425 lines
62 KiB
XML
425 lines
62 KiB
XML
<document id="D6B3618EC9D8A08F2C78A82E4DEE8244" ID-CLB-Dataset="9412" ID-DOI="10.11646/zootaxa.4885.3.5" ID-GBIF-Dataset="fd568ba9-4fd4-4f51-9a5f-42061140c137" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296773" ID-ZooBank="AFE72E8B-A590-4B89-96A1-880C2D2AE14B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606681170164" checkinUser="plazi" docAuthor="Salazar-Silva, Patricia, López-Sánchez, Daniel A. & Salazar-Vallejo, Sergio I." docDate="2020" docId="03EC4719FFC0E54866D0F9A0FF306596" docLanguage="en" docName="zootaxa.4885.3.5.pdf" docOrigin="Zootaxa 4885 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chaetacanthus ornatus Salazar-Silva & López-Sánchez & Salazar-Vallejo 2020, n. sp." docType="treatment" docVersion="7" lastPageNumber="411" masterDocId="FFD53F61FFCCE5586647FF96FFDD6766" masterDocTitle="Revision of Chaetacanthus Seidler, 1922 (Annelida, Phyllodocida, Polynoidae)" masterLastPageNumber="422" masterPageNumber="395" pageNumber="407" updateTime="1698846816447" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2C3EE90C7D121C52FCD2BC64CACEA233">Revision of Chaetacanthus Seidler, 1922 (Annelida, Phyllodocida, Polynoidae)</mods:title>
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<mods:namePart id="A9A60FE6FFC76C9BB6273E4DEEE5F65D">Salazar-Silva, Patricia</mods:namePart>
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<mods:namePart id="8A0FF08CE998E7969B06DC8A861E1FB2">López-Sánchez, Daniel A.</mods:namePart>
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<mods:namePart id="0D483E23497AB8C8BA6CA36435849DBD">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="32D9B57F6B8C6E2953F318836048C5D4">Zootaxa</mods:title>
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<mods:date id="0E2F466223C32BBC33A7D1B3DC827C3F">2020</mods:date>
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<paragraph id="8BFAF60FFFC0E55466D0F9A0FE216137" blockId="12.[151,967,1590,1689]" box="[151,508,1590,1617]" pageId="12" pageNumber="407">
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<taxonomicName id="4C458D8CFFC0E55466D0F9A0FE6F6137" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[151,434,1590,1617]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
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<emphasis id="B9312A1DFFC0E55466D0F9A0FE6F6137" bold="true" box="[151,434,1590,1617]" italics="true" pageId="12" pageNumber="407">Chaetacanthus ornatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2029766FFC0E55467FEF9A1FE216137" box="[441,508,1591,1617]" pageId="12" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8BFAF60FFFC0E55466D0F9CDFC1A6113" blockId="12.[151,967,1590,1689]" box="[151,967,1627,1653]" pageId="12" pageNumber="407">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C35FA584FFC0E55466D0F9E9FE4A61FF" box="[151,407,1663,1689]" pageId="12" pageNumber="407" type="description">
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<paragraph id="8BFAF60FFFC0E55466D0F9E9FE4A61FF" blockId="12.[151,967,1590,1689]" box="[151,407,1663,1689]" pageId="12" pageNumber="407">
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<figureCitation id="137EEA8AFFC0E55466D0F9E9FEE661FF" box="[151,315,1663,1689]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="13.[151,250,1608,1632]" captionStartId-1="14.[151,250,626,650]" captionStartId-2="14.[151,250,1880,1904]" captionTargetBox-0="[158,1430,181,1584]" captionTargetBox-1="[170,1417,181,602]" captionTargetBox-2="[170,1418,710,1856]" captionTargetId-0="figure-21@13.[157,1430,181,1584]" captionTargetId-1="figure-30@14.[170,1418,181,602]" captionTargetId-2="figure-69@14.[170,1418,710,1856]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="14" captionText-0="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." captionText-1="FIGURE 11. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Pharynx everted, left lateral view. B. Same, frontal view. Scale bars: A: 0.45 mm; B: 0.50 mm." captionText-2="FIGURE 12. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formaline, elytra in dorsal view. A. Left elytron from first pair. B. Left elytron of middle segment. C. Left elytron from middle segment. D. Left elytron, from posterior segment. Scale bars: A, B: 0.6 mm; C: 0.8 mm; D: 0.7 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4296793" figureDoi-1="http://doi.org/10.5281/zenodo.4296795" figureDoi-2="http://doi.org/10.5281/zenodo.4296797" httpUri-0="https://zenodo.org/record/4296793/files/figure.png" httpUri-1="https://zenodo.org/record/4296795/files/figure.png" httpUri-2="https://zenodo.org/record/4296797/files/figure.png" pageId="12" pageNumber="407">Figures 10–12</figureCitation>
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,
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<figureCitation id="137EEA8AFFC0E5546701F9E9FE8261FF" box="[326,351,1663,1689]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="12" pageNumber="407">13</figureCitation>
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A–C
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35FA584FFC0E55466D0F955FB03619C" pageId="12" pageNumber="407" type="reference_group">
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<paragraph id="8BFAF60FFFC0E55466D0F955FBF961BD" blockId="12.[151,1246,1731,1786]" box="[151,1060,1731,1755]" pageId="12" pageNumber="407">
