treatments-xml/data/03/F1/A2/03F1A22EFF9ACB22FF728397FC06FBD3.xml
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<document id="49E879FDAE73F0E39C52752708FEA510" ID-DOI="10.11646/zootaxa.5260.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7794920" ID-ZooBank="7171477F-DB75-4CF6-9507-3329F5D7A9F7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680527871645" checkinUser="plazi" docAuthor="Blake, James A. &amp; Maciolek, Nancy J." docDate="2023" docId="03F1A22EFF9ACB22FF728397FC06FBD3" docLanguage="en" docName="zootaxa.5260.1.1.pdf" docOrigin="Zootaxa 5260 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heterospio dibranchiata Blake &amp; Maciolek 2023, new species" docType="treatment" docUuid="1FB493ED-53AC-4828-BE77-784002C767DD" docUuidSource="ZooBank" docVersion="6" lastPageNumber="27" masterDocId="FFC8DA56FF8CCB38FFE58038FFEAFFA3" masterDocTitle="New species and records of Heterospio (Annelida, Longosomatidae) from continental shelf, slope and abyssal depths of the Atlantic Ocean, Pacific Ocean, Indian Ocean and adjacent seas" masterLastPageNumber="74" masterPageNumber="1" pageNumber="23" updateTime="1699481072705" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="58E0431D09C6C2E150B66770E9166ED8">New species and records of Heterospio (Annelida, Longosomatidae) from continental shelf, slope and abyssal depths of the Atlantic Ocean, Pacific Ocean, Indian Ocean and adjacent seas</mods:title>
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<mods:namePart id="7F647FC93C83CE036F7C2D829D871A21">Blake, James A.</mods:namePart>
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<mods:namePart id="D38BB48DDCC3271C50F1FB0A7556A67F">Maciolek, Nancy J.</mods:namePart>
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<taxonomicName id="4C5868BBFF9ACB2EFF728397FE54FC6A" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[151,446,942,969]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="22" pageNumber="23" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF9ACB2EFF728397FE54FC6A" bold="true" box="[151,446,942,969]" italics="true" pageId="22" pageNumber="23">Heterospio dibranchiata</emphasis>
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<paragraph id="8BE71338FF9ACB2EFF7283EBFEF7FC4E" blockId="22.[151,968,942,1041]" box="[151,285,979,1005]" pageId="22" pageNumber="23">
<figureCitation id="13630FBDFF9ACB2EFF7283EBFEF7FC4E" box="[151,285,979,1005]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="24.[151,250,1944,1968]" captionStartId-1="25.[151,250,1936,1960]" captionTargetBox-0="[153,1435,181,1920]" captionTargetBox-1="[196,1391,200,1898]" captionTargetId-0="figure-18@24.[153,1435,181,1920]" captionTargetId-1="figure-17@25.[174,1412,181,1912]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." captionText-1="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi-0="http://doi.org/10.5281/zenodo.7794936" figureDoi-1="http://doi.org/10.5281/zenodo.7794938" httpUri-0="https://zenodo.org/record/7794936/files/figure.png" httpUri-1="https://zenodo.org/record/7794938/files/figure.png" pageId="22" pageNumber="23">Figures 89</figureCitation>
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urn:lsid:zoobank.org:act:
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<emphasis id="B92CCF2AFF9ACB2EFF728406FE96FBFB" bold="true" box="[151,380,1086,1112]" pageId="22" pageNumber="23">Material examined</emphasis>
. (
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)
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of
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, off
<collectingRegion id="499CDDDAFF9ACB2EFCD38406FC44FBFB" box="[822,942,1086,1112]" country="United States of America" name="Louisiana" pageId="22" pageNumber="23">Louisiana</collectingRegion>
.
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,
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coll.
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,
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Chief Scientist.
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M-5S,
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, BC,
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(
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;
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M-1S
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(2,
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M-2S
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,
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, BC, 28.738.738°N,
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,
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1
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(
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<location id="8E8745E3FF9ACB2EFB988492FB41FB67" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFB988492FB41FB67" box="[1149,1195,1194,1220]" latitude="28.737" longLatPrecision="73" longitude="-88.826" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-3S
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,
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<collectingDate id="EFA2CC10FF9ACB2EFAE78493FA72FB66" box="[1282,1432,1195,1221]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
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, BC,
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(
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)
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<emphasis id="B92CCF2AFF9ACB2EFCBF84F6FC3AFB4B" bold="true" box="[858,976,1230,1256]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFCBF84F6FC62FB4B" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFCBF84F6FC62FB4B" box="[858,904,1230,1256]" latitude="28.733" longLatPrecision="73" longitude="-88.826" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-4S
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,
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<collectingDate id="EFA2CC10FF9ACB2EFC3984F7FB85FB4A" box="[988,1135,1231,1257]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
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, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFB4984F7FAF1FB4A" box="[1196,1307,1231,1257]" degrees="28.733" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.733">28.733°N</geoCoordinate>
,
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,
<quantity id="4CA0BEDDFF9ACB2EFF7284CBFF0AFAAE" box="[151,224,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="9.08" pageId="22" pageNumber="23" unit="m" value="908.0">
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,
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(
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)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFDCE84CAFF12FA92" collectingDate="2008-11-10" collectionCode="MCZ" elevation="867" latitude="28.742" location="Sta." longLatPrecision="73" longitude="-88.822" pageId="22" pageNumber="23" specimenCode="MCZ 163733" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFDCE84CAFD49FAAF" bold="true" box="[555,675,1266,1292]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFDCE84CAFDB2FAAF" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFDCE84CAFDB2FAAF" box="[555,600,1266,1292]" latitude="28.742" longLatPrecision="73" longitude="-88.822" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-1E
</emphasis>
,
<date id="FFE635F8FF9ACB2EFD5484CBFCACFAAE" box="[689,838,1267,1293]" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFD5484CBFCACFAAE" box="[689,838,1267,1293]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFC6184CBFC19FAAE" box="[900,1011,1267,1293]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFC1B84CBFB9AFAAE" box="[1022,1136,1267,1293]" degrees="88.822" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.822">88.822°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFB9E84CBFB2EFAAE" box="[1147,1220,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="8.67" pageId="22" pageNumber="23" unit="m" value="867.0">
<elevation id="0075F40BFF9ACB2EFB9E84CBFB2EFAAE" box="[1147,1220,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="8.67" pageId="22" pageNumber="23" unit="m" value="867.0">867 m</elevation>
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,
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1
<emphasis id="B92CCF2AFF9ACB2EFB0184CBFAA5FAAE" bold="true" box="[1252,1359,1267,1293]" pageId="22" pageNumber="23">paratype</emphasis>
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(
<specimenCode id="DBFEBB43FF9ACB2EFABB84CBFF1AFA92" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163733</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFEE1852EFB01FA92" box="[260,1259,1302,1329]" collectingDate="2008-11-10" collectionCode="MCZ" elevation="867" latitude="28.742" location="Sta." longLatPrecision="73" longitude="-88.821" pageId="22" pageNumber="23" specimenCode="MCZ 163734" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFEE1852EFE95FA93" bold="true" box="[260,383,1302,1328]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFEE1852EFED8FA93" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFEE1852EFED8FA93" box="[260,306,1302,1328]" latitude="28.742" longLatPrecision="73" longitude="-88.821" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-2E
</emphasis>
,
<date id="FFE635F8FF9ACB2EFE68852FFDCEFA92" box="[397,548,1303,1329]" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFE68852FFDCEFA92" box="[397,548,1303,1329]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFD86852FFD39FA92" box="[611,723,1303,1329]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFD3A852FFCBBFA92" box="[735,849,1303,1329]" degrees="88.821" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.821">88.821°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFCB8852FFC4DFA92" box="[861,935,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="8.67" pageId="22" pageNumber="23" unit="m" value="867.0">
<elevation id="0075F40BFF9ACB2EFCB8852FFC4DFA92" box="[861,935,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="8.67" pageId="22" pageNumber="23" unit="m" value="867.0">867 m</elevation>
