treatments-xml/data/03/ED/FB/03EDFB556D1DFF90FBF3C2341AD4FDA6.xml

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<mods:title id="8D367D72E3AF49BAD6D3D47AAE00F789">Lethrus (Scelolethrus) golestanicus (Coleoptera: Geotrupidae: Lethrinae), a new species from Iran</mods:title>
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<paragraph id="8BFB4A436D1DFF93FBF3C2341BDDF861" blockId="0.[151,1317,1917,2016]" box="[151,927,1917,1944]" pageId="0" pageNumber="585">
<heading id="D0B3FD2F6D1DFF93FBF3C2341BDDF861" bold="true" box="[151,927,1917,1944]" fontSize="11" level="1" pageId="0" pageNumber="585" reason="1">
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<taxonomicName id="4C4431C06D1DFF93FBF3C2341B40F861" ID-CoL="8RXWW" authority="Kral &amp; Hillert" authorityName="Kral &amp; Hillert" authorityYear="2021" box="[151,770,1917,1944]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="species" species="golestanicus" status="sp. nov." subGenus="Scelolethrus">
<emphasis id="B93096516D1DFF93FBF3C23418B4F861" bold="true" box="[151,246,1917,1944]" italics="true" pageId="0" pageNumber="585">Lethrus</emphasis>
(
<emphasis id="B93096516D1DFF93FA63C23419DFF861" bold="true" box="[263,413,1917,1944]" italics="true" pageId="0" pageNumber="585">Scelolethrus</emphasis>
) 
<emphasis id="B93096516D1DFF93FACAC2341A06F861" bold="true" box="[430,580,1917,1944]" italics="true" pageId="0" pageNumber="585">golestanicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
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<subSubSection id="C35E19C86D1DFF93FBF3C2ED19E9F845" box="[151,427,1956,1980]" pageId="0" pageNumber="585" type="description">
<paragraph id="8BFB4A436D1DFF93FBF3C2ED19E9F845" blockId="0.[151,1317,1917,2016]" box="[151,427,1956,1980]" pageId="0" pageNumber="585">
(
<figureCitation id="137F56C66D1DFF93FBFBC2ED196EF845" box="[159,300,1956,1980]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–7" captionStart-2="FIGURES 8–13" captionStartId-0="1.[151,264,1624,1648]" captionStartId-1="2.[151,264,1105,1129]" captionStartId-2="3.[151,264,1939,1963]" captionTargetBox-0="[319,1284,382,1584]" captionTargetBox-1="[305,1283,182,1067]" captionTargetBox-2="[216,1374,634,1904]" captionTargetId-0="figure-111@1.[303,1285,378,1598]" captionTargetId-1="figure-19@2.[303,1285,181,1081]" captionTargetId-2="figure-247@3.[213,1375,631,1914]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 1–3. Habitus of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 1, holotype, ♂, dorsal view; 2, al- lotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale." captionText-1="FIGURES 4–7. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." captionText-2="FIGURES 8–13. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 8–10, head, dorsal view; 11–13, head, left lateral view. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5577371" figureDoi-1="http://doi.org/10.5281/zenodo.5577375" figureDoi-2="http://doi.org/10.5281/zenodo.5577377" httpUri-0="https://zenodo.org/record/5577371/files/figure.png" httpUri-1="https://zenodo.org/record/5577375/files/figure.png" httpUri-2="https://zenodo.org/record/5577377/files/figure.png" pageId="0" pageNumber="585">Figs. 1–8, 11</figureCitation>
, 
<figureCitation id="137F56C66D1DFF93FA5CC2ED1917F845" box="[312,341,1956,1980]" captionStart="FIGURES 14–16" captionStartId="4.[151,264,713,737]" captionTargetBox="[261,1321,188,676]" captionTargetId="figure-19@4.[258,1329,181,689]" captionTargetPageId="4" captionText="FIGURES 14–16. Lethrus (Scelolethrus) spp., parameres, dorsal view. 14, L. (S.) golestanicus Král &amp; Hillert, new species; 15, L. (S.) mithras Reitter, 1904 (Iran, Bojnurd); 16, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577381" httpUri="https://zenodo.org/record/5577381/files/figure.png" pageId="0" pageNumber="585">14</figureCitation>
, 
<figureCitation id="137F56C66D1DFF93FA04C2ED19E1F845" box="[352,419,1956,1980]" captionStart="FIGURES 17–18" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[377,1210,836,1938]" captionTargetId="figure-75@4.[377,1210,833,1939]" captionTargetPageId="4" captionText="FIGURES 17–18. Type locality of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. Iran, Golestan province, N of Qareh Makher, 13.iv.2016 (photo by David Král)." figureDoi="http://doi.org/10.5281/zenodo.5577383" httpUri="https://zenodo.org/record/5577383/files/figure.png" pageId="0" pageNumber="585">17–18</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BFB4A436D1DFF93FBF3C2811D67F819" blockId="0.[151,1317,1917,2016]" box="[151,1317,1991,2016]" pageId="0" pageNumber="585">
<emphasis id="B93096516D1DFF93FBF3C2811972F826" box="[151,304,1991,2016]" inLineHeading="true" pageId="0" pageNumber="585" reason="1">
<emphasis id="B93096516D1DFF93FBF3C281196BF826" bold="true" box="[151,297,1991,2016]" pageId="0" pageNumber="585">
<typeStatus id="54FFF4E16D1DFF93FBF3C281188DF819" box="[151,207,1992,2016]" pageId="0" pageNumber="585">Type</typeStatus>
locality
</emphasis>
.
