263 lines
31 KiB
XML
263 lines
31 KiB
XML
<document id="856D5ACD7B6D7C0977B5DE856984CF4D" ID-DOI="10.11646/phytotaxa.252.1.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13674915" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725405123220" checkinUser="felipe" docAuthor="Sang, Julia & Kiew, Ruth" docDate="2016" docId="03A8A444FF82C467FF08FD8BFD5DF8A5" docLanguage="en" docName="phytotaxa.252.1.2.pdf" docOrigin="Phytotaxa 252 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.252.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia edgariana S. Julia & Kiew 2016, spec. nov." docType="treatment" docVersion="2" lastPageNumber="26" masterDocId="FF91DC3CFF8AC46EFF80FFDBFFE9FFA6" masterDocTitle="Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species" masterLastPageNumber="30" masterPageNumber="17" pageNumber="25" updateTime="1725426159086" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="1F5AC48CDB1FA03A0FE86BB135FA40C9">
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<mods:title id="94FC4EF20B8312E98F5C36B6042DE0AB">Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species</mods:title>
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<mods:name id="59C14BEC8D8A15B65732D7A23A163EEA" type="personal">
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<mods:roleTerm id="8F06B444650B47584491BCCF3579EA49">Author</mods:roleTerm>
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<mods:namePart id="3D1425BB7DB7C316E87D86F33870CC54">Sang, Julia</mods:namePart>
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<mods:nameIdentifier id="4855297E83D296DCEA5C081794283764" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
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<mods:name id="F590BB6D82A2C7B97B98B98378B791D3" type="personal">
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<mods:role id="12DAF42A1556332FEB14E398447A9A6F">
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<mods:roleTerm id="AE0DE1893B33EA8949C433F3C6C7BE14">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="57A3A56F85045CAFF70F42941A868C5D">Kiew, Ruth</mods:namePart>
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<mods:typeOfResource id="47740F9AC2124B6F00B4E5F8120FECEB">text</mods:typeOfResource>
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<mods:titleInfo id="3BFFF73E8B0130C0D8CFF6413D671520">
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<mods:title id="293CE19EA98DB313AFAD9CDDC00A0EF2">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="880C868C5417B441F980D9F51BA8005E">
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<mods:date id="234816A9EF31C58ABDCE6AC9E4791B41">2016</mods:date>
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<mods:detail id="B9207FBD23C3F79DC1E282C2014CCDCE" type="pubDate">
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<mods:number id="C44775351ED98A8563C530CC8E08783E">2016-03-08</mods:number>
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<mods:detail id="F405A33E0C91A554D6CC708334069465" type="volume">
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<mods:number id="A910DC2F8EB03FC37188D661EFFDB208">252</mods:number>
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<mods:number id="C90527095BA96B0629B0E2478BC260E7">1</mods:number>
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<mods:start id="BC65CC75C9102BF5196659F9EEF26ABD">17</mods:start>
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<mods:end id="58F9BC39C1E8BDD809D295256AEC25F5">30</mods:end>
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<mods:url id="7689AE0C76406BE4228E265B2B8C1720">http://dx.doi.org/10.11646/phytotaxa.252.1.2</mods:url>
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<mods:classification id="046FFE224D54594DCFB205AD32A670EC">journal article</mods:classification>
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<mods:identifier id="FB0C38E0BC0EDAFAF9FBDA2B1E4AAB78" type="DOI">10.11646/phytotaxa.252.1.2</mods:identifier>
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<mods:identifier id="A22840182653B94AEF26E6907CCE98E3" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="05640D244EC42D34EAB7A642D7415EB8" type="Zenodo-Dep">13674915</mods:identifier>
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<treatment id="03A8A444FF82C467FF08FD8BFD5DF8A5" LSID="urn:lsid:plazi:treatment:03A8A444FF82C467FF08FD8BFD5DF8A5" httpUri="http://treatment.plazi.org/id/03A8A444FF82C467FF08FD8BFD5DF8A5" lastPageId="9" lastPageNumber="26" pageId="8" pageNumber="25">
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<subSubSection id="C31B46D9FF82C466FF08FD8BFCCDFDCC" box="[136,804,591,618]" pageId="8" pageNumber="25" type="nomenclature">