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,
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1
<emphasis id="B92CCF2AFF9ACB2EFC2F852FFBDFFA92" bold="true" box="[970,1077,1303,1329]" pageId="22" pageNumber="23">paratype</emphasis>
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(
<specimenCode id="DBFEBB43FF9ACB2EFBA1852FFB09FA92" box="[1092,1251,1303,1329]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163734</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFB12852EFC2EFAF6" collectingDate="2008-11-10" collectionCode="MCZ" elevation="868" latitude="28.742" location="Sta." longLatPrecision="490" longitude="-88.82" pageId="22" pageNumber="23" specimenCode="MCZ 163735" specimenCount="2" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFB12852EFA98FA93" bold="true" box="[1271,1394,1302,1328]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFB12852EFACFFA93" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFB12852EFACFFA93" box="[1271,1317,1302,1328]" latitude="28.742" longLatPrecision="490" longitude="-88.82" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-3E
</emphasis>
,
<date id="FFE635F8FF9ACB2EFA65852FFEE3FAF6" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFA65852FFEE3FAF6" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFEA28503FE5CFAF6" box="[327,438,1339,1365]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFE248503FDCCFAF6" box="[449,550,1339,1365]" degrees="88.82" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="-88.82">88.82°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFDD48503FD90FAF6" box="[561,634,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="22" pageNumber="23" unit="m" value="868.0">
<elevation id="0075F40BFF9ACB2EFDD48503FD90FAF6" box="[561,634,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="22" pageNumber="23" unit="m" value="868.0">868 m</elevation>
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,
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2
<emphasis id="B92CCF2AFF9ACB2EFD7F8503FCFAFAF6" bold="true" box="[666,784,1339,1365]" pageId="22" pageNumber="23">paratypes</emphasis>
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(
<specimenCode id="DBFEBB43FF9ACB2EFCFB8503FC57FAF6" box="[798,957,1338,1365]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163735</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFC2A8502FDDAFADA" collectingDate="2008-11-10" collectionCode="MCZ" elevation="891" latitude="28.742" location="Sta." longLatPrecision="73" longitude="-88.815" pageId="22" pageNumber="23" specimenCode="MCZ 163736" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFC2A8502FBA3FAF7" bold="true" box="[975,1097,1338,1364]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFC2A8502FC17FAF7" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFC2A8502FC17FAF7" box="[975,1021,1338,1364]" latitude="28.742" longLatPrecision="73" longitude="-88.815" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-4E
</emphasis>
,
<date id="FFE635F8FF9ACB2EFBB38503FB01FAF6" box="[1110,1259,1339,1365]" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFBB38503FB01FAF6" box="[1110,1259,1339,1365]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFACC8503FA72FAF6" box="[1321,1432,1339,1365]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFF728566FF14FADA" box="[151,254,1374,1401]" degrees="88.815" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.815">88.815W</geoCoordinate>
<quantity id="4CA0BEDDFF9ACB2EFEEC8567FEBEFADA" box="[265,340,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="8.91" pageId="22" pageNumber="23" unit="m" value="891.0">
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</quantity>
(1,
<specimenCode id="DBFEBB43FF9ACB2EFE628567FDC2FADA" box="[391,552,1375,1401]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163736</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFDDA8566FEA2FA3E" collectingDate="2008-11-09" collectionCode="MCZ" elevation="921" latitude="28.742" location="Sta." longLatPrecision="73" longitude="-88.805" pageId="22" pageNumber="23" specimenCode="MCZ 163737" specimenCount="3" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFDDA8566FD56FADA" bold="true" box="[575,700,1374,1401]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFDDA8566FD87FADB" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFDDA8566FD87FADB" box="[575,621,1374,1400]" latitude="28.742" longLatPrecision="73" longitude="-88.805" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-5E
</emphasis>
,
<date id="FFE635F8FF9ACB2EFD2B8567FC81FADA" box="[718,875,1375,1401]" pageId="22" pageNumber="23" value="2008-11-09">
<collectingDate id="EFA2CC10FF9ACB2EFD2B8567FC81FADA" box="[718,875,1375,1401]" pageId="22" pageNumber="23" value="2008-11-09">09 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFC548567FBCAFADA" box="[945,1056,1375,1401]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFBCA8566FB48FADA" box="[1071,1186,1374,1401]" degrees="88.805" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.805">88.805°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFB548567FB14FADA" box="[1201,1278,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="22" pageNumber="23" unit="m" value="921.0">
<elevation id="0075F40BFF9ACB2EFB548567FB14FADA" box="[1201,1278,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="22" pageNumber="23" unit="m" value="921.0">921 m</elevation>
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,
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3
<emphasis id="B92CCF2AFF9ACB2EFAC38567FA76FADA" bold="true" box="[1318,1436,1375,1401]" pageId="22" pageNumber="23">paratypes</emphasis>
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(
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)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFEB385BAFB0EFA3E" box="[342,1252,1410,1437]" collectingDate="2008-11-10" collectionCode="MCZ" elevation="825" latitude="28.746" location="Sta." longLatPrecision="73" longitude="-88.826" pageId="22" pageNumber="23" specimenCode="MCZ 163738" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFEB385BAFE39FA3F" bold="true" box="[342,467,1410,1436]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFEB385BAFE6EFA3F" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFEB385BAFE6EFA3F" box="[342,388,1410,1436]" latitude="28.746" longLatPrecision="73" longitude="-88.826" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-2N
</emphasis>
,
<date id="FFE635F8FF9ACB2EFE0185BBFD95FA3E" box="[484,639,1411,1437]" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFE0185BBFD95FA3E" box="[484,639,1411,1437]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFD2685BBFCD8FA3E" box="[707,818,1411,1437]" degrees="28.746" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.746">28.746°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFCA585BBFC5EFA3E" box="[832,948,1411,1437]" degrees="88.826" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.826">88.826°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFC2485BAFBE1FA3E" box="[961,1035,1410,1437]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="22" pageNumber="23" unit="m" value="825.0">
<elevation id="0075F40BFF9ACB2EFC2485BAFBE1FA3E" box="[961,1035,1410,1437]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="22" pageNumber="23" unit="m" value="825.0">825 m</elevation>
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(1,
<specimenCode id="DBFEBB43FF9ACB2EFBD985BBFB36FA3E" box="[1084,1244,1411,1437]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163738</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFB1785BAFC2DFA62" collectingDate="2008-11-10" collectionCode="MCZ" elevation="821" latitude="28.752" location="Sta." longLatPrecision="73" longitude="-88.826" pageId="22" pageNumber="23" specimenCode="MCZ 163739" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFB1785BAFA85FA3F" bold="true" box="[1266,1391,1410,1436]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFB1785BAFACAFA3F" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFB1785BAFACAFA3F" box="[1266,1312,1410,1436]" latitude="28.752" longLatPrecision="73" longitude="-88.826" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-4N
</emphasis>
,
<date id="FFE635F8FF9ACB2EFA6585BBFEE3FA62" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFA6585BBFEE3FA62" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFEA2859EFE5CFA62" box="[327,438,1446,1473]" degrees="28.752" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.752">28.752°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFE24859FFDDEFA62" box="[449,564,1447,1473]" degrees="88.826" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.826">88.826°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFDDA859FFD6DFA62" box="[575,647,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="8.209999999999999" pageId="22" pageNumber="23" unit="m" value="821.0">
<elevation id="0075F40BFF9ACB2EFDDA859FFD6DFA62" box="[575,647,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="8.209999999999999" pageId="22" pageNumber="23" unit="m" value="821.0">821 m</elevation>
</quantity>
,
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1
<emphasis id="B92CCF2AFF9ACB2EFD42859FFCF8FA62" bold="true" box="[679,786,1447,1473]" pageId="22" pageNumber="23">paratype</emphasis>
</specimenCount>
(
<specimenCode id="DBFEBB43FF9ACB2EFCC4859FFC55FA62" box="[801,959,1447,1473]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163739</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFC37859EFD5FFA46" collectingDate="2008-11-10" collectionCode="MCZ" elevation="850" latitude="28.742" location="Sta." longLatPrecision="73" longitude="-88.831" pageId="22" pageNumber="23" specimenCode="MCZ 163740" specimenCount="2" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFC37859EFBBEFA63" bold="true" box="[978,1108,1446,1472]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFC37859EFBEAFA63" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFC37859EFBEAFA63" box="[978,1024,1446,1472]" latitude="28.742" longLatPrecision="73" longitude="-88.831" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-3W
</emphasis>
,