</emphasis>
<collectingCountry id="F3530AD36D1DFF93FA52C2811926F826" box="[310,356,1992,2015]" name="Iran" pageId="0" pageNumber="585">Iran</collectingCountry>
, 
<collectingRegion id="498084A16D1DFF93FA0AC2811A76F819" box="[366,564,1992,2016]" country="Iran" name="Golestan" pageId="0" pageNumber="585">Golestan province</collectingRegion>
, N of 
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, 
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<geoCoordinate id="EE702C846D1DFF93F81BC2811B99F819" box="[895,987,1992,2016]" degrees="54" direction="east" minutes="59" orientation="longitude" pageId="0" pageNumber="585" precision="925" value="54.983334">54°59’E</geoCoordinate>
, 
<quantity id="4CBCE7A66D1DFF93F881C2811C5EF826" box="[997,1052,1992,2016]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="585" unit="m" value="40.0">
<elevation id="0069AD706D1DFF93F881C2811C5EF826" box="[997,1052,1992,2016]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="585" unit="m" value="40.0">40 m</elevation>
</quantity>
[a.s.l.] (see 
<figureCitation id="137F56C66D1DFF93FFFAC2811D5FF819" box="[1182,1309,1992,2016]" captionStart="FIGURES 17–18" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[377,1210,836,1938]" captionTargetId="figure-75@4.[377,1210,833,1939]" captionTargetPageId="4" captionText="FIGURES 17–18. Type locality of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. Iran, Golestan province, N of Qareh Makher, 13.iv.2016 (photo by David Král)." figureDoi="http://doi.org/10.5281/zenodo.5577383" httpUri="https://zenodo.org/record/5577383/files/figure.png" pageId="0" pageNumber="585">Figs. 17–18</figureCitation>
)
</paragraph>
<paragraph id="8BFB4A436D1CFF92FBF3C5DE1CCBFF0E" blockId="1.[151,1437,151,355]" pageId="1" pageNumber="586">
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<emphasis id="B93096516D1CFF92FBF3C5DE1976FF56" bold="true" box="[151,308,151,175]" pageId="1" pageNumber="586">Type material</emphasis>
.
</emphasis>
<emphasis id="B93096516D1CFF92FA25C5DE1A6FFF56" bold="true" box="[321,557,151,175]" pageId="1" pageNumber="586">
Iran, 
<collectingRegion id="498084A16D1CFF92FA1BC5DE1A6AFF56" box="[383,552,151,175]" country="Iran" name="Golestan" pageId="1" pageNumber="586">Golestan Prov.</collectingRegion>
:
</emphasis>
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<typeStatus id="54FFF4E16D1CFF92F957C5DE1ADBFF56" box="[563,665,151,175]" pageId="1" pageNumber="586" type="holotype">Holotype</typeStatus>
, 
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(
<collectionCode id="ED55D2866D1CFF92F9A7C5DE1B56FF56" box="[707,788,151,175]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="586">NMPC</collectionCode>
), “ 
<collectingCountry id="F3530AD36D1CFF92F84AC5DE1B36FF56" box="[814,884,151,175]" name="Iran" pageId="1" pageNumber="586">IRAN</collectingCountry>
, 
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| N of QAREH MAKHER | 
<geoCoordinate id="EE702C846D1CFF92FE24C5DE1DDFFF56" box="[1344,1437,151,175]" degrees="37" direction="north" minutes="32" orientation="latitude" pageId="1" pageNumber="586" precision="925" value="37.533333">37°32‘N</geoCoordinate>
<geoCoordinate id="EE702C846D1CFF92FBF3C5F218B1FF2A" box="[151,243,187,211]" degrees="54" direction="east" minutes="59" orientation="longitude" pageId="1" pageNumber="586" precision="925" value="54.983334">54°59‘E</geoCoordinate>
, 
<quantity id="4CBCE7A66D1CFF92FB9BC5F21975FF2A" box="[255,311,187,211]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="586" unit="m" value="40.0">
<elevation id="0069AD706D1CFF92FB9BC5F21975FF2A" box="[255,311,187,211]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="586" unit="m" value="40.0">40 m</elevation>
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| 
<date id="FFFA6C836D1CFF92FA28C5F219FDFF2A" box="[332,447,187,211]" pageId="1" pageNumber="586" value="2016-04-13">
<collectingDate id="EFBE956B6D1CFF92FA28C5F219FDFF2A" box="[332,447,187,211]" pageId="1" pageNumber="586" value="2016-04-13">13.iv.2016</collectingDate>
</date>
, David Král lgt. [printed]”
</materialsCitation>
. 
<materialsCitation id="3B2C401E6D1CFF92F98BC5F518BAFF0E" ID-GBIF-Occurrence="3390811301" collectingDate="2016-04-13" collectionCode="NMPC" country="Iran" elevation="40" latitude="37.533333" location="Golestan prov." longLatPrecision="1181" longitude="54.983334" pageId="1" pageNumber="586" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Golestan" typeStatus="paratype">
<typeStatus id="54FFF4E16D1CFF92F98BC5F51B19FF2D" box="[751,859,188,212]" pageId="1" pageNumber="586" type="paratype">Paratypes</typeStatus>
: 
<typeStatus id="54FFF4E16D1CFF92F802C5F21BFDFF2A" box="[870,959,187,211]" pageId="1" pageNumber="586" type="allotype">allotype</typeStatus>
, 
<specimenCount id="9D4281CA6D1CFF92F8AEC5F51B9CFF2A" box="[970,990,188,211]" pageId="1" pageNumber="586" type="female">♀</specimenCount>
and 
<specimenCount id="9D4281CA6D1CFF92FF71C5F21C16FF2A" box="[1045,1108,187,211]" pageId="1" pageNumber="586" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="9D4281CA6D1CFF92FF05C5F21CDDFF2A" box="[1121,1183,187,211]" pageId="1" pageNumber="586" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED55D2866D1CFF92FFCAC5F21CBDFF2A" box="[1198,1279,187,211]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="586">NMPC</collectionCode>
), same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B2C401E6D1CFF92FA67C5A91AECFF0E" ID-GBIF-Occurrence="3390811302" box="[259,686,223,247]" collectionCode="OHCB" collectorName="Hillert" country="Iran" pageId="1" pageNumber="586" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Golestan" typeStatus="paratype">
<specimenCount id="9D4281CA6D1CFF92FA67C5A91903FF0E" box="[259,321,224,247]" pageId="1" pageNumber="586" type="male">7 ♂♂</specimenCount>
, 
<specimenCount id="9D4281CA6D1CFF92FA28C59619CAFF0E" box="[332,392,223,247]" pageId="1" pageNumber="586" type="female">6 ♀♀</specimenCount>
(
<collectionCode id="ED55D2866D1CFF92FAF2C59619A7FF0E" box="[406,485,223,247]" pageId="1" pageNumber="586">OHCB</collectionCode>
), but O. 
