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<document id="1F6D34375A1553CC67EBF0F744ED1D57" ID-CLB-Dataset="2764" ID-DOI="10.11646/zootaxa.5087.1.5" ID-GBIF-Dataset="fc6ee7f7-704f-4356-b63d-be4ff718763a" ID-ISSN="1175-5326" ID-Zenodo-Dep="5819774" ID-ZooBank="40518AD3-6B53-408C-919D-466C81038C6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641371269439" checkinUser="plazi" docAuthor="Cao, Tong, Watson, Gillian W., Hodgson, Chris J., Jing, Qi &amp; Feng, Ji-Nian" docDate="2022" docId="03BE3D79FFEFD429FF53FF5DFB5E2AB7" docLanguage="en" docName="zootaxa.5087.1.5.pdf" docOrigin="Zootaxa 5087 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coccus nanningensis Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng 2022, sp. n." docType="treatment" docVersion="7" lastPageNumber="122" masterDocId="FF874501FFE7D423FFC4FFCAFFCE2D30" masterDocTitle="The genera Coccus and Prococcus (Hemiptera: Coccomorpha: Coccidae) in China with two new combinations and descriptions of two new species" masterLastPageNumber="128" masterPageNumber="112" pageNumber="120" updateTime="1699295769707" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="43976625FD199C9E0BCD6986FFAD097B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E31A43A7228EFDB5C7D4A8B4A5F74BCB">
<mods:title id="DFB5A2696607FC848C9E724D6467123D">The genera Coccus and Prococcus (Hemiptera: Coccomorpha: Coccidae) in China with two new combinations and descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name id="08D4DF109531229F573F5C5BAFB8F7DF" type="personal">
<mods:role id="17F4C2ED1BB9B5984576ECB5123A616C">
<mods:roleTerm id="616F1F07D704D0B32AD6316BF8BB713D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81435AB9B9FDF763A4785DF5DA47EE30">Cao, Tong</mods:namePart>
<mods:nameIdentifier id="F984E5A8A19A6AA1FB147C6F527DA03B" type="ORCID">0000-0002-6738-9976</mods:nameIdentifier>
<mods:affiliation id="EE8D15C17ADDE6A57DB32F4E4A6474C9">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi Province, 712100, China. &amp; amycaotong @ 163. com; https: // orcid. org / 0000 - 0002 - 6738 - 9976</mods:affiliation>
</mods:name>
<mods:name id="EB277B796367F198614CD55917F4341C" type="personal">
<mods:role id="DD241F95BDF28F289BF81ECF90C5B045">
<mods:roleTerm id="49446F224C460038A49747FE68971D7E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B48D366407131047A16BABFD8A4B2EB">Watson, Gillian W.</mods:namePart>
<mods:affiliation id="D59814FBDF599B6ED7A0CD9B78ECCA07">Department of Life Sciences, the Natural History Museum, London, SW 7 5 BD, U. K.</mods:affiliation>
</mods:name>
<mods:name id="09DAB258BF04AE5D057BB48668F3725D" type="personal">
<mods:role id="725A436EE3173E4E48577C76619BD638">
<mods:roleTerm id="DA30DD31FA671E948B3D18400E10DC6D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="80782928C8B1C5D9828A1A8E16002A7F">Hodgson, Chris J.</mods:namePart>
<mods:nameIdentifier id="E90B8E5657C3CBE5307EB5CE977A09A3" type="ORCID">0000-0002-9073-1485</mods:nameIdentifier>
<mods:affiliation id="9B895A25E3AE21BB665F00626B48B87A">Department of Biodiversity and Biological Systematics, The National Museum of Wales, Cardiff, Wales, CF 10 3 NP, U. K. hodgsoncj @ cardiff. ac. uk; https: // orcid. org / 0000 - 0002 - 9073 - 1485</mods:affiliation>
<mods:nameIdentifier id="F7B5511D95D12086973A488D7290627B" type="email">hodgsoncj@cardiff.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="6459708C0AD0EEFD13B478CB590E4D6B" type="personal">
<mods:role id="C443F3764FC34829907D2A9AAB4BCC56">
<mods:roleTerm id="4B17DE617B3D61ABAB600AE7DC7C2EE5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81F36EE546EC8B03FCF1A0A2DC0A3319">Jing, Qi</mods:namePart>
<mods:nameIdentifier id="8A30CD0816EE226B275C5B2026EE76A7" type="ORCID">0000-0002-7865-7761</mods:nameIdentifier>
<mods:affiliation id="72BA45D0AC2FA73E3CDC0B5AD551048F">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi Province, 712100, China. &amp; 2018050190 @ nwafu. edu. cn; https: // orcid. org / 0000 - 0002 - 7865 - 7761</mods:affiliation>