<date id="FFE635F8FF9ACB2EFB87859FFB1CFA62" box="[1122,1270,1447,1473]" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFB87859FFB1CFA62" box="[1122,1270,1447,1473]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFAD1859FFA73FA62" box="[1332,1433,1447,1473]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFF7285F3FEE0FA46" box="[151,266,1483,1509]" degrees="88.831" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.831">88.831°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFEF285F2FE88FA46" box="[279,354,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="22" pageNumber="23" unit="m" value="850.0">
<elevation id="0075F40BFF9ACB2EFEF285F2FE88FA46" box="[279,354,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="22" pageNumber="23" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<specimenCount id="9D5ED8B1FF9ACB2EFE8A85F3FE16FA46" box="[367,508,1483,1509]" pageId="22" pageNumber="23" type="generic" typeStatus="paratypes">
2
<emphasis id="B92CCF2AFF9ACB2EFE6385F3FE16FA46" bold="true" box="[390,508,1483,1509]" pageId="22" pageNumber="23">paratypes</emphasis>
</specimenCount>
(
<specimenCode id="DBFEBB43FF9ACB2EFDE885F3FD47FA46" box="[525,685,1483,1509]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163740</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFD2785F2FE58F9AA" collectingDate="2008-11-10" collectionCode="MCZ" elevation="839" latitude="28.742" location="Sta." longLatPrecision="73" longitude="-88.837" pageId="22" pageNumber="23" specimenCode="MCZ 163741" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFD2785F2FCACFA47" bold="true" box="[706,838,1482,1508]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFD2785F2FD1AFA47" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFD2785F2FD1AFA47" box="[706,752,1482,1508]" latitude="28.742" longLatPrecision="73" longitude="-88.837" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
M-4W
</emphasis>
,
<date id="FFE635F8FF9ACB2EFCB385F3FC05FA46" box="[854,1007,1483,1509]" pageId="22" pageNumber="23" value="2008-11-10">
<collectingDate id="EFA2CC10FF9ACB2EFCB385F3FC05FA46" box="[854,1007,1483,1509]" pageId="22" pageNumber="23" value="2008-11-10">10 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFBD485F3FB4AFA46" box="[1073,1184,1483,1509]" degrees="28.742" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.742">28.742°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFB4B85F3FAC3FA46" box="[1198,1321,1483,1509]" degrees="88.837" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.837">88.837° W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFAD385F3FA6BFA46" box="[1334,1409,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="22" pageNumber="23" unit="m" value="839.0">
<elevation id="0075F40BFF9ACB2EFAD385F3FA6BFA46" box="[1334,1409,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="22" pageNumber="23" unit="m" value="839.0">839 m</elevation>
</quantity>
,
<specimenCount id="9D5ED8B1FF9ACB2EFA6B85F3FEE8F9AA" pageId="22" pageNumber="23" type="generic" typeStatus="paratype">
1
<emphasis id="B92CCF2AFF9ACB2EFF7285D7FEE8F9AA" bold="true" box="[151,258,1519,1545]" pageId="22" pageNumber="23">paratype</emphasis>
</specimenCount>
(
<specimenCode id="DBFEBB43FF9ACB2EFEF585D7FE40F9AA" box="[272,426,1519,1545]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163741</specimenCode>
)
</materialsCitation>
.—
<materialsCitation id="3B301965FF9ACB2EFE3085D6FD4CF98E" collectingDate="2008-11-11" collectionCode="MCZ" collectorName="J. A. Blake &amp; Chief Scientist. Sta." elevation="340" latitude="29.182" location="Virgo Platform Survey" longLatPrecision="73" longitude="-88.164" pageId="22" pageNumber="23" specimenCode="MCZ 163742" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFE3085D6FD02F9AB" bold="true" box="[469,744,1518,1544]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFE3085D6FD09F9AB" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFE3085D6FD09F9AB" box="[469,739,1518,1544]" latitude="29.182" longLatPrecision="73" longitude="-88.164" name="Virgo Platform Survey" pageId="22" pageNumber="23" stateProvince="Mexico">Virgo Platform Survey</location>
,
</emphasis>
coll.
<collectorName id="26AD76EEFF9ACB2EFCC085D7FC76F9AA" box="[805,924,1519,1545]" pageId="22" pageNumber="23">J.A. Blake</collectorName>
,
<collectorName id="26AD76EEFF9ACB2EFC4385D7FB6CF9AB" box="[934,1158,1518,1545]" pageId="22" pageNumber="23">
Chief Scientist.
<emphasis id="B92CCF2AFF9ACB2EFBBD85D6FB6CF9AB" bold="true" box="[1112,1158,1518,1544]" pageId="22" pageNumber="23">Sta.</emphasis>
</collectorName>
V-1E,
<date id="FFE635F8FF9ACB2EFB3185D7FA8CF9AA" box="[1236,1382,1519,1545]" pageId="22" pageNumber="23" value="2008-11-11">
<collectingDate id="EFA2CC10FF9ACB2EFB3185D7FA8CF9AA" box="[1236,1382,1519,1545]" pageId="22" pageNumber="23" value="2008-11-11">11 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFF72862BFEECF98E" box="[151,262,1555,1581]" degrees="29.182" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="29.182">29.182°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFEF4862BFE6EF98E" box="[273,388,1555,1581]" degrees="88.164" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.164">88.164°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFE6A862BFE3CF98E" box="[399,470,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="22" pageNumber="23" unit="m" value="340.0">
<elevation id="0075F40BFF9ACB2EFE6A862BFE3CF98E" box="[399,470,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="22" pageNumber="23" unit="m" value="340.0">340 m</elevation>
</quantity>
(1,
<specimenCode id="DBFEBB43FF9ACB2EFDE7862BFD74F98E" box="[514,670,1555,1581]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163742</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFD54862AFE84F9F2" collectingDate="2008-11-11" collectionCode="MCZ" elevation="340" latitude="29.185" location="Sta." longLatPrecision="73" longitude="-88.172" pageId="22" pageNumber="23" specimenCode="MCZ 167743" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFD54862AFCC1F98F" bold="true" box="[689,811,1554,1580]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFD54862AFD35F98F" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFD54862AFD35F98F" box="[689,735,1554,1580]" latitude="29.185" longLatPrecision="73" longitude="-88.172" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
V-1W
</emphasis>
,
<date id="FFE635F8FF9ACB2EFCDD862BFC26F98E" box="[824,972,1555,1581]" pageId="22" pageNumber="23" value="2008-11-11">
<collectingDate id="EFA2CC10FF9ACB2EFCDD862BFC26F98E" box="[824,972,1555,1581]" pageId="22" pageNumber="23" value="2008-11-11">11 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFBEF862AFB93F98E" box="[1034,1145,1554,1581]" degrees="29.185" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="29.185">29.185°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFB61862BFB1DF98E" box="[1156,1271,1555,1581]" degrees="88.172" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.172">88.172°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFAE7862BFAA3F98E" box="[1282,1353,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="22" pageNumber="23" unit="m" value="340.0">
<elevation id="0075F40BFF9ACB2EFAE7862BFAA3F98E" box="[1282,1353,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="22" pageNumber="23" unit="m" value="340.0">340 m</elevation>
</quantity>
(1 post end
<specimenCode id="DBFEBB43FF9ACB2EFF2D860FFE8CF9F2" box="[200,358,1591,1617]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 167743</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFE9E860EFB07F9F2" box="[379,1261,1590,1617]" collectingDate="2008-11-11" collectionCode="MCZ" elevation="335" latitude="29.185" location="Sta." longLatPrecision="73" longitude="-88.171" pageId="22" pageNumber="23" specimenCode="MCZ 163744" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFE9E860EFE05F9F3" bold="true" box="[379,495,1590,1616]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFE9E860EFE43F9F3" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFE9E860EFE43F9F3" box="[379,425,1590,1616]" latitude="29.185" longLatPrecision="73" longitude="-88.171" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
V-1N
</emphasis>
,
<date id="FFE635F8FF9ACB2EFE1B860FFD7FF9F2" box="[510,661,1591,1617]" pageId="22" pageNumber="23" value="2008-11-11">
<collectingDate id="EFA2CC10FF9ACB2EFE1B860FFD7FF9F2" box="[510,661,1591,1617]" pageId="22" pageNumber="23" value="2008-11-11">11 Nov 2008</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFD33860EFCAFF9F2" box="[726,837,1590,1617]" degrees="29.185" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="29.185">29.185°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFCB7860FFC2FF9F2" box="[850,965,1591,1617]" degrees="88.171" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.171">88.171°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFC37860EFBF0F9F2" box="[978,1050,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="3.35" pageId="22" pageNumber="23" unit="m" value="335.0">
<elevation id="0075F40BFF9ACB2EFC37860EFBF0F9F2" box="[978,1050,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="3.35" pageId="22" pageNumber="23" unit="m" value="335.0">335 m</elevation>
</quantity>
(1,
<specimenCode id="DBFEBB43FF9ACB2EFBAC860FFB0FF9F2" box="[1097,1253,1591,1617]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163744</specimenCode>
)
</materialsCitation>
.—
<materialsCitation id="3B301965FF9ACB2EFAEA860FFED7F93A" collectingDate="2009-06-05" collectionCode="MCZ" collectorName="P. A. Neubert &amp; Chief Scientist. Sta." elevation="533" latitude="28.872" location="Matterhorn Gas Pipeline Survey" longLatPrecision="489" longitude="-88.88" pageId="22" pageNumber="23" specimenCode="MCZ 163745" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFAEA860FFE67F9D7" bold="true" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFAEA860FFE6DF9D7" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFAEA860FFE6DF9D7" latitude="28.872" longLatPrecision="489" longitude="-88.88" name="Matterhorn Gas Pipeline Survey" pageId="22" pageNumber="23" stateProvince="Mexico">Matterhorn Gas Pipeline Survey</location>
,
</emphasis>
coll.