<collectorName id="26B12F956D1CFF92F925C5961AC5FF0E" box="[577,647,223,247]" pageId="1" pageNumber="586">Hillert</collectorName>
lgt
</materialsCitation>
., 
<materialsCitation id="3B2C401E6D1CFF92F9DAC5961CC6FF0E" ID-GBIF-Occurrence="3390811304" box="[702,1156,223,247]" collectionCode="SJCP" collectorName="Stanislav Jakl" country="Iran" pageId="1" pageNumber="586" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Golestan" typeStatus="paratype">
<specimenCount id="9D4281CA6D1CFF92F9DAC5961ABEFF0E" box="[702,764,223,247]" pageId="1" pageNumber="586" type="male">8 ♂♂</specimenCount>
, 
<specimenCount id="9D4281CA6D1CFF92F863C5961B01FF0E" box="[775,835,223,247]" pageId="1" pageNumber="586" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED55D2866D1CFF92F835C5961BD2FF0E" box="[849,912,223,247]" pageId="1" pageNumber="586">SJCP</collectionCode>
), but 
<collectorName id="26B12F956D1CFF92F8AFC5961C1DFF0E" box="[971,1119,223,247]" pageId="1" pageNumber="586">Stanislav Jákl</collectorName>
lgt
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E19C86D1CFF91FBA3C44A18BBF830" lastPageId="2" lastPageNumber="587" pageId="1" pageNumber="586" type="description">
<paragraph id="8BFB4A436D1CFF92FBA3C44A19CCFE9A" blockId="1.[151,1437,151,355]" pageId="1" pageNumber="586">
<emphasis id="B93096516D1CFF92FBA3C44A1A4AFEE2" box="[199,520,259,283]" inLineHeading="true" pageId="1" pageNumber="586" reason="1">
<emphasis id="B93096516D1CFF92FBA3C44A1991FEE2" bold="true" box="[199,467,259,283]" pageId="1" pageNumber="586">
Description of 
<typeStatus id="54FFF4E16D1CFF92FA15C44A1991FEE2" box="[369,467,259,283]" pageId="1" pageNumber="586" type="holotype">holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D4281CA6D1CFF92FA80C44A19B8FEE2" box="[484,506,259,283]" pageId="1" pageNumber="586" type="male">♂</specimenCount>
).
</emphasis>
Maximally developed (hyperthelic) male with well developed ventral mandible processes (
<figureCitation id="137F56C66D1CFF92FB8DC46E1912FEC6" box="[233,336,295,319]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[319,1284,382,1584]" captionTargetId="figure-111@1.[303,1285,378,1598]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitus of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 1, holotype, ♂, dorsal view; 2, al- lotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577371" httpUri="https://zenodo.org/record/5577371/files/figure.png" pageId="1" pageNumber="586">Figs. 1, 3</figureCitation>
). Oblong, strongly convex; dorsal surface black, claws black-brown; setation black; moderately shiny, except of opaque head.
</paragraph>
<caption id="DF3B1ACB6D1CFF92FBF3C3111B0CF96D" ID-DOI="http://doi.org/10.5281/zenodo.5577371" ID-Zenodo-Dep="5577371" httpUri="https://zenodo.org/record/5577371/files/figure.png" pageId="1" pageNumber="586" startId="1.[151,264,1624,1648]" targetBox="[319,1284,382,1584]" targetPageId="1">
<paragraph id="8BFB4A436D1CFF92FBF3C3111B0CF96D" blockId="1.[151,1436,1624,1684]" pageId="1" pageNumber="586">
<emphasis id="B93096516D1CFF92FBF3C3111979F989" bold="true" box="[151,315,1624,1648]" pageId="1" pageNumber="586">FIGURES 1–3.</emphasis>
Habitus of 
<taxonomicName id="4C4431C06D1CFF92FAD7C3111BF6F989" authority="Kral &amp; Hillert" authorityName="Kral &amp; Hillert" authorityYear="2021" box="[435,948,1624,1648]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="golestanicus" status="sp. nov." subGenus="Scelolethrus">
<emphasis id="B93096516D1CFF92FAD7C3111A42F989" box="[435,512,1624,1648]" italics="true" pageId="1" pageNumber="586">Lethrus</emphasis>
(
<emphasis id="B93096516D1CFF92F96BC3111ACEF989" box="[527,652,1624,1648]" italics="true" pageId="1" pageNumber="586">Scelolethrus</emphasis>
) 
<emphasis id="B93096516D1CFF92F9F8C3111B59F989" box="[668,795,1624,1648]" italics="true" pageId="1" pageNumber="586">golestanicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="A2032B2A6D1CFF92F8DAC3101C79F989" box="[958,1083,1624,1648]" pageId="1" pageNumber="586" rank="species">new species</taxonomicNameLabel>
. 1, holotype, ♂, dorsal view; 2, allotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale.