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<treatmentCitationGroup id="AB55D121FFC0E55466D0F955FBF961BD" box="[151,1060,1731,1755]" pageId="12" pageNumber="407">
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<treatmentCitation id="0AE4D01EFFC0E55466D0F955FDBB61BD" author="Monro, C. C. A." box="[151,614,1731,1755]" page="558" pageId="12" pageNumber="407" year="1928">
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<taxonomicName id="4C458D8CFFC0E55466D0F955FDBB61BD" ID-CoL="69MLG" authority=": Monro 1928: 558" authorityName="Monro" authorityPageNumber="558" authorityYear="1928" box="[151,614,1731,1755]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="magnificus">
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<emphasis id="B9312A1DFFC0E55466D0F955FE7861BD" box="[151,421,1731,1755]" italics="true" pageId="12" pageNumber="407">Chaetacanthus magnificus</emphasis>
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:
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<bibRefCitation id="EFD48BFEFFC0E55467F5F952FDBB61BD" author="Monro, C. C. A." box="[434,614,1731,1755]" pageId="12" pageNumber="407" pagination="553 - 576" refId="ref14860" refString="Monro, C. C. A. (1928) Polychaeta of the families Polynoidae and Acoetidae from the vicinity of the Panama Canal, collected by Dr. C. Crossland and Dr. Th. Mortensen. Zoological Journal of the Linnean Society, London, 36, 553 - 576. https: // doi. org / 10.1111 / j. 1096 - 3642.1928. tb 02209. x" type="journal article" year="1928">Monro 1928: 558</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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||
<treatmentCitation id="0AE4D01EFFC0E5546436F952FC8761BD" author="Hartman, O." box="[625,858,1731,1755]" page="28" pageId="12" pageNumber="407" year="1939">
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<bibRefCitation id="EFD48BFEFFC0E5546436F952FC8761BD" author="Hartman, O." box="[625,858,1731,1755]" pageId="12" pageNumber="407" pagination="1 - 156" refId="ref13886" refString="Hartman, O. (1939 a) Polychaetous annelids, 1. Aphroditidae to Pisionidae. Allan Hancock Pacific Expeditions, 7, 1 - 156, pls. 1 - 28." type="journal article" year="1939">Hartman 1939a: 28–29</bibRefCitation>
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</treatmentCitation>
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||
(
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<emphasis id="B9312A1DFFC0E554652EF952FC5261BD" box="[873,911,1732,1755]" italics="true" pageId="12" pageNumber="407">non</emphasis>
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||
<bibRefCitation id="EFD48BFEFFC0E55465D2F955FBC561BD" author="Grube, A. E." box="[917,1048,1731,1755]" pageId="12" pageNumber="407" pagination="46 - 72" refId="ref13845" refString="Grube, A. E. (1876) Bemerkungen ʾ ber die Familie der Aphroditeen (Gruppe Polyonina, Acoetea, Polylepidea). Jahres-Bericht der Schlesischen Gesellschaft f ̡ r vaterlandische Cultur, 1875, 46 - 72." type="journal article" year="1876">Grube, 1876</bibRefCitation>
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).
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BFAF60FFFC0E55466D0F974FB03619C" blockId="12.[151,1246,1731,1786]" box="[151,1246,1762,1786]" pageId="12" pageNumber="407">
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<treatmentCitationGroup id="AB55D121FFC0E55466D0F974FB03619C" box="[151,1246,1762,1786]" pageId="12" pageNumber="407">
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<treatmentCitation id="0AE4D01EFFC0E55466D0F974FCFD619C" author="Hernandez-Moreno, P." box="[151,800,1762,1786]" page="32" pageId="12" pageNumber="407" year="2011">
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||
<taxonomicName id="4C458D8CFFC0E55466D0F974FCFD619C" ID-CoL="69MLG" authority=": Hernandez-Moreno 2011: 28 - 32" authorityName=": Hernandez-Moreno" authorityPageNumber="28 - 32" authorityYear="2011" box="[151,800,1762,1786]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="magnificus">
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||
<emphasis id="B9312A1DFFC0E55466D0F974FEF2619C" box="[151,303,1762,1786]" italics="true" pageId="12" pageNumber="407">Chaetacanthus</emphasis>
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||
cf.
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<emphasis id="B9312A1DFFC0E5546711F974FE18619C" box="[342,453,1762,1786]" italics="true" pageId="12" pageNumber="407">magnificus</emphasis>
|
||
:
|
||
<bibRefCitation id="EFD48BFEFFC0E5546795F974FCFD619C" author="Hernandez-Moreno, P." box="[466,800,1762,1786]" pageId="12" pageNumber="407" refId="ref14187" refString="Hernandez-Moreno, P. (2011) Polinoidos (Polychaeta: Polynoidae) del Pacifico sur de Mexico. Tesis Profesional, Universidad del Mar, Puerto Angel, Oaxaca, 147 pp." type="book" year="2011">Hernández-Moreno 2011: 28–32</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
|
||
,
|
||
<figureCitation id="137EEA8AFFC0E554656DF974FCB8619C" box="[810,869,1762,1786]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9</figureCitation>
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||
A–O, Pls. 5D, 6F (non
|
||