<collectorName id="26AD76EEFF9ACB2EFE2E8663FDB7F9D6" box="[459,605,1627,1653]" pageId="22" pageNumber="23">P.A. Neubert</collectorName>
,
<collectorName id="26AD76EEFF9ACB2EFD8D8663FCA2F9D7" box="[616,840,1626,1653]" pageId="22" pageNumber="23">
Chief Scientist.
<emphasis id="B92CCF2AFF9ACB2EFCFE8662FCA2F9D7" bold="true" box="[795,840,1626,1652]" pageId="22" pageNumber="23">Sta.</emphasis>
</collectorName>
2C,
<date id="FFE635F8FF9ACB2EFC9A8662FBE0F9D6" box="[895,1034,1626,1653]" pageId="22" pageNumber="23" value="2009-06-05">
<collectingDate id="EFA2CC10FF9ACB2EFC9A8662FBE0F9D6" box="[895,1034,1626,1653]" pageId="22" pageNumber="23" value="2009-06-05">05 Jun 2009</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFBA28663FB5DF9D6" box="[1095,1207,1627,1653]" degrees="28.872" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.872">28.872°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFB248663FACCF9D6" box="[1217,1318,1627,1653]" degrees="88.88" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="-88.88">88.88°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFAD48662FA92F9D6" box="[1329,1400,1626,1653]" metricMagnitude="2" metricUnit="m" metricValue="5.33" pageId="22" pageNumber="23" unit="m" value="533.0">
<elevation id="0075F40BFF9ACB2EFAD48662FA92F9D6" box="[1329,1400,1626,1653]" metricMagnitude="2" metricUnit="m" metricValue="5.33" pageId="22" pageNumber="23" unit="m" value="533.0">533 m</elevation>
</quantity>
(3,
<specimenCode id="DBFEBB43FF9ACB2EFF728647FEDFF93A" box="[151,309,1662,1689]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163745</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFEAC8646FB7CF93A" box="[329,1174,1662,1689]" collectingDate="2009-06-04" collectionCode="MCZ" elevation="282" latitude="28.872" location="Sta." longLatPrecision="73" longitude="-88.941" pageId="22" pageNumber="23" specimenCode="MCZ 163746" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFEAC8646FE4BF93B" bold="true" box="[329,417,1662,1688]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFEAC8646FE9DF93B" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFEAC8646FE9DF93B" box="[329,375,1662,1688]" latitude="28.872" longLatPrecision="73" longitude="-88.941" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
6C
</emphasis>
,
<date id="FFE635F8FF9ACB2EFE548647FDD5F93A" box="[433,575,1663,1689]" pageId="22" pageNumber="23" value="2009-06-04">
<collectingDate id="EFA2CC10FF9ACB2EFE548647FDD5F93A" box="[433,575,1663,1689]" pageId="22" pageNumber="23" value="2009-06-04">04 Jun 2009</collectingDate>
</date>
, BC,
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,
<geoCoordinate id="EE6C75FFFF9ACB2EFD1E8647FC84F93A" box="[763,878,1663,1689]" degrees="88.941" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.941">88.941°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFC9F8647FC28F93A" box="[890,962,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="22" pageNumber="23" unit="m" value="282.0">
<elevation id="0075F40BFF9ACB2EFC9F8647FC28F93A" box="[890,962,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="22" pageNumber="23" unit="m" value="282.0">282 m</elevation>
</quantity>
(1,
<specimenCode id="DBFEBB43FF9ACB2EFC148647FB64F93A" box="[1009,1166,1663,1689]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163746</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFB468646FD02F91E" collectingDate="2009-06-04" collectionCode="MCZ" elevation="241" latitude="28.863" location="Sta." longLatPrecision="73" longitude="-88.984" pageId="22" pageNumber="23" specimenCode="MCZ 163747" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFB468646FB11F93B" bold="true" box="[1187,1275,1662,1688]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFB468646FB3AF93B" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFB468646FB3AF93B" box="[1187,1232,1662,1688]" latitude="28.863" longLatPrecision="73" longitude="-88.984" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
8C
</emphasis>
,
<date id="FFE635F8FF9ACB2EFAEF8647FA72F93A" box="[1290,1432,1663,1689]" pageId="22" pageNumber="23" value="2009-06-04">
<collectingDate id="EFA2CC10FF9ACB2EFAEF8647FA72F93A" box="[1290,1432,1663,1689]" pageId="22" pageNumber="23" value="2009-06-04">04 Jun 2009</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFF29869BFED1F91E" box="[204,315,1699,1725]" degrees="28.863" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.863">28.863°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFEAC869BFE51F91E" box="[329,443,1699,1725]" degrees="88.984" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.984">88.984°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFE2C869BFDF8F91E" box="[457,530,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.41" pageId="22" pageNumber="23" unit="m" value="241.0">
<elevation id="0075F40BFF9ACB2EFE2C869BFDF8F91E" box="[457,530,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.41" pageId="22" pageNumber="23" unit="m" value="241.0">241 m</elevation>
</quantity>
(1,
<specimenCode id="DBFEBB43FF9ACB2EFDA7869BFD0AF91E" box="[578,736,1699,1725]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163747</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B301965FF9ACB2EFD10869AFED1F942" collectingDate="2009-06-04" collectionCode="MCZ" elevation="229" latitude="28.897" location="Sta." longLatPrecision="73" longitude="-88.946" pageId="22" pageNumber="23" specimenCode="MCZ 163748" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFD10869AFCA4F91F" bold="true" box="[757,846,1698,1724]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFD10869AFCC9F91F" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFD10869AFCC9F91F" box="[757,803,1698,1724]" latitude="28.897" longLatPrecision="73" longitude="-88.946" name="Sta." pageId="22" pageNumber="23" stateProvince="Mexico">Sta.</location>
9C
</emphasis>
,
<date id="FFE635F8FF9ACB2EFCBB869BFC04F91E" box="[862,1006,1699,1725]" pageId="22" pageNumber="23" value="2009-06-04">
<collectingDate id="EFA2CC10FF9ACB2EFCBB869BFC04F91E" box="[862,1006,1699,1725]" pageId="22" pageNumber="23" value="2009-06-04">04 Jun 2009</collectingDate>
</date>
, BC,
<geoCoordinate id="EE6C75FFFF9ACB2EFBD5869BFB75F91E" box="[1072,1183,1699,1725]" degrees="28.897" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.897">28.897°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFB49869BFACAF91E" box="[1196,1312,1699,1725]" degrees="88.946" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.946">88.946°W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFAC9869BFA9FF91E" box="[1324,1397,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="23" unit="m" value="229.0">
<elevation id="0075F40BFF9ACB2EFAC9869BFA9FF91E" box="[1324,1397,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="23" unit="m" value="229.0">229 m</elevation>
</quantity>
(2,
<specimenCode id="DBFEBB43FF9ACB2EFF7286FFFED9F942" box="[151,307,1735,1761]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 163748</specimenCode>
)
</materialsCitation>
.—
<materialsCitation id="3B301965FF9ACB2EFEB886FFFD76F8A6" collectingDate="2008-07-10" collectionCode="MCZ" collectorName="J. A. Blake" elevation="364" latitude="28.821" location="Matterhorn Oil Export Pipeline Survey" longLatPrecision="73" longitude="-88.985" pageId="22" pageNumber="23" specimenCode="MCZ 154127" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B92CCF2AFF9ACB2EFEB886FFFCD0F943" bold="true" box="[349,826,1734,1761]" pageId="22" pageNumber="23">
<location id="8E8745E3FF9ACB2EFEB886FFFCD0F943" LSID="urn:lsid:plazi:treatment:03F1A22EFF9ACB22FF728397FC06FBD3:8E8745E3FF9ACB2EFEB886FFFCD0F943" box="[349,826,1734,1761]" latitude="28.821" longLatPrecision="73" longitude="-88.985" name="Matterhorn Oil Export Pipeline Survey" pageId="22" pageNumber="23" stateProvince="Mexico">Matterhorn Oil Export Pipeline Survey</location>
</emphasis>
, coll.