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<paragraph id="8BBE1552FF82C466FF08FD8BFCCDFDCC" blockId="8.[136,804,591,618]" box="[136,804,591,618]" pageId="8" pageNumber="25">
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<heading id="D0F6A23EFF82C466FF08FD8BFCCDFDCC" box="[136,804,591,618]" fontSize="11" level="2" pageId="8" pageNumber="25" reason="3">
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<emphasis id="B975C940FF82C466FF08FD8BFF74FDCC" bold="true" box="[136,157,592,618]" italics="true" pageId="8" pageNumber="25">7.</emphasis>
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<taxonomicName id="4C016ED1FF82C466FF24FD8BFDDDFDCC" ID-CoL="7SFGZ" authority="S.Julia & Kiew" authorityName="S. Julia & Kiew" authorityYear="2016" box="[164,564,591,618]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="25" phylum="Tracheophyta" rank="species" species="edgariana" status="sp. nov.">
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<emphasis id="B975C940FF82C466FF24FD8BFE68FDCC" bold="true" box="[164,385,591,618]" italics="true" pageId="8" pageNumber="25">Begonia edgariana</emphasis>
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S.Julia & Kiew
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</taxonomicName>
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,
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<emphasis id="B975C940FF82C466FDBFFD8AFD44FDCC" box="[575,685,593,618]" italics="true" pageId="8" pageNumber="25">
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<taxonomicNameLabel id="A246743BFF82C466FDBFFD8AFD44FDCC" box="[575,685,593,618]" pageId="8" pageNumber="25" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="133A09D7FF82C466FD3BFD8AFCF2FDCC" box="[699,795,593,618]" captionStart="FIGURE 5" captionStartId="9.[136,229,1377,1398]" captionTargetBox="[345,1245,195,1349]" captionTargetId="figure-14@9.[339,1249,190,1352]" captionTargetPageId="9" captionText="FIGURE 5. Begonia edgariana. A. Habit (green form); B. Leaves of bronzy-dark green form; C. Stipule; D. Male flowers; E. Female flower; F. Female flower (dull red form); G. Female flower (white-tinged pink form) showing tepals and pistil; H. Mature fruit; I. Cross section of ovary. All photos from SFC 2758 by Julia Sang (A–H) and Ling Chea Yiing (I)." figureDoi="http://doi.org/10.5281/zenodo.13674929" httpUri="https://zenodo.org/record/13674929/files/figure.png" pageId="8" pageNumber="25">Figure 5</figureCitation>
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||
)
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||
</heading>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C31B46D9FF82C466FF08FDAEFE9EFD28" box="[136,375,629,654]" pageId="8" pageNumber="25" type="description">
|
||
<paragraph id="8BBE1552FF82C466FF08FDAEFE9EFD28" blockId="8.[136,375,629,654]" box="[136,375,629,654]" pageId="8" pageNumber="25">
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||
<emphasis id="B975C940FF82C466FF08FDAEFF36FD28" bold="true" box="[136,223,629,654]" pageId="8" pageNumber="25">Section</emphasis>
|
||
<taxonomicName id="4C016ED1FF82C466FF66FDAEFE9EFD28" box="[230,375,629,654]" class="Magnoliopsida" family="Begoniaceae" genus="Petermannia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="25" phylum="Tracheophyta" rank="genus">
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<emphasis id="B975C940FF82C466FF66FDAEFE9EFD28" box="[230,375,629,654]" italics="true" pageId="8" pageNumber="25">Petermannia</emphasis>
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||
</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C31B46D9FF82C466FF08FD1BFB75FC2F" pageId="8" pageNumber="25" type="diagnosis">
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<paragraph id="8BBE1552FF82C466FF08FD1BFB75FC2F" blockId="8.[136,1452,704,977]" pageId="8" pageNumber="25">
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<emphasis id="B975C940FF82C466FF08FD1BFF1AFD73" bold="true" box="[136,243,704,725]" pageId="8" pageNumber="25">Diagnosis:</emphasis>
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||
—It most resembles
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<taxonomicName id="4C016ED1FF82C466FE46FD1BFC46FD73" authority="S. Julia & Kiew (2015: 95)" authorityName="S. Julia & Kiew" authorityPageNumber="95" authorityYear="2015" box="[454,943,704,725]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="25" phylum="Tracheophyta" rank="species" species="matangensis">
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<emphasis id="B975C940FF82C466FE46FD1BFD71FD73" box="[454,664,704,725]" italics="true" pageId="8" pageNumber="25">Begonia matangensis</emphasis>
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S.Julia &
