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<document id="6D95593DC8B1A1BF40F7418C141B99EF" ID-CLB-Dataset="29763" ID-DOI="10.11646/zootaxa.4442.2.6" ID-GBIF-Dataset="6ab5cadd-8428-4e3f-a5c6-b909cee2f89f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303149" ID-ZooBank="C02CFBF9-F96B-45C5-9793-E9E4145FBC55" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530533632922" checkinUser="plazi" docAuthor="Volynkin, Anton V. &amp; László, Gyula M." docDate="2018" docId="03BC87C6FFB6FFE6E4B7F88AFB8F9ED8" docLanguage="en" docName="zootaxa.4442.2.6.pdf" docOrigin="Zootaxa 4442 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tumicla tsonga Volynkin &amp; László 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="304" masterDocId="FF85FFBEFFB1FFEDE420FFA2FFEF9900" masterDocTitle="On the taxonomy of the genus Tumicla Wallengren, 1863 with description of two new species from Mozambique (Lepidoptera: Erebidae: Arctiinae)" masterLastPageNumber="306" masterPageNumber="293" pageNumber="300" updateTime="1698552150249" updateUser="ExternalLinkService">
<mods:mods id="D0282F8707FA5832E5C5FEEC41647ADB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A2382F8B0331B6A99E1E6915EFDD2395">On the taxonomy of the genus Tumicla Wallengren, 1863 with description of two new species from Mozambique (Lepidoptera: Erebidae: Arctiinae)</mods:title>
</mods:titleInfo>
<mods:name id="0A5AB9EA5ABBDF1F12465A696DE3C637" type="personal">
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<mods:namePart id="2577AC069C415A84E38F8BE7FCF6C7DE">Volynkin, Anton V.</mods:namePart>
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<mods:namePart id="06FB1FD8FAA074FBA58770FC7ED8B21F">László, Gyula M.</mods:namePart>
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<mods:title id="5828C4DF2F10E1779735B8BBD7186158">Zootaxa</mods:title>
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<mods:date id="BD3C36DF075570BB08AB6870A3917F27">2018</mods:date>
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<mods:number id="622ABD958D1B62C9E9FAA9A092E48FF6">2018-07-02</mods:number>
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<mods:classification id="5ACE01BE1AC3920C5981029B2E8A8AA0">journal article</mods:classification>
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<mods:identifier id="3DCB3F7F4EC36762430288EBC5AC6B5B" type="DOI">10.11646/zootaxa.4442.2.6</mods:identifier>
<mods:identifier id="F8B8186EACF239B8F0C6FB773820BA04" type="GBIF-Dataset">6ab5cadd-8428-4e3f-a5c6-b909cee2f89f</mods:identifier>
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<treatment id="03BC87C6FFB6FFE6E4B7F88AFB8F9ED8" ID-DOI="http://doi.org/10.5281/zenodo.6485648" ID-GBIF-Taxon="146044090" ID-Zenodo-Dep="6485648" LSID="urn:lsid:plazi:treatment:03BC87C6FFB6FFE6E4B7F88AFB8F9ED8" httpUri="http://treatment.plazi.org/id/03BC87C6FFB6FFE6E4B7F88AFB8F9ED8" lastPageId="11" lastPageNumber="304" pageId="7" pageNumber="300">
<subSubSection id="C30F655BFFB6FFEAE4B7F88AFEB09E63" pageId="7" pageNumber="300" type="nomenclature">
<paragraph id="8BAA36D0FFB6FFEAE4B7F88AFD419E42" blockId="7.[151,686,1832,1891]" box="[151,686,1832,1858]" pageId="7" pageNumber="300">
<heading id="D0E281BCFFB6FFEAE4B7F88AFD419E42" bold="true" box="[151,686,1832,1858]" fontSize="11" level="1" pageId="7" pageNumber="300" reason="1">
<emphasis id="B961EAC2FFB6FFEAE4B7F88AFD419E42" bold="true" box="[151,686,1832,1858]" pageId="7" pageNumber="300">
<taxonomicName id="4C154D53FFB6FFEAE4B7F88AFDAB9E42" ID-CoL="BFXRK" authority="Volynkin &amp; Laszlo" authorityName="Volynkin &amp; László" authorityYear="2018" box="[151,580,1832,1858]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="300" phylum="Arthropoda" rank="species" species="tsonga" status="sp. nov.">