<collectorName id="26AD76EEFF9ACB2EFC6786FFFC16F942" box="[898,1020,1735,1761]" pageId="22" pageNumber="23">J.A. Blake</collectorName>
,
<collectorName id="26AD76EEFF9ACB2EFBEC86FFFB58F942" box="[1033,1202,1735,1761]" pageId="22" pageNumber="23">Chief Scientist</collectorName>
,
<collectorName id="26AD76EEFF9ACB2EFB5A86FFFB00F942" box="[1215,1258,1735,1761]" pageId="22" pageNumber="23">Sta.</collectorName>
4,
<date id="FFE635F8FF9ACB2EFAF586FFFA72F942" box="[1296,1432,1735,1761]" pageId="22" pageNumber="23" value="2008-07-10">
<collectingDate id="EFA2CC10FF9ACB2EFAF586FFFA72F942" box="[1296,1432,1735,1761]" pageId="22" pageNumber="23" value="2008-07-10">10 Jul 2008</collectingDate>
</date>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFF7286D3FEECF8A6" box="[151,262,1771,1797]" degrees="28.821" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="28.821">28.821°N</geoCoordinate>
,
<geoCoordinate id="EE6C75FFFF9ACB2EFEF786D2FE93F8A6" box="[274,377,1770,1797]" degrees="88.985" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="-88.985">88.985W</geoCoordinate>
,
<quantity id="4CA0BEDDFF9ACB2EFE6186D3FE21F8A6" box="[388,459,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="3.6399999999999997" pageId="22" pageNumber="23" unit="m" value="364.0">
<elevation id="0075F40BFF9ACB2EFE6186D3FE21F8A6" box="[388,459,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="3.6399999999999997" pageId="22" pageNumber="23" unit="m" value="364.0">364 m</elevation>
</quantity>
(1,
<specimenCode id="DBFEBB43FF9ACB2EFE1D86D3FD7EF8A6" box="[504,660,1770,1797]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ 154127</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34240B3FF9ACB2FFF228736FAAFF836" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BE71338FF9ACB2FFF228736FD21FEC6" blockId="22.[151,1437,1086,2013]" lastBlockId="23.[151,1437,150,2013]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="B92CCF2AFF9ACB2EFF228736FEBBF88B" bold="true" box="[199,337,1806,1832]" pageId="22" pageNumber="23">Description</emphasis>
. An elongate, threadlike species.
<typeStatus id="54E3AD9AFF9ACB2EFD298737FCDEF88A" box="[716,820,1807,1833]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
(MCZ 163727) complete, with 27 setigers (9 thoracic, 15 abdominal, 3 on posterior section),
<quantity id="4CA0BEDDFF9ACB2EFDAF870BFD44F8EE" box="[586,686,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="22" pageNumber="23" unit="mm" value="14.9">14.9 mm</quantity>
long,
<quantity id="4CA0BEDDFF9ACB2EFD13870BFCA6F8EE" box="[758,844,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="0.1">0.1 mm</quantity>
wide across narrow thoracic setigers,
<quantity id="4CA0BEDDFF9ACB2EFB1F870BFAB5F8EE" box="[1274,1375,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="22" pageNumber="23" unit="mm" value="0.26">0.26 mm</quantity>
wide across middle abdominal segments, and 0.22 wide across bulbous posterior section (
<figureCitation id="13630FBDFF9ACB2EFBB7876FFB47F8D2" box="[1106,1197,1879,1905]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="22" pageNumber="23">Fig. 8A</figureCitation>
). A smaller complete specimen (MCZ 163744) with 28 setigers (9 thoracic, 15 abdominal, 4 on posterior section),
<quantity id="4CA0BEDDFF9ACB2EFB718743FB1DF836" box="[1172,1271,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="7.27" pageId="22" pageNumber="23" unit="mm" value="7.27">7.27 mm</quantity>
long,
<quantity id="4CA0BEDDFF9ACB2EFADF8743FA77F836" box="[1338,1437,1915,1941]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="mm" value="0.08">0.08 mm</quantity>
wide across thoracic setigers,
<quantity id="4CA0BEDDFF9ACB2EFE0787A7FDAFF81A" box="[482,581,1951,1977]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="mm" value="0.12">0.12 mm</quantity>
wide across abdominal segments, and
<quantity id="4CA0BEDDFF9ACB2EFC1687A7FBBCF81A" box="[1011,1110,1951,1977]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="22" pageNumber="23" unit="mm" value="0.07">0.07 mm</quantity>
across posterior section. One small complete juvenile also present (see below).
<typeStatus id="54E3AD9AFF9ACB2EFD2487FBFCC4F87E" box="[705,814,1987,2013]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
and other specimens incomplete. Large
<typeStatus id="54E3AD9AFF9ACB2EFB0A87FBFAA5F87E" box="[1263,1359,1987,2013]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
(MCZ 163735) with 13 setigers,
<quantity id="4CA0BEDDFF9BCB2FFE2780AFFDC2FF12" box="[450,552,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="23" pageNumber="24" unit="mm" value="23.3">23.3 mm</quantity>
long,
<quantity id="4CA0BEDDFF9BCB2FFD9780AFFD32FF12" box="[626,728,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="24" unit="mm" value="0.21">0.21 mm</quantity>
wide across thoracic setigers, and
<quantity id="4CA0BEDDFF9BCB2FFB8C80AFFB25FF12" box="[1129,1231,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="23" pageNumber="24" unit="mm" value="0.71">0.71 mm</quantity>
across abdominal segments; small
<typeStatus id="54E3AD9AFF9BCB2FFEB78083FE58FF76" box="[338,434,187,213]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
(MCZ 163728) with 13 setigers,
<quantity id="4CA0BEDDFF9BCB2FFCCF8082FC64FF76" box="[810,910,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.71" pageId="23" pageNumber="24" unit="mm" value="5.71">5.71 mm</quantity>
long, 0.15 across thoracic setigers, 0.12 across abdominal segments. Body divided into thoracic region with nine setigers, setigers 16 short, slightly wider than long; setigers 79 becoming progressively longer with setiger 7 about 2 times longer than setiger 6, setiger 8 about 3.5 times longer than setiger 6; setiger 9 greatly elongated, about 9 times longer than setiger 6. Color in alcohol opaque white to light tan; pigment entirely absent.
