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<document id="1B888AC6021EA460906D8C1B7F91AD5C" ID-CLB-Dataset="4037" ID-DOI="10.5852/ejt.2021.774.1537" ID-GBIF-Dataset="c7b35919-a8bf-4baa-aae3-f029e93332f6" ID-ISSN="2118-9773" ID-Zenodo-Dep="5567094" ID-ZooBank="68C21186-889C-4F61-8DF3-5752A8EA35AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634142345964" checkinUser="felipe" docAuthor="Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi &amp; Faille, Arnaud" docDate="2021" docId="03CC663BFFAFFF87FF7DA49AFAF9FCA8" docLanguage="en" docName="ejt-774_135-154.pdf" docOrigin="European Journal of Taxonomy 774" docSource="http://dx.doi.org/10.5852/ejt.2021.774.1537" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Guiodytes Tian 2013" docType="key" docVersion="6" lastPageNumber="153" masterDocId="FFF51E43FFBEFF95FFC0A04DFF8EFFFA" masterDocTitle="Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini)" masterLastPageNumber="154" masterPageNumber="135" pageNumber="152" updateTime="1699459911858" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9078AA407EA6268108721CB7C9CA7B40">Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini)</mods:title>
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<mods:namePart id="70D423F840C9F9E5EB498982959EDB09">Huang, Sunbin</mods:namePart>
<mods:nameIdentifier id="56388D24E8A03BED656332CBDA11AB82" type="LSID">B465D2B4-4B8F-4C75-BCF8-4CF07E48B7BF</mods:nameIdentifier>
<mods:affiliation id="1E29C680D0EC8D04935106081614DA4D">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, 510642 Guangzhou, China. &amp; Mécanismes adaptatifs et évolution (MECADEV), UMR 7179 CNRS-MNHN, Muséum national d’histoire naturelle, CP 50, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="033788CEBD28DD341A4F1DDB53847536" type="email">huangsunbin@163.com</mods:nameIdentifier>
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<mods:name id="A4EFFE2CD26C5616A98136F4AE35B0A9" type="personal">
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<mods:namePart id="DE2306569EF73FDB31431DB765DAB4ED">Zhou, Jiajun</mods:namePart>
<mods:nameIdentifier id="6FFF65D2D7C1B00D3C08A2F1882C6AAE" type="LSID">0B581FD2-8D89-4501-9B59-81BFC76D8FC7</mods:nameIdentifier>
<mods:affiliation id="856B5091C648130FCDF4E382FFF3A70C">Zhejiang Forest Survey, Planning and Design Company, 310020 Hangzhou, China.</mods:affiliation>
<mods:nameIdentifier id="F520BFBE689C2AE8917245273B19680F" type="email">cnwaters@foxmail.com</mods:nameIdentifier>
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<mods:roleTerm id="84F2A9F97E5240A2CA81C3EA34750298">Author</mods:roleTerm>
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<mods:namePart id="646BF7C94EEC68E04054FE9CE7960800">Tian, Mingyi</mods:namePart>
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<mods:affiliation id="733F780CF1D11DA81ADFFFEB3F12805C">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, 510642 Guangzhou, China.</mods:affiliation>
<mods:nameIdentifier id="40154B7E64D2DE951DCBC06A603C23CD" type="email">mytian@scau.edu.cn</mods:nameIdentifier>
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<mods:namePart id="0530275E12A0A03B6B97DBD9789FCF53">Faille, Arnaud</mods:namePart>
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<mods:affiliation id="5D4462ED4CFF870D4CFF1A5F3D0AC29D">Department of Entomology, Stuttgart State Museum of Natural History, 70191 Stuttgart, Germany.</mods:affiliation>
<mods:nameIdentifier id="48F3CC5081245335D0C8C87F7F0250E0" type="email">arnaud.faille@smns-bw.de</mods:nameIdentifier>
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<mods:title id="63F579CA1C8DCC9057AD5099EF3D5CBE">European Journal of Taxonomy</mods:title>
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<mods:date id="FFD1D8E25C14E35ABDD836F2908F7549">2021</mods:date>