<bibRefCitation id="EFD48BFEFFC0E5546217F974FB0E619C" author="Grube, A. E." box="[1104,1235,1762,1786]" pageId="12" pageNumber="407" pagination="46 - 72" refId="ref13845" refString="Grube, A. E. (1876) Bemerkungen ʾ ber die Familie der Aphroditeen (Gruppe Polyonina, Acoetea, Polylepidea). Jahres-Bericht der Schlesischen Gesellschaft f ̡ r vaterlandische Cultur, 1875, 46 - 72." type="journal article" year="1876">Grube, 1876</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E55466D0F8BEFBC76000" pageId="12" pageNumber="407" type="materials_examined">
|
||
<paragraph id="8BFAF60FFFC0E55466D0F8BEFBC76000" blockId="12.[151,1436,1832,2039]" pageId="12" pageNumber="407">
|
||
<emphasis id="B9312A1DFFC0E55466D0F8BEFDDA6024" bold="true" box="[151,519,1832,1858]" pageId="12" pageNumber="407">Type material. Eastern Pacific.</emphasis>
|
||
<materialsCitation id="3B2DFC52FFC0E5546449F8BEFBC76000" ID-GBIF-Occurrence="2989844315" collectingDate="2004-07-04" collectionCode="ECOSUR" collectorName="S. Garcia" country="Mexico" location="Puerto Escondido" pageId="12" pageNumber="407" specimenCode="ECOSUR 233" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
|
||
<emphasis id="B9312A1DFFC0E5546449F8BEFDA66024" bold="true" box="[526,635,1832,1858]" pageId="12" pageNumber="407">
|
||
<typeStatus id="54FE48ADFFC0E5546449F8BEFDA66024" box="[526,635,1832,1858]" pageId="12" pageNumber="407" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCode id="DBE35E74FFC0E55464C6F8BEFCF96024" box="[641,804,1832,1858]" collectionCode="ECOSUR" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="12" pageNumber="407" type="Museum">ECOSUR 233</specimenCode>
|
||
,
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||
<location id="8E9AA0D4FFC0E5546569F8BFFC2A6024" LSID="urn:lsid:plazi:treatment:03EC4719FFC0E54866D0F9A0FF306596:8E9AA0D4FFC0E5546569F8BFFC2A6024" box="[814,1015,1832,1859]" country="Mexico" name="Puerto Escondido" pageId="12" pageNumber="407" stateProvince="Oaxaca">Puerto Escondido</location>
|
||
,
|
||
<collectingRegion id="498138EDFFC0E5546245F8BEFB846024" box="[1026,1113,1832,1858]" country="Mexico" name="Oaxaca" pageId="12" pageNumber="407">Oaxaca</collectingRegion>
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,
|
||
<collectingCountry id="F352B69FFFC0E5546223F8BEFB616024" box="[1124,1212,1832,1858]" name="Mexico" pageId="12" pageNumber="407">México</collectingCountry>
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||
, 10, associated with algae on thorny bivalve (
|
||
<taxonomicName id="4C458D8CFFC0E55467E8F8DAFD086000" authority="Sowerby, 1847" authorityName="Sowerby" authorityYear="1847" box="[431,725,1868,1894]" class="Bivalvia" family="Spondylidae" genus="Spondylus" kingdom="Animalia" order="Pectinoida" pageId="12" pageNumber="407" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9312A1DFFC0E55467E8F8DAFDFF6000" box="[431,546,1868,1894]" italics="true" pageId="12" pageNumber="407">Spondylus</emphasis>
|
||
<bibRefCitation id="EFD48BFEFFC0E554646EF8DAFD086000" author="Sowerby, G. B." box="[553,725,1868,1894]" pageId="12" pageNumber="407" pagination="86 - 88" refId="ref15911" refString="Sowerby, G. B. (1847) Descriptions of several new species of Spondylus. Proceedings of the Zoological Society of London, 15, 86 - 88." type="journal article" year="1847">Sowerby, 1847</bibRefCitation>
|
||
</taxonomicName>
|
||
),
|
||
<date id="FFFBD0CFFFC0E55464AFF8DAFCBA6000" box="[744,871,1868,1894]" pageId="12" pageNumber="407" value="2004-07-04">
|
||
<collectingDate id="EFBF2927FFC0E55464AFF8DAFCBA6000" box="[744,871,1868,1894]" pageId="12" pageNumber="407" value="2004-07-04">4 Jul. 2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B093D9FFC0E5546535F8DAFC016000" box="[882,988,1868,1894]" pageId="12" pageNumber="407">S. García</collectorName>
|
||
, coll.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E5546680F8E6FF106091" pageId="12" pageNumber="407" type="diagnosis">
|
||
<paragraph id="8BFAF60FFFC0E5546680F8E6FF106091" blockId="12.[151,1436,1832,2039]" pageId="12" pageNumber="407">
|
||
<emphasis id="B9312A1DFFC0E5546680F8E6FE9F60EC" bold="true" box="[199,322,1904,1930]" pageId="12" pageNumber="407">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C458D8CFFC0E5546700F8E6FE3260EC" authorityName="Seidler" authorityYear="1922" box="[327,495,1904,1930]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="genus">
|
||
<emphasis id="B9312A1DFFC0E5546700F8E6FE3260EC" box="[327,495,1904,1930]" italics="true" pageId="12" pageNumber="407">Chaetacanthus</emphasis>
|
||
</taxonomicName>
|
||
with branchial filament digitiform with septate appearance; anterior eyes area bulky; elytral macrotubercles globular, amber-colour, sclerotized in large honeycomb like groups, outer area with fusiform macrotubercles distally blunt; numerous microtubercles dark, conical with thick spines on overlapping medially area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3AA687FFC1E55566D0F9DEFDA361C1" ID-DOI="http://doi.org/10.5281/zenodo.4296793" ID-Zenodo-Dep="4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="13" pageNumber="408" startId="13.[151,250,1608,1632]" targetBox="[158,1430,181,1584]" targetPageId="13">
|
||
<paragraph id="8BFAF60FFFC1E55566D0F9DEFDA361C1" blockId="13.[151,1437,1608,1704]" pageId="13" pageNumber="408">
|
||