<mods:nameIdentifier id="C2C8279C575A050DCACE903B012262B2" type="email">2018050190@nwafu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="32EE1222E234ADDAD07B8B72069B726D" type="personal">
<mods:role id="EBD62D228D72BD0AEAF769898BDF0E8B">
<mods:roleTerm id="88154242715B281209C3E534F1259014">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="84E9278447D0F4FA1206731EC7294ABB">Feng, Ji-Nian</mods:namePart>
<mods:nameIdentifier id="C3DA415E26798A23F087B0A22DC5A108" type="ORCID">0000-0002-3994-1348</mods:nameIdentifier>
<mods:affiliation id="3D892467F83E6DFEBCCF052A88F88776">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi Province, 712100, China. &amp; jinianf @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 3994 - 1348</mods:affiliation>
<mods:nameIdentifier id="0C25880C95553896BA02233BCEBF2D17" type="email">jinianf@nwsuaf.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8249CCD731700C1F3469FB59CC710786">text</mods:typeOfResource>
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<mods:title id="3BAAF76BA37746E7C9F47CAE3D26F40A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7144DB40D1E42328D08811314DC00C50">
<mods:date id="B8B6226F8C3576F5F52A23907B137D3B">2022</mods:date>
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<mods:number id="D218653190FA6F775360082E56257E60">2022-01-05</mods:number>
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<mods:number id="063DE60E8783F92D2A86D29E05F948AF">5087</mods:number>
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<mods:classification id="1481DECB42CE08CC532AE324F2116BE6">journal article</mods:classification>
<mods:identifier id="39751C3D05D370F8A0238BB59EE1ED7C" type="CLB-Dataset">2764</mods:identifier>
<mods:identifier id="8C22AE8DB1FB8AA66C1A4422EFB5AB62" type="DOI">10.11646/zootaxa.5087.1.5</mods:identifier>
<mods:identifier id="D2EC959AB77A1E9ABEA14DF3AF18184C" type="GBIF-Dataset">fc6ee7f7-704f-4356-b63d-be4ff718763a</mods:identifier>
<mods:identifier id="C6AE519C2A0A4FA07123EFD755A45A36" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BE3D79FFEFD429FF53FF5DFB5E2AB7" ID-DOI="http://doi.org/10.5281/zenodo.5828007" ID-GBIF-Taxon="191928624" ID-Zenodo-Dep="5828007" LSID="urn:lsid:plazi:treatment:03BE3D79FFEFD429FF53FF5DFB5E2AB7" httpUri="http://treatment.plazi.org/id/03BE3D79FFEFD429FF53FF5DFB5E2AB7" lastPageId="10" lastPageNumber="122" pageId="8" pageNumber="120">
<subSubSection id="C30DDFE4FFEFD42BFF53FF5DFD472D81" box="[151,649,151,177]" pageId="8" pageNumber="120" type="nomenclature">
<paragraph id="8BA88C6FFFEFD42BFF53FF5DFD472D81" blockId="8.[151,649,151,214]" box="[151,649,151,177]" pageId="8" pageNumber="120">
<heading id="D0E03B03FFEFD42BFF53FF5DFD472D81" bold="true" box="[151,649,151,177]" fontSize="11" level="1" pageId="8" pageNumber="120" reason="1">
<emphasis id="B963507DFFEFD42BFF53FF5DFD472D81" bold="true" box="[151,649,151,177]" pageId="8" pageNumber="120">
<taxonomicName id="4C17F7ECFFEFD42BFF53FF5DFDF52D81" authority="Cao &amp; Feng" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[151,571,151,177]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="120" phylum="Arthropoda" rank="species" species="nanningensis" status="sp. nov.">
<emphasis id="B963507DFFEFD42BFF53FF5DFE542D81" bold="true" box="[151,410,151,177]" italics="true" pageId="8" pageNumber="120">Coccus nanningensis</emphasis>
Cao &amp; Feng
</taxonomicName>
,
<taxonomicNameLabel id="A250ED06FFEFD42BFD83FF5DFD472D81" box="[583,649,151,177]" pageId="8" pageNumber="120" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30DDFE4FFEFD42BFF53FF71FEFD2DE6" box="[151,307,187,214]" pageId="8" pageNumber="120" type="description">
<paragraph id="8BA88C6FFFEFD42BFF53FF71FEFD2DE6" blockId="8.[151,649,151,214]" box="[151,307,187,214]" pageId="8" pageNumber="120">
(