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<bibRefCitation id="EF9068A3FF82C466FC89FD1BFC46FD73" author="Kiew, R. & Julia, S. & Rimi, R. & Joffree, A. A." box="[777,943,704,725]" pageId="8" pageNumber="25" refId="ref9437" refString="Kiew, R., Julia, S., Rimi, R. & Joffree, A. A. (2015) A Guide to Begonias of Borneo. Natural History Publications (Kota Kinabalu), 293 pp." type="book" year="2015">Kiew (2015: 95)</bibRefCitation>
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</taxonomicName>
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||
in its habit, its dense hispid indumentums and its subopposite velvety leaves but it is different in its broader leaves about twice as long as wide (vs. leaves about three times longer than wide in
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<taxonomicName id="4C016ED1FF82C466FEBDFCD3FE39FCBB" authorityName="S. Julia & Kiew" authorityYear="2015" box="[317,464,776,797]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="25" phylum="Tracheophyta" rank="species" species="matangensis">
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<emphasis id="B975C940FF82C466FEBDFCD3FE39FCBB" box="[317,464,776,797]" italics="true" pageId="8" pageNumber="25">B. matangensis</emphasis>
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||
</taxonomicName>
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||
), its female flowers with dimorphic tepals with acute apex (vs. isomorphic tepals with pointed apex), style
|
||
<quantity id="4CF9B8B7FF82C466FF70FCF7FED6FCE7" box="[240,319,812,833]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="25" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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||
long, more than half of the length of the tepals with only a shallow Y-shaped stigma (vs. style ca.
|
||
<quantity id="4CF9B8B7FF82C466FB66FCF7FAF5FCE7" box="[1254,1308,812,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="25" unit="mm" value="2.0">2 mm</quantity>
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long, less than half of the tepal length and a with wide Y-shaped stigma) and a broader capsule, triangular in outline and 10–15 ×
|
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<quantity id="4CF9B8B7FF82C466FA93FC8BFA93FCC3" box="[1299,1402,848,869]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="8" pageNumber="25" unit="mm" value="12.5" valueMax="14.0" valueMin="11.0">11–14 mm</quantity>
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with wider wings,
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<quantity id="4CF9B8B7FF82C466FEBFFCAFFE67FC2F" box="[319,398,884,905]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="8" pageNumber="25" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
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wide (vs. an oblong capsule ca. 10 ×
|
||
<quantity id="4CF9B8B7FF82C466FD79FCAFFCD9FC2F" box="[761,816,884,905]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="25" unit="mm" value="8.0">8 mm</quantity>
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with narrower wings,
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<quantity id="4CF9B8B7FF82C466FB89FCAFFBB1FC2F" box="[1033,1112,884,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="25" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
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wide).
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||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C31B46D9FF82C466FF08FC43FC32F894" pageId="8" pageNumber="25" type="materials_examined">
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<paragraph id="8BBE1552FF82C466FF08FC43FB26FC77" blockId="8.[136,1452,704,977]" pageId="8" pageNumber="25">
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<emphasis id="B975C940FF82C466FF08FC43FF2BFC0B" bold="true" box="[136,194,920,941]" pageId="8" pageNumber="25">Type:</emphasis>
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—
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||
<materialsCitation id="3B691F0FFF82C466FF5BFC43FB25FC77" collectingDate="2015-08-02" collectionCode="SAR, BRUN, KEP, SNP" country="Malaysia" county="Borneo" elevation="222" latitude="1.3166666" location="Batang Ai National Park" longLatPrecision="21" longitude="112.066666" municipality="Lubok Antu District" pageId="8" pageNumber="25" specimenCode="SFC 2758" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">
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<collectingCountry id="F31655C2FF82C466FF5BFC43FEB3FC0B" box="[219,346,920,941]" name="Malaysia" pageId="8" pageNumber="25">MALAYSIA</collectingCountry>
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.