<emphasis id="B9312A1DFFC1E55566D0F9DEFEC16106" bold="true" box="[151,284,1608,1632]" pageId="13" pageNumber="408">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="4C458D8CFFC1E555676CF9DEFDC46106" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[299,537,1608,1632]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="408" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC1E555676CF9DEFDC46106" box="[299,537,1608,1632]" italics="true" pageId="13" pageNumber="408">Chaetacanthus ornatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9312A1DFFC1E5556464F9DEFD826106" bold="true" box="[547,607,1608,1632]" pageId="13" pageNumber="408">
|
||
<taxonomicNameLabel id="A2029766FFC1E5556464F9DEFD826106" box="[547,607,1608,1632]" pageId="13" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35FA584FFC1E5576680F940FB0A631B" lastPageId="15" lastPageNumber="410" pageId="13" pageNumber="408" type="description">
|
||
<paragraph id="8BFAF60FFFC1E5556680F940FAEA608B" blockId="13.[151,1437,1750,2029]" pageId="13" pageNumber="408">
|
||
<emphasis id="B9312A1DFFC1E5556680F940FE846196" bold="true" box="[199,345,1750,1776]" pageId="13" pageNumber="408">Description.</emphasis>
|
||
<typeStatus id="54FE48ADFFC1E5556725F940FE176196" box="[354,458,1750,1776]" pageId="13" pageNumber="408" type="holotype">Holotype</typeStatus>
|
||
(ECOSUR 233) 2.9 cm long, 1.1 cm wide, 26 segments (
|
||
<figureCitation id="137EEA8AFFC1E555622DF940FB086196" box="[1130,1237,1750,1776]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="13" pageNumber="408">Fig. 10A</figureCitation>
|
||
). Prostomium bilobed, facial tubercle small, round. Two pairs of dark eyes; anterior pair on widest prostomial bulky area; posterior pair near posterior prostomial margin; posterior eyes much smaller than anterior ones (
|
||
<figureCitation id="137EEA8AFFC1E555622BF888FB0E605E" box="[1132,1235,1822,1848]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="13" pageNumber="408">Fig. 10B</figureCitation>
|
||
). Median antenna with thick, long ceratophore inserted frontally between prostomial lobes, style long, 1.5 times longer than prostomial length, thick, subdistally clavate, tip filiform, surface smooth, with pigmentation rings medially and subdistally. Lateral antennae with ceratophores inserted terminally on frontal prostomial border, brownish, styles similar in shape and pigmentation to median antenna. Palps robust, dark, taper into thin tips, surface with long rows of papillae. Pharynx everted with soft marginal papillae, 12 dorsally and 13 ventrally (
|
||
<figureCitation id="137EEA8AFFC1E5556278F844FB16608B" box="[1087,1227,2002,2029]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="13" pageNumber="408">Figs 10A, B</figureCitation>
|
||
;
|
||
<figureCitation id="137EEA8AFFC1E5556290F844FAF4608B" box="[1239,1321,2002,2029]" captionStart="FIGURE 11" captionStartId="14.[151,250,626,650]" captionTargetBox="[170,1417,181,602]" captionTargetId="figure-30@14.[170,1418,181,602]" captionTargetPageId="14" captionText="FIGURE 11. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Pharynx everted, left lateral view. B. Same, frontal view. Scale bars: A: 0.45 mm; B: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.4296795" httpUri="https://zenodo.org/record/4296795/files/figure.png" pageId="13" pageNumber="408">11A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3AA687FFC2E55666D0FDE4FCBF65C8" ID-DOI="http://doi.org/10.5281/zenodo.4296795" ID-Zenodo-Dep="4296795" httpUri="https://zenodo.org/record/4296795/files/figure.png" pageId="14" pageNumber="409" startId="14.[151,250,626,650]" targetBox="[170,1417,181,602]" targetPageId="14">
|
||
<paragraph id="8BFAF60FFFC2E55666D0FDE4FCBF65C8" blockId="14.[151,1436,626,686]" pageId="14" pageNumber="409">
|
||
<emphasis id="B9312A1DFFC2E55666D0FDE4FEC465EC" bold="true" box="[151,281,626,650]" pageId="14" pageNumber="409">FIGURE 11</emphasis>
|
||
.
|
||
<taxonomicName id="4C458D8CFFC2E5566761FDE4FDCF65EC" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[294,530,626,650]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="409" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC2E5566761FDE4FDCF65EC" box="[294,530,626,650]" italics="true" pageId="14" pageNumber="409">Chaetacanthus ornatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFC2E556645EFDE4FD8F65EC" box="[537,594,626,650]" pageId="14" pageNumber="409" rank="species">
|
||
<emphasis id="B9312A1DFFC2E556645EFDE4FD9165EC" bold="true" box="[537,588,626,650]" pageId="14" pageNumber="409">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype (ECOSUR 233), fixed in formalin solution. A. Pharynx everted, left lateral view. B. Same, frontal view. Scale bars: A: 0.45 mm; B: 0.50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3AA687FFC2E55666D0F8CEFD4D60DE" ID-DOI="http://doi.org/10.5281/zenodo.4296797" ID-Zenodo-Dep="4296797" httpUri="https://zenodo.org/record/4296797/files/figure.png" pageId="14" pageNumber="409" startId="14.[151,250,1880,1904]" targetBox="[170,1418,710,1856]" targetPageId="14">
|
||
<paragraph id="8BFAF60FFFC2E55666D0F8CEFD4D60DE" blockId="14.[151,1437,1880,1977]" pageId="14" pageNumber="409">
|
||
<emphasis id="B9312A1DFFC2E55666D0F8CEFEC46016" bold="true" box="[151,281,1880,1904]" pageId="14" pageNumber="409">FIGURE 12</emphasis>
|
||
.