</paragraph>
<paragraph id="8BE71338FF9BCB2FFF228157FC7FFD26" blockId="23.[151,1437,150,2013]" pageId="23" pageNumber="24">
Pre-setiger region short, as long as first two thoracic setigers (
<figureCitation id="13630FBDFF9BCB2FFC448157FBCAFE2A" box="[929,1056,367,393]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">Fig. 9AC</figureCitation>
). Prostomium narrow, triangular, tapering to narrow apex (
<figureCitation id="13630FBDFF9BCB2FFE5581ABFDC1FE0E" box="[432,555,403,429]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">Fig. 9AC</figureCitation>
); eyespots absent; nuchal organs distinct slits on posterior lateral margins (
<figureCitation id="13630FBDFF9BCB2FFA8A81ABFF35FE72" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">Fig. 9AB</figureCitation>
). Dorsal surface with narrow ridge or crest extending from prostomium mid-dorsally over peristomium (
<figureCitation id="13630FBDFF9BCB2FFA8A818FFF56FE56" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">Fig. 9B</figureCitation>
). Peristomium a single ring, interrupted dorsally by dorsal ridge (
<figureCitation id="13630FBDFF9BCB2FFC4D81E3FBCCFE56" box="[936,1062,475,501]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">Fig. 9AB</figureCitation>
). Mouth narrow vertical opening bordered anteriorly by three short narrow lobes, laterally by two large protruding lobulated lobes that could be mistaken for protruding proboscis, and posteriorly by 79 lobes connecting to elongate narrow ridge extending posteriorly over peristomium and setigers 12 (
<figureCitation id="13630FBDFF9BCB2FFD59827FFCFDFDC2" box="[700,791,583,609]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">Fig. 9C</figureCitation>
). Dorsal tentacles observed on
<specimenCount id="9D5ED8B1FF9BCB2FFB66827FFAC9FDC2" box="[1155,1315,583,609]" pageId="23" pageNumber="24" type="generic">one specimen</specimenCount>
, arising in groove or notch lateral to dorsal crest; tentacles thickened filaments.
</paragraph>
<paragraph id="8BE71338FF9BCB2FFF2282B7FBD1FD52" blockId="23.[151,1437,150,2013]" pageId="23" pageNumber="24">
Branchiae present on setigers 23 (
<figureCitation id="13630FBDFF9BCB2FFDB282B7FD34FD0A" box="[599,734,655,681]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="23" pageNumber="24">Figs. 8AB</figureCitation>
,
<figureCitation id="13630FBDFF9BCB2FFD0E82B7FCC5FD0A" box="[747,815,655,681]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">9AB</figureCitation>
) on all specimens examined except smallest juveniles, where present only on setiger 2; branchiae long, thin (
<figureCitation id="13630FBDFF9BCB2FFD1A828BFCB2FD6E" box="[767,856,691,717]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="23" pageNumber="24">Fig. 8B</figureCitation>
), rounded in cross section, tapering to rounded tip; each with narrow ciliated groove; internal blood vessel extends along entire length.
</paragraph>
<paragraph id="8BE71338FF9BCB2FFF2282C3FC64FC6A" blockId="23.[151,1437,150,2013]" pageId="23" pageNumber="24">All parapodia biramous with setal fascicles arising from near anterior edge of segment. All thoracic setigers slightly flattened dorsally with parapodia weakly inflated and elevated over dorsum and bearing setal fascicles in tight bundles. Abdominal setigers 10 and following round in cross section and bearing setae in transverse rows encircling each anterior end of segment. Abdominal parapodia from setiger 10 narrow elongate lobes, becoming thicker and more prominent from setigers 1112; noto- and neuropodia with distinct dorsal and ventral gaps between setal fascicles; lateral gaps between noto- and neuropodia narrower.</paragraph>
<paragraph id="8BE71338FF9BCB2FFF2283EBFABDFA92" blockId="23.[151,1437,150,2013]" pageId="23" pageNumber="24">
Thoracic notopodia of setigers 19 with 1015 long capillaries in spreading fascicles; capillaries of neuropodia more numerous, with up to 2025 setae arranged in tight, dense fascicle. Noto- and neuropodia of setiger 10 with capillaries arranged in two elongate transverse rows with distinct dorsal and lateral gaps. Setiger 11 with notosetae of both rows all capillaries; neuropodia with anterior row of acicular spines and posterior row with capillaries; setiger 12 with both noto- and neuropodia with anterior row of acicular spines and posterior row of capillaries. Setiger 13 and all subsequent abdominal setigers (to setiger
<quantity id="4CA0BEDDFF9BCB2FFD5684BFFD01FB02" box="[691,747,1159,1185]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="23" pageNumber="24" unit="in" value="24.0">24 in</quantity>
<typeStatus id="54E3AD9AFF9BCB2FFD1584BFFCB9FB02" box="[752,851,1159,1185]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
) with both noto- and neuropodia with anterior row of acicular spines and posterior row of narrower spines with long aristate tip. Acicular spines thick, tapering to narrow pointed tip (
<figureCitation id="13630FBDFF9BCB2FFEC184F7FE43FB4A" box="[292,425,1231,1257]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="23" pageNumber="24">Figs. 8DE</figureCitation>
,
<figureCitation id="13630FBDFF9BCB2FFE5D84F7FE34FB4A" box="[440,478,1231,1257]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">9D</figureCitation>
); aristate spines narrower than acicular spines, with long, tapering tip (
<figureCitation id="13630FBDFF9BCB2FFAF684F7FA72FB4A" box="[1299,1432,1231,1257]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="23" pageNumber="24">Figs. 8DE</figureCitation>
,
<figureCitation id="13630FBDFF9BCB2FFF7284CBFF36FAAE" box="[151,220,1267,1293]" captionStart="FIGURE 9" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[196,1391,200,1898]" captionTargetId="figure-17@25.[174,1412,181,1912]" captionTargetPageId="25" captionText="FIGURE 9. Heterospio dibranchiata n. sp. A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini.A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727)." figureDoi="http://doi.org/10.5281/zenodo.7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="23" pageNumber="24">9EG</figureCitation>
). Far posterior bulbous section of
<typeStatus id="54E3AD9AFF9BCB2FFD8984CBFD27FAAE" box="[620,717,1267,1293]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
oval-shaped (
<figureCitation id="13630FBDFF9BCB2FFC9584CBFC20FAAE" box="[880,970,1267,1293]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="23" pageNumber="24">Fig. 8C</figureCitation>
), with terminal anal opening; with three parapodia, each bearing 12 hooked spines in each ramus; each spine short, with curved pointed tip (
<figureCitation id="13630FBDFF9BCB2FFB10852FFAA0FA92" box="[1269,1354,1303,1329]" captionStart="FIGURE 8" captionStartId="24.[151,250,1944,1968]" captionTargetBox="[153,1435,181,1920]" captionTargetId="figure-18@24.[153,1435,181,1920]" captionTargetPageId="24" captionText="FIGURE 8. Heterospio dibranchiata n. sp. Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="23" pageNumber="24">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8BE71338FF9BCB2FFF228502FF08F942" blockId="23.[151,1437,150,2013]" pageId="23" pageNumber="24">
<emphasis id="B92CCF2AFF9BCB2FFF228502FE29FAF6" bold="true" box="[199,451,1338,1365]" pageId="23" pageNumber="24">Juvenile morphology</emphasis>
. A complete juvenile with 28 setigers (MCZ 163745) with nine thoracic, 16 abdominal, and three posterior setigers;
<quantity id="4CA0BEDDFF9BCB2FFE3D8567FDD7FADA" box="[472,573,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="4.94" pageId="23" pageNumber="24" unit="mm" value="4.94">4.94 mm</quantity>
long and
<quantity id="4CA0BEDDFF9BCB2FFD4A8567FCFEFADA" box="[687,788,1375,1401]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="24" unit="mm" value="0.06">0.06 mm</quantity>
across anterior setigers; body consistently narrow except for bulbous posterior end. A pair of long branchiae present on setiger 2 but absent on setiger 3. Setigers 7, 8, and 9 longer than individual setigers 16, but setiger 9 not as long proportionately as in adults; all abdominal segments of a relatively equal length.A second complete juvenile from Sta. 6C (MCZ 163746) with 28 setigers: nine thoracic, 15 abdominal, and four posterior, measuring
<quantity id="4CA0BEDDFF9BCB2FFD8085D7FD50F9AA" box="[613,698,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="23" pageNumber="24" unit="mm" value="3.3">3.3 mm</quantity>
long,
<quantity id="4CA0BEDDFF9BCB2FFD1B85D7FC8AF9AA" box="[766,864,1519,1545]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="23" pageNumber="24" unit="mm" value="0.73">0.73 mm</quantity>
across thoracic setigers,
<quantity id="4CA0BEDDFF9BCB2FFB9685D7FB3FF9AA" box="[1139,1237,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="23" pageNumber="24" unit="mm" value="1.22">1.22 mm</quantity>
across abdominal setigers, and 0.91 across the posterior section. This second juvenile with abdominal segments contracted, thus appearing shorter. Posterior bulbous section of juveniles similar in appearance to that of adults. Transverse setal fascicles of abdominal segments greatly reduced in juveniles, restricted to simple noto- and neuropodia with fewer setae (46 spines per fascicle), but have same arrangement of acicular spines and aristate spines in two rows as in adults from setiger 13. Short curved hooks in posterior region of same morphology, arrangement, and number as in adults.