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<collectingCounty id="62DF6DDEFF82C466FEE2FC43FE42FC0B" box="[354,427,920,941]" pageId="8" pageNumber="25">Borneo</collectingCounty>
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,
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<collectingRegion id="49C5DBB0FF82C466FE34FC43FDE3FC0B" box="[436,522,920,941]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="25">Sarawak</collectingRegion>
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.
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<collectingRegion id="49C5DBB0FF82C466FD92FC43FD81FC0B" box="[530,616,920,941]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="25">Sarawak</collectingRegion>
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.
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<collectingMunicipality id="6BDA8F28FF82C466FDEFFC43FCDAFC0B" box="[623,819,920,941]" pageId="8" pageNumber="25">Lubok Antu District</collectingMunicipality>
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:
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<location id="8EDE4389FF82C466FCBBFC43FBCFFC0B" LSID="urn:lsid:plazi:treatment:03A8A444FF82C467FF08FD8BFD5DF8A5:8EDE4389FF82C466FCBBFC43FBCFFC0B" box="[827,1062,920,941]" country="Malaysia" county="Borneo" latitude="1.3166666" longLatPrecision="21" longitude="112.066666" municipality="Lubok Antu District" name="Batang Ai National Park" pageId="8" pageNumber="25" stateProvince="Sarawak">Batang Ai National Park</location>
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,
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<location id="8EDE4389FF82C466FBAFFC43FAF8FC0B" LSID="urn:lsid:plazi:treatment:03A8A444FF82C467FF08FD8BFD5DF8A5:8EDE4389FF82C466FBAFFC43FAF8FC0B" box="[1071,1297,920,941]" country="Malaysia" county="Borneo" latitude="1.3166666" longLatPrecision="21" longitude="112.066666" municipality="Lubok Antu District" name="Sungai Bebiyong Besai" pageId="8" pageNumber="25" stateProvince="Sarawak">Sungai Bebiyong Besai</location>
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,
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<geoCoordinate id="EE357395FF82C466FA9AFC43FA6DFC0B" box="[1306,1412,920,941]" degrees="1" direction="north" minutes="19" orientation="latitude" pageId="8" pageNumber="25" precision="15" seconds="00" value="1.3166666">1°19’00”N</geoCoordinate>
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and
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<geoCoordinate id="EE357395FF82C466FF3DFC67FED4FC77" box="[189,317,956,977]" degrees="112" direction="east" minutes="04" orientation="longitude" pageId="8" pageNumber="25" precision="15" seconds="00" value="112.066666">112°04’00”E</geoCoordinate>
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,
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<quantity id="4CF9B8B7FF82C466FEC7FC67FE6FFC77" box="[327,390,956,977]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" pageId="8" pageNumber="25" unit="m" value="222.0">
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<elevation id="002CF261FF82C466FEC7FC67FE6FFC77" box="[327,390,956,977]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" pageId="8" pageNumber="25" unit="m" value="222.0">222 m</elevation>
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</quantity>
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,
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||
<date id="FFBF3392FF82C466FE10FC67FDF7FC77" box="[400,542,956,977]" pageId="8" pageNumber="25" value="2015-08-02">
|
||
<collectingDate id="EFFBCA7AFF82C466FE10FC67FDF7FC77" box="[400,542,956,977]" pageId="8" pageNumber="25" value="2015-08-02">2 August 2015</collectingDate>
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</date>
|
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,
|
||
<emphasis id="B975C940FF82C466FDA8FC67FD1CFC77" box="[552,757,956,977]" italics="true" pageId="8" pageNumber="25">
|
||
Julia et al.