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<treatment id="03CC663BFFAFFF87FF7DA49AFAF9FCA8" ID-DOI="http://doi.org/10.5281/zenodo.5572587" ID-GBIF-Taxon="188646001" ID-Zenodo-Dep="5572587" LSID="urn:lsid:zoobank.org:act:EA3D7117-F8F3-4F45-848A-9E3879ABCBF8" httpUri="http://treatment.plazi.org/id/03CC663BFFAFFF87FF7DA49AFAF9FCA8" lastPageId="18" lastPageNumber="153" pageId="17" pageNumber="152">
<subSubSection id="C37F84A6FFAFFF84FF7DA49AFC48FB09" box="[189,966,1238,1267]" pageId="17" pageNumber="152" type="nomenclature">
<paragraph id="8BDAD72DFFAFFF84FF7DA49AFC48FB09" blockId="17.[189,966,1238,1267]" box="[189,966,1238,1267]" pageId="17" pageNumber="152">
<emphasis id="B9110B3FFFAFFF84FF7DA49AFC48FB09" bold="true" box="[189,966,1238,1267]" pageId="17" pageNumber="152">
<emphasis id="B9110B3FFFAFFF84FF7DA49AFE46FB09" bold="true" box="[189,456,1238,1267]" italics="true" pageId="17" pageNumber="152">Key to the species of</emphasis>
<taxonomicName id="4C65ACAEFFAFFF84FE10A49BFDD2FB09" ID-CoL="8KX4J" authorityName="Tian" authorityYear="2013" box="[464,604,1238,1267]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="154" phylum="Arthropoda" rank="genus">Guiodytes</taxonomicName>
<emphasis id="B9110B3FFFAFFF84FDA4A49BFC48FB09" bold="true" box="[612,966,1238,1267]" italics="true" pageId="17" pageNumber="152">
(after
<bibRefCitation id="EFF4AADCFFAFFF84FD75A49BFCB5FB08" author="Tian M. Y." box="[693,827,1238,1266]" pageId="17" pageNumber="152" pagination="355 - 362" refId="ref7089" refString="Tian M. Y. 2014. New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae). Zootaxa 3861 (4): 355 - 362. https: // doi. org / 10.11646 / zootaxa. 3861.4.5" type="journal article" year="2014">Tian 2014</bibRefCitation>
, modified)
</emphasis>
</emphasis>
</paragraph>
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<subSubSection id="C37F84A6FFAFFF87FF7DA546FAF9FCA8" lastPageId="18" lastPageNumber="153" pageId="17" pageNumber="152" type="key">
<key id="FFF4801CFFAFFF87FF7DA546FAF9FCA8" lastPageId="18" lastPageNumber="153" pageId="17" pageNumber="152">
<keyStep id="3091CF88FFAFFF84FF7DA546FAF9FA02" pageId="17" pageNumber="152">
<paragraph id="8BDAD72DFFAFFF84FF7DA546FAF9FA91" blockId="17.[189,1399,1291,1941]" pageId="17" pageNumber="152">
<keyLead id="30947418FFAFFF84FF7DA546FAF9FA91" pageId="17" pageNumber="152">
1. Labrum not or slightly emarginate; pronotum with fore angles not or slightly protruding (
<figureCitation id="135ECBA8FFAFFF84FACDA546FAD8FADF" box="[1293,1366,1291,1317]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[218,1333,303,1800]" captionTargetId="figure-16@4.[189,1398,295,1822]" captionTargetPageId="4" captionText="Fig. 3. Head and pronotum of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU)." figureDoi="http://doi.org/10.5281/zenodo.5567100" httpUri="https://zenodo.org/record/5567100/files/figure.png" pageId="17" pageNumber="152">Figs 3</figureCitation>
A- E); elytral interval three with four or five dorsal setiferous pores, elytral interval five without dorsal setiferous pores (
<figureCitation id="135ECBA8FFAFFF84FE73A51CFE74FA91" box="[435,506,1361,1387]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="17" pageNumber="152">Fig. 4</figureCitation>
) .................................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BDAD72DFFAFFF84FF01A538FAF9FA02" blockId="17.[189,1399,1291,1941]" pageId="17" pageNumber="152">
<keyLead id="30947418FFAFFF84FF01A538FAF9FA02" pageId="17" pageNumber="152">
– Labrum conspicuously emarginate as a semicircular incision; pronotum with fore angles strongly protruding (
<figureCitation id="135ECBA8FFAFFF84FEB8A5DAFE40FA48" box="[376,462,1431,1458]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[218,1333,303,1800]" captionTargetId="figure-16@4.[189,1398,295,1822]" captionTargetPageId="4" captionText="Fig. 3. Head and pronotum of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU)." figureDoi="http://doi.org/10.5281/zenodo.5567100" httpUri="https://zenodo.org/record/5567100/files/figure.png" pageId="17" pageNumber="152">Fig. 3F</figureCitation>