</paragraph>
<paragraph id="8BE71338FF9BCB2FFF2286D3FDCAF8D2" blockId="23.[151,1437,150,2013]" pageId="23" pageNumber="24">
These observations suggest that full segmental complement of 2728 or more setigers of
<taxonomicName id="4C5868BBFF9BCB2FFB6986D3FA77F8A6" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[1164,1437,1771,1797]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF9BCB2FFB6986D3FA77F8A6" box="[1164,1437,1771,1797]" italics="true" pageId="23" pageNumber="24">Heterospio dibranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF9BCB2FFF728737FF3DF88A" box="[151,215,1807,1833]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B92CCF2AFF9BCB2FFF728737FF3AF88A" bold="true" box="[151,208,1807,1833]" pageId="23" pageNumber="24">n. sp</emphasis>
.
</taxonomicNameLabel>
develop early in juveniles and that subsequent growth involves elongation, enlargement, and elaboration of individual segments and parapodia as well as an increase in number of setae. It is likely that this same growth pattern occurs in all species of
<taxonomicName id="4C5868BBFF9BCB2FFE78876FFDF2F8D3" authorityName="Ehlers" authorityYear="1874" box="[413,536,1879,1904]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="23" pageNumber="24" phylum="Annelida" rank="genus">
<emphasis id="B92CCF2AFF9BCB2FFE78876FFDF2F8D3" box="[413,536,1879,1904]" italics="true" pageId="23" pageNumber="24">Heterospio</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE71338FF9BCB2FFF228743FAAFF836" blockId="23.[151,1437,150,2013]" box="[199,1349,1914,1941]" pageId="23" pageNumber="24">
<emphasis id="B92CCF2AFF9BCB2FFF228743FE3EF837" bold="true" box="[199,468,1914,1941]" pageId="23" pageNumber="24">Methyl Green staining</emphasis>
. Body stains uniformly; weak staining of pre-setigerous region not prominent.
</paragraph>
</subSubSection>
<subSubSection id="C34240B3FF9BCB22FF2287A7FC63FD01" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BE71338FF9BCB22FF2287A7FC63FD01" blockId="23.[151,1437,150,2013]" lastBlockId="26.[151,1437,151,1136]" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
<emphasis id="B92CCF2AFF9BCB2FFF2287A7FEDEF81A" bold="true" box="[199,308,1951,1977]" pageId="23" pageNumber="24">Remarks</emphasis>
.
<taxonomicName id="4C5868BBFF9BCB2FFEDB87A7FDA7F81A" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[318,589,1951,1977]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF9BCB2FFEDB87A7FDA7F81A" box="[318,589,1951,1977]" italics="true" pageId="23" pageNumber="24">Heterospio dibranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF9BCB2FFDB587A7FD61F81A" box="[592,651,1951,1977]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B92CCF2AFF9BCB2FFDB587A7FD69F81A" bold="true" box="[592,643,1951,1977]" pageId="23" pageNumber="24">n.sp</emphasis>
.
</taxonomicNameLabel>
is unusual within the genus in having only two pairs of branchiae(setigers 23); these branchiae are remarkably long, and when complete are as long as one-third or more of the body length. Other unusual characters include: (1) a conical prostomium with a narrow anterior apex and a posterior extension or caruncle that extends over the peristomium as a dorsal crest merging with setiger 1, (2) a single peristomial ring instead of two where the curved nuchal organs can be seen immediately anterior to the peristomium on the posterior lateral margin of the prostomium, (3) a notch on the anterior border of the peristomium where the dorsal tentacles are attached, (4) an unusual oral apparatus where a narrow vertical oral opening is bordered laterally by two large protruding lobes resembling a protruding pharynx, and (5) only six short thoracic segments with setigers 7, 8, and 9 becoming increasingly longer, and setiger 9 is as long as setiger 10 of other species. Other characters are more typical of known species, such as the first nine setigers (thorax) having widely spaced and well-defined noto- and neuropodia with setal fascicles in setiger 10 that transition to setae arranged in transverse rows across each noto- and neuropodia, nearly encircling the body; when spines are present, these encircling rows of setae resemble the cinctures of spines found in cirratulids of the genus
<taxonomicName id="4C5868BBFF96CB22FD3381C2FCB0FDB7" box="[726,858,506,532]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="26" pageNumber="27" phylum="Annelida" rank="genus">
<emphasis id="B92CCF2AFF96CB22FD3381C2FCB0FDB7" box="[726,858,506,532]" italics="true" pageId="26" pageNumber="27">Chaetozone</emphasis>
</taxonomicName>
. The hooked spines of the posterior bulbous section are similar to those reported for
<taxonomicName id="4C5868BBFF96CB22FE1A8226FD70FD94" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[511,666,541,567]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="B92CCF2AFF96CB22FE1A8226FD70FD94" box="[511,666,541,567]" italics="true" pageId="26" pageNumber="27">H. hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF96CB22FD448225FD0AFD9B" box="[673,736,541,568]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B92CCF2AFF96CB22FD448225FD33FD94" bold="true" box="[673,729,541,567]" pageId="26" pageNumber="27">n. sp</emphasis>
.
</taxonomicNameLabel>
and others in this study. Another species in this study found to have only two pairs of branchiae and a single peristomial ring is
<taxonomicName id="4C5868BBFF96CB22FC96827AFC3AFDF8" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[883,976,577,603]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="alata" status="sp. nov.">
<emphasis id="B92CCF2AFF96CB22FC96827AFC3AFDF8" box="[883,976,577,603]" italics="true" pageId="26" pageNumber="27">H. alata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF96CB22FC328279FBFDFDF8" box="[983,1047,577,603]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B92CCF2AFF96CB22FC328279FBE5FDF8" bold="true" box="[983,1039,577,603]" pageId="26" pageNumber="27">n. sp</emphasis>
.
</taxonomicNameLabel>
from off
<collectingCountry id="F34F53A8FF96CB22FB618279FB39FDF8" box="[1156,1235,577,603]" name="Brunei" pageId="26" pageNumber="27">Brunei</collectingCountry>
, Island of Borneo (see below). However
<taxonomicName id="4C5868BBFF96CB22FE7F825DFE12FDDD" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[410,504,612,638]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="alata" status="sp. nov.">
<emphasis id="B92CCF2AFF96CB22FE7F825DFE12FDDD" box="[410,504,612,638]" italics="true" pageId="26" pageNumber="27">H. alata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF96CB22FDE4825CFDA8FDDC" box="[513,578,612,639]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B92CCF2AFF96CB22FDE4825CFDD0FDDD" bold="true" box="[513,570,612,638]" pageId="26" pageNumber="27">n. sp</emphasis>
.
</taxonomicNameLabel>
is one of only four known species to have neuropodial hooks on setiger 1, whereas
<taxonomicName id="4C5868BBFF96CB22FF1F82B0FE47FD01" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[250,429,648,674]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF96CB22FF1F82B0FE47FD01" box="[250,429,648,674]" italics="true" pageId="26" pageNumber="27">H. dibranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF96CB22FE5182B0FE19FD01" box="[436,499,648,674]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B92CCF2AFF96CB22FE5182B0FE01FD01" bold="true" box="[436,491,648,674]" pageId="26" pageNumber="27">n. sp</emphasis>
.
</taxonomicNameLabel>
has only capillaries in that position.
</paragraph>
</subSubSection>
<caption id="DF2743B0FF94CB20FF7287A0FE49F85B" ID-DOI="http://doi.org/10.5281/zenodo.7794936" ID-Zenodo-Dep="7794936" httpUri="https://zenodo.org/record/7794936/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1944,1968]" targetBox="[153,1435,181,1920]" targetPageId="24">
<paragraph id="8BE71338FF94CB20FF7287A0FE49F85B" blockId="24.[151,1436,1944,2040]" pageId="24" pageNumber="25">
<emphasis id="B92CCF2AFF94CB20FF7287A0FEE5F813" bold="true" box="[151,271,1944,1968]" pageId="24" pageNumber="25">FIGURE 8</emphasis>
.