<figureCitation id="132C90EAFFEFD42BFF5BFF71FF282DE5" box="[159,230,187,213]" captionStart="FIGURE 4" captionStartId="9.[151,250,1883,1907]" captionTargetBox="[174,1405,193,1855]" captionTargetId="figure-19@9.[164,1423,181,1859]" captionTargetPageId="9" captionText="FIGURE 4. Adult female of Coccus nanningensis sp. n. Abbreviations: AGF ano-genital fold; ANP anal plates; ANT antenna; DA body derm appearance; DMD dorsal microduct; DS dorsal setae; DT dorsal tubercles; LG tibio-tarsal joint of hind leg; MS marginal setae; MDP multilocular disc pores; POP preopercular pore; SDP spiracular disc pores; SP spiracle; SSP stigmatic spines; VS ventral setae." figureDoi="http://doi.org/10.5281/zenodo.5819786" httpUri="https://zenodo.org/record/5819786/files/figure.png" pageId="8" pageNumber="120">Figs 4</figureCitation>
and
<figureCitation id="132C90EAFFEFD42BFED8FF71FEE42DE6" box="[284,298,187,214]" captionStart="FIGURE 5" captionStartId="10.[151,250,1038,1062]" captionTargetBox="[216,1372,201,1006]" captionTargetId="figure-17@10.[201,1386,181,1014]" captionTargetPageId="10" captionText="FIGURE 5. A photograph of the type slides of Coccus cephalotaxus Cao &amp; Feng sp. n. (bottom) and C. nanningensis Cao &amp; Feng sp. n. (top) to facilitate future recognition. The holotype specimen on each slide is ringed in blue." figureDoi="http://doi.org/10.5281/zenodo.5819788" httpUri="https://zenodo.org/record/5819788/files/figure.png" pageId="8" pageNumber="120">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30DDFE4FFEFD42BFF53FEC9FAED2C9D" pageId="8" pageNumber="120" type="materials_examined">
<paragraph id="8BA88C6FFFEFD42BFF53FEC9FAED2C9D" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF53FEC9FE4D2C2D" bold="true" box="[151,387,259,285]" pageId="8" pageNumber="120">Material examined.</emphasis>
<materialsCitation id="3B7F8632FFEFD42BFE4FFEC9FC512C55" ID-GBIF-Occurrence="3427697304" collectingDate="2019-08-10" collectionCode="NWAFU" country="China" location="Nanning" municipality="Nanning" pageId="8" pageNumber="120" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi Province" typeStatus="holotype">
<emphasis id="B963507DFFEFD42BFE4FFEC9FE362C2D" bold="true" box="[395,504,259,285]" pageId="8" pageNumber="120">
<typeStatus id="54AC32CDFFEFD42BFE4FFEC9FE362C2D" box="[395,504,259,285]" pageId="8" pageNumber="120" type="holotype">Holotype</typeStatus>
</emphasis>
adult
<emphasis id="B963507DFFEFD42BFD86FECFFD082C2D" bold="true" box="[578,710,259,288]" pageId="8" pageNumber="120">
<specimenCount id="9D1147E6FFEFD42BFD86FECFFD972C10" box="[578,601,261,288]" pageId="8" pageNumber="120" type="female">♀</specimenCount>
:
<collectingCountry id="F300CCFFFFEFD42BFDADFEC9FD082C2D" box="[617,710,259,285]" name="China" pageId="8" pageNumber="120">CHINA</collectingCountry>
</emphasis>
,
<collectingRegion id="49D3428DFFEFD42BFD13FEC9FC662C2D" box="[727,936,259,285]" country="China" name="Guangxi" pageId="8" pageNumber="120">Guangxi Province</collectingRegion>
: label:
<collectingMunicipality id="6BCC1615FFEFD42BFC3FFEC9FB902C2D" box="[1019,1118,259,285]" pageId="8" pageNumber="120">Nanning</collectingMunicipality>
, Guangxi Province, China / Guangxi University of Finance and Economics /
<date id="FFA9AAAFFFEFD42BFD00FEEDFC822C71" box="[708,844,295,321]" pageId="8" pageNumber="120" value="2019-08-10">
<collectingDate id="EFED5347FFEFD42BFD00FEEDFC822C71" box="[708,844,295,321]" pageId="8" pageNumber="120" value="2019-08-10">10.viii.2019</collectingDate>
</date>
on
<taxonomicName id="4C17F7ECFFEFD42BFCBCFEE2FB402C71" box="[888,1166,295,321]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="120" phylum="Tracheophyta" rank="species" species="carica">
<emphasis id="B963507DFFEFD42BFCBCFEE2FBC92C71" box="[888,1031,296,321]" italics="true" pageId="8" pageNumber="120">Ficus carica</emphasis>
(Moraceae)
</taxonomicName>
/ Cao Tong (
<collectionCode id="ED0614AAFFEFD42BFAD9FEEDFA4B2C71" box="[1309,1413,295,321]" pageId="8" pageNumber="120">NWAFU</collectionCode>
) /
<taxonomicName id="4C17F7ECFFEFD42BFF53FE86FDD12C56" authority="Cao &amp; Feng" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[151,543,331,358]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="120" phylum="Arthropoda" rank="species" species="nanningensis">
<emphasis id="B963507DFFEFD42BFF53FE86FE482C55" box="[151,390,332,358]" italics="true" pageId="8" pageNumber="120">Coccus nanningensis</emphasis>
Cao &amp; Feng
</taxonomicName>
; holotype on the left side, circled
</materialsCitation>
.