</paragraph>
</caption>
<paragraph id="8BFB4A436D1CFF92FBA3C38D1C1BF821" blockId="1.[151,1437,1732,2008]" pageId="1" pageNumber="586">
<emphasis id="B93096516D1CFF92FBA3C38D1943F925" box="[199,257,1732,1756]" italics="true" pageId="1" pageNumber="586">Head</emphasis>
(
<figureCitation id="137F56C66D1CFF92FA76C38D1922F925" box="[274,352,1732,1756]" captionStart="FIGURES 4–7" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 4–7. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="1" pageNumber="586">Figs. 5</figureCitation>
, 
<figureCitation id="137F56C66D1CFF92FA0AC38D19E5F925" box="[366,423,1732,1756]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 8–13. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 8–10, head, dorsal view; 11–13, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="1" pageNumber="586">8, 11</figureCitation>
). Dorsal surface with large sized, coarse, superficial, regularly spaced punctures, posteriorly punctures becoming rather sparser and larger than in clypeus. Keels separating genae with eyes distinct, almost parallel. Frontal tubercles and medial impression only slightly developed; frontoclypeal suture only slightly indicated. Genae sized as usually, with lateral margins emarginate and rather divergent anteriad, anterior angles approximately rectangular, posterior angles rounded. Pleurostomal processes conspicuous, conical, rounded apically, clearly exceeding beyond ventrolateral mandible outline (
<figureCitation id="137F56C66D1CFF92FAA0C2311A52F869" box="[452,528,1912,1936]" captionStart="FIGURES 4–7" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 4–7. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="1" pageNumber="586">Figs. 5</figureCitation>
, 
<figureCitation id="137F56C66D1CFF92F979C2311A7BF869" box="[541,569,1912,1936]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 8–13. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 8–10, head, dorsal view; 11–13, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="1" pageNumber="586">11</figureCitation>
). Clypeus transverse, trapezoidal with anterior angles rounded. Labrum bilobed, asymmetrical, right lobe remarkably more developed; surface rugosely and coarsely, shallowly and sparsely punctate, each puncture bearing short recumbent seta; anterior margin with dense row of long setae.
</paragraph>
<caption id="DF3B1ACB6D1FFF91FBF3C1181BAEFB74" ID-DOI="http://doi.org/10.5281/zenodo.5577375" ID-Zenodo-Dep="5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="2" pageNumber="587" startId="2.[151,264,1105,1129]" targetBox="[305,1283,182,1067]" targetPageId="2">
<paragraph id="8BFB4A436D1FFF91FBF3C1181BAEFB74" blockId="2.[151,1436,1105,1165]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBF3C1181978FB90" bold="true" box="[151,314,1105,1129]" pageId="2" pageNumber="587">FIGURES 4–7.</emphasis>
Holotype of 
<taxonomicName id="4C4431C06D1FFF91FAD8C1181BF7FB90" authority="Kral &amp; Hillert" authorityName="Kral &amp; Hillert" authorityYear="2021" box="[444,949,1105,1130]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="golestanicus" status="sp. nov." subGenus="Scelolethrus">
<emphasis id="B93096516D1FFF91FAD8C1181A4BFB90" box="[444,521,1105,1129]" italics="true" pageId="2" pageNumber="587">Lethrus</emphasis>
(
<emphasis id="B93096516D1FFF91F973C1181AD6FB90" box="[535,660,1105,1129]" italics="true" pageId="2" pageNumber="587">Scelolethrus</emphasis>
) 
<emphasis id="B93096516D1FFF91F9C6C1181B63FB90" box="[674,801,1105,1129]" italics="true" pageId="2" pageNumber="587">golestanicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="A2032B2A6D1FFF91F8DAC11B1C7BFB90" box="[958,1081,1105,1129]" pageId="2" pageNumber="587" rank="species">new species</taxonomicNameLabel>
. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale.
</paragraph>
</caption>
<paragraph id="8BFB4A436D1FFF91FBA3C1F41A43FABB" blockId="2.[151,1437,1213,2030]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C1F4197AFB2C" box="[199,312,1213,1237]" italics="true" pageId="2" pageNumber="587">Mandibles</emphasis>
almost symmetrical, in dorsal aspect with weakly emarginate dorsolateral margin, rounded apically, just before half length from basis with distinct oblique keel, reaching mandibular outline; in lateral aspect without longitudinal keel. Ventral mandibular processes distinct, directed obliquely anteroventrally and pointed in lateral aspect, clearly visible in dorsal aspect (
<figureCitation id="137F56C66D1FFF91FA28C06019F1FAB8" box="[332,435,1321,1345]" captionStart="FIGURES 4–7" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 4–7. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="2" pageNumber="587">Figs. 4–5</figureCitation>
, 
<figureCitation id="137F56C66D1FFF91FAD9C06319B6FABB" box="[445,500,1322,1346]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 8–13. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 8–10, head, dorsal view; 11–13, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="2" pageNumber="587">8, 11</figureCitation>
).
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C0071899F9E0" blockId="2.[151,1437,1213,2030]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C0071973FA9C" box="[199,305,1358,1381]" italics="true" pageId="2" pageNumber="587">Pronotum</emphasis>
(
<figureCitation id="137F56C66D1FFF91FA24C00419E6FA9F" box="[320,420,1357,1382]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[319,1284,382,1584]" captionTargetId="figure-111@1.[303,1285,378,1598]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitus of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 1, holotype, ♂, dorsal view; 2, al- lotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577371" httpUri="https://zenodo.org/record/5577371/files/figure.png" pageId="2" pageNumber="587">Figs. 1, 3</figureCitation>
) transverse, distinctly broader than base of elytra, broadest just behind middle; margin entirely bordered, slightly crenulate in anterior parts. Anterior angles weakly but distinctly projecting anterolaterad, with angulate outline; lateral margin approximately weakly emarginate anteriorly, than straight to round posterior angle; basal margin straight. Punctation of dorsal surface simple, consisting of deep, sparsely and irregularly distributed punctures, separated by approximately two to four their diameters discally; surface near lateral margins considerably shagreened and alutaceous.
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C36C1BF6F9C4" blockId="2.[151,1437,1213,2030]" box="[199,948,1573,1597]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C36C1972F9C4" box="[199,304,1573,1597]" italics="true" pageId="2" pageNumber="587">Scutellum</emphasis>
considerably short, almost twice as wider as long, glabrous.