<taxonomicName id="4C5868BBFF94CB20FEF887A1FDFDF813" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[285,535,1944,1968]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="24" pageNumber="25" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF94CB20FEF887A1FDFDF813" box="[285,535,1944,1968]" italics="true" pageId="24" pageNumber="25">Heterospio dibranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF94CB20FDFA87A0FDB3F813" box="[543,601,1944,1968]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B92CCF2AFF94CB20FDFA87A0FDB9F813" bold="true" box="[543,595,1944,1968]" pageId="24" pageNumber="25">n. sp</emphasis>
.
</taxonomicNameLabel>
Holotype (MCZ 163727): A, entire worm; B, anterior end, right lateral view; C, posterior end, right lateral view; D, subuluncini setiger 16; E, subuluncini and acicular spine setiger 13; F, posterior hook. AC, stained with Shirlastain A.
</paragraph>
</caption>
<caption id="DF2743B0FF95CB21FF7287A8FF19F853" ID-DOI="http://doi.org/10.5281/zenodo.7794938" ID-Zenodo-Dep="7794938" httpUri="https://zenodo.org/record/7794938/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1936,1960]" targetBox="[196,1391,200,1898]" targetPageId="25">
<paragraph id="8BE71338FF95CB21FF7287A8FF19F853" blockId="25.[151,1437,1936,2032]" pageId="25" pageNumber="26">
<emphasis id="B92CCF2AFF95CB21FF7287A8FEFAF80B" bold="true" box="[151,272,1936,1960]" pageId="25" pageNumber="26">FIGURE 9</emphasis>
.
<taxonomicName id="4C5868BBFF95CB21FEFB87A9FDF0F80B" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[286,538,1936,1960]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="25" pageNumber="26" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF95CB21FEFB87A9FDF0F80B" box="[286,538,1936,1960]" italics="true" pageId="25" pageNumber="26">Heterospio dibranchiata</emphasis>
</taxonomicName>
<emphasis id="B92CCF2AFF95CB21FDC687A8FDB4F80B" bold="true" box="[547,606,1936,1960]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A21F7251FF95CB21FDC687A8FDB4F80B" box="[547,606,1936,1960]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A, anterior end, left lateral view; B, anterior end, dorsal view; C, anterior end, ventral view; D, acicular spines; EG, subuluncini. A, C, paratype (MCZ 163728), B, paratype (163735), DG, holotype (MCZ 163727).
</paragraph>
</caption>
<subSubSection id="C34240B3FF96CB22FF228293FE1DFBA7" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="8BE71338FF96CB22FF228293FDD3FCAF" blockId="26.[151,1437,151,1136]" pageId="26" pageNumber="27">
<emphasis id="B92CCF2AFF96CB22FF228293FECBFD66" bold="true" box="[199,289,683,709]" pageId="26" pageNumber="27">Biology</emphasis>
. The presence of four complete specimens in these samples is unusual for a species of
<taxonomicName id="4C5868BBFF96CB22FB148294FA86FD66" authorityName="Ehlers" authorityYear="1874" box="[1265,1388,684,709]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="genus">
<emphasis id="B92CCF2AFF96CB22FB148294FA86FD66" box="[1265,1388,684,709]" italics="true" pageId="26" pageNumber="27">Heterospio</emphasis>
</taxonomicName>
and likely represents the careful manner in which they were collected; gentle handling minimized fragmentation and damage (see Materials and methods).
</paragraph>
<paragraph id="8BE71338FF96CB22FF22832EFE1DFBA7" blockId="26.[151,1437,151,1136]" pageId="26" pageNumber="27">
<taxonomicName id="4C5868BBFF96CB22FF22832EFE33FC8C" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[199,473,789,815]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF96CB22FF22832EFE33FC8C" box="[199,473,789,815]" italics="true" pageId="26" pageNumber="27">Heterospio dibranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF96CB22FE3B832EFDF6FC93" box="[478,540,790,816]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B92CCF2AFF96CB22FE3B832EFDFEFC93" bold="true" box="[478,532,790,816]" pageId="26" pageNumber="27">n. sp</emphasis>
.
</taxonomicNameLabel>
was widely distributed in the various survey samples at the platform and pipeline locations ranging from
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241
<emphasis id="B92CCF2AFF96CB22FE2A8301FE37FCF0" bold="true" box="[463,477,825,851]" pageId="26" pageNumber="27"></emphasis>
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</quantity>
. About a third of the specimens (10) were collected from a single sample at Station M-5S at a depth of
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, the deepest location from which the species was collected. Sediment grain size at Sta. M-5S was sand (1.86%), silt (34.00%), and clay (64.14%), with TOC at 1.24%. Station M-5S (
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) was the only location where
<taxonomicName id="4C5868BBFF96CB22FE3A839CFD79FC1E" authority="Blake &amp; Maciolek, 2023" authorityName="Blake &amp; Maciolek" authorityYear="2023" box="[479,659,931,957]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="dibranchiata" status="sp. nov.">
<emphasis id="B92CCF2AFF96CB22FE3A839CFD79FC1E" box="[479,659,931,957]" italics="true" pageId="26" pageNumber="27">H. dibranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7251FF96CB22FD7E839CFD36FC1D" box="[667,732,932,958]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B92CCF2AFF96CB22FD7E839CFD3EFC1D" bold="true" box="[667,724,932,958]" pageId="26" pageNumber="27">n. sp</emphasis>
.
</taxonomicNameLabel>
was among the top 10 species. The station was dominated by aplacophoran molluscs, polychaetes including cirratulids, paraonids, and spionids, and a thyasirid bivalve mollusc (Blake
<emphasis id="B92CCF2AFF96CB22FF0C83D3FECBFBA7" box="[233,289,1002,1028]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C34240B3FF96CB22FF228436FD54FBE8" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8BE71338FF96CB22FF228436FD54FBE8" blockId="26.[151,1437,151,1136]" pageId="26" pageNumber="27">
<emphasis id="B92CCF2AFF96CB22FF228436FEA2FB8B" bold="true" box="[199,328,1038,1064]" pageId="26" pageNumber="27">Etymology</emphasis>
. The epithet
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<emphasis id="B92CCF2AFF96CB22FE328436FD82FB8B" box="[471,616,1038,1064]" italics="true" pageId="26" pageNumber="27">dibranchiata</emphasis>
</taxonomicName>
is from the Greek,
<emphasis id="B92CCF2AFF96CB22FCDE8436FCBBFB8B" box="[827,849,1038,1064]" italics="true" pageId="26" pageNumber="27">di</emphasis>
-, for two or double and
<emphasis id="B92CCF2AFF96CB22FBBF8436FB28FB8B" box="[1114,1218,1038,1064]" italics="true" pageId="26" pageNumber="27">branchos</emphasis>
for gill in reference to this species having only two pair of branchiae.
</paragraph>
</subSubSection>
<subSubSection id="C34240B3FF96CB22FF22846DFC06FBD3" box="[199,1004,1109,1136]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BE71338FF96CB22FF22846DFC06FBD3" blockId="26.[151,1437,151,1136]" box="[199,1004,1109,1136]" pageId="26" pageNumber="27">
<emphasis id="B92CCF2AFF96CB22FF22846DFEB3FBCC" bold="true" box="[199,345,1109,1135]" pageId="26" pageNumber="27">Distribution</emphasis>
.
<collectingCountry id="F34F53A8FF96CB22FE82846DFE73FBCC" box="[359,409,1109,1135]" name="United States of America" pageId="26" pageNumber="27">U.S.</collectingCountry>
northern Gulf of Mexico, off
<collectingRegion id="499CDDDAFF96CB22FD0C846DFCB0FBCC" box="[745,858,1109,1135]" country="United States of America" name="Louisiana" pageId="26" pageNumber="27">Louisiana</collectingRegion>
,
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241
<specimenCount id="9D5ED8B1FF96CB22FC7B846DFC02FBD3" box="[926,1000,1109,1136]" pageId="26" pageNumber="27" type="generic">955 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>