<emphasis id="B961EAC2FFB6FFEAE4B7F88AFEBE9E42" bold="true" box="[151,337,1832,1858]" italics="true" pageId="7" pageNumber="300">Tumicla tsonga</emphasis>
Volynkin &amp; László
</taxonomicName>
,
<taxonomicNameLabel id="A25257B9FFB6FFEAE671F88AFD419E42" box="[593,686,1832,1858]" pageId="7" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36D0FFB6FFEAE4B7F8E8FEB09E63" blockId="7.[151,686,1832,1891]" box="[151,351,1866,1891]" pageId="7" pageNumber="300">
(
<figureCitation id="132E2A55FFB6FFEAE4BFF8E8FEEB9E63" box="[159,260,1866,1891]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1885,1908]" captionTargetBox="[165,1435,196,1854]" captionTargetId="figure@5.[151,1436,195,1864]" captionTargetPageId="5" captionText="FIGURES 1–8. Tumicla spp.: adults. 1, T. arktos, holotype ♂, MoZambique (ANHRT); 2, T. arktos, paratype ♀, MoZambique (ANHRT); 3, T. tsonga, holotype ♂, MoZambique (ANHRT); 4, T. tsonga, paratype ♂, MoZambique (ANHRT); 5, T. tsonga, paratype ♀, MoZambique (ANHRT); 6, T. doa, holotype ♀, [TanZania] Kwedikundi (ZMB); 7, T. dimidiata, holotype ♀, Democratic Republic of the Congo (©NHMUK); 8, T. mutabilis, holotype ♀, [TanZania] D.O. Africa (ZMB)." httpUri="https://zenodo.org/record/1303151/files/figure.png" pageId="7" pageNumber="300">Figs 3–5</figureCitation>
,
<figureCitation id="132E2A55FFB6FFEAE52FF8E9FEC29E63" box="[271,301,1867,1891]" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="7" pageNumber="300">18</figureCitation>
,
<figureCitation id="132E2A55FFB6FFEAE519F8E8FEB99E63" box="[313,342,1866,1891]" captionStart="FIGURES 26–33" captionStartId="9.[151,264,1473,1496]" captionTargetBox="[154,1436,194,1438]" captionTargetId="figure@9.[151,1436,193,1452]" captionTargetPageId="9" captionText="FIGURES 26–33. Tumicla spp.: female genitalia. 26, T. tsonga, paratype, MoZambique, slide AV3036 (ANHRT); 27, T. arktos, paratype, MoZambique, slide AV3044 (ANHRT); 28, T. doa, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30, T. similis, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31, T. mutabilis, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32, T. dimidiata, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK)." httpUri="https://zenodo.org/record/1303160/files/figure.png" pageId="7" pageNumber="300">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F655BFFB6FFE5E4B7F82FFD129EE2" lastPageId="8" lastPageNumber="301" pageId="7" pageNumber="300" type="materials_examined">
<paragraph id="8BAA36D0FFB6FFEAE4B7F82FFCB19EEE" blockId="7.[151,1436,1933,2030]" pageId="7" pageNumber="300">
<materialsCitation id="3B7D3C8DFFB6FFEAE4B7F82FFCD69ECA" ID-GBIF-Occurrence="1852084083" collectingDate="2017-05-25" collectingDateMax="2017-05-30" collectingDateMin="2017-05-25" country="Mozambique" elevation="15" latitude="-26.44961" location="Dune Forest" longLatPrecision="1" longitude="32.924946" municipality="Ponta Milibangalala" pageId="7" pageNumber="300" specimenCount="1" specimenCount-male="1" stateProvince="Maputo" typeStatus="holotype">
<emphasis id="B961EAC2FFB6FFEAE4B7F82FFEAD9EA6" bold="true" box="[151,322,1933,1958]" pageId="7" pageNumber="300">
<typeStatus id="54AE8872FFB6FFEAE4B7F82FFF3D9EA6" box="[151,210,1933,1958]" pageId="7" pageNumber="300">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54AE8872FFB6FFEAE576F82FFE519EA6" box="[342,446,1933,1958]" pageId="7" pageNumber="300" type="holotype">Holotype</typeStatus>
(