<materialsCitation id="3B7F8632FFEFD42BFC6FFE81FAD12C9D" ID-GBIF-Occurrence="3427697307" pageId="8" pageNumber="120" typeStatus="paratype">
<emphasis id="B963507DFFEFD42BFC6FFE81FB912C58" bold="true" box="[939,1119,331,360]" pageId="8" pageNumber="120">
<typeStatus id="54AC32CDFFEFD42BFC6FFE81FBED2C55" box="[939,1059,331,357]" pageId="8" pageNumber="120" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D1147E6FFEFD42BFBE8FE87FB972C58" box="[1068,1113,333,360]" pageId="8" pageNumber="120" type="female">♀♀</specimenCount>
:
</emphasis>
<specimenCount id="9D1147E6FFEFD42BFBA3FE81FB782C55" box="[1127,1206,331,357]" pageId="8" pageNumber="120" type="adult">1 adult</specimenCount>
female on the same slide with
<typeStatus id="54AC32CDFFEFD42BFECDFEA5FEA02CB9" box="[265,366,367,393]" pageId="8" pageNumber="120" type="holotype">holotype</typeStatus>
, in the lower right corner, plus 1 slide with
<specimenCount id="9D1147E6FFEFD42BFC8DFEA5FC5B2CB9" box="[841,917,367,393]" pageId="8" pageNumber="120" type="adult">2 adult</specimenCount>
females, and another 2 slides each with
<specimenCount id="9D1147E6FFEFD42BFA94FEA5FA522CB9" box="[1360,1436,367,393]" pageId="8" pageNumber="120" type="adult">3 adult</specimenCount>
females, collection data same as
<typeStatus id="54AC32CDFFEFD42BFDC2FE59FDA62C9D" box="[518,616,403,429]" pageId="8" pageNumber="120" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED0614AAFFEFD42BFDB3FE59FD2C2C9D" box="[631,738,403,429]" pageId="8" pageNumber="120">NWAFU</collectionCode>
). All the slide labels (
<figureCitation id="132C90EAFFEFD42BFC16FE59FBDB2C9E" box="[978,1045,403,430]" captionStart="FIGURE 5" captionStartId="10.[151,250,1038,1062]" captionTargetBox="[216,1372,201,1006]" captionTargetId="figure-17@10.[201,1386,181,1014]" captionTargetPageId="10" captionText="FIGURE 5. A photograph of the type slides of Coccus cephalotaxus Cao &amp; Feng sp. n. (bottom) and C. nanningensis Cao &amp; Feng sp. n. (top) to facilitate future recognition. The holotype specimen on each slide is ringed in blue." figureDoi="http://doi.org/10.5281/zenodo.5819788" httpUri="https://zenodo.org/record/5819788/files/figure.png" pageId="8" pageNumber="120">Fig. 5</figureCitation>
) are written in Chinese
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30DDFE4FFEFD42BFF03FE7DFC582A7D" pageId="8" pageNumber="120" type="description">
<paragraph id="8BA88C6FFFEFD42BFF03FE7DFD132CE2" blockId="8.[151,1437,259,2013]" box="[199,733,439,466]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03FE7DFE9F2CE1" bold="true" box="[199,337,439,465]" pageId="8" pageNumber="120">Description</emphasis>
(data taken from all
<specimenCount id="9D1147E6FFEFD42BFDF8FE7DFD1B2CE2" box="[572,725,439,466]" pageId="8" pageNumber="120" type="generic">10 specimens</specimenCount>
)
</paragraph>
<paragraph id="8BA88C6FFFEFD42BFF03FE11FEA02F29" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03FE11FE742CC5" bold="true" box="[199,442,475,501]" pageId="8" pageNumber="120">Adult female in life.</emphasis>
Body oval to subcircular, yellowish-brown to pale brown, with symmetrical dark spot present on dorsum.
</paragraph>
<paragraph id="8BA88C6FFFEFD42BFF03FDE9FD4E2F51" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03FDE9FDD42F0D" bold="true" box="[199,538,547,573]" pageId="8" pageNumber="120">Slide-mounted adult female.</emphasis>
Body elongate oval to broadly oval; 1.5–2.0 mm long, 1.0–
<quantity id="4CEF218AFFEFD42BFB01FDE9FAD42F0E" box="[1221,1306,547,574]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="120" unit="mm" value="1.5">1.5 mm</quantity>
wide. Anal cleft approximately 1/7–1/6 of body length.
</paragraph>
<paragraph id="8BA88C6FFFEFD42BFF03FDA6FA4E2EDD" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03FDA6FEE72FB5" box="[199,297,620,645]" italics="true" pageId="8" pageNumber="120">Dorsum.</emphasis>
Derm with small circular clear cell-like areolations distributed over entire dorsum, becoming well developed on older females. Dorsal microducts present singly in areolations. Setae setose, fine and pointed, each with a well-developed basal socket, scattered, each about 8–13 μm long. Submarginal tubercles present on anterior half of body only (with 1 or 2 on each side, numbering 0‒2 between anterior stigmatic clefts on head, 0 or 1 between anterior and posterior stigmatic clefts on each side), absent from abdominal region. Dorsal tubular ducts absent. Preopercular pores absent. Anal plates each triangular, together quadrate, 111–144 μm long, 82–92 μm wide, width of anal plates together slightly longer than their length, anterolateral margin 108–114 μm long, posterolateral margin 109–119 μm long, length of posterior margin almost equal to anterior margin, outer angle slightly obtuse; each plate with a well-developed supporting bar, and a robust discal seta with a pointed apex, 41–57 μm long; also 3 apical setae, each 15–17 μm long.Anogenital fold with 3 pairs of anterior margin setae, each 51–63 μm long, and 3 pairs of lateral marginal setae, each 41–68 μm long. Anal ring subcircular, bearing 8 anal ring setae. Eyespots not found.
</paragraph>
<paragraph id="8BA88C6FFFEFD42BFF03FC32FC0F2802" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03FC32FEEC2921" box="[199,290,1016,1041]" italics="true" pageId="8" pageNumber="120">Margin.</emphasis>
Marginal setae lengths variable, 22–67 μm long, in 1 or 2 rows, with well-developed basal sockets, structure highly varied, mostly rather long and extremely robust, but some with apices branched or bifurcate, others either conical, straight or curved with pointed apices (hard to distinguish from the stigmatic spine), or with apices swollen, sagittate or spatulate, or setose, extremely slender, fine and sharply pointed; with 39–45 setae between anterior stigmatic clefts, 10–12 setae on each side between anterior and posterior stigmatic clefts, and 22–25 setae between the posterior stigmatic cleft and anal cleft. Stigmatic clefts shallow but distinct, each cleft containing 3 slender, tapering and bluntly spinose stigmatic spines, with well-developed basal sockets; median spine longest, 108–126 μm long, about 2.5–6.5 times as long as a lateral spine, each 21–47 μm long; (
<specimenCount id="9D1147E6FFEFD42BFBA6FB39FB33283D" box="[1122,1277,1267,1293]" pageId="8" pageNumber="120" type="generic">one specimen</specimenCount>
with one cleft containing 1 stigmatic spine, this only as long as a normal lateral spine).