|
||
<taxonomicName id="4C458D8CFFC2E5566761F8CEFDCC6016" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[294,529,1880,1904]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="409" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC2E5566761F8CEFDCC6016" box="[294,529,1880,1904]" italics="true" pageId="14" pageNumber="409">Chaetacanthus ornatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFC2E556645FF8CFFD8D6016" box="[536,592,1881,1905]" pageId="14" pageNumber="409" rank="species">
|
||
<emphasis id="B9312A1DFFC2E556645FF8CFFD976017" bold="true" box="[536,586,1881,1905]" pageId="14" pageNumber="409">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype (ECOSUR 233), fixed in formaline, elytra in dorsal view. A. Left elytron from first pair. B. Left elytron of middle segment. C. Left elytron from middle segment. D. Left elytron, from posterior segment. Scale bars: A, B: 0.6 mm; C: 0.8 mm; D: 0.7 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFAF60FFFC3E5576680FF01FDF5667F" blockId="15.[151,1437,151,1151]" pageId="15" pageNumber="410">
|
||
Tentacular segment not visible dorsally. Tentaculophores basally thick, inserted laterally to prostomium. Tentacular cirri with similar shape and pigmentation as median antenna. Segment two projected anteriorly over prostomium as a small round lobe (
|
||
<figureCitation id="137EEA8AFFC3E557679FFF4AFDE36790" box="[472,574,220,246]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="15" pageNumber="410">Fig. 10B</figureCitation>
|
||
). First pair of elytrophores long, not covering tentaculophores. Segment three narrower than successive segments.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC3E5576680FEB7FB8C656D" blockId="15.[151,1437,151,1151]" pageId="15" pageNumber="410">
|
||
All elytra with fringe of abundant, long papillae, and a bundle of long papillae in posterior inner margin (
|
||
<figureCitation id="137EEA8AFFC3E5576328FEB7FF6F6638" captionStart="FIGURE 12" captionStartId="14.[151,250,1880,1904]" captionTargetBox="[170,1418,710,1856]" captionTargetId="figure-69@14.[170,1418,710,1856]" captionTargetPageId="14" captionText="FIGURE 12. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formaline, elytra in dorsal view. A. Left elytron from first pair. B. Left elytron of middle segment. C. Left elytron from middle segment. D. Left elytron, from posterior segment. Scale bars: A, B: 0.6 mm; C: 0.8 mm; D: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296797" httpUri="https://zenodo.org/record/4296797/files/figure.png" pageId="15" pageNumber="410">Fig. 12</figureCitation>
|
||
A–D). First pair of elytra with abundant hemispherical microtubercles along its surface, each with surface spinulose; globular macrotubercles arranged in a curved J-shaped row, smaller along posterior border (
|
||
<figureCitation id="137EEA8AFFC3E55762ADFEF1FA8C66E7" box="[1258,1361,359,385]" captionStart="FIGURE 12" captionStartId="14.[151,250,1880,1904]" captionTargetBox="[170,1418,710,1856]" captionTargetId="figure-69@14.[170,1418,710,1856]" captionTargetPageId="14" captionText="FIGURE 12. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formaline, elytra in dorsal view. A. Left elytron from first pair. B. Left elytron of middle segment. C. Left elytron from middle segment. D. Left elytron, from posterior segment. Scale bars: A, B: 0.6 mm; C: 0.8 mm; D: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296797" httpUri="https://zenodo.org/record/4296797/files/figure.png" pageId="15" pageNumber="410">Fig. 12A</figureCitation>
|
||
) some pedunculated. From third segment, elytra bear globular macrotubercles with minute spines, mainly along posterior border, and sclerotized on central area forming a honeycomb patch (
|
||
<figureCitation id="137EEA8AFFC3E55765CDFE3AFC0466A0" box="[906,985,428,454]" captionStart="FIGURE 12" captionStartId="14.[151,250,1880,1904]" captionTargetBox="[170,1418,710,1856]" captionTargetId="figure-69@14.[170,1418,710,1856]" captionTargetPageId="14" captionText="FIGURE 12. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formaline, elytra in dorsal view. A. Left elytron from first pair. B. Left elytron of middle segment. C. Left elytron from middle segment. D. Left elytron, from posterior segment. Scale bars: A, B: 0.6 mm; C: 0.8 mm; D: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296797" httpUri="https://zenodo.org/record/4296797/files/figure.png" pageId="15" pageNumber="410">Fig. 12</figureCitation>
|
||
B–D). Towards dorsal midline, each elytron has dark microtubercles conical with thick peaks; fusiform macrotubercles long, thin, with short spines along their surface; along outer region numerous fusiform macrotubercles distally rounded.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC3E5576680FD82FCC765DF" blockId="15.[151,1437,151,1151]" pageId="15" pageNumber="410">
|
||
Parapodia biramous; branchial filaments digitate, abundant, some appearing septate (
|
||
<figureCitation id="137EEA8AFFC3E55762E4FD82FAD26548" box="[1187,1295,532,558]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="15" pageNumber="410">Fig. 10C</figureCitation>
|
||
). Notopodia short with acicular tips emerging. Neuropodia distally truncate, acicular tips slightly projected. Elytrophores wide, dorsal tubercles inconspicuous. Dorsal cirri short, not reaching neurochaetal tips, distally clavate with cirrophores short, swollen. Ventral cirri thick, short with filiform tips. Nephridial papillae present from segment 11, cylindrical, long. Pygidium with two anal cirri, similar to dorsal cirri.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC3E5576680FD57FDB56425" blockId="15.[151,1437,151,1151]" pageId="15" pageNumber="410">
|
||
Notochaetae abundant, thin, short, as long as half neurochaetal length, tips filiform, surface with rows of denticles (
|
||
<figureCitation id="137EEA8AFFC3E55766A2FD72FE926598" box="[229,335,740,766]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="15" pageNumber="410">Fig. 10C</figureCitation>
|
||
). Neurochaetae dark amber (
|
||
<figureCitation id="137EEA8AFFC3E55764D8FD72FCD56598" box="[671,776,740,766]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="15" pageNumber="410">Fig. 10C</figureCitation>
|
||
), thick, slightly curved, slender along first two segments (
|
||
<figureCitation id="137EEA8AFFC3E55766E7FC91FED86447" box="[160,261,775,801]" captionStart="FIGURE 10" captionStartId="13.[151,250,1608,1632]" captionTargetBox="[158,1430,181,1584]" captionTargetId="figure-21@13.[157,1430,181,1584]" captionTargetPageId="13" captionText="FIGURE 10. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233), fixed in formalin solution. A. Dorsal view, pharynx everted. B. Anterior end, dorsal view pharynx everted. C. Left parapodium, anterior view (inset: segment 3, neurochaetal tip). Scale bars: A: 2.8 mm; B: 0.2 mm; C: 0.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4296793" httpUri="https://zenodo.org/record/4296793/files/figure.png" pageId="15" pageNumber="410">Fig. 10C</figureCitation>
|
||