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C3001B48F97C" blockId="2.[151,1437,1213,2030]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C3001948F998" box="[199,266,1609,1633]" italics="true" pageId="2" pageNumber="587">Elytra</emphasis>
(
<figureCitation id="137F56C66D1FFF91FA73C300193AF99B" box="[279,376,1609,1634]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[319,1284,382,1584]" captionTargetId="figure-111@1.[303,1285,378,1598]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitus of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 1, holotype, ♂, dorsal view; 2, al- lotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577371" httpUri="https://zenodo.org/record/5577371/files/figure.png" pageId="2" pageNumber="587">Figs. 1, 3</figureCitation>
) almost semicircular, apices not prominent, each apex forming independent arc. Epipleuron strongly narrowed apicad, epipleural keel not reaching elytral apex.
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C3DB1BABF934" blockId="2.[151,1437,1213,2030]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C3DB18BBF950" box="[199,249,1682,1705]" italics="true" pageId="2" pageNumber="587">Legs</emphasis>
(
<figureCitation id="137F56C66D1FFF91FA6EC3D81936F953" box="[266,372,1681,1706]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[319,1284,382,1584]" captionTargetId="figure-111@1.[303,1285,378,1598]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitus of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 1, holotype, ♂, dorsal view; 2, al- lotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577371" httpUri="https://zenodo.org/record/5577371/files/figure.png" pageId="2" pageNumber="587">Figs. 1, 3</figureCitation>
). Profemora with transversal edge anteroventrally; protibia with row of eight gradually proximad diminishing exterior denticles, and with row of tubercles on ventromedial edge.
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C3901BBAF90B" blockId="2.[151,1437,1213,2030]" box="[199,1016,1753,1778]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C390196DF908" box="[199,303,1753,1777]" italics="true" pageId="2" pageNumber="587">Aedeagus</emphasis>
with inner edge of parameres angulate subapically (
<figureCitation id="137F56C66D1FFF91F838C3901B81F908" box="[860,963,1753,1777]" captionStart="FIGURES 4–7" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 4–7. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="2" pageNumber="587">Figs. 6–7</figureCitation>
, 
<figureCitation id="137F56C66D1FFF91F8AAC3931BAEF90B" box="[974,1004,1754,1778]" captionStart="FIGURES 14–16" captionStartId="4.[151,264,713,737]" captionTargetBox="[261,1321,188,676]" captionTargetId="figure-19@4.[258,1329,181,689]" captionTargetPageId="4" captionText="FIGURES 14–16. Lethrus (Scelolethrus) spp., parameres, dorsal view. 14, L. (S.) golestanicus Král &amp; Hillert, new species; 15, L. (S.) mithras Reitter, 1904 (Iran, Bojnurd); 16, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577381" httpUri="https://zenodo.org/record/5577381/files/figure.png" pageId="2" pageNumber="587">14</figureCitation>
).
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C3B71C00F8EC" blockId="2.[151,1437,1213,2030]" box="[199,1090,1789,1814]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C3B71937F8EF" bold="true" box="[199,373,1790,1814]" inLineHeading="true" pageId="2" pageNumber="587" reason="1">Measurements.</emphasis>
Body length 14.5–18.5 mm; 
<typeStatus id="54FFF4E16D1FFF91F9CAC3B41B49F8EC" box="[686,779,1789,1813]" pageId="2" pageNumber="587" type="holotype">holotype</typeStatus>
18.5 mm; 
<typeStatus id="54FFF4E16D1FFF91F81BC3B41B97F8EC" box="[895,981,1789,1813]" pageId="2" pageNumber="587" type="allotype">allotype</typeStatus>
18.5 mm.
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C26819EEF8A4" blockId="2.[151,1437,1213,2030]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C26819F1F8C0" box="[199,435,1825,1850]" inLineHeading="true" pageId="2" pageNumber="587" reason="1">
<emphasis id="B93096516D1FFF91FBA3C26819EEF8C3" bold="true" box="[199,428,1825,1850]" pageId="2" pageNumber="587">Variability in males</emphasis>
.
</emphasis>
Mandible processes in medium developed and underdeveloped (hypothelic) males shorter, weaker, angulate apically.
</paragraph>
<paragraph id="8BFB4A436D1FFF91FBA3C22318BBF830" blockId="2.[151,1437,1213,2030]" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C2231966F87B" bold="true" box="[199,292,1898,1922]" pageId="2" pageNumber="587">Females</emphasis>
(
<figureCitation id="137F56C66D1FFF91FA56C2201933F87B" box="[306,369,1897,1922]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1624,1648]" captionTargetBox="[319,1284,382,1584]" captionTargetId="figure-111@1.[303,1285,378,1598]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitus of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 1, holotype, ♂, dorsal view; 2, al- lotype, ♀, dorsal view; 3, holotype, ♂, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577371" httpUri="https://zenodo.org/record/5577371/files/figure.png" pageId="2" pageNumber="587">Fig. 2</figureCitation>
) differ from males as follows: exterior outline of mandibles almost straight, in apical quarter rounded in dorsal aspect; ventral mandibular process absent; protibia broader, row of tubercles on ventromedial edge less pronounced.
</paragraph>
</subSubSection>
<subSubSection id="C35E19C86D1FFF90FBA3C29C1B7AFE52" lastPageId="3" lastPageNumber="588" pageId="2" pageNumber="587" type="diagnosis">
<paragraph id="8BFB4A436D1FFF90FBA3C29C1B7AFE52" blockId="2.[151,1437,1213,2030]" lastBlockId="3.[151,1437,151,607]" lastPageId="3" lastPageNumber="588" pageId="2" pageNumber="587">
<emphasis id="B93096516D1FFF91FBA3C29C1983F814" box="[199,449,2005,2029]" inLineHeading="true" pageId="2" pageNumber="587" reason="1">
<emphasis id="B93096516D1FFF91FBA3C29C19FBF814" bold="true" box="[199,441,2005,2029]" pageId="2" pageNumber="587">Differential diagnosis</emphasis>
.