</paragraph>
<paragraph id="8BA88C6FFFEFD42BFF03FAF6FC582A7D" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03FAF6FEDD2865" box="[199,275,1340,1365]" italics="true" pageId="8" pageNumber="120">Venter.</emphasis>
Derm membranous. Antennae each with 7 segments, total antennal length 355–383 μm, segment III longest, 79–83 μm long; lengths of other segments in μm: I, 46–54; II, 42–45; IV, 54–71; V, 33–37; VI, 32–39; and VII, 61–81. Setal distribution on antenna: scape 3; pedicel 2; III, 0; IV, 2 or 3; V &amp; VI, each segment with 1 fleshy seta and 1 hair-like seta; VII with 3 fleshy setae, 3 stiff setae and 2 hair-like setae. Inter-antennal setae numbering 3 or 4 pairs, comprising 1 pair of long outer setae, each 64–66 μm long, and 2 or 3 pairs of short inner setae, each 11–15 μm long. Long pregenital setae numbering 2 pairs (
<specimenCount id="9D1147E6FFEFD42BFCE3FA25FC6B2B39" box="[807,933,1519,1545]" pageId="8" pageNumber="120" type="generic">1 specimen</specimenCount>
with 3 pregenital setae on one side but other side with only 2, a total of 5 setae on 3 segments), each seta 79–83 μm long. Other ventral setae setose, fine, sparsely distributed over entire venter, each seta 9–17 μm long. Submarginal setae not observed. Legs well developed, each with a tibio-tarsal articulation but no articulatory sclerosis; tibia 115–122 μm long, longer than tarsus; tarsus 85–97 μm long. Claw without a denticle; claw digitules similar, each broad and expanded at apex, about 30–31 μm long. Tarsal digitules longer than claw digitules, slender, knobbed, expanded at apex, each about 44–50 μm long. Spiracles normal, without sclerotic plates. Spiracular disc pores mostly each with 5 loculi in outer ring, occasionally 4 or 6; spiracular pore bands narrow, each 2 or 3 rows wide. Anterior spiracular pore band with 15–23 pores, posterior spiracular pore band with 20–33 pores. Multilocular disc pores each primarily with 7 loculi, occasionally with 8‒10, restricted to genital area (segment VII). Ventral tubular ducts absent.
</paragraph>
</subSubSection>
<subSubSection id="C30DDFE4FFEFD42BFF03F89DFB442AA5" pageId="8" pageNumber="120" type="etymology">
<paragraph id="8BA88C6FFFEFD42BFF03F89DFB442AA5" blockId="8.[151,1437,259,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03F89DFE802A41" bold="true" box="[199,334,1879,1905]" pageId="8" pageNumber="120">Etymology.</emphasis>
The species epithet
<taxonomicName id="4C17F7ECFFEFD42BFDF4F892FD0A2A41" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[560,708,1880,1905]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="120" phylum="Arthropoda" rank="species" species="nanningensis">
<emphasis id="B963507DFFEFD42BFDF4F892FD0A2A41" box="[560,708,1880,1905]" italics="true" pageId="8" pageNumber="120">nanningensis</emphasis>
</taxonomicName>
is formed from the name of the place where this new species was collected, i.e., the city of Nanning, combined with the Latin suffix -
<emphasis id="B963507DFFEFD42BFC48F8B6FC0A2AA5" box="[908,964,1916,1941]" italics="true" pageId="8" pageNumber="120">ensis</emphasis>
, meaning ‘from’.
</paragraph>
</subSubSection>
<subSubSection id="C30DDFE4FFEFD42BFF03F855FC372A8A" box="[199,1017,1951,1978]" pageId="8" pageNumber="120" type="biology_ecology">
<paragraph id="8BA88C6FFFEFD42BFF03F855FC372A8A" blockId="8.[151,1437,259,2013]" box="[199,1017,1951,1978]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03F855FE852A89" bold="true" box="[199,331,1951,1977]" pageId="8" pageNumber="120">Host plant.</emphasis>
The species was collected from the leaves of
<taxonomicName id="4C17F7ECFFEFD42BFC89F86AFC142A89" box="[845,986,1952,1977]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="120" phylum="Tracheophyta" rank="species" species="carica">
<emphasis id="B963507DFFEFD42BFC89F86AFC142A89" box="[845,986,1952,1977]" italics="true" pageId="8" pageNumber="120">Ficus carica</emphasis>
</taxonomicName>
L.