, inset), markedly larger and thicker thereafter; median segments with upper bundle with shorter chaetae, subdistally with scarce spines, tips entire.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC3E5576680FCDAFB0A631B" blockId="15.[151,1437,151,1151]" pageId="15" pageNumber="410">
|
||
<emphasis id="B9312A1DFFC3E5576680FCDAFEE26400" bold="true" box="[199,319,844,870]" pageId="15" pageNumber="410">Variation.</emphasis>
|
||
Elytral variation will be better understood in a comparison between
|
||
<taxonomicName id="4C458D8CFFC3E5576206FCDAFB6A6400" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[1089,1207,844,870]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC3E5576206FCDAFB6A6400" box="[1089,1207,844,870]" italics="true" pageId="15" pageNumber="410">C. ornatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFC3E55762F9FCDBFB276401" box="[1214,1274,845,871]" pageId="15" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<emphasis id="B9312A1DFFC3E5576376FCDAFE3764EE" italics="true" pageId="15" pageNumber="410">
|
||
<taxonomicName id="4C458D8CFFC3E5576376FCDAFF0764EE" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="brasiliensis">C. brasiliensis</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C458D8CFFC3E55766A2FCF9FE3764EE" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[229,490,879,905]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="ornatus">Chaetacanthus ornatus</taxonomicName>
|
||
</emphasis>
|
||
has swollen macrotubercles and larger honeycomb-like groups than those present in
|
||
<taxonomicName id="4C458D8CFFC3E55766D0FC04FEEA64CD" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[151,311,913,940]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFC3E55766D0FC04FEEA64CD" box="[151,311,913,940]" italics="true" pageId="15" pageNumber="410">C. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
. In first elytra,
|
||
<taxonomicName id="4C458D8CFFC3E55767A7FC04FD8B64CD" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[480,598,914,940]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="ornatus">
|
||
<emphasis id="B9312A1DFFC3E55767A7FC04FD8B64CD" box="[480,598,914,940]" italics="true" pageId="15" pageNumber="410">C. ornatus</emphasis>
|
||
</taxonomicName>
|
||
macrotubercles are about twice as large and mostly arranged in a single row (
|
||
<figureCitation id="137EEA8AFFC3E55766E7FC22FEDA64A8" box="[160,263,948,974]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410">Fig. 13A</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C458D8CFFC3E55767D9FC22FD9D64A8" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[414,576,948,974]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFC3E55767D9FC22FD9D64A8" box="[414,576,948,974]" italics="true" pageId="15" pageNumber="410">C. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
they are smaller, arranged in a row with 1–2 macrotubercles (
|
||
<figureCitation id="137EEA8AFFC3E55762B9FC22FABA64A8" box="[1278,1383,948,974]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410">Fig. 13D</figureCitation>
|
||
). By segment
|
||
<quantity id="4CBD5BEAFFC3E55766BCFC41FEF86497" box="[251,293,983,1009]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="15" pageNumber="410" unit="in" value="9.0">9 in</quantity>
|
||
<emphasis id="B9312A1DFFC3E557676CFC41FE756496" box="[299,424,983,1009]" italics="true" pageId="15" pageNumber="410">
|
||
<taxonomicName id="4C458D8CFFC3E557676CFC41FE796496" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[299,420,983,1009]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="ornatus">C. ornatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the honeycomb groups are 3–4 times larger and occupy about ¼ elytral surface (
|
||
<figureCitation id="137EEA8AFFC3E557636EFC41FA536497" box="[1321,1422,983,1009]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410">Fig. 13B</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C458D8CFFC3E5576750FC6CFE646375" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[279,441,1017,1044]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFC3E5576750FC6CFE646375" box="[279,441,1017,1044]" italics="true" pageId="15" pageNumber="410">C. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
they extend for about
|
||
<superScript id="7C305B47FFC3E55764FFFC6FFD1D6361" attach="right" box="[696,704,1017,1031]" fontSize="6" pageId="15" pageNumber="410">1</superScript>
|
||
/
|
||
<subScript id="17C1F44AFFC3E557648FFB9AFD0D637C" attach="left" box="[712,720,1036,1050]" fontSize="6" pageId="15" pageNumber="410">6</subScript>
|
||
of elytral surface and macrotubercles are smaller (
|
||
<figureCitation id="137EEA8AFFC3E557634CFC6FFAAD6375" box="[1291,1392,1017,1043]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410">Fig. 13E</figureCitation>
|
||
). In segment 24, the individual macrotubercles in
|
||
<taxonomicName id="4C458D8CFFC3E55764D7FB8AFCDB6350" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[656,774,1052,1078]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="ornatus">
|
||
<emphasis id="B9312A1DFFC3E55764D7FB8AFCDB6350" box="[656,774,1052,1078]" italics="true" pageId="15" pageNumber="410">C. ornatus</emphasis>
|
||
</taxonomicName>
|
||
have enlarged about 3–4 times more than in segment 9, and now they extend for about
|
||
<superScript id="7C305B47FFC3E5576786FBA8FE14632A" attach="right" box="[449,457,1086,1100]" fontSize="6" pageId="15" pageNumber="410">1</superScript>
|
||
/
|
||
<subScript id="17C1F44AFFC3E5576796FBC7FE046339" attach="left" box="[465,473,1105,1119]" fontSize="6" pageId="15" pageNumber="410">3</subScript>
|
||
elytral surface (
|
||
<figureCitation id="137EEA8AFFC3E55764CAFBA9FD2E633F" box="[653,755,1087,1113]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410">Fig. 13C</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C458D8CFFC3E55765C2FBA9FBFB633F" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[901,1062,1087,1113]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFC3E55765C2FBA9FBFB633F" box="[901,1062,1087,1113]" italics="true" pageId="15" pageNumber="410">C. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
the macrotubercles have doubled their size and expanded over elytra such that now they occupy about ¼ elytral surface (
|
||
<figureCitation id="137EEA8AFFC3E5576220FBF5FB16631B" box="[1127,1227,1123,1149]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. A–C. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. D–F. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410">Fig. 13F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3AA687FFC3E55766D0F8ECFC0C60BF" ID-DOI="http://doi.org/10.5281/zenodo.4296799" ID-Zenodo-Dep="4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="15" pageNumber="410" startId="15.[151,250,1914,1938]" targetBox="[174,1414,1177,1887]" targetPageId="15">
|
||
<paragraph id="8BFAF60FFFC3E55766D0F8ECFC0C60BF" blockId="15.[151,1436,1914,2010]" pageId="15" pageNumber="410">
|
||
<emphasis id="B9312A1DFFC3E55766D0F8ECFEC660F4" bold="true" box="[151,283,1914,1938]" pageId="15" pageNumber="410">FIGURE 13</emphasis>
|
||
. A–C.