</emphasis>
<taxonomicName id="4C4431C06D1FFF91FAA2C29C1B99F814" authority="Kral &amp; Hillert" authorityName="Kral &amp; Hillert" authorityYear="2021" box="[454,987,2005,2030]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="golestanicus" status="sp. nov." subGenus="Scelolethrus">
<emphasis id="B93096516D1FFF91FAA2C29C1A55F814" box="[454,535,2005,2029]" italics="true" pageId="2" pageNumber="587">Lethrus</emphasis>
(
<emphasis id="B93096516D1FFF91F942C29C1AE8F814" box="[550,682,2005,2029]" italics="true" pageId="2" pageNumber="587">Scelolethrus</emphasis>
) 
<emphasis id="B93096516D1FFF91F9DDC29C1B7CF814" box="[697,830,2005,2029]" italics="true" pageId="2" pageNumber="587">golestanicus</emphasis>
Král &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="A2032B2A6D1FFF91F881C29F1C27F814" box="[997,1125,2005,2029]" pageId="2" pageNumber="587" rank="species">new species</taxonomicNameLabel>
is classified in the 
<emphasis id="B93096516D1FFF91FE49C29F1D7EF814" box="[1325,1340,2006,2029]" italics="true" pageId="2" pageNumber="587">L</emphasis>
. 
<taxonomicName id="4C4431C06D1FFF90FE2DC29F188DFF56" authorityName="Kraatz" authorityYear="1883" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" lastPageId="3" lastPageNumber="588" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="sulcatus" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1FFF91FE36C29F1D1DF814" box="[1362,1375,2006,2029]" italics="true" pageId="2" pageNumber="587">S</emphasis>
.) 
<emphasis id="B93096516D1FFF90FE10C29C188DFF56" italics="true" lastPageId="3" lastPageNumber="588" pageId="2" pageNumber="587">sulcatus</emphasis>
</taxonomicName>
species group due to the unique shape of the endophallus, which contains a sclerotized arrow that points obliquely backwards (
<emphasis id="B93096516D1EFF90FA7DC5F21976FF2A" box="[281,308,187,211]" italics="true" pageId="3" pageNumber="588">cf.</emphasis>
<bibRefCitation id="EFD537B26D1EFF90FA5FC5F21A65FF2A" author="Nikolajev, G. V." box="[315,551,187,211]" pageId="3" pageNumber="588" refId="ref3030" refString="Nikolajev, G. V. (2003 b) Zhuki-kravchiki (Scarabaeidae, Geotrupinae, Lethrini): biologiya, sistematika, rasprostranenie, opredelitel'. Lethrini (Coleoptera: Scarabaeidae, Geotrupinae): biology, systematics, distribution, keys. Kazak universiteti, Almaty, 254 pp. [in Russian, English summary]" type="book" year="2003">Nikolajev 2003b: 237</bibRefCitation>
, figs. 176: 1–3). The species 
<emphasis id="B93096516D1EFF90F807C5F51B30FF2A" box="[867,882,188,211]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90F8E4C5F21CD6FF2A" authority="Reitter, 1904" authorityName="Reitter" authorityYear="1904" box="[896,1172,187,211]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="mithras" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90F8EDC5F21BD4FF2B" box="[905,918,187,210]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90F8C9C5F21BBCFF2A" box="[941,1022,187,211]" italics="true" pageId="3" pageNumber="588">mithras</emphasis>
Reitter, 1904
</taxonomicName>
, 
<emphasis id="B93096516D1EFF90FFFBC5F51CECFF2A" box="[1183,1198,188,211]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FFD8C5F2195EFF0E" authority="Kral &amp; Olexa, 1996" authorityName="Kral &amp; Olexa" authorityYear="1996" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="tekkensis" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FFA1C5F21C90FF2B" box="[1221,1234,187,210]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90FF8DC5F21D0BFF2A" box="[1257,1353,187,211]" italics="true" pageId="3" pageNumber="588">tekkensis</emphasis>
<bibRefCitation id="EFD537B26D1EFF90FE35C5F2195EFF0E" author="Kral, D. &amp; Olexa, A." pageId="3" pageNumber="588" pagination="49 - 65" refId="ref2922" refString="Kral, D. &amp; Olexa, A. (1996) New and otherwise noteworthy Lethrus species from Central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana, 4, 49 - 65." type="journal article" year="1996">Král &amp; Olexa, 1996</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B93096516D1EFF90FA42C5A91977FF0E" box="[294,309,224,247]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FA26C5961B45FF0E" authority="Gusakov &amp; Klimenko, 2011" authorityName="Gusakov &amp; Klimenko" authorityYear="2011" box="[322,775,223,247]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="percutens" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FA2FC596191AFF0F" box="[331,344,223,246]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90FA0AC5A91997FF0E" box="[366,469,224,247]" italics="true" pageId="3" pageNumber="588">percutens</emphasis>
<bibRefCitation id="EFD537B26D1EFF90FABFC5961B45FF0E" author="Gusakov, A. A. &amp; Klimenko, A. A." box="[475,775,223,247]" pageId="3" pageNumber="588" pagination="3 - 8" refId="ref2795" refString="Gusakov, A. A. &amp; Klimenko, A. A. (2011) Novyy vid roda Lethrus Scopoli, 1777 (podroda Scelolethrus Semenov, 1892) iz Irana (Coleoptera: Scarabaeidae: Geotrupinae). A new species of the genus Lethrus Scopoli, 1777 (subgenus Scelolethrus Semenov, 1892) from Iran (Coleoptera: Scarabaeidae: Geotrupinae). Eversmannia, 25 - 26, 3 - 8. [in Russian, English abstract]" type="journal article" year="2011">Gusakov &amp; Klimenko, 2011</bibRefCitation>
</taxonomicName>
and also our new species posses conspicuous conical pleurostomal processes that clearly extend beyond the ventrolateral mandible outline (
<figureCitation id="137F56C66D1EFF90F886C44A1C6FFEE2" box="[994,1069,259,283]" captionStart="FIGURES 4–7" captionStartId="2.[151,264,1105,1129]" captionTargetBox="[305,1283,182,1067]" captionTargetId="figure-19@2.[303,1285,181,1081]" captionTargetPageId="2" captionText="FIGURES 4–7. Holotype of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. 4, mandibles, dorsal view; 5, head, left lateral view; 6, parameres, dorsal view; 7, parameres, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577375" httpUri="https://zenodo.org/record/5577375/files/figure.png" pageId="3" pageNumber="588">Figs. 5</figureCitation>
, 
<figureCitation id="137F56C66D1EFF90FF5DC44A1C3FFEE2" box="[1081,1149,259,283]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1939,1963]" captionTargetBox="[216,1374,634,1904]" captionTargetId="figure-247@3.[213,1375,631,1914]" captionTargetPageId="3" captionText="FIGURES 8–13. 8, 11, Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species; 9, 12, L. (S.) mithras Reitter, 1904, Iran (Bojnurd); 10, 13, L. (S.) tekkensis Král &amp; Olexa, 1996 (Turkmenistan, Kara Kala [= Garrygala or Maktymguly]). 8–10, head, dorsal view; 11–13, head, left lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5577377" httpUri="https://zenodo.org/record/5577377/files/figure.png" pageId="3" pageNumber="588">11–13</figureCitation>
, 
<emphasis id="B93096516D1EFF90FFEDC44A1CDEFEE2" box="[1161,1180,259,283]" italics="true" pageId="3" pageNumber="588">cf</emphasis>
. 
