" refString="Saint Laurent, M. de (1970) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. V. Trichopagurus de Saint Laurent. (Crustaces Decapodes Paguridae). VI. Conclusion. Bulletin du Museum national d'Histoire naturelle, Paris, (2) 42, 210 - 222." type="journal article">Saint Laurent (1970)</bibRefCitation>
illustrated a shallowly divided gill lamella. However, our examination of the
<typeStatus id="54C413BAFF96F96F361664E819B56CA8" box="[538,641,964,990]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
and additional specimens has shown that the gill lamellae of
<taxonomicName id="4C7FD69BFF96F96F352464C019316B73" box="[296,517,1004,1029]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF96F96F352464C019316B73" box="[296,517,1004,1029]" italics="true" pageId="2" pageNumber="3">T. trichophthalmus</emphasis>
</taxonomicName>
are actually entire, lacking distal division. Most gill lamellae, however, are abruptly narrowed distally, and the partially overlapping distal parts of a few gill lamellae might give the incorrect impression that each lamella is divided distally.
</paragraph>
<paragraph id="8BC0AD18FF96F96F35346348181A6940" blockId="2.[264,1324,284,1590]" pageId="2" pageNumber="3">
It is worth comparing
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<emphasis id="B90B710AFF96F96F3644634819C16B0B" box="[584,757,1124,1149]" italics="true" pageId="2" pageNumber="3">Trichopagurus</emphasis>
</taxonomicName>
with
<taxonomicName id="4C7FD69BFF96F96F3732634818E06B0B" box="[830,980,1124,1149]" class="Malacostraca" family="Paguridae" genus="Anapagrides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF96F96F3732634818E06B0B" box="[830,980,1124,1149]" italics="true" pageId="2" pageNumber="3">Anapagrides</emphasis>
</taxonomicName>
de
<bibRefCitation id="EFEED0E9FF96F96F300863481A776BD0" author="Saint" pageId="2" pageNumber="20" refString="Saint Laurent-Dechance, M. de (1966) Remarques sur la classification de la famille des Paguridae et sur la position systematique d' Iridopagurus de Saint Laurent. Diagnose d' Anapagrides gen. nov. Bulletin du Museum national d'Histoire naturelle, Paris, (2) 38, 257 - 265." type="journal article" year="1966">Saint Laurent­Dechancé, 1966</bibRefCitation>
and
<taxonomicName id="4C7FD69BFF96F96F357663A018086BD0" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[378,828,1164,1190]" class="Malacostraca" family="Paguridae" genus="Catapagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF96F96F357663A019276BD3" box="[378,531,1164,1189]" italics="true" pageId="2" pageNumber="3">Catapagurus</emphasis>
<bibRefCitation id="EFEED0E9FF96F96F361663A018086BD0" author="Milne-Edwards" box="[538,828,1164,1190]" lastPageId="2" lastPageNumber="20" pageId="2" pageNumber="19" refString="Milne-Edwards, A. (1880) Reports on the result of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, 78, 79, by the United States Coast Survey steamer, ' Blake', Lieut. Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. VIII Etudes preliminaires sur les Crustaces. Bulletin of the Museum of Comparative Zoology, 8, 1 - 68." type="journal article" year="1880">A. Milne­Edwards, 1880</bibRefCitation>
</taxonomicName>
(rediagnosed by
<bibRefCitation id="EFEED0E9FF96F96F300863A01FEF6BD0" author="McLaughlin" box="[1028,1243,1164,1190]" pageId="2" pageNumber="19" refString="McLaughlin, P. A. (2004) Reappraisal of the hermit crab genera Catapagurus A. Milne-Edward and Hemipagurus Smith (Crustacea: Decapoda: Anomura: Paguridae), with the description of a new species. Zootaxa, 433, 1 - 16." type="journal article" year="2004">McLaughlin 2004</bibRefCitation>
), as all are characterized by 11 pairs of biserial gills, strongly unequal chelipeds, single row of scales in the propodal rasp on the fourth pereopod, and some development of a right sexual tube of male.
<taxonomicName id="4C7FD69BFF96F96F35B0622819666A6B" box="[444,594,1284,1309]" class="Malacostraca" family="Paguridae" genus="Anapagrides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF96F96F35B0622819666A6B" box="[444,594,1284,1309]" italics="true" pageId="2" pageNumber="3">Anapagrides</emphasis>
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primarily differs from
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<emphasis id="B90B710AFF96F96F376162281F2E6A6B" box="[877,1050,1284,1309]" italics="true" pageId="2" pageNumber="3">Trichopagurus</emphasis>
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in the structure of the right sexual tube. In
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<emphasis id="B90B710AFF96F96F3605620019AB6A33" box="[521,671,1324,1349]" italics="true" pageId="2" pageNumber="3">Anapagrides</emphasis>
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, the right sexual tube is short (about 0.51.2 times the length of the coxa of the fifth pereopod) and is posteriorly directed.
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<emphasis id="B90B710AFF96F96F302762781FF06A1B" box="[1067,1220,1364,1389]" italics="true" pageId="2" pageNumber="3">Catapagurus</emphasis>
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is easily distinguished from
<taxonomicName id="4C7FD69BFF96F96F35E0625019AD6AE3" box="[492,665,1404,1429]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF96F96F35E0625019AD6AE3" box="[492,665,1404,1429]" italics="true" pageId="2" pageNumber="3">Trichopagurus</emphasis>
</taxonomicName>
by the broadly rounded rostral lobe, the possession of a ventral row of very small spinules on the ambulatory dactyli, instead of rather strong corneous spines, and the presence of paired female gonopores. In addition,
<taxonomicName id="4C7FD69BFF96F96F305662E01FC76A93" box="[1114,1267,1484,1509]" class="Malacostraca" family="Paguridae" genus="Catapagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90B710AFF96F96F305662E01FC76A93" box="[1114,1267,1484,1509]" italics="true" pageId="2" pageNumber="3">Catapagurus</emphasis>
</taxonomicName>
species occasionally have elongate ocular acicles and usually a well developed preungual process on the dactylus of the fourth pereopod.
</paragraph>
</subSubSection>
</treatment>
</document>