</paragraph>
</subSubSection>
<subSubSection id="C30DDFE4FFEFD42BFF03F809FD6C2AED" box="[199,674,1987,2013]" pageId="8" pageNumber="120" type="distribution">
<paragraph id="8BA88C6FFFEFD42BFF03F809FD6C2AED" blockId="8.[151,1437,259,2013]" box="[199,674,1987,2013]" pageId="8" pageNumber="120">
<emphasis id="B963507DFFEFD42BFF03F809FEAF2AED" bold="true" box="[199,353,1987,2013]" pageId="8" pageNumber="120">Distribution.</emphasis>
<collectingCountry id="F300CCFFFFEFD42BFEA3F809FE0A2AED" box="[359,452,1987,2013]" name="China" pageId="8" pageNumber="120">CHINA</collectingCountry>
:
<collectingRegion id="49D3428DFFEFD42BFE0BF809FD532AED" box="[463,669,1987,2013]" country="China" name="Guangxi" pageId="8" pageNumber="120">Guangxi Province</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="DF68DCE7FFEED42AFF53F891FE5F2AEF" ID-DOI="http://doi.org/10.5281/zenodo.5819786" ID-Zenodo-Dep="5819786" httpUri="https://zenodo.org/record/5819786/files/figure.png" pageId="9" pageNumber="121" startId="9.[151,250,1883,1907]" targetBox="[174,1405,193,1855]" targetPageId="9">
<paragraph id="8BA88C6FFFEED42AFF53F891FE5F2AEF" blockId="9.[151,1436,1883,2015]" pageId="9" pageNumber="121">
<emphasis id="B963507DFFEED42AFF53F891FEDC2A43" bold="true" box="[151,274,1883,1907]" pageId="9" pageNumber="121">FIGURE 4.</emphasis>
Adult female of
<taxonomicName id="4C17F7ECFFEED42AFE78F896FD5D2A43" authority="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng, 2022" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[444,659,1884,1907]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="121" phylum="Arthropoda" rank="species" species="nanningensis" status="sp. nov.">
<emphasis id="B963507DFFEED42AFE78F896FD5D2A43" box="[444,659,1884,1907]" italics="true" pageId="9" pageNumber="121">Coccus nanningensis</emphasis>
</taxonomicName>
<emphasis id="B963507DFFEED42AFD5CF896FD1F2A44" bold="true" box="[664,721,1884,1908]" pageId="9" pageNumber="121">
<taxonomicNameLabel id="A250ED06FFEED42AFD5CF896FD1F2A44" box="[664,721,1884,1908]" pageId="9" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Abbreviations:
<emphasis id="B963507DFFEED42AFCB7F891FC662A43" bold="true" box="[883,936,1883,1907]" pageId="9" pageNumber="121">AGF</emphasis>
ano-genital fold;
<emphasis id="B963507DFFEED42AFB9FF891FB402A43" bold="true" box="[1115,1166,1883,1907]" pageId="9" pageNumber="121">ANP</emphasis>
anal plates;
<emphasis id="B963507DFFEED42AFAC8F891FA8F2A43" bold="true" box="[1292,1345,1883,1907]" pageId="9" pageNumber="121">ANT</emphasis>
antenna;
<emphasis id="B963507DFFEED42AFF53F8B5FF722AA7" bold="true" box="[151,188,1919,1943]" pageId="9" pageNumber="121">DA</emphasis>
body derm appearance;
<emphasis id="B963507DFFEED42AFE77F84AFE212AA8" bold="true" box="[435,495,1920,1944]" pageId="9" pageNumber="121">DMD</emphasis>
dorsal microduct;
<emphasis id="B963507DFFEED42AFD68F8B5FD022AA7" bold="true" box="[684,716,1919,1943]" pageId="9" pageNumber="121">DS</emphasis>
dorsal setae;
<emphasis id="B963507DFFEED42AFC92F84AFCB72AA8" bold="true" box="[854,889,1920,1944]" pageId="9" pageNumber="121">DT</emphasis>
dorsal tubercles;
<emphasis id="B963507DFFEED42AFBEFF8B5FB9E2AA7" bold="true" box="[1067,1104,1919,1943]" pageId="9" pageNumber="121">LG</emphasis>
tibio-tarsal joint of hind leg;
<emphasis id="B963507DFFEED42AFAB2F8B5FA522AA7" bold="true" box="[1398,1436,1919,1943]" pageId="9" pageNumber="121">MS</emphasis>
marginal setae;
<emphasis id="B963507DFFEED42AFEFEF86EFEBC2A8C" bold="true" box="[314,370,1956,1980]" pageId="9" pageNumber="121">MDP</emphasis>
multilocular disc pores;
<emphasis id="B963507DFFEED42AFDB1F869FD682A8B" bold="true" box="[629,678,1955,1979]" pageId="9" pageNumber="121">POP</emphasis>
preopercular pore;
<emphasis id="B963507DFFEED42AFCB5F869FC512A8B" bold="true" box="[881,927,1955,1979]" pageId="9" pageNumber="121">SDP</emphasis>
spiracular disc pores;
<emphasis id="B963507DFFEED42AFB4EF869FB662A8B" bold="true" box="[1162,1192,1955,1979]" pageId="9" pageNumber="121">SP</emphasis>
spiracle;
<emphasis id="B963507DFFEED42AFAC9F869FAF92A8B" bold="true" box="[1293,1335,1955,1979]" pageId="9" pageNumber="121">SSP</emphasis>
stigmatic spines;
<emphasis id="B963507DFFEED42AFF20F80DFECA2AEF" bold="true" box="[228,260,1991,2015]" pageId="9" pageNumber="121">VS</emphasis>
ventral setae.