); elytral interval three with nine dorsal setiferous pores, elytral interval five with six dorsal setiferous pores (
<figureCitation id="135ECBA8FFAFFF84FDECA5F6FD0DFA2F" box="[556,643,1467,1493]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="17" pageNumber="152">Fig. 4F</figureCitation>
) (Longzhou: Zhuzhu Dong cave) ............................................... .................................................................................................
<taxonomicName id="4C65ACAEFFAFFF84FC79A593FA9BFA02" authority="Tian &amp; Zhou" authorityName="Tian &amp; Zhou" authorityYear="2021" box="[953,1301,1502,1528]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="152" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. nov.">
<emphasis id="B9110B3FFFAFFF84FC79A593FBF8FA02" bold="true" box="[953,1142,1502,1528]" pageId="17" pageNumber="152">
<emphasis id="B9110B3FFFAFFF84FC79A593FC40FA02" bold="true" box="[953,974,1502,1528]" italics="true" pageId="17" pageNumber="152">G</emphasis>
.
<emphasis id="B9110B3FFFAFFF84FC1DA593FBF8FA02" bold="true" box="[989,1142,1502,1528]" italics="true" pageId="17" pageNumber="152">inexpectatus</emphasis>
</emphasis>
Tian &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="A222B644FFAFFF84FADCA593FAF9FA02" box="[1308,1399,1502,1528]" pageId="17" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="3091CF88FFAFFF84FF7DA652FAF9F9A7" pageId="17" pageNumber="152">
<paragraph id="8BDAD72DFFAFFF84FF7DA652FAF9F9C3" blockId="17.[189,1399,1291,1941]" box="[189,1399,1567,1594]" pageId="17" pageNumber="152">
<keyLead id="30947418FFAFFF84FF7DA652FAF9F9C3" box="[189,1399,1567,1594]" pageId="17" pageNumber="152">
2. Elytral interval three with four dorsal setiferous pores (
<figureCitation id="135ECBA8FFAFFF84FCB0A652FC7FF9C3" box="[880,1009,1567,1593]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="17" pageNumber="152">Figs 4B, C</figureCitation>
) ................................................ 3
</keyLead>
</paragraph>
<paragraph id="8BDAD72DFFAFFF84FF01A60EFAF9F9A7" blockId="17.[189,1399,1291,1941]" box="[193,1399,1602,1629]" pageId="17" pageNumber="152">
<keyLead id="30947418FFAFFF84FF01A60EFAF9F9A7" box="[193,1399,1602,1629]" pageId="17" pageNumber="152">
– Elytral interval three with five dorsal setiferous pores (
<figureCitation id="135ECBA8FFAFFF84FCABA60FFB82F9A7" box="[875,1036,1602,1629]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="17" pageNumber="152">Fig. 4A, D–E</figureCitation>
) ............................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3091CF88FFAFFF84FF7DA6C9FAF9F86E" pageId="17" pageNumber="152">
<paragraph id="8BDAD72DFFAFFF84FF7DA6C9FAF9F8F2" blockId="17.[189,1399,1291,1941]" pageId="17" pageNumber="152">
<keyLead id="30947418FFAFFF84FF7DA6C9FAF9F8F2" pageId="17" pageNumber="152">
3. Body smaller (
<quantity id="4C9D7AC8FFAFFF84FE5DA6C9FD8EF964" box="[413,512,1668,1695]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="17" pageNumber="152" unit="mm" value="5.2">5.2 mm</quantity>
); head stouter, labrum seven-setose, palpomeres slightly stouter, antennae shorter and stouter; entire lateral margin of elytra crenulated throughout, striae composed of larger punctures, interval six faintly carinated near base (
<figureCitation id="135ECBA8FFAFFF84FCFBA687FC18F91E" box="[827,918,1738,1764]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="17" pageNumber="152">Fig. 4B</figureCitation>
) (Longzhou: Paoma Dong cave)............ ........................................................................................................................