<bibRefCitation id="EFD537B26D1EFF90FFCEC44A18ABFEC6" author="Gusakov, A. A. &amp; Klimenko, A. A." pageId="3" pageNumber="588" pagination="3 - 8" refId="ref2795" refString="Gusakov, A. A. &amp; Klimenko, A. A. (2011) Novyy vid roda Lethrus Scopoli, 1777 (podroda Scelolethrus Semenov, 1892) iz Irana (Coleoptera: Scarabaeidae: Geotrupinae). A new species of the genus Lethrus Scopoli, 1777 (subgenus Scelolethrus Semenov, 1892) from Iran (Coleoptera: Scarabaeidae: Geotrupinae). Eversmannia, 25 - 26, 3 - 8. [in Russian, English abstract]" type="journal article" year="2011">Gusakov &amp; Klimenko, 2011: 7</bibRefCitation>
, fig. 5). Other species in this group 
<emphasis id="B93096516D1EFF90F909C4611A3EFEC6" box="[621,636,296,319]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90F9EEC46E1BBAFEC6" authority="Kral &amp; Olexa, 1996" authorityName="Kral &amp; Olexa" authorityYear="1996" box="[650,1016,295,319]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="crypticus" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90F9F7C46E1AE2FEC7" box="[659,672,295,318]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90F9D3C4611B5BFEC6" box="[695,793,296,319]" italics="true" pageId="3" pageNumber="588">crypticus</emphasis>
<bibRefCitation id="EFD537B26D1EFF90F844C46E1BBAFEC6" author="Kral, D. &amp; Olexa, A." box="[800,1016,295,319]" pageId="3" pageNumber="588" pagination="49 - 65" refId="ref2922" refString="Kral, D. &amp; Olexa, A. (1996) New and otherwise noteworthy Lethrus species from Central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana, 4, 49 - 65." type="journal article" year="1996">Král &amp; Olexa, 1996</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B93096516D1EFF90FF60C4611C51FEC6" box="[1028,1043,296,319]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FF45C46E1D13FEC6" authority="Gusakov, 2017" authorityName="Gusakov" authorityYear="2017" box="[1057,1361,295,319]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="nazarovi" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FF4DC46E1C74FEC7" box="[1065,1078,295,318]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90FF2AC4611CE8FEC6" box="[1102,1194,296,319]" italics="true" pageId="3" pageNumber="588">nazarovi</emphasis>
Gusakov, 2017
</taxonomicName>
and 
<emphasis id="B93096516D1EFF90FEE3C4611DD4FEC6" box="[1415,1430,296,319]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FBF3C402195FFE9A" authorityName="Kraatz" authorityYear="1883" box="[151,285,331,355]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="sulcatus" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FBC4C40218EFFE9B" box="[160,173,331,354]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90FBA2C402195FFE9A" box="[198,285,331,355]" italics="true" pageId="3" pageNumber="588">sulcatus</emphasis>
</taxonomicName>
have rounded pleurostomal processes not exceeding the ventrolateral mandible outline (
<emphasis id="B93096516D1EFF90FF80C4021CB5FE9A" box="[1252,1271,331,355]" italics="true" pageId="3" pageNumber="588">cf</emphasis>
. 
<bibRefCitation id="EFD537B26D1EFF90FE63C40218BAFE7E" author="Kral, D. &amp; Olexa, A." pageId="3" pageNumber="588" pagination="49 - 65" refId="ref2922" refString="Kral, D. &amp; Olexa, A. (1996) New and otherwise noteworthy Lethrus species from Central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana, 4, 49 - 65." type="journal article" year="1996">Král &amp; Olexa 1996: 61</bibRefCitation>
, fig. 25 or 
<bibRefCitation id="EFD537B26D1EFF90FA08C4261A1AFE7E" author="Nikolajev, G. V." box="[364,600,367,391]" pageId="3" pageNumber="588" refId="ref3030" refString="Nikolajev, G. V. (2003 b) Zhuki-kravchiki (Scarabaeidae, Geotrupinae, Lethrini): biologiya, sistematika, rasprostranenie, opredelitel'. Lethrini (Coleoptera: Scarabaeidae, Geotrupinae): biology, systematics, distribution, keys. Kazak universiteti, Almaty, 254 pp. [in Russian, English summary]" type="book" year="2003">Nikolajev 2003b: 204</bibRefCitation>
, fig. 152: 3). The key below can be used to distinguish the new species, 
<emphasis id="B93096516D1EFF90FE3EC4391D2BFE7E" box="[1370,1385,368,391]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FE12C42618AAFE52" authorityName="Reitter" authorityYear="1904" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="mithras" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FE1BC4261DCEFE7F" box="[1407,1420,367,390]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90FBF3C4DA18AAFE52" box="[151,232,403,427]" italics="true" pageId="3" pageNumber="588">mithras</emphasis>
</taxonomicName>
, 
<emphasis id="B93096516D1EFF90FB92C4DD1947FE52" box="[246,261,404,427]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FA76C4DA19E4FE52" authorityName="Gusakov &amp; Klimenko" authorityYear="2011" box="[274,422,403,427]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="percutens" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FA7FC4DA196AFE53" box="[283,296,403,426]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90FA5BC4DD19E4FE52" box="[319,422,404,427]" italics="true" pageId="3" pageNumber="588">percutens</emphasis>
</taxonomicName>
and 
<emphasis id="B93096516D1EFF90FABDC4DD19AAFE52" box="[473,488,404,427]" italics="true" pageId="3" pageNumber="588">L</emphasis>
. 