|
||
<specimenCode id="DBA7BD29FF82C466FD14FC67FD1CFC77" box="[660,757,956,977]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="8" pageNumber="25">SFC 2758</specimenCode>
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="54BAABF0FF82C466FC81FC67FCBEFC77" box="[769,855,956,977]" pageId="8" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="ED108D97FF82C466FCDDFC67FC7AFC77" box="[861,915,956,977]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="8" pageNumber="25" type="Herbarium">SAR</collectionCode>
|
||
!;
|
||
<typeStatus id="54BAABF0FF82C466FC20FC67FC06FC77" box="[928,1007,956,977]" pageId="8" pageNumber="25" type="isotype">isotypes</typeStatus>
|
||
<collectionCode id="ED108D97FF82C466FC75FC67FBD6FC77" box="[1013,1087,956,977]" country="Brunei" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12726" lsid="urn:lsid:biocol.org:col:12726" name="Brunei Forestry Centre" pageId="8" pageNumber="25" type="Herbarium">BRUN</collectionCode>
|
||
!,
|
||
<collectionCode id="ED108D97FF82C466FBCBFC67FB96FC77" box="[1099,1151,956,977]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="8" pageNumber="25" type="Herbarium">KEP</collectionCode>
|
||
!,
|
||
<collectionCode id="ED108D97FF82C466FB0DFC67FB28FC77" box="[1165,1217,956,977]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15856" lsid="urn:lsid:biocol.org:col:15856" name="Sabah Parks" pageId="8" pageNumber="25" type="Herbarium">SNP</collectionCode>
|
||
!)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF82C466FF08FBDAFC32F894" blockId="8.[136,1452,1025,2058]" pageId="8" pageNumber="25">
|
||
Erect begonia
|
||
<quantity id="4CF9B8B7FF82C466FEAEFBDAFE76FBBC" box="[302,415,1025,1050]" metricMagnitude="-1" metricUnit="m" metricValue="3.55" metricValueMax="4.5" metricValueMin="2.6" pageId="8" pageNumber="25" unit="cm" value="35.5" valueMax="45.0" valueMin="26.0">26–45 cm</quantity>
|
||
tall. Indumentum of stem, stipules and petioles densely hispid, hairs white,
|
||
<quantity id="4CF9B8B7FF82C466FA8BFBDAFA83FBBC" box="[1291,1386,1025,1050]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="8" pageNumber="25" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
|
||
long. Stem branched, dark brown, succulent, slender,
|
||
<quantity id="4CF9B8B7FF82C466FD1AFBFEFD1FFB98" box="[666,758,1061,1086]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="25" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
thick, internodes (0.7–)
|
||
<quantity id="4CF9B8B7FF82C466FB80FBFEFBB9FB98" box="[1024,1104,1061,1086]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="8" pageNumber="25" unit="cm" value="5.5" valueMax="8.0" valueMin="3.0">3–8 cm</quantity>
|
||
long, thicker at nodes. Stipules reddish brown, 6–12 ×
|
||
<quantity id="4CF9B8B7FF82C466FE0BFB92FE03FBC4" box="[395,490,1097,1122]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="25" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
, margin entire, keeled, apex acuminate, seta to
|
||
<quantity id="4CF9B8B7FF82C466FC79FB92FBD3FBC4" box="[1017,1082,1097,1122]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="25" unit="mm" value="6.0">6 mm</quantity>
|
||
long, caducous. Leaves alternate below, upper leaves sub-opposite, pairs distant, held horizontally, not oblique; petiole brownish,
|
||
<quantity id="4CF9B8B7FF82C466FB68FBB6FA81FB20" box="[1256,1384,1133,1158]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="8" pageNumber="25" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 cm</quantity>
|
||
long, slightly grooved above; lamina plain dark green above, paler beneath, velvety, densely covered in appressed hairs ca.