|
||
<taxonomicName id="4C458D8CFFC3E5576721F8ECFD8E60F4" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[358,595,1914,1938]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC3E5576721F8ECFD8E60F4" box="[358,595,1914,1938]" italics="true" pageId="15" pageNumber="410">Chaetacanthus ornatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9312A1DFFC3E557641BF8ECFD4A60F4" bold="true" box="[604,663,1914,1938]" pageId="15" pageNumber="410">
|
||
<taxonomicNameLabel id="A2029766FFC3E557641BF8ECFD4A60F4" box="[604,663,1914,1938]" pageId="15" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (ECOSUR 233) elytra in dorsal view. D–F.
|
||
<taxonomicName id="4C458D8CFFC3E557628CF8ECFABD60F4" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[1227,1376,1914,1938]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="410" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFC3E557628CF8ECFABD60F4" box="[1227,1376,1914,1938]" italics="true" pageId="15" pageNumber="410">C. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35FA584FFDCE5486680FF00FA4867B2" pageId="16" pageNumber="411" type="etymology">
|
||
<paragraph id="8BFAF60FFFDCE5486680FF00FA4867B2" blockId="16.[151,1437,150,752]" pageId="16" pageNumber="411">
|
||
<emphasis id="B9312A1DFFDCE5486680FF00FE9567D6" bold="true" box="[199,328,150,176]" pageId="16" pageNumber="411">Etymology</emphasis>
|
||
. The specific epithet is derived from the Latin word
|
||
<taxonomicName id="4C458D8CFFDCE5486580FF01FBC067D6" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[967,1053,151,176]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="ornatus">
|
||
<emphasis id="B9312A1DFFDCE5486580FF01FBC067D6" box="[967,1053,151,176]" italics="true" pageId="16" pageNumber="411">ornatus</emphasis>
|
||
</taxonomicName>
|
||
(masculine), meaning furnished, equipped, adorned, or decorated, in reference to the large honeycomb group of macrotubercles on elytral surfaces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFDCE5486680FF48FA9E65E2" pageId="16" pageNumber="411" type="discussion">
|
||
<paragraph id="8BFAF60FFFDCE5486680FF48FF0266B6" blockId="16.[151,1437,150,752]" pageId="16" pageNumber="411">
|
||
<emphasis id="B9312A1DFFDCE5486680FF48FEE6679E" bold="true" box="[199,315,222,248]" pageId="16" pageNumber="411">Remarks.</emphasis>
|
||
Members of
|
||
<taxonomicName id="4C458D8CFFDCE5486795FF48FD05679E" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[466,728,222,248]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFDCE5486795FF48FD05679E" box="[466,728,222,248]" italics="true" pageId="16" pageNumber="411">Chaetacanthus ornatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFDCE5486498FF49FCC6679F" box="[735,795,223,249]" pageId="16" pageNumber="411" rank="species">n. sp.</taxonomicNameLabel>
|
||
were compared with the
|
||
<typeStatus id="54FE48ADFFDCE548627EFF49FBB4679F" box="[1081,1129,223,249]" pageId="16" pageNumber="411">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C458D8CFFDCE54862B5FF49FA49679E" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[1266,1428,222,249]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFDCE54862B5FF49FA49679E" box="[1266,1428,222,249]" italics="true" pageId="16" pageNumber="411">C. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C458D8CFFDCE54866D0FE95FED2667A" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[151,271,259,285]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="ornatus" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFDCE54866D0FE95FED2667A" box="[151,271,259,285]" italics="true" pageId="16" pageNumber="411">C. ornatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFDCE5486750FE95FE89667B" box="[279,340,259,285]" pageId="16" pageNumber="411" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs because its dorsal cirri are short, the facial tubercle is short and round, not elongate. The globular tubercles present on first pair of elytra are similar in shape to those of middle and posterior elytra; each is globular with spinous surface. Further,
|
||
<taxonomicName id="4C458D8CFFDCE548642AFEDDFD386602" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[621,741,331,357]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="ornatus">
|
||
<emphasis id="B9312A1DFFDCE548642AFEDDFD386602" box="[621,741,331,357]" italics="true" pageId="16" pageNumber="411">C. ornatus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C458D8CFFDCE548653DFEDDFBCB6602" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" baseAuthorityName="de Quatrefages" baseAuthorityYear="1866" box="[890,1046,330,357]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="brasiliensis">
|
||
<emphasis id="B9312A1DFFDCE548653DFEDDFBCB6602" box="[890,1046,330,357]" italics="true" pageId="16" pageNumber="411">C brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
because the macrotubercles of the central patch are not depressed and the patch is wider, covering a region larger than the elytrophoral plug. The long fusiform tubercles are distally blunt, without a sharp tip, and the microtubercles along the inner surface have thicker peaks.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFDCE5486680FE4DFA9E65E2" blockId="16.[151,1437,150,752]" pageId="16" pageNumber="411">
|
||
<bibRefCitation id="EFD48BFEFFDCE5486680FE4DFEB96692" author="Monro, C. C. A." box="[199,356,474,501]" pageId="16" pageNumber="411" pagination="553 - 576" refId="ref14860" refString="Monro, C. C. A. (1928) Polychaeta of the families Polynoidae and Acoetidae from the vicinity of the Panama Canal, collected by Dr. C. Crossland and Dr. Th. Mortensen. Zoological Journal of the Linnean Society, London, 36, 553 - 576. https: // doi. org / 10.1111 / j. 1096 - 3642.1928. tb 02209. x" type="journal article" year="1928">Monro (1928)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C458D8CFFDCE5486795FE4DFDB16692" authorityName=": Hernandez-Moreno" authorityYear="2011" box="[466,620,474,501]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="magnificus">
|
||
<emphasis id="B9312A1DFFDCE5486795FE4DFDB16692" box="[466,620,474,501]" italics="true" pageId="16" pageNumber="411">C. magnificus</emphasis>
|
||
</taxonomicName>
|
||
from Coiba and Gorgona Islands, in the Eastern Pacific, and indicated his accounts were the first records for the Caribbean species into the Pacific Ocean. Monro materials were not examined, but they are regarded as conspecific with
|
||
<taxonomicName id="4C458D8CFFDCE54864E0FDB5FCFC655A" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[679,801,547,573]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="ornatus">
|
||
<emphasis id="B9312A1DFFDCE54864E0FDB5FD67655B" box="[679,698,547,573]" italics="true" pageId="16" pageNumber="411">C</emphasis>
|
||
.