<figureCitation id="132E2A55FFB6FFEAE5F2F82FFDF09EA6" box="[466,543,1933,1958]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1885,1908]" captionTargetBox="[165,1435,196,1854]" captionTargetId="figure@5.[151,1436,195,1864]" captionTargetPageId="5" captionText="FIGURES 1–8. Tumicla spp.: adults. 1, T. arktos, holotype ♂, MoZambique (ANHRT); 2, T. arktos, paratype ♀, MoZambique (ANHRT); 3, T. tsonga, holotype ♂, MoZambique (ANHRT); 4, T. tsonga, paratype ♂, MoZambique (ANHRT); 5, T. tsonga, paratype ♀, MoZambique (ANHRT); 6, T. doa, holotype ♀, [TanZania] Kwedikundi (ZMB); 7, T. dimidiata, holotype ♀, Democratic Republic of the Congo (©NHMUK); 8, T. mutabilis, holotype ♀, [TanZania] D.O. Africa (ZMB)." httpUri="https://zenodo.org/record/1303151/files/figure.png" pageId="7" pageNumber="300">Figs 3</figureCitation>
,
<figureCitation id="132E2A55FFB6FFEAE611F82CFDBF9EA6" box="[561,592,1934,1958]" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="7" pageNumber="300">18</figureCitation>
):
<specimenCount id="9D13FD59FFB6FFEAE64AF82CFD6E9EA5" box="[618,641,1934,1957]" pageId="7" pageNumber="300" type="male">♂</specimenCount>
,
<collectingCountry id="F3027640FFB6FFEAE6B3F82FFCC59EA6" box="[659,810,1933,1958]" name="Mozambique" pageId="7" pageNumber="300">Mozambique</collectingCountry>
,
<elevation id="0038D1E3FFB6FFEAE719F82FFC819EA6" box="[825,878,1933,1958]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="300" unit="m" value="15.0">
<quantity id="4CED9B35FFB6FFEAE719F82FFC819EA6" box="[825,878,1933,1958]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="300" unit="m" value="15.0">15m</quantity>
</elevation>
,
<collectingRegion id="49D1F832FFB6FFEAE75EF82CFC3A9EA5" box="[894,981,1934,1957]" country="Mozambique" name="Maputo" pageId="7" pageNumber="300">Maputo</collectingRegion>
Special Reserve,
<collectingMunicipality id="6BCEACAAFFB6FFEAE08EF82CFA779EA6" box="[1198,1432,1933,1958]" pageId="7" pageNumber="300">Ponta Milibangalala</collectingMunicipality>
, (
<location id="8ECA600BFFB6FFEAE4BEF813FEDB9EC9" LSID="urn:lsid:plazi:treatment:03BC87C6FFB6FFE6E4B7F88AFB8F9ED8:8ECA600BFFB6FFEAE4BEF813FEDB9EC9" box="[158,308,1969,1993]" country="Mozambique" latitude="-26.44961" longLatPrecision="1" longitude="32.924946" municipality="Ponta Milibangalala" name="Dune Forest" pageId="7" pageNumber="300" stateProvince="Maputo">Dune Forest</location>
),
<geoCoordinate id="EE215017FFB6FFEAE56BF813FE0F9ECA" box="[331,480,1969,1994]" direction="south" orientation="latitude" pageId="7" pageNumber="300" precision="1" value="-26.44961">26°26'58.6''S</geoCoordinate>
,
<geoCoordinate id="EE215017FFB6FFEAE5D1F813FD679ECA" box="[497,648,1969,1994]" direction="east" orientation="longitude" pageId="7" pageNumber="300" precision="1" value="32.924946">32°55'29.8''E</geoCoordinate>
,
<collectingDate id="EFEFE9F8FFB6FFEAE6B8F813FCD69ECA" box="[664,825,1969,1994]" pageId="7" pageNumber="300" value="2017-05-25" valueMax="2017-05-30" valueMin="2017-05-25">
<date id="FFAB1010FFB6FFEAE6B8F813FCD69ECA" box="[664,825,1969,1994]" pageId="7" pageNumber="300" value="2017-05-25" valueMax="2017-05-30" valueMin="2017-05-25">25–30.V.2017</date>
</collectingDate>
</materialsCitation>
.
<collectingMethod id="52544EC7FFB6FFEAE768F813FBC79EC9" box="[840,1064,1969,1994]" pageId="7" pageNumber="300">Actinic Light Trap</collectingMethod>
. Aristophanous, M., Laszlo, G., Miles, W., Vetina, A. leg., slide No.: AV3035♂ (coll. ANHRT).
</paragraph>
<caption id="DF6A6658FFB9FFE5E4B7F9D4FAC89FCA" httpUri="https://zenodo.org/record/1303158/files/figure.png" pageId="8" pageNumber="301" startId="8.[151,264,1654,1677]" targetBox="[187,1426,194,1623]" targetPageId="8">
<paragraph id="8BAA36D0FFB9FFE5E4B7F9D4FAC89FCA" blockId="8.[151,1436,1654,1738]" pageId="8" pageNumber="301">
<emphasis id="B961EAC2FFB9FFE5E4B7F9D4FEA19F8C" bold="true" box="[151,334,1654,1677]" pageId="8" pageNumber="301">FIGURES 22–25</emphasis>
.