<taxonomicName id="4C65ACAEFFAFFF84FBACA6A0FAF9F8F2" ID-CoL="6L5K7" authority="Tian, 2013" authorityName="Tian" authorityYear="2013" box="[1132,1399,1773,1800]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="152" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="B9110B3FFFAFFF84FBACA6A0FB7FF8F2" bold="true" box="[1132,1265,1773,1800]" italics="true" pageId="17" pageNumber="152">G. bedosae</emphasis>
<bibRefCitation id="EFF4AADCFFAFFF84FB38A6A0FAF9F8F2" author="Tian M. Y." box="[1272,1399,1773,1800]" pageId="17" pageNumber="152" pagination="113 - 120" refId="ref7041" refString="Tian M. Y. 2013. Occurrence of troglobitic clivinines in China (Insecta: Coleoptera: Carabidae). Journal of Cave and Karst Studies 75 (2): 113 - 120. https: // doi. org / 10.4311 / 2011 LSC 0226" type="journal article" year="2013">Tian, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDAD72DFFAFFF84FF01A75CFAF9F86E" blockId="17.[189,1399,1291,1941]" pageId="17" pageNumber="152">
<keyLead id="30947418FFAFFF84FF01A75CFAF9F86E" pageId="17" pageNumber="152">
– Body larger (
<quantity id="4C9D7AC8FFAFFF84FE42A75CFE44F8D1" box="[386,458,1809,1835]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="152" unit="mm" value="7.0">7 mm</quantity>
); head slender, labrum five-setose, palpomeres slender, antennae longer and more slender; lateral margin of elytra crenulated on basal half, smooth on apical half, striae composed of smaller punctures, interval six strongly carinated near base (
<figureCitation id="135ECBA8FFAFFF84FC6CA71AFB88F88B" box="[940,1030,1879,1905]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="17" pageNumber="152">Fig. 4C</figureCitation>
) (Du’an: three caves) ............. .........................................................................................................................