|
||
<quantity id="4CF9B8B7FF82C466FF08FB6EFF21FB68" box="[136,200,1205,1230]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="25" unit="mm" value="2.0">2 mm</quantity>
|
||
long, less dense beneath, in life papery and soft (not fleshy), matt, slightly asymmetric, obovate, slightly falcate, 6–14 ×
|
||
<quantity id="4CF9B8B7FF82C466FF61FB02FE8AFB54" box="[225,355,1241,1266]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.5" metricValueMin="2.5" pageId="8" pageNumber="25" unit="cm" value="4.5" valueMax="6.5" valueMin="2.5">2.5–6.5 cm</quantity>
|
||
, broad side
|
||
<quantity id="4CF9B8B7FF82C466FE6EFB02FDBEFB54" box="[494,599,1241,1266]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="8" pageNumber="25" unit="cm" value="2.75" valueMax="4.0" valueMin="1.5">1.5–4 cm</quantity>
|
||
wide, base unequal, slightly lobed on narrow side, rounded on broad side, basal lobe rounded,
|
||
<quantity id="4CF9B8B7FF82C466FEE8FB26FE0DFAB0" box="[360,484,1277,1302]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="8" pageNumber="25" unit="cm" value="0.5" valueMax="0.8" valueMin="0.2">0.2–0.8 cm</quantity>
|
||
long, margin ciliate, minutely dentate in the lower half, becoming more deeply dentate towards the apex and appearing scalloped, apex acuminate, acumen
|
||
<quantity id="4CF9B8B7FF82C466FC17FAFAFBE8FA9C" box="[919,1025,1313,1338]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="25" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
|
||
long; venation pinnate, 4–6 veins on either side of the midrib, concolorous on both surfaces, densely pilose, 1–2 veins in basal lobe, prominent on both surfaces, more so beneath. Inflorescences on short shoots below the subopposite leaf, protogynous, densely pilose, female flower single or occasionally in a pair, male flowers in umbellate clusters with individual flowers subtended by a bract. Bracts and bracteoles similar to stipules, persistent. Male flowers with white pedicels, sometimes dull red,
|
||
<quantity id="4CF9B8B7FF82C466FF08FA0EFF0FFA48" box="[136,230,1493,1518]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="8" pageNumber="25" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
|
||
long; tepals 2, white or greenish white at the centre, sometimes dull red, densely pilose outside, hairs white, glabrous inside, oval, 4–5 ×
|
||
<quantity id="4CF9B8B7FF82C466FE46FA22FDE0F9B4" box="[454,521,1529,1554]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="25" unit="mm" value="3.0">3 mm</quantity>
|
||
, margin entire, apex acute; stamens 14–16, cluster conical, stalked to
|
||
<quantity id="4CF9B8B7FF82C466FA95FA22FA82F9B4" box="[1301,1387,1529,1554]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="25" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long; filaments whitish, ca.
|
||
<quantity id="4CF9B8B7FF82C466FEFBF9C6FE52F990" box="[379,443,1565,1590]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="25" unit="mm" value="1.0">1 mm</quantity>
|
||
long; anthers pale yellow, obovate, 1–1.2 ×
|
||
<quantity id="4CF9B8B7FF82C466FC23F9C6FC13F990" box="[931,1018,1565,1590]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="25" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, apex emarginate. Female flowers with greenish white pedicels
|
||
<quantity id="4CF9B8B7FF82C466FE18F99AFE1FF9FC" box="[408,502,1601,1626]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="8" pageNumber="25" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 mm</quantity>
|
||
long, sometimes reddish, sparsely hairy; ovary greenish yellow, sometimes reddish, sparsely hairy, oblong, 11–15 ×
|
||
<quantity id="4CF9B8B7FF82C466FE75F9BEFD9BF9D8" box="[501,626,1637,1662]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="25" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
|
||
, wings 3, equal, locules 3, placentas 2 per locule; tepals 5, pale greenish yellow to white tinged pink or reddish, sparsely pilose outside, glabrous inside, lanceolate; outer 2 tepals 6–9 ×