</paragraph>
</caption>
<caption id="DF68DCE7FFEDD429FF53FBC4FB65297A" ID-DOI="http://doi.org/10.5281/zenodo.5819788" ID-Zenodo-Dep="5819788" httpUri="https://zenodo.org/record/5819788/files/figure.png" pageId="10" pageNumber="122" startId="10.[151,250,1038,1062]" targetBox="[216,1372,201,1006]" targetPageId="10">
<paragraph id="8BA88C6FFFEDD429FF53FBC4FB65297A" blockId="10.[151,1437,1038,1099]" pageId="10" pageNumber="122">
<emphasis id="B963507DFFEDD429FF53FBC4FEDA2916" bold="true" box="[151,276,1038,1062]" pageId="10" pageNumber="122">FIGURE 5.</emphasis>
A photograph of the type slides of
<taxonomicName id="4C17F7ECFFEDD429FDBAFBC5FC142917" authority="Cao &amp; Feng" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[638,986,1038,1063]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="cephalotaxus" status="sp. nov.">
<emphasis id="B963507DFFEDD429FDBAFBC5FC982916" box="[638,854,1038,1062]" italics="true" pageId="10" pageNumber="122">Coccus cephalotaxus</emphasis>
Cao &amp; Feng
</taxonomicName>
<emphasis id="B963507DFFEDD429FC25FBC4FBD52916" bold="true" box="[993,1051,1038,1062]" pageId="10" pageNumber="122">
<taxonomicNameLabel id="A250ED06FFEDD429FC25FBC4FBD52916" box="[993,1051,1038,1062]" pageId="10" pageNumber="122" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(bottom) and
<taxonomicName id="4C17F7ECFFEDD429FB69FBC5FF04297B" authority="Cao &amp; Feng" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="nanningensis" status="sp. nov.">
<emphasis id="B963507DFFEDD429FB69FBC5FA9C2916" box="[1197,1362,1039,1062]" italics="true" pageId="10" pageNumber="122">C. nanningensis</emphasis>
Cao &amp; Feng
</taxonomicName>
<emphasis id="B963507DFFEDD429FF14FBF8FEC7297A" bold="true" box="[208,265,1074,1098]" pageId="10" pageNumber="122">
<taxonomicNameLabel id="A250ED06FFEDD429FF14FBF8FEC7297A" box="[208,265,1074,1098]" pageId="10" pageNumber="122" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(top) to facilitate future recognition. The holotype specimen on each slide is ringed in blue.
</paragraph>
</caption>
<subSubSection id="C30DDFE4FFEDD429FF03FBB2FB5E2AB7" pageId="10" pageNumber="122" type="discussion">
<paragraph id="8BA88C6FFFEDD429FF03FBB2FBF929EA" blockId="10.[151,1437,1144,1927]" pageId="10" pageNumber="122">
<emphasis id="B963507DFFEDD429FF03FBB2FE8129A2" bold="true" box="[199,335,1144,1170]" pageId="10" pageNumber="122">Comments.</emphasis>
The adult female of
<taxonomicName id="4C17F7ECFFEDD429FDFAFBB3FD3A29A2" authority="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng, 2022" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[574,756,1145,1171]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="nanningensis" status="sp. nov.">
<emphasis id="B963507DFFEDD429FDFAFBB3FD3A29A2" box="[574,756,1145,1171]" italics="true" pageId="10" pageNumber="122">C. nanningensis</emphasis>
</taxonomicName>
<emphasis id="B963507DFFEDD429FD38FBB3FCF329A3" bold="true" box="[764,829,1145,1171]" pageId="10" pageNumber="122">
<taxonomicNameLabel id="A250ED06FFEDD429FD38FBB3FCF329A3" box="[764,829,1145,1171]" pageId="10" pageNumber="122" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is morphologically similar to two African species,
<taxonomicName id="4C17F7ECFFEDD429FA47FBB3FE2E2986" authority="Hodgson, 1967" authorityName="Hodgson" authorityYear="1967" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="inyangombae">
<emphasis id="B963507DFFEDD429FA47FBB3FEE02986" italics="true" pageId="10" pageNumber="122">C. inyangombae</emphasis>
<bibRefCitation id="EF86F19EFFEDD429FEF1FB56FE2E2986" author="Hodgson, C. J." box="[309,480,1180,1206]" pageId="10" pageNumber="122" pagination="1 - 22" refId="ref12432" refString="Hodgson, C. J. (1967) Notes on Rhodesian Coccidae (Homoptera: Coccoidea): Part 1: The genera Coccus, Parasaissetia, Saissetia and a new genus Mashona. Arnoldia, Rhodesia, 3, 1 - 22." type="journal article" year="1967">Hodgson, 1967</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C17F7ECFFEDD429FDD2FB57FC842987" authority="De Lotto, 1957" authorityName="De Lotto" authorityYear="1957" box="[534,842,1180,1207]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="B963507DFFEDD429FDD2FB57FD5B2986" box="[534,661,1180,1207]" italics="true" pageId="10" pageNumber="122">C. sordidus</emphasis>
<bibRefCitation id="EF86F19EFFEDD429FD59FB57FC842987" author="De Lotto, G." box="[669,842,1180,1207]" pageId="10" pageNumber="122" pagination="295 - 314" refId="ref11782" refString="De Lotto, G. (1957) On some Ethiopian species of the genus Coccus (Homoptera: Coccoidea: Coccidae). Journal of the Entomological Society of southern Africa, 20 (2), 295 - 314." type="journal article" year="1957">De Lotto, 1957</bibRefCitation>
</taxonomicName>
, both of which also lack tubular ducts from both dorsum and venter, and have marginal setae mostly with branched or fimbriate apices.