|
||
<emphasis id="B9312A1DFFDCE548648FFDB5FCFC655A" box="[712,801,547,572]" italics="true" pageId="16" pageNumber="411">ornatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9312A1DFFDCE5486566FDB5FCF8655A" box="[801,805,547,572]" italics="true" pageId="16" pageNumber="411">.</emphasis>
|
||
<bibRefCitation id="EFD48BFEFFDCE548656CFDB5FC36655A" author="Hartman, O." box="[811,1003,546,573]" pageId="16" pageNumber="411" pagination="1 - 156" refId="ref13886" refString="Hartman, O. (1939 a) Polychaetous annelids, 1. Aphroditidae to Pisionidae. Allan Hancock Pacific Expeditions, 7, 1 - 156, pls. 1 - 28." type="journal article" year="1939">Hartman (1939a)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C458D8CFFDCE548621CFDB5FB2A655A" authorityName=": Hernandez-Moreno" authorityYear="2011" box="[1115,1271,546,573]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="411" phylum="Annelida" rank="species" species="magnificus">
|
||
<emphasis id="B9312A1DFFDCE548621CFDB5FB2A655A" box="[1115,1271,546,573]" italics="true" pageId="16" pageNumber="411">C. magnificus</emphasis>
|
||
</taxonomicName>
|
||
for the Pacific with specimens from
|
||
<collectingCountry id="F352B69FFFDCE54867CFFDD0FE3A6506" box="[392,487,582,608]" name="Ecuador" pageId="16" pageNumber="411">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="F352B69FFFDCE54867B5FDD0FD966506" box="[498,587,582,608]" name="Panama" pageId="16" pageNumber="411">Panamá</collectingCountry>
|
||
and
|
||
<collectingCountry id="F352B69FFFDCE54864C7FDD0FD056506" box="[640,728,582,608]" name="Mexico" pageId="16" pageNumber="411">México</collectingCountry>
|
||
, The specimens from Socorro Island,
|
||
<collectingRegion id="498138EDFFDCE5486239FDD0FB0D6506" box="[1150,1232,582,608]" country="Mexico" name="Colima" pageId="16" pageNumber="411">Colima</collectingRegion>
|
||
(sta. 132-34), and from Petatlan Bay,
|
||
<collectingRegion id="498138EDFFDCE5486728FDFCFE0965E2" box="[367,468,618,644]" country="Mexico" name="Guerrero" pageId="16" pageNumber="411">Guerrero</collectingRegion>
|
||
(sta. 264-34) require corroboration, but might also belong to this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFDCE5486680FD18FCDD65CE" box="[199,768,654,681]" pageId="16" pageNumber="411" type="materials_examined">
|
||
<paragraph id="8BFAF60FFFDCE5486680FD18FCDD65CE" blockId="16.[151,1437,150,752]" box="[199,768,654,681]" pageId="16" pageNumber="411">
|
||
<materialsCitation id="3B2DFC52FFDCE5486680FD18FD2165CE" ID-GBIF-Occurrence="2989844302" box="[199,764,654,681]" country="Mexico" location="Puerto Escondido" pageId="16" pageNumber="411" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
|
||
<emphasis id="B9312A1DFFDCE5486680FD18FEBB65CE" bold="true" box="[199,358,654,680]" pageId="16" pageNumber="411">
|
||
<typeStatus id="54FE48ADFFDCE5486680FD18FEDF65CE" box="[199,258,654,680]" pageId="16" pageNumber="411">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8E9AA0D4FFDCE548672BFD19FDE865CE" LSID="urn:lsid:plazi:treatment:03EC4719FFC0E54866D0F9A0FF306596:8E9AA0D4FFDCE548672BFD19FDE865CE" box="[364,565,654,681]" country="Mexico" name="Puerto Escondido" pageId="16" pageNumber="411" stateProvince="Oaxaca">Puerto Escondido</location>
|
||
,
|
||
<collectingRegion id="498138EDFFDCE5486406FD18FD4565CE" box="[577,664,654,680]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="411">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry id="F352B69FFFDCE54864E4FD18FD2165CE" box="[675,764,654,680]" name="Mexico" pageId="16" pageNumber="411">México</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFDCE5486680FD24FF306596" pageId="16" pageNumber="411" type="distribution">
|
||
<paragraph id="8BFAF60FFFDCE5486680FD24FF306596" blockId="16.[151,1437,150,752]" pageId="16" pageNumber="411">
|
||
<emphasis id="B9312A1DFFDCE5486680FD24FEBC65AA" bold="true" box="[199,353,690,716]" pageId="16" pageNumber="411">Distribution.</emphasis>
|
||
Puerto Escondido,
|
||
<collectingRegion id="498138EDFFDCE548647AFD24FD4965AA" box="[573,660,690,716]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="411">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry id="F352B69FFFDCE54864D8FD24FD2565AA" box="[671,760,690,716]" name="Mexico" pageId="16" pageNumber="411">México</collectingCountry>
|
||
; probably also present in Coiba,
|
||
<collectingCountry id="F352B69FFFDCE548622CFD25FB1A65AB" box="[1131,1223,691,717]" name="Panama" pageId="16" pageNumber="411">Panama</collectingCountry>
|
||
; Isla Gorgona,
|
||
<collectingCountry id="F352B69FFFDCE5486335FD24FF356596" name="Colombia" pageId="16" pageNumber="411">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |