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<document id="D3B33DC394734D8B1274566B1F0DC132" ID-DOI="10.5281/zenodo.276689" ID-GBIF-Dataset="b6781430-d288-4cb4-b178-db6192a61dde" ID-ISSN="1175-5326" ID-Zenodo-Dep="276689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460230557710" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2011" docId="03BC87EAFFB0317DFF22FD800543F80C" docLanguage="en" docName="zt02753p041.pdf" docOrigin="Zootaxa 2753" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hadrobolus Cook 1897" docType="treatment" docVersion="9" lastPageNumber="30" masterDocId="FF85FF92FFAD3160FFB5FFF00761FFF9" masterDocTitle="East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="30" updateTime="1698238933756" updateUser="plazi">
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<mods:title id="46808D580AC8FCB575F3E6B07841011D">East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="D4462D830AFA5B05101669869D4ACC34">Enghoff, Henrik</mods:namePart>
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<mods:date id="D598BF32EC728AD90EAA578D8464B3BF">2011</mods:date>
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<mods:identifier id="A564B308F5FDB0C1EDDB9AFA3FEDDEE2" type="DOI">10.5281/zenodo.276689</mods:identifier>
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<treatment id="03BC87EAFFB0317DFF22FD800543F80C" ID-DOI="http://doi.org/10.5281/zenodo.5612670" ID-GBIF-Taxon="119387219" ID-Zenodo-Dep="5612670" LSID="urn:lsid:plazi:treatment:03BC87EAFFB0317DFF22FD800543F80C" httpUri="http://treatment.plazi.org/id/03BC87EAFFB0317DFF22FD800543F80C" lastPageNumber="30" pageId="29" pageNumber="30">
<subSubSection id="C30F6577FFB0317DFF22FD800570FD73" box="[151,529,624,651]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BAA36FCFFB0317DFF22FD800570FD73" blockId="29.[151,529,624,651]" box="[151,529,624,651]" pageId="29" pageNumber="30">
<heading id="D0E28190FFB0317DFF22FD800570FD73" bold="true" box="[151,529,624,651]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B961EAEEFFB0317DFF22FD800570FD73" bold="true" box="[151,529,624,651]" pageId="29" pageNumber="30">
Genus
<taxonomicName id="4C154D7FFFB0317DFF5BFD810570FD73" ID-CoL="7PHY3" authority="Cook, 1897" authorityName="Cook" authorityYear="1897" box="[238,529,624,651]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAEEFFB0317DFF5BFD81061FFD72" bold="true" box="[238,382,625,651]" italics="true" pageId="29" pageNumber="30">Hadrobolus</emphasis>
<bibRefCitation id="EF844B0DFFB0317DFE31FD800570FD73" author="Cook" box="[388,529,624,650]" pageId="29" pageNumber="40" refString="Cook, O. F. (1897) New relatives of Spirobolus giganteus. Brandtia, 18, 73 - 75." type="journal article" year="1897">Cook, 1897</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFB0317DFF22FD430476FCFE" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BAA36FCFFB0317DFF22FD4305EEFD31" blockId="29.[151,655,690,712]" box="[151,655,690,712]" pageId="29" pageNumber="30">
<typeStatus id="54AE885EFFB0317DFF22FD4307A9FD31" box="[151,200,691,712]" pageId="29" pageNumber="30">Type</typeStatus>
species:
<taxonomicName id="4C154D7FFFB0317DFE93FD4205EEFD31" authority="Peters, 1855" authorityName="Peters" authorityYear="1855" box="[294,655,690,712]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicollis">
<emphasis id="B961EAEEFFB0317DFE93FD42056DFD31" box="[294,524,690,712]" italics="true" pageId="29" pageNumber="30">Spirobolus crassicollis</emphasis>
<bibRefCitation id="EF844B0DFFB0317DFDA6FD4305EEFD31" author="Peters" box="[531,655,690,712]" lastPageId="29" lastPageNumber="41" pageId="29" pageNumber="40" refString="Peters, W. (1855) Uber die Myriapoden im Allgemeinen und insbesondere uber die in Mossambique beobachteten Arten diser Familie. Monatsbericht der Koniglichen Akademia der Wissenschaften in Berlin, 75, 75 - 85." type="journal article" year="1855">Peters, 1855</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FD000476FCFE" blockId="29.[151,882,751,810]" box="[151,791,751,776]" pageId="29" pageNumber="30">
<emphasis id="B961EAEEFFB0317DFF22FD000600FCF1" bold="true" box="[151,353,751,776]" pageId="29" pageNumber="30">Included species.</emphasis>
<taxonomicName id="4C154D7FFFB0317DFEDDFD000476FCFE" authority="Peters, 1855" authorityName="Peters" authorityYear="1855" box="[360,791,751,776]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicollis">
<emphasis id="B961EAEEFFB0317DFEDDFD000512FCF1" box="[360,627,752,776]" italics="true" pageId="29" pageNumber="30">Hadrobolus crassicollis</emphasis>
(
<bibRefCitation id="EF844B0DFFB0317DFD37FD1F046EFCFE" author="Peters" box="[642,783,751,775]" lastPageId="29" lastPageNumber="41" pageId="29" pageNumber="40" refString="Peters, W. (1855) Uber die Myriapoden im Allgemeinen und insbesondere uber die in Mossambique beobachteten Arten diser Familie. Monatsbericht der Koniglichen Akademia der Wissenschaften in Berlin, 75, 75 - 85." type="journal article" year="1855">Peters, 1855</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFB0317DFF22FCE10543F80C" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8BAA36FCFFB0317DFF22FCE10413FCD0" blockId="29.[151,882,751,810]" box="[151,882,785,810]" pageId="29" pageNumber="30">
<emphasis id="B961EAEEFFB0317DFF22FCE10670FCD3" bold="true" box="[151,273,785,810]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
A genus of East African Pachybolini characterized by
</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FCBF0537FC9F" blockId="29.[151,1290,846,1113]" box="[151,598,846,870]" pageId="29" pageNumber="30">- no coxal lobes on male legs 6 and 7</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FC820455FBED" blockId="29.[151,1290,846,1113]" pageId="29" pageNumber="30">- one pair of ventral macrosetae on male legs from pair 5 onward - a distinct mesal process ca. at midlength of posterior gonopod - no striated membranous lamellae at tip of posterior gonopod - no mesal process, no free solenomerite, no distal complex of parallel ridges on posterior gonopod - no long meso-distal process on anterior gonopod coxite</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FBEF047FFBCF" blockId="29.[151,1290,846,1113]" box="[151,798,1054,1078]" pageId="29" pageNumber="30">- no overlapping lobes on anterior gonopod telopodites</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FBB106BCFBA0" blockId="29.[151,1290,846,1113]" box="[151,477,1089,1113]" pageId="29" pageNumber="30">(female characters unknown)</paragraph>
<paragraph id="8BAA36FCFFB0317DFF72FB720663FB19" blockId="29.[151,1436,1154,1248]" pageId="29" pageNumber="30">
<bibRefCitation id="EF844B0DFFB0317DFF72FB720634FB63" author="Cook" box="[199,341,1154,1178]" pageId="29" pageNumber="40" refString="Cook, O. F. (1899) African Diplopoda of the genus Pachybolus. Proceedings of the United States National Museum, 21, 657 - 666." type="journal article" year="1899">Cook (1899)</bibRefCitation>
listed a number of characters separating
<taxonomicName id="4C154D7FFFB0317DFCABFB7304C2FB62" box="[798,931,1155,1179]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAEEFFB0317DFCABFB7304C2FB62" box="[798,931,1155,1179]" italics="true" pageId="29" pageNumber="30">Hadrobolus</emphasis>
</taxonomicName>
from his “
<emphasis id="B961EAEEFFB0317DFBAFFB7303CDFB62" box="[1050,1196,1155,1179]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="4C154D7FFFB0317DFBAFFB7303C4FB62" box="[1050,1189,1155,1179]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Pachybolus</taxonomicName>
”
</emphasis>
(which also included species now classified in
<taxonomicName id="4C154D7FFFB0317DFE0EFB55052FFB44" box="[443,590,1189,1213]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAEEFFB0317DFE0EFB55052FFB44" box="[443,590,1189,1213]" italics="true" pageId="29" pageNumber="30">Pelmatojulus</emphasis>
</taxonomicName>
). In addition to the characteristic mesal process of the posterior gonopod, these are:
</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FAFA03D9FAD8" blockId="29.[151,1437,1289,1660]" box="[151,1208,1289,1314]" pageId="29" pageNumber="30">- sternum of anterior gonopods very broad and short, being scarcely produced in the middle</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FADD06B1FA9F" blockId="29.[151,1437,1289,1660]" pageId="29" pageNumber="30">
- telopodites of anterior gonopods widely separated (“in
<taxonomicName id="4C154D7FFFB0317DFC80FADC04D9FABD" box="[821,952,1324,1348]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAEEFFB0317DFC80FADC04D9FABD" box="[821,952,1324,1348]" italics="true" pageId="29" pageNumber="30">Pachybolus</emphasis>
</taxonomicName>
they are crowded mesad by the flagella [= posterior gonopods]...”)
</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FA820647FA55" blockId="29.[151,1437,1289,1660]" pageId="29" pageNumber="30">
- the last podomere (tarsus) of male legs greatly elongated in comparison with the penultimate podomere (tibia) (
<figureCitation id="132E2A79FFB0317DFF78FA64067EFA55" box="[205,287,1428,1452]" captionStart="FIGURES 58 – 72" captionStartId="30.[151,264,1818,1841]" captionTargetBox="[234,1355,193,1817]" captionTargetId="figure@30.[232,1356,183,1817]" captionTargetPageId="30" captionText="FIGURES 58 – 72. Plate LII from Cook (1899), relabelled. 58 – 65. Pachybolus tectus. 58: head and first six body rings, lateral view, 59: telson: lateral view, 60: male leg, 61: gonopods, anterior view, 62: do: posterior view, 63: do: lateral view, 64: posterior gonopod, posterior-lateral view, 65: do., anterior-mesal view. 66 – 72. Hadrobolus crassicollis. 66: gonopods, anterior view, 67: do., posterior view, 68: do., lateral view, 69: posterior gonopods, anterior-lateral view, 70: do., posterior-mesal view, 71: male leg, 72: first six body rings." httpUri="https://zenodo.org/record/276710/files/figure.png" pageId="29" pageNumber="30">Fig. 71</figureCitation>
)
</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FA4706B2FA08" blockId="29.[151,1437,1289,1660]" pageId="29" pageNumber="30">- the ventral pad of the male tarsus oblong and extending the whole length of the podomere, the subterminal setae being farther apart</paragraph>
<paragraph id="8BAA36FCFFB0317DFF22FA0D0550F985" blockId="29.[151,1437,1289,1660]" pageId="29" pageNumber="30">
- the ventral pad “fleshy throughout, the sides being scarcely chitinized and the surface being convex or irregularly shrunken” as opposed to the condition in
<taxonomicName id="4C154D7FFFB0317DFD69F9EE048BF9CF" authority="Cook" authorityName="Cook" box="[732,1002,1566,1590]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAEEFFB0317DFD69F9EE043EF9CF" box="[732,863,1566,1590]" italics="true" pageId="29" pageNumber="30">Pachybolus</emphasis>
sensu Cook
</taxonomicName>
where “the sides of the pad are much more prominent, strongly chitinized, and dark coloured, while the surface is also of firm texture and always uniformly and strongly concave.
</paragraph>
<paragraph id="8BAA36FCFFB0317DFF72F9560543F80C" blockId="29.[151,1437,1701,2037]" pageId="29" pageNumber="30">
In the light of the variability seen between the genera recognised here, the two gonopodal characters listed by
<bibRefCitation id="EF844B0DFFB0317DFF22F9370644F926" author="Cook" box="[151,293,1735,1759]" pageId="29" pageNumber="40" refString="Cook, O. F. (1899) African Diplopoda of the genus Pachybolus. Proceedings of the United States National Museum, 21, 657 - 666." type="journal article" year="1899">Cook (1899)</bibRefCitation>
are insignificant. The tarsus of
<taxonomicName id="4C154D7FFFB0317DFD3CF93804F4F919" box="[649,917,1736,1760]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicollis">
<emphasis id="B961EAEEFFB0317DFD3CF93804F4F919" box="[649,917,1736,1760]" italics="true" pageId="29" pageNumber="30">Hadrobolus crassicollis</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A79FFB0317DFC16F937048DF919" box="[931,1004,1735,1760]" captionStart="FIGURES 4 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[157,1430,195,1890]" captionTargetId="figure@5.[151,1436,193,1898]" captionTargetPageId="5" captionText="FIGURES 4 – 8. Male legs, SEM. 4: Crurifarcimen vagans n. gen., n. sp., paratype, tarsus, lateral view. 5: C. vagans, paratype, tarsus, ventral view. 6: Epibolus pulchripes, tarsus, lateral view. 7: Hadrobolus crassicollis, holotype, ventral-lateral view. 8: C. vagans, paratype, surface structure of ventral pad. Scales 1 mm (5, 7), 0.1 mm (4, 6), 0.01 mm (8)." httpUri="https://zenodo.org/record/276692/files/figure.png" pageId="29" pageNumber="30">Figs 7</figureCitation>
,
<figureCitation id="132E2A79FFB0317DFC4DF9370374F926" box="[1016,1045,1735,1759]" captionStart="FIGURES 58 – 72" captionStartId="30.[151,264,1818,1841]" captionTargetBox="[234,1355,193,1817]" captionTargetId="figure@30.[232,1356,183,1817]" captionTargetPageId="30" captionText="FIGURES 58 – 72. Plate LII from Cook (1899), relabelled. 58 – 65. Pachybolus tectus. 58: head and first six body rings, lateral view, 59: telson: lateral view, 60: male leg, 61: gonopods, anterior view, 62: do: posterior view, 63: do: lateral view, 64: posterior gonopod, posterior-lateral view, 65: do., anterior-mesal view. 66 – 72. Hadrobolus crassicollis. 66: gonopods, anterior view, 67: do., posterior view, 68: do., lateral view, 69: posterior gonopods, anterior-lateral view, 70: do., posterior-mesal view, 71: male leg, 72: first six body rings." httpUri="https://zenodo.org/record/276710/files/figure.png" pageId="29" pageNumber="30">71</figureCitation>
) is indeed quite long (and the tibia ventrally quite short), but not strikingly different from what is seen in the other genera (
<figureCitation id="132E2A79FFB0317DFBC0F91A03BAF8FB" box="[1141,1243,1770,1794]" captionStart="FIGURES 4 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[157,1430,195,1890]" captionTargetId="figure@5.[151,1436,193,1898]" captionTargetPageId="5" captionText="FIGURES 4 – 8. Male legs, SEM. 4: Crurifarcimen vagans n. gen., n. sp., paratype, tarsus, lateral view. 5: C. vagans, paratype, tarsus, ventral view. 6: Epibolus pulchripes, tarsus, lateral view. 7: Hadrobolus crassicollis, holotype, ventral-lateral view. 8: C. vagans, paratype, surface structure of ventral pad. Scales 1 mm (5, 7), 0.1 mm (4, 6), 0.01 mm (8)." httpUri="https://zenodo.org/record/276692/files/figure.png" pageId="29" pageNumber="30">Figs 4–6</figureCitation>
,
<figureCitation id="132E2A79FFB0317DFB52F91A0266F8FB" box="[1255,1287,1770,1794]" captionStart="FIGURES 58 – 72" captionStartId="30.[151,264,1818,1841]" captionTargetBox="[234,1355,193,1817]" captionTargetId="figure@30.[232,1356,183,1817]" captionTargetPageId="30" captionText="FIGURES 58 – 72. Plate LII from Cook (1899), relabelled. 58 – 65. Pachybolus tectus. 58: head and first six body rings, lateral view, 59: telson: lateral view, 60: male leg, 61: gonopods, anterior view, 62: do: posterior view, 63: do: lateral view, 64: posterior gonopod, posterior-lateral view, 65: do., anterior-mesal view. 66 – 72. Hadrobolus crassicollis. 66: gonopods, anterior view, 67: do., posterior view, 68: do., lateral view, 69: posterior gonopods, anterior-lateral view, 70: do., posterior-mesal view, 71: male leg, 72: first six body rings." httpUri="https://zenodo.org/record/276710/files/figure.png" pageId="29" pageNumber="30">60</figureCitation>
). The ventral pad in
<taxonomicName id="4C154D7FFFB0317DFF52F8FE06EAF8DC" box="[231,395,1805,1829]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicollis">
<emphasis id="B961EAEEFFB0317DFF52F8FE06EAF8DC" box="[231,395,1805,1829]" italics="true" pageId="29" pageNumber="30">H. crassicollis</emphasis>
</taxonomicName>
is slightly, but not dramatically longer than that of
<emphasis id="B961EAEEFFB0317DFC54F8FD054AF8B1" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="4C154D7FFFB0317DFC54F8FD0386F8DC" authority="Gabolus" authorityName="Gabolus" box="[993,1255,1805,1829]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Pelmatojulus, Gabolus</taxonomicName>
,
<taxonomicName id="4C154D7FFFB0317DFB41F8FD06C0F8B1" authority="Hyperbolus, Parabolus" authorityName="Hyperbolus, Parabolus" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Crurifarcimen, Hyperbolus, Parabolus</taxonomicName>
,
<taxonomicName id="4C154D7FFFB0317DFE18F8C0054AF8B1" box="[429,555,1840,1864]" class="Diplopoda" family="Pachybolidae" genus="Madabolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Madabolus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C154D7FFFB0317DFDD1F8C005A9F8B1" box="[612,712,1840,1864]" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
<emphasis id="B961EAEEFFB0317DFDD1F8C005A9F8B1" box="[612,712,1840,1864]" italics="true" pageId="29" pageNumber="30">Epibolus</emphasis>
</taxonomicName>
. The drawing of the pads in
<taxonomicName id="4C154D7FFFB0317DFBA6F8C003BEF8B1" box="[1043,1247,1840,1864]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="tectus">
<emphasis id="B961EAEEFFB0317DFBA6F8C003BEF8B1" box="[1043,1247,1840,1864]" italics="true" pageId="29" pageNumber="30">Pachybolus tectus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF844B0DFFB0317DFAB9F8C002FDF8B1" author="Cook" box="[1292,1436,1840,1864]" pageId="29" pageNumber="40" refString="Cook, O. F. (1899) African Diplopoda of the genus Pachybolus. Proceedings of the United States National Museum, 21, 657 - 666." type="journal article" year="1899">Cook (1899)</bibRefCitation>
(
<figureCitation id="132E2A79FFB0317DFF2AF8A20790F893" box="[159,241,1874,1898]" captionStart="FIGURES 58 – 72" captionStartId="30.[151,264,1818,1841]" captionTargetBox="[234,1355,193,1817]" captionTargetId="figure@30.[232,1356,183,1817]" captionTargetPageId="30" captionText="FIGURES 58 – 72. Plate LII from Cook (1899), relabelled. 58 – 65. Pachybolus tectus. 58: head and first six body rings, lateral view, 59: telson: lateral view, 60: male leg, 61: gonopods, anterior view, 62: do: posterior view, 63: do: lateral view, 64: posterior gonopod, posterior-lateral view, 65: do., anterior-mesal view. 66 – 72. Hadrobolus crassicollis. 66: gonopods, anterior view, 67: do., posterior view, 68: do., lateral view, 69: posterior gonopods, anterior-lateral view, 70: do., posterior-mesal view, 71: male leg, 72: first six body rings." httpUri="https://zenodo.org/record/276710/files/figure.png" pageId="29" pageNumber="30">Fig. 60</figureCitation>
) shows a much shorter pad, so maybe it is
<taxonomicName id="4C154D7FFFB0317DFD6FF8A3043CF892" box="[730,861,1875,1899]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B961EAEEFFB0317DFD6FF8A3043CF892" box="[730,861,1875,1899]" italics="true" pageId="29" pageNumber="30">Pachybolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525795FFB0317DFCD1F8A304E8F893" box="[868,905,1875,1898]" pageId="29" pageNumber="30" sensu="stricto">s.s.</taxonomicNameLabel>
that differs markedly here. The ventral surface of the pad in
<taxonomicName id="4C154D7FFFB0317DFE9DF88606A8F874" box="[296,457,1909,1933]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicollis">
<emphasis id="B961EAEEFFB0317DFE9DF88606A8F874" box="[296,457,1909,1933]" italics="true" pageId="29" pageNumber="30">H. crassicollis</emphasis>
</taxonomicName>
is indeed particularly soft and convex, where in the other genera this surface is concave and delimited by a sharp rim, the difference is, however, not at all obvious in the scanning electron microscope (
<figureCitation id="132E2A79FFB0317DFF2AF84A0666F82B" box="[159,263,1978,2002]" captionStart="FIGURES 4 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[157,1430,195,1890]" captionTargetId="figure@5.[151,1436,193,1898]" captionTargetPageId="5" captionText="FIGURES 4 – 8. Male legs, SEM. 4: Crurifarcimen vagans n. gen., n. sp., paratype, tarsus, lateral view. 5: C. vagans, paratype, tarsus, ventral view. 6: Epibolus pulchripes, tarsus, lateral view. 7: Hadrobolus crassicollis, holotype, ventral-lateral view. 8: C. vagans, paratype, surface structure of ventral pad. Scales 1 mm (5, 7), 0.1 mm (4, 6), 0.01 mm (8)." httpUri="https://zenodo.org/record/276692/files/figure.png" pageId="29" pageNumber="30">Figs 4–8</figureCitation>
). The significance of these differences cannot be properly evaluated until fresh material of
<taxonomicName id="4C154D7FFFB0317DFAA2F84B02FDF82A" box="[1303,1436,1979,2003]" class="Diplopoda" family="Pachybolidae" genus="Hadrobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAEEFFB0317DFAA2F84B02FDF82A" box="[1303,1436,1979,2003]" italics="true" pageId="29" pageNumber="30">Hadrobolus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D7FFFB0317DFF73F82D0628F80C" box="[198,329,2013,2037]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B961EAEEFFB0317DFF73F82D0628F80C" box="[198,329,2013,2037]" italics="true" pageId="29" pageNumber="30">Pachybolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525795FFB0317DFEE5F82E0612F80C" box="[336,371,2014,2037]" pageId="29" pageNumber="30" sensu="stricto">s.s.</taxonomicNameLabel>
can be studied.
</paragraph>
</subSubSection>
</treatment>
</document>
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