</paragraph>
<paragraph id="8BA88C6FFFEDD429FF03FB2EFD142B2F" blockId="10.[151,1437,1144,1927]" pageId="10" pageNumber="122">
The new species can be separated from
<taxonomicName id="4C17F7ECFFEDD429FD4BFB2FFC8629CE" authorityName="Hodgson" authorityYear="1967" box="[655,840,1252,1279]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="inyangombae">
<emphasis id="B963507DFFEDD429FD4BFB2FFC8629CE" box="[655,840,1252,1279]" italics="true" pageId="10" pageNumber="122">C. inyangombae</emphasis>
</taxonomicName>
as follows (data for
<taxonomicName id="4C17F7ECFFEDD429FBF3FB2FFB3E29CE" authorityName="Hodgson" authorityYear="1967" box="[1079,1264,1252,1279]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="inyangombae">
<emphasis id="B963507DFFEDD429FBF3FB2FFB3E29CE" box="[1079,1264,1252,1279]" italics="true" pageId="10" pageNumber="122">C. inyangombae</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF86F19EFFEDD429FAF3FB2EFF2C2813" author="Hodgson, C. J." pageId="10" pageNumber="122" pagination="1 - 22" refId="ref12432" refString="Hodgson, C. J. (1967) Notes on Rhodesian Coccidae (Homoptera: Coccoidea): Part 1: The genera Coccus, Parasaissetia, Saissetia and a new genus Mashona. Arnoldia, Rhodesia, 3, 1 - 22." type="journal article" year="1967">Hodgson (1967)</bibRefCitation>
given in brackets): (i) dorsal setae setose, extremely fine and pointed, sparsely scattered (robust and pointed, quite numerous); (ii) anal plates each with a robust discal seta with pointed apex (without discal setae); (iii) anogenital fold with 6 anterior margin setae (with 4 anterior margin setae); (iv) marginal setae mainly rather long and highly variable in structure, some with apices branched or bifurcate; some conical, straight or curved with apices pointed; some with apices swollen, sagittate or spatulate; some setose, slender, fine and sharply pointed, but mostly without fimbriate apices (all rather long with fimbriate apices); (v) marginal setae between anterior and posterior stigmatic clefts numbering 10‒12 on each side (5‒7 on each side); (vi) antenna with 7 segments (with 8 segments); and (vii) pregenital setae mainly numbering 2 pairs (3 pairs).
</paragraph>
<paragraph id="8BA88C6FFFEDD429FF03F9E3FB5E2AB7" blockId="10.[151,1437,1144,1927]" pageId="10" pageNumber="122">
<taxonomicName id="4C17F7ECFFEDD429FF03F9E3FE7B2B72" authority="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng, 2022" authorityName="Cao &amp; Watson &amp; Hodgson &amp; Jing &amp; Feng" authorityYear="2022" box="[199,437,1577,1603]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="nanningensis" status="sp. nov.">
<emphasis id="B963507DFFEDD429FF03F9E3FE7B2B72" box="[199,437,1577,1603]" italics="true" pageId="10" pageNumber="122">Coccus nanningensis</emphasis>
</taxonomicName>
<emphasis id="B963507DFFEDD429FE79F9E3FE332B73" bold="true" box="[445,509,1577,1603]" pageId="10" pageNumber="122">
<taxonomicNameLabel id="A250ED06FFEDD429FE79F9E3FE332B73" box="[445,509,1577,1603]" pageId="10" pageNumber="122" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be separated from
<taxonomicName id="4C17F7ECFFEDD429FCC1F9E3FC4B2B72" authorityName="De Lotto" authorityYear="1957" box="[773,901,1576,1603]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="B963507DFFEDD429FCC1F9E3FC4B2B72" box="[773,901,1576,1603]" italics="true" pageId="10" pageNumber="122">C. sordidus</emphasis>
</taxonomicName>
as follows (data for
<taxonomicName id="4C17F7ECFFEDD429FBB4F9E3FB3F2B72" authorityName="De Lotto" authorityYear="1957" box="[1136,1265,1576,1603]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="122" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="B963507DFFEDD429FBB4F9E3FB3F2B72" box="[1136,1265,1576,1603]" italics="true" pageId="10" pageNumber="122">C. sordidus</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF86F19EFFEDD429FAF3F9E3FF2C2B57" author="De Lotto, G." pageId="10" pageNumber="122" pagination="295 - 314" refId="ref11782" refString="De Lotto, G. (1957) On some Ethiopian species of the genus Coccus (Homoptera: Coccoidea: Coccidae). Journal of the Entomological Society of southern Africa, 20 (2), 295 - 314." type="journal article" year="1957">De Lotto (1957)</bibRefCitation>
given in brackets): (i) body oval to subcircular, 1.5–2.0 mm long, 1.0–
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wide (elongate and asymmetric, up to
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long and up to
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wide); (ii) anal plates each with a robust discal seta with a pointed apex (without discal setae); (iii) anogenital fold with 6 anterior margin setae (with 4 anterior margin setae); (iv) marginal setae mainly rather long and highly variable in structure, some with apices branched or bifurcate; some conical, straight or curved with apices pointed; some with apices swollen, sagittate or spatulate; some setose, slender, fine and sharply pointed, but mostly without fimbriate apices (all rather long with slightly fimbriate apices); (v) antenna with 7 segments (with 8 segments); (vi) pregenital setae mostly numbering 2 pairs (3 pairs); (vii) leg without a tibiotarsal articulatory sclerosis (with a tibio-tarsal articulatory sclerosis); and (viii) multilocular disc pores restricted to genital area only (resent around genital area and on preceding 2 or 3 abdominal segments).
</paragraph>
</subSubSection>
</treatment>
</document>
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