<taxonomicName id="4C4431C06D1EFF90FA91C4DA1AC0FE52" authorityName="Kral &amp; Olexa" authorityYear="1996" box="[501,642,403,427]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="tekkensis" subGenus="Scelolethrus">
(
<emphasis id="B93096516D1EFF90FA9AC4DA1A49FE53" box="[510,523,403,426]" italics="true" pageId="3" pageNumber="588">S</emphasis>
.) 
<emphasis id="B93096516D1EFF90F946C4DA1AC0FE52" box="[546,642,403,427]" italics="true" pageId="3" pageNumber="588">tekkensis</emphasis>
</taxonomicName>
from each other.
</paragraph>
</subSubSection>
<subSubSection id="C35E19C86D1EFF90FBA3C4FE1B4AFDC2" pageId="3" pageNumber="588" type="etymology">
<paragraph id="8BFB4A436D1EFF90FBA3C4FE196DFE0A" blockId="3.[151,1437,151,607]" pageId="3" pageNumber="588">
<emphasis id="B93096516D1EFF90FBA3C4FE190BFE36" box="[199,329,439,463]" inLineHeading="true" pageId="3" pageNumber="588" reason="1">
<emphasis id="B93096516D1EFF90FBA3C4FE1900FE36" bold="true" box="[199,322,439,463]" pageId="3" pageNumber="588">Etymology</emphasis>
.
</emphasis>
Toponymic; the specific name 
<emphasis id="B93096516D1EFF90F9F1C4FE1B58FE36" box="[661,794,439,463]" italics="true" pageId="3" pageNumber="588">golestanicus</emphasis>
reflects the 
<collectingRegion id="498084A16D1EFF90F8FFC4FE1C1FFE36" box="[923,1117,439,463]" country="Iran" name="Golestan" pageId="3" pageNumber="588">Golestan province</collectingRegion>
in 
<collectingCountry id="F3530AD36D1EFF90FF1AC4F11CEBFE36" box="[1150,1193,440,463]" name="Iran" pageId="3" pageNumber="588">Iran</collectingCountry>
where the new species was collected.
</paragraph>
<paragraph id="8BFB4A436D1EFF90FBA3C4B61B4AFDC2" blockId="3.[151,1437,151,607]" pageId="3" pageNumber="588">
<emphasis id="B93096516D1EFF90FBA3C4B619A8FDEE" box="[199,490,511,535]" inLineHeading="true" pageId="3" pageNumber="588" reason="1">
<emphasis id="B93096516D1EFF90FBA3C4B619A1FDEE" bold="true" box="[199,483,511,535]" pageId="3" pageNumber="588">Collecting circumstances</emphasis>
.
</emphasis>
The 
<typeStatus id="54FFF4E16D1EFF90F945C7491A0DFDE1" box="[545,591,512,536]" pageId="3" pageNumber="588">type</typeStatus>
series was collected on slow loess slopes, mainly on pastures and uncultivated fields with very poor herbaceous vegetation (
<figureCitation id="137F56C66D1EFF90F91DC76A1ABEFDC2" box="[633,764,547,571]" captionStart="FIGURES 17–18" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[377,1210,836,1938]" captionTargetId="figure-75@4.[377,1210,833,1939]" captionTargetPageId="4" captionText="FIGURES 17–18. Type locality of Lethrus (Scelolethrus) golestanicus Král &amp; Hillert, new species. Iran, Golestan province, N of Qareh Makher, 13.iv.2016 (photo by David Král)." figureDoi="http://doi.org/10.5281/zenodo.5577383" httpUri="https://zenodo.org/record/5577383/files/figure.png" pageId="3" pageNumber="588">Figs. 17–18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E19C86D1EFF90FBA3C70E1AD4FDA6" box="[199,662,583,607]" pageId="3" pageNumber="588" type="distribution">
<paragraph id="8BFB4A436D1EFF90FBA3C70E1AD4FDA6" blockId="3.[151,1437,151,607]" box="[199,662,583,607]" pageId="3" pageNumber="588">
<emphasis id="B93096516D1EFF90FBA3C70E191BFDA6" box="[199,345,583,607]" inLineHeading="true" pageId="3" pageNumber="588" reason="1">
<emphasis id="B93096516D1EFF90FBA3C70E1910FDA6" bold="true" box="[199,338,583,607]" pageId="3" pageNumber="588">Distribution</emphasis>
.
</emphasis>
Northwestern 
<collectingRegion id="498084A16D1EFF90FA9CC70E1A18FDA6" box="[504,602,583,607]" country="Iran" name="Golestan" pageId="3" pageNumber="588">Golestan</collectingRegion>
, 
<collectingCountry id="F3530AD36D1EFF90F900C7011AD0FDA6" box="[612,658,584,607]" name="Iran" pageId="3" pageNumber="588">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>