|
||
<quantity id="4CF9B8B7FF82C466FA02F952FF5FF960" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="25" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
, margin entire, apex acute; inner 3 tepals 6–8 ×
|
||
<quantity id="4CF9B8B7FF82C466FD67F976FCB4F960" box="[743,861,1709,1734]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="8" pageNumber="25" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
|
||
, styles 3, pale green, anchor-shaped,
|
||
<quantity id="4CF9B8B7FF82C466FA8BF976FA83F960" box="[1291,1386,1709,1734]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="25" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
long, divided to base, papillose forming a continuous twisted band. Capsules single, axillary, hispid, 10–15 ×
|
||
<quantity id="4CF9B8B7FF82C466FAABF90AFA4EF94C" box="[1323,1447,1745,1770]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="8" pageNumber="25" unit="mm" value="12.5" valueMax="14.0" valueMin="11.0">11–14 mm</quantity>
|
||
, triangular in outline, narrowed proximally, expanded distally to a sharp point, locules 3, wings 3, equal, wings
|
||
<quantity id="4CF9B8B7FF82C466FAD0F92EFA45F8A8" box="[1360,1452,1781,1806]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="8" pageNumber="25" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
|
||
wide, papery, dehiscing between the locule and wing; pedicel
|
||
<quantity id="4CF9B8B7FF82C466FCBEF8C2FC72F894" box="[830,923,1817,1842]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="8" pageNumber="25" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B46D9FF82C466FF3DF8E6FD26F838" pageId="8" pageNumber="25" type="etymology">
|
||
<paragraph id="8BBE1552FF82C466FF3DF8E6FD26F838" blockId="8.[136,1452,1025,2058]" pageId="8" pageNumber="25">
|
||
<emphasis id="B975C940FF82C466FF3DF8E6FEA1F8F0" bold="true" box="[189,328,1853,1878]" pageId="8" pageNumber="25">Etymology:</emphasis>
|
||
—Named after Edgar Irmscher (1887–1968), a German botanist expert in
|
||
<taxonomicName id="4C016ED1FF82C466FB2AF8E6FAEEF8F0" box="[1194,1287,1853,1878]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="25" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B975C940FF82C466FB2AF8E6FAEEF8F0" box="[1194,1287,1853,1878]" italics="true" pageId="8" pageNumber="25">Begonia</emphasis>
|
||
</taxonomicName>
|
||
, who in 1953 described 11
|
||
<taxonomicName id="4C016ED1FF82C466FEA0F8BAFE3EF8DC" box="[288,471,1889,1914]" pageId="8" pageNumber="25">
|
||
<emphasis id="B975C940FF82C466FEA0F8BAFE94F8DC" box="[288,381,1889,1914]" italics="true" pageId="8" pageNumber="25">Begonia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
collected by Odoardo Beccari from
|
||
<collectingCountry id="F31655C2FF82C466FCFBF8BAFC32F8DC" box="[891,987,1889,1914]" name="Malaysia" pageId="8" pageNumber="25">Sarawak</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9068A3FF82C466FC6BF8BAFB71F8DC" author="Irmscher, E." box="[1003,1176,1889,1914]" pageId="8" pageNumber="25" pagination="469 - 509" refId="ref9397" refString="Irmscher, E. (1953) Neue Begoniaceen, von O. Beccari in Malesian Gesammlt. Webbia 9 (2): 469 - 509. http: // dx. doi. org / 10.1080 / 00837792.1954.10669621" type="journal article" year="1953">Irmscher, 1953</bibRefCitation>
|
||
) and who explained the growth pattern of begonias with subopposite leaves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B46D9FF82C467FF3DF872FD5DF8A5" lastPageId="9" lastPageNumber="26" pageId="8" pageNumber="25" type="distribution">
|
||
<paragraph id="8BBE1552FF82C466FF3DF872FF2AF840" blockId="8.[136,1452,1025,2058]" pageId="8" pageNumber="25">
|
||
<emphasis id="B975C940FF82C466FF3DF872FEB0F864" bold="true" box="[189,345,1961,1986]" pageId="8" pageNumber="25">Distribution:</emphasis>
|
||
—
|
||
<collectingCountry id="F31655C2FF82C466FEF6F872FDE3F864" box="[374,522,1961,1986]" name="Malaysia" pageId="8" pageNumber="25">MALAYSIA</collectingCountry>
|
||
. Borneo.