<taxonomicName id="4C65ACAEFFAFFF84FBA9A737FAF9F86E" ID-CoL="3HLDV" authority="Tian, 2013" authorityName="Tian" authorityYear="2013" box="[1129,1399,1914,1941]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="152" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="B9110B3FFFAFFF84FBA9A737FB7FF86F" bold="true" box="[1129,1265,1914,1941]" italics="true" pageId="17" pageNumber="152">G. cavicola</emphasis>
<bibRefCitation id="EFF4AADCFFAFFF84FB38A737FAF9F86E" author="Tian M. Y." box="[1272,1399,1914,1940]" pageId="17" pageNumber="152" pagination="113 - 120" refId="ref7041" refString="Tian M. Y. 2013. Occurrence of troglobitic clivinines in China (Insecta: Coleoptera: Carabidae). Journal of Cave and Karst Studies 75 (2): 113 - 120. https: // doi. org / 10.4311 / 2011 LSC 0226" type="journal article" year="2013">Tian, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3091CF88FFACFF87FF7DA144FAF9FDE0" pageId="18" pageNumber="153">
<paragraph id="8BDAD72DFFACFF87FF7DA144FAF9FE90" blockId="18.[189,1399,265,850]" pageId="18" pageNumber="153">
<keyLead id="30947418FFACFF87FF7DA144FAF9FE90" pageId="18" pageNumber="153">
4. Body larger (
<quantity id="4C9D7AC8FFACFF87FE42A144FD9BFED9" box="[386,533,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="9.35" metricValueMax="9.5" metricValueMin="9.2" pageId="18" pageNumber="153" unit="mm" value="9.35" valueMax="9.5" valueMin="9.2">9.2–9.5 mm</quantity>
); frontal carinae long and nearly parallel-sided; pronotum wider than long; elytra with striae smaller and denser punctured, elytral intervals six to eight carinated near base, four and five not carinated (
<figureCitation id="135ECBA8FFACFF87FE37A102FDDCFE93" box="[503,594,335,361]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="18" pageNumber="153">Fig. 4A</figureCitation>
) ........................................................................................................ 5
</keyLead>
</paragraph>
<paragraph id="8BDAD72DFFACFF87FF01A13EFAF9FDE0" blockId="18.[189,1399,265,850]" pageId="18" pageNumber="153">
<keyLead id="30947418FFACFF87FF01A13EFAF9FDE0" pageId="18" pageNumber="153">
– Body smaller and more slender (
<quantity id="4C9D7AC8FFACFF87FDB7A13FFD55FE77" box="[631,731,370,397]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="153" unit="mm" value="4.5">4.5 mm</quantity>
); frontal carinae short and not parallel-sided; pronotum longer than wide; elytra with striae composed of larger and sparser punctures, elytral intervals six to eight carinated throughout, four and five carinated near base (
<figureCitation id="135ECBA8FFACFF87FC2BA1F4FBC9FE29" box="[1003,1095,441,467]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="18" pageNumber="153">Fig. 4D</figureCitation>
) (Huanjiang: Dapo Dong cave) .................................................................................................................................................... ...........................................................................................................
<taxonomicName id="4C65ACAEFFACFF87FBC1A1B2FA9BFDE3" authority="Huang &amp; Faille" authorityName="Huang &amp; Faille" authorityYear="2021" box="[1025,1301,511,538]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="153" phylum="Arthropoda" rank="species" species="weii" status="sp. nov.">
<emphasis id="B9110B3FFFACFF87FBC1A1B2FBD9FDE3" bold="true" box="[1025,1111,511,537]" italics="true" pageId="18" pageNumber="153">G. weii</emphasis>
Huang &amp; Faille
</taxonomicName>
<taxonomicNameLabel id="A222B644FFACFF87FADCA24DFAF9FDE0" box="[1308,1399,512,538]" pageId="18" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3091CF88FFACFF87FF7DA20CFAF9FCA8" pageId="18" pageNumber="153">
<paragraph id="8BDAD72DFFACFF87FF7DA20CFAF9FD3F" blockId="18.[189,1399,265,850]" pageId="18" pageNumber="153">
<keyLead id="30947418FFACFF87FF7DA20CFAF9FD3F" pageId="18" pageNumber="153">
5. Body more depigmented, of light brown colour; pronotum shorter and stouter, with remarkably emarginate anterior margin (
<figureCitation id="135ECBA8FFACFF87FDF8A229FD18FD84" box="[568,662,612,638]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[218,1333,303,1800]" captionTargetId="figure-16@4.[189,1398,295,1822]" captionTargetPageId="4" captionText="Fig. 3. Head and pronotum of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU)." figureDoi="http://doi.org/10.5281/zenodo.5567100" httpUri="https://zenodo.org/record/5567100/files/figure.png" pageId="18" pageNumber="153">Fig. 3A</figureCitation>
); elytral intervals slightly vaulted, elytral striae broad and well marked, with large isolated punctures (
<figureCitation id="135ECBA8FFACFF87FD78A2CAFC9BFD5B" box="[696,789,647,673]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="18" pageNumber="153">Fig. 4A</figureCitation>
) (Huanjiang: Ji Dong cave, formerly called Shuiku Dong) .........................................................................................................
<taxonomicName id="4C65ACAEFFACFF87FB84A2E7FAF9FD3F" ID-CoL="3HLDW" authority="Tian, 2014" authorityName="Tian" authorityYear="2014" box="[1092,1399,682,709]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="153" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="B9110B3FFFACFF87FB84A2E7FB7FFD3F" bold="true" box="[1092,1265,682,709]" italics="true" pageId="18" pageNumber="153">G. deharvengi</emphasis>
<bibRefCitation id="EFF4AADCFFACFF87FB38A2E7FAF9FD3F" author="Tian M. Y." box="[1272,1399,682,709]" pageId="18" pageNumber="153" pagination="355 - 362" refId="ref7089" refString="Tian M. Y. 2014. New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae). Zootaxa 3861 (4): 355 - 362. https: // doi. org / 10.11646 / zootaxa. 3861.4.5" type="journal article" year="2014">Tian, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDAD72DFFACFF87FF01A283FAF9FCA8" blockId="18.[189,1399,265,850]" pageId="18" pageNumber="153">
<keyLead id="30947418FFACFF87FF01A283FAF9FCA8" pageId="18" pageNumber="153">
– Body less depigmented, of reddish-brown colour; pronotum longer and more slender, with anterior margin almost not emarginate (
<figureCitation id="135ECBA8FFACFF87FD92A2BCFD22FCF1" box="[594,684,753,779]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[218,1333,303,1800]" captionTargetId="figure-16@4.[189,1398,295,1822]" captionTargetPageId="4" captionText="Fig. 3. Head and pronotum of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU)." figureDoi="http://doi.org/10.5281/zenodo.5567100" httpUri="https://zenodo.org/record/5567100/files/figure.png" pageId="18" pageNumber="153">Fig. 3E</figureCitation>
); elytral intervals rather strongly vaulted, elytral striae narrow and deep, with small isolated punctures (
<figureCitation id="135ECBA8FFACFF87FD07A359FCAEFCD4" box="[711,800,788,814]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[370,1206,272,1843]" captionTargetId="figure-12@5.[349,1239,265,1857]" captionTargetPageId="5" captionText="Fig. 4. Left elytron of species of Guiodytes Tian, 2013. A. G. deharvengi Tian, 2014. B. G. bedosae Tian, 2013. C. G. cavicola Tian, 2013. D. G. weii Huang &amp; Faille sp. nov., holotype, ♂ (SCAU). E. G. yueliangensis Huang &amp; Tian sp. nov., holotype, ♂ (SCAU). F. G. inexpectatus Tian &amp; Zhou sp. nov., holotype, ♀ (SCAU). Dorsal pores indicated by white points." figureDoi="http://doi.org/10.5281/zenodo.5567102" httpUri="https://zenodo.org/record/5567102/files/figure.png" pageId="18" pageNumber="153">Fig. 4E</figureCitation>
) (Huanjiang: Huang Dong cave) .......................... .............................................................................................
<taxonomicName id="4C65ACAEFFACFF87FC5BA37AFA9BFCAB" authority="Huang &amp; Tian" authorityName="Huang &amp; Tian" authorityYear="2021" box="[923,1301,823,850]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="153" phylum="Arthropoda" rank="species" species="yueliangensis" status="sp. nov.">
<emphasis id="B9110B3FFFACFF87FC5BA37AFBEBFCA8" bold="true" box="[923,1125,823,850]" italics="true" pageId="18" pageNumber="153">G. yueliangensis</emphasis>
Huang &amp; Tian
</taxonomicName>
<taxonomicNameLabel id="A222B644FFACFF87FADCA375FAF9FCA8" box="[1308,1399,824,850]" pageId="18" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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