<taxonomicName id="4C154D53FFB9FFE5E57CF9D4FE419F8C" authorityName="Wallengren" authorityYear="1863" box="[348,430,1654,1676]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC2FFB9FFE5E57CF9D4FE419F8C" box="[348,430,1654,1676]" italics="true" pageId="8" pageNumber="301">Tumicla</emphasis>
</taxonomicName>
spp.: male genitalia. 22,
<taxonomicName id="4C154D53FFB9FFE5E690F9DAFCC79F8C" authorityName="Kuhne" authorityYear="2007" box="[688,808,1654,1677]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B961EAC2FFB9FFE5E690F9DAFCC79F8C" box="[688,808,1654,1677]" italics="true" pageId="8" pageNumber="301">T. mutabilis</emphasis>
</taxonomicName>
, paratype, Uganda, slide BMNH(E) Arct-6052 (©NHMUK); 23,
<taxonomicName id="4C154D53FFB9FFE5E49DF934FEDF9FAB" authorityName="Kuhne" authorityYear="2007" box="[189,304,1686,1707]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="B961EAC2FFB9FFE5E49DF934FEDF9FAB" box="[189,304,1686,1707]" italics="true" pageId="8" pageNumber="301">T. gigantea</emphasis>
</taxonomicName>
, paratype, Urundi, slide AV2897 (ZMB); 24,
<taxonomicName id="4C154D53FFB9FFE5E724F934FC6D9FAB" authorityName="Wallengren" authorityYear="1860" box="[772,898,1686,1707]" class="Insecta" family="Erebidae" genus="Setina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="B961EAC2FFB9FFE5E724F934FC6D9FAB" box="[772,898,1686,1707]" italics="true" pageId="8" pageNumber="301">T. sagenaria</emphasis>
</taxonomicName>
, paralectotype, [South Africa] Caffraria slide NHRS 11085 (photo by T. Malm, ©2017NHRS); 25,
<taxonomicName id="4C154D53FFB9FFE5E64CF917FD059FCA" authorityName="Wallengren" authorityYear="1860" box="[620,746,1717,1738]" class="Insecta" family="Erebidae" genus="Setina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="B961EAC2FFB9FFE5E64CF917FD059FCA" box="[620,746,1717,1738]" italics="true" pageId="8" pageNumber="301">T. sagenaria</emphasis>
</taxonomicName>
, South Africa, slide BMNH(E) Arct-5861 (©NHMUK).
</paragraph>
</caption>
<paragraph id="8BAA36D0FFB9FFE5E4E7F953FD129EE2" blockId="8.[151,1437,1777,2018]" pageId="8" pageNumber="301">
<materialsCitation id="3B7D3C8DFFB9FFE5E4E7F953FBE29E09" ID-GBIF-Occurrence="1852084077" box="[199,1037,1777,1802]" country="Mozambique" location="Maputo" pageId="8" pageNumber="301" specimenCount="13" specimenCount-female="11" specimenCount-male="2" stateProvince="Maputo" typeStatus="paratype">
<typeStatus id="54AE8872FFB9FFE5E4E7F953FED09E0A" box="[199,319,1777,1802]" pageId="8" pageNumber="301" type="paratype">
<emphasis id="B961EAC2FFB9FFE5E4E7F953FED09E0A" bold="true" box="[199,319,1777,1802]" pageId="8" pageNumber="301">Paratypes</emphasis>
</typeStatus>
<specimenCount id="9D13FD59FFB9FFE5E566F950FE9C9E09" box="[326,371,1778,1802]" pageId="8" pageNumber="301" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D13FD59FFB9FFE5E55FF950FE589E09" box="[383,439,1778,1802]" pageId="8" pageNumber="301" type="female">11 ♀</specimenCount>
in total from
<collectingCountry id="F3027640FFB9FFE5E673F953FD059E0A" box="[595,746,1777,1802]" name="Mozambique" pageId="8" pageNumber="301">Mozambique</collectingCountry>
,
<collectingRegion id="49D1F832FFB9FFE5E6D5F950FCA39E09" box="[757,844,1778,1801]" country="Mozambique" name="Maputo" pageId="8" pageNumber="301">Maputo</collectingRegion>
Special Reserve
</materialsCitation>
:
<materialsCitation id="3B7D3C8DFFB9FFE5E037F950FDC39E52" ID-GBIF-Occurrence="1852084076" collectingDate="2017-06-09" collectingDateMax="2017-06-12" collectingDateMin="2017-06-09" collectorName="Aristophanous &amp; Laszlo &amp; Miles &amp; Vetina, A. &amp; No." country="Mozambique" elevation="11" latitude="-26.289999" location="Sand Forest" longLatPrecision="20" longitude="32.7625" pageId="8" pageNumber="301" specimenCount="2" specimenCount-female="2" stateProvince="Maputo" typeStatus="paratype">
<specimenCount id="9D13FD59FFB9FFE5E037F950FBAD9E09" box="[1047,1090,1778,1802]" pageId="8" pageNumber="301" type="female">1 ♀</specimenCount>
,
<elevation id="0038D1E3FFB9FFE5E070F950FB6C9E0A" box="[1104,1155,1778,1802]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="301" unit="m" value="11.0">
<quantity id="4CED9B35FFB9FFE5E070F950FB6C9E0A" box="[1104,1155,1778,1802]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="301" unit="m" value="11.0">11m</quantity>
</elevation>
, forest clearing campsite (
<location id="8ECA600BFFB9FFE5E4BEF8B7FEC49E2E" LSID="urn:lsid:plazi:treatment:03BC87C6FFB6FFE6E4B7F88AFB8F9ED8:8ECA600BFFB9FFE5E4BEF8B7FEC49E2E" box="[158,299,1813,1838]" country="Mozambique" latitude="-26.289999" longLatPrecision="20" longitude="32.7625" name="Sand Forest" pageId="8" pageNumber="301" stateProvince="Maputo">Sand Forest</location>
),
<geoCoordinate id="EE215017FFB9FFE5E51FF8B7FE509E2E" box="[319,447,1813,1838]" direction="south" orientation="latitude" pageId="8" pageNumber="301" precision="15" value="-26.289999">26°17'24''S</geoCoordinate>
,
<geoCoordinate id="EE215017FFB9FFE5E5ECF8B7FDA09E2E" box="[460,591,1813,1838]" direction="east" orientation="longitude" pageId="8" pageNumber="301" precision="15" value="32.7625">32°45'45''E</geoCoordinate>
,
<collectingDate id="EFEFE9F8FFB9FFE5E67BF8B4FD169E2E" box="[603,761,1814,1838]" pageId="8" pageNumber="301" value="2017-06-09" valueMax="2017-06-12" valueMin="2017-06-09">
<date id="FFAB1010FFB9FFE5E67BF8B4FD169E2E" box="[603,761,1814,1838]" pageId="8" pageNumber="301" value="2017-06-09" valueMax="2017-06-12" valueMin="2017-06-09">9–12.VI.2017</date>
</collectingDate>
,
<collectingMethod id="52544EC7FFB9FFE5E726F8B4FC569E2D" box="[774,953,1813,1838]" pageId="8" pageNumber="301">MV Light Trap</collectingMethod>
.
<collectorName id="26E05306FFB9FFE5E7E4F8B7FB859E2E" box="[964,1130,1813,1838]" pageId="8" pageNumber="301">Aristophanous</collectorName>
, M.,
<collectorName id="26E05306FFB9FFE5E086F8B7FB1C9E2E" box="[1190,1267,1813,1838]" pageId="8" pageNumber="301">Laszlo</collectorName>
, G.,
<collectorName id="26E05306FFB9FFE5E107F8B7FA859E2E" box="[1319,1386,1813,1838]" pageId="8" pageNumber="301">Miles</collectorName>
, W.,
<collectorName id="26E05306FFB9FFE5E4B7F89BFEE89E52" box="[151,263,1849,1874]" pageId="8" pageNumber="301">Vetina, A.</collectorName>
leg., slide
<collectorName id="26E05306FFB9FFE5E5A0F898FE429E51" box="[384,429,1850,1873]" pageId="8" pageNumber="301">No.</collectorName>
: AV3036
<specimenCount id="9D13FD59FFB9FFE5E63FF89BFDC39E52" box="[543,556,1849,1874]" pageId="8" pageNumber="301" type="female">♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B7D3C8DFFB9FFE5E617F898FB419E51" ID-GBIF-Occurrence="1852084157" box="[567,1198,1849,1874]" collectingDate="2017-06-09" collectingDateMax="2017-06-12" collectingDateMin="2017-06-09" collectorName="Aristophanous &amp; Laszlo &amp; Miles &amp; Vetina, A. &amp; No." country="Mozambique" elevation="11" latitude="-26.289999" location="Sand Forest" longLatPrecision="20" longitude="32.7625" pageId="8" pageNumber="301" specimenCount="1" specimenCount-female="1" stateProvince="Maputo" typeStatus="paratype">
<specimenCount id="9D13FD59FFB9FFE5E617F898FD8E9E51" box="[567,609,1850,1874]" pageId="8" pageNumber="301" type="female">1 ♀</specimenCount>
, same data,
<collectingMethod id="52544EC7FFB9FFE5E6CBF89BFB419E51" box="[747,1198,1849,1874]" pageId="8" pageNumber="301">but collected with an Actinic Light Trap</collectingMethod>
</materialsCitation>
;
<materialsCitation id="3B7D3C8DFFB9FFE5E098F898FCA29E9A" ID-GBIF-Occurrence="1852084159" collectingDate="2016-11-21" collectingDateMax="2016-11-30" collectingDateMin="2016-11-21" collectorName="Aristophanous &amp; Cristovao &amp; Laszlo, G. &amp; Miles, W. &amp; Nos" country="Mozambique" elevation="22" latitude="-26.503944" location="Sand Thicket" longLatPrecision="1" longitude="32.716557" pageId="8" pageNumber="301" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="West" typeStatus="paratype">
<specimenCount id="9D13FD59FFB9FFE5E098F898FB0C9E51" box="[1208,1251,1850,1874]" pageId="8" pageNumber="301" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D13FD59FFB9FFE5E0CFF898FAF79E51" box="[1263,1304,1850,1874]" pageId="8" pageNumber="301" type="female">1 ♀</specimenCount>
,
<elevation id="0038D1E3FFB9FFE5E106F898FAB59E52" box="[1318,1370,1850,1874]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="301" unit="m" value="22.0">
<quantity id="4CED9B35FFB9FFE5E106F898FAB59E52" box="[1318,1370,1850,1874]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="301" unit="m" value="22.0">22m</quantity>
</elevation>
,
<collectingRegion id="49D1F832FFB9FFE5E145F898FA739E51" box="[1381,1436,1850,1873]" country="Tanzania" name="Mjini Magharibi" pageId="8" pageNumber="301">West</collectingRegion>
Gate, (
<location id="8ECA600BFFB9FFE5E4C4F8FFFE6E9E76" LSID="urn:lsid:plazi:treatment:03BC87C6FFB6FFE6E4B7F88AFB8F9ED8:8ECA600BFFB9FFE5E4C4F8FFFE6E9E76" box="[228,385,1885,1910]" country="Mozambique" latitude="-26.503944" longLatPrecision="1" longitude="32.716557" name="Sand Thicket" pageId="8" pageNumber="301" stateProvince="West">Sand Thicket</location>
),
<geoCoordinate id="EE215017FFB9FFE5E5B6F8FFFDC39E76" box="[406,556,1885,1910]" direction="south" orientation="latitude" pageId="8" pageNumber="301" precision="1" value="-26.503944">26°30'14.2''S</geoCoordinate>
,
<geoCoordinate id="EE215017FFB9FFE5E619F8FFFD3F9E76" box="[569,720,1885,1910]" direction="east" orientation="longitude" pageId="8" pageNumber="301" precision="1" value="32.716557">32°42'59.6''E</geoCoordinate>
,
<collectingDate id="EFEFE9F8FFB9FFE5E6FEF8FFFC649E76" box="[734,907,1885,1910]" pageId="8" pageNumber="301" value="2016-11-21" valueMax="2016-11-30" valueMin="2016-11-21">
<date id="FFAB1010FFB9FFE5E6FEF8FFFC649E76" box="[734,907,1885,1910]" pageId="8" pageNumber="301" value="2016-11-21" valueMax="2016-11-30" valueMin="2016-11-21">21–30.XI.2016</date>
</collectingDate>
.
<collectingMethod id="52544EC7FFB9FFE5E7B8F8FFFBF99E75" box="[920,1046,1885,1910]" pageId="8" pageNumber="301">Light Trap</collectingMethod>
.
<collectorName id="26E05306FFB9FFE5E002F8FFFB279E76" box="[1058,1224,1885,1910]" pageId="8" pageNumber="301">Aristophanous</collectorName>
, M.,
<collectorName id="26E05306FFB9FFE5E126F8FFFA999E76" box="[1286,1398,1885,1910]" pageId="8" pageNumber="301">Cristóvão</collectorName>
, J.,
<collectorName id="26E05306FFB9FFE5E4B7F823FEE19E9A" box="[151,270,1921,1946]" pageId="8" pageNumber="301">Laszlo, G.</collectorName>
,
<collectorName id="26E05306FFB9FFE5E53DF823FE609E99" box="[285,399,1921,1946]" pageId="8" pageNumber="301">Miles, W.</collectorName>
leg., slide
<collectorName id="26E05306FFB9FFE5E635F820FDA99E99" box="[533,582,1922,1945]" pageId="8" pageNumber="301">Nos</collectorName>
: AV3265
<specimenCount id="9D13FD59FFB9FFE5E697F823FD259E9A" box="[695,714,1921,1946]" pageId="8" pageNumber="301" type="male">♂</specimenCount>
, AV2979
<specimenCount id="9D13FD59FFB9FFE5E760F820FCA29E9A" box="[832,845,1922,1946]" pageId="8" pageNumber="301" type="female">♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B7D3C8DFFB9FFE5E77DF820FD169EE2" ID-GBIF-Occurrence="1852084081" collectingDate="2016-12-07" collectingDateMax="2016-12-09" collectingDateMin="2016-12-07" collectionCode="ANHRT, NHMM" collectorName="Aristophanous, M. &amp; Cristovao, J. &amp; Laszlo, G. &amp; Miles, W. &amp; No." country="Mozambique" elevation="9" latitude="-26.32664" location="Mangrove - Woodland Mosaic" longLatPrecision="1" longitude="32.70992" municipality="Mangrove Camp" pageId="8" pageNumber="301" specimenCount="9" specimenCount-female="9" stateProvince="West" typeStatus="paratype">
<specimenCount id="9D13FD59FFB9FFE5E77DF820FC649E99" box="[861,907,1922,1946]" pageId="8" pageNumber="301" type="female">8 ♀</specimenCount>
,
<elevation id="0038D1E3FFB9FFE5E7BDF820FC2B9E9A" box="[925,964,1922,1946]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="301" unit="m" value="9.0">
<quantity id="4CED9B35FFB9FFE5E7BDF820FC2B9E9A" box="[925,964,1922,1946]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="301" unit="m" value="9.0">9m</quantity>
</elevation>
,
<collectingMunicipality id="6BCEACAAFFB9FFE5E7F3F820FB7A9E9A" box="[979,1173,1922,1946]" pageId="8" pageNumber="301">Mangrove Camp</collectingMunicipality>
(
<location id="8ECA600BFFB9FFE5E087F823FF019EBE" LSID="urn:lsid:plazi:treatment:03BC87C6FFB6FFE6E4B7F88AFB8F9ED8:8ECA600BFFB9FFE5E087F823FF019EBE" country="Mozambique" latitude="-26.32664" longLatPrecision="1" longitude="32.70992" municipality="Mangrove Camp" name="Mangrove - Woodland Mosaic" pageId="8" pageNumber="301" stateProvince="West">Mangrove–Woodland Mosaic</location>
),
<geoCoordinate id="EE215017FFB9FFE5E523F807FE779EBE" box="[259,408,1957,1982]" direction="south" orientation="latitude" pageId="8" pageNumber="301" precision="1" value="-26.32664">26°19'35.9''S</geoCoordinate>
,
<geoCoordinate id="EE215017FFB9FFE5E585F807FDD39EBE" box="[421,572,1957,1982]" direction="east" orientation="longitude" pageId="8" pageNumber="301" precision="1" value="32.70992">32°42'35.7''E</geoCoordinate>
,
<collectingDate id="EFEFE9F8FFB9FFE5E66AF807FD0B9EBE" box="[586,740,1957,1982]" pageId="8" pageNumber="301" value="2016-12-07" valueMax="2016-12-09" valueMin="2016-12-07">
<date id="FFAB1010FFB9FFE5E66AF807FD0B9EBE" box="[586,740,1957,1982]" pageId="8" pageNumber="301" value="2016-12-07" valueMax="2016-12-09" valueMin="2016-12-07">7–9.XII.2016</date>
</collectingDate>
. MV.
<collectorName id="26E05306FFB9FFE5E70AF807FC109EBD" box="[810,1023,1957,1982]" pageId="8" pageNumber="301">Aristophanous, M.</collectorName>
,
<collectorName id="26E05306FFB9FFE5E02DF807FB729EBD" box="[1037,1181,1957,1982]" pageId="8" pageNumber="301">Cristóvão, J.</collectorName>
,
<collectorName id="26E05306FFB9FFE5E08BF807FACF9EBE" box="[1195,1312,1957,1982]" pageId="8" pageNumber="301">Laszlo, G.</collectorName>
,
<collectorName id="26E05306FFB9FFE5E10EF807FA729EBD" box="[1326,1437,1957,1982]" pageId="8" pageNumber="301">Miles, W.</collectorName>
leg., slide
<collectorName id="26E05306FFB9FFE5E52AF868FED89EE1" box="[266,311,1994,2017]" pageId="8" pageNumber="301">No.</collectorName>
: AV2980
<specimenCount id="9D13FD59FFB9FFE5E587F868FE5B9EE2" box="[423,436,1994,2018]" pageId="8" pageNumber="301" type="female">♀</specimenCount>
(coll.
<collectionCode id="ED04AE15FFB9FFE5E5DCF868FDB49EE2" box="[508,603,1994,2018]" pageId="8" pageNumber="301">ANHRT</collectionCode>
and
<collectionCode id="ED04AE15FFB9FFE5E6B1F868FD1E9EE2" box="[657,753,1994,2018]" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="8" pageNumber="301">NHMM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6A6658FFB8FFE4E4B7FA63FBAC9F71" httpUri="https://zenodo.org/record/1303160/files/figure.png" pageId="9" pageNumber="302" startId="9.[151,264,1473,1496]" targetBox="[154,1436,194,1438]" targetPageId="9">
<paragraph id="8BAA36D0FFB8FFE4E4B7FA63FBAC9F71" blockId="9.[151,1436,1473,1649]" pageId="9" pageNumber="302">
<emphasis id="B961EAC2FFB8FFE4E4B7FA63FEA39CD6" bold="true" box="[151,332,1473,1496]" pageId="9" pageNumber="302">FIGURES 26–33</emphasis>
.
<taxonomicName id="4C154D53FFB8FFE4E578FA63FE469CD7" authorityName="Wallengren" authorityYear="1863" box="[344,425,1473,1495]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC2FFB8FFE4E578FA63FE469CD7" box="[344,425,1473,1495]" italics="true" pageId="9" pageNumber="302">Tumicla</emphasis>
</taxonomicName>
spp.: female genitalia. 26,
<taxonomicName id="4C154D53FFB8FFE4E699FA60FCFA9CD7" authorityName="Volynkin &amp; László" authorityYear="2018" box="[697,789,1474,1495]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="tsonga">
<emphasis id="B961EAC2FFB8FFE4E699FA60FCFA9CD7" box="[697,789,1474,1495]" italics="true" pageId="9" pageNumber="302">T. tsonga</emphasis>
</taxonomicName>
, paratype, MoZambique, slide AV3036 (ANHRT); 27,
<taxonomicName id="4C154D53FFB8FFE4E11EFA60FA7A9CD7" authorityName="Volynkin &amp; László" authorityYear="2018" box="[1342,1429,1473,1495]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="arktos">
<emphasis id="B961EAC2FFB8FFE4E11EFA60FA7A9CD7" box="[1342,1429,1473,1495]" italics="true" pageId="9" pageNumber="302">T. arktos</emphasis>
</taxonomicName>
, paratype, MoZambique, slide AV3044 (ANHRT); 28,
<taxonomicName id="4C154D53FFB8FFE4E698FA43FD1A9CF6" authorityName="Kuhne" authorityYear="2007" box="[696,757,1504,1526]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="B961EAC2FFB8FFE4E698FA43FD1A9CF6" box="[696,757,1504,1526]" italics="true" pageId="9" pageNumber="302">T. doa</emphasis>
</taxonomicName>
, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29,
<taxonomicName id="4C154D53FFB8FFE4E49CF9A2FEE49F15" authorityName="Durante" authorityYear="2008" box="[188,267,1536,1557]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B961EAC2FFB8FFE4E49CF9A2FEE49F15" box="[188,267,1536,1557]" italics="true" pageId="9" pageNumber="302">T. rosea</emphasis>
</taxonomicName>
, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30,
<taxonomicName id="4C154D53FFB8FFE4E006F9A2FB6E9F14" authorityName="Durante" authorityYear="2008" box="[1062,1153,1534,1557]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B961EAC2FFB8FFE4E006F9A2FB6E9F14" box="[1062,1153,1534,1557]" italics="true" pageId="9" pageNumber="302">T. similis</emphasis>
</taxonomicName>
, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31,
<taxonomicName id="4C154D53FFB8FFE4E67CF9BCFD3D9F33" authorityName="Kuhne" authorityYear="2007" box="[604,722,1565,1587]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B961EAC2FFB8FFE4E67CF9BCFD3D9F33" box="[604,722,1565,1587]" italics="true" pageId="9" pageNumber="302">T. mutabilis</emphasis>
</taxonomicName>
, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32,
<taxonomicName id="4C154D53FFB8FFE4E5CBF99FFD839F52" authorityName="Durante" authorityYear="2008" box="[491,620,1596,1618]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B961EAC2FFB8FFE4E5CBF99FFD839F52" box="[491,620,1596,1618]" italics="true" pageId="9" pageNumber="302">T. dimidiata</emphasis>
</taxonomicName>
, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33,
<taxonomicName id="4C154D53FFB8FFE4E56EF9FEFE4F9F71" authorityName="Durante" authorityYear="2008" box="[334,416,1627,1649]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="B961EAC2FFB8FFE4E56EF9FEFE4F9F71" box="[334,416,1627,1649]" italics="true" pageId="9" pageNumber="302">T. rubea</emphasis>
</taxonomicName>
, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK).
</paragraph>
</caption>
<subSubSection id="