|
||
<collectingRegion id="49C5DBB0FF82C466FDF6F872FD33F864" box="[630,730,1961,1986]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="25">Sarawak</collectingRegion>
|
||
. Endemic in
|
||
<collectingRegion id="49C5DBB0FF82C466FCEEF872FC38F864" box="[878,977,1961,1986]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="25">Sarawak</collectingRegion>
|
||
, known only from the Batang Ai National Park.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF82C466FF3DF82AFAB4F7AC" blockId="8.[136,1452,1025,2058]" box="[189,1373,2033,2058]" pageId="8" pageNumber="25">
|
||
<emphasis id="B975C940FF82C466FF3DF82AFECAF7AC" bold="true" box="[189,291,2033,2058]" pageId="8" pageNumber="25">Habitat:</emphasis>
|
||
—Lowland mixed dipterocarp forest, to
|
||
<quantity id="4CF9B8B7FF82C466FD67F82AFCC7F7AC" box="[743,814,2033,2058]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" pageId="8" pageNumber="25" unit="m" value="222.0">222 m</quantity>
|
||
elevation on steep or gentle slopes in deep shade.
|
||
</paragraph>
|
||
<caption id="DF7E45DAFF83C467FF08FABAFC03FA18" ID-DOI="http://doi.org/10.5281/zenodo.13674929" ID-Zenodo-Dep="13674929" httpUri="https://zenodo.org/record/13674929/files/figure.png" pageId="9" pageNumber="26" startId="9.[136,229,1377,1398]" targetBox="[345,1245,195,1349]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BBE1552FF83C467FF08FABAFC03FA18" blockId="9.[136,1452,1377,1470]" pageId="9" pageNumber="26">
|
||
<emphasis id="B975C940FF83C467FF08FABAFF16FAD0" bold="true" box="[136,255,1377,1398]" pageId="9" pageNumber="26">FIGURE 5.</emphasis>
|
||
<emphasis id="B975C940FF83C467FE86FABAFE2EFAD0" box="[262,455,1377,1398]" italics="true" pageId="9" pageNumber="26">
|
||
<taxonomicName id="4C016ED1FF83C467FE86FABAFE2AFAD0" authorityName="S. Julia & Kiew" authorityYear="2016" box="[262,451,1377,1398]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="26" phylum="Tracheophyta" rank="species" species="edgariana">Begonia edgariana</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
A. Habit (green form); B. Leaves of bronzy-dark green form; C. Stipule; D. Male flowers; E. Female flower; F. Female flower (dull red form); G. Female flower (white-tinged pink form) showing tepals and pistil; H. Mature fruit; I. Cross section of ovary. All photos from SFC 2758 by Julia Sang (A–H) and Ling Chea Yiing (I).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBE1552FF83C467FF3DFA34FD80F91D" blockId="9.[136,1452,1519,1795]" pageId="9" pageNumber="26">
|
||
<emphasis id="B975C940FF83C467FF3DFA34FEE0F9AE" bold="true" box="[189,265,1519,1544]" pageId="9" pageNumber="26">Notes:</emphasis>
|
||
—This species is common where it occurs. Two colour forms of leaves and flowers were observed in the field: (i) plants with green or bronzy-dark green leaves and (ii) male flowers usually white or occasionally dull red and the female flowers usually pale green or white tinged pink, occasionally dull red. However, the leaf colour is not associated with flower colour, thus plants with bronzy leaves may have white flowers and those with green leaves have red flowers. The species is monoecious but because the male phase does not overlap with the female, individual plants tend to have either female or male flowers.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF83C467FF3DF91CFD5DF8A5" blockId="9.[136,1452,1519,1795]" pageId="9" pageNumber="26">
|
||
<emphasis id="B975C940FF83C467FF3DF91CFDCDF946" bold="true" box="[189,548,1735,1760]" pageId="9" pageNumber="26">Proposed conservation status:</emphasis>
|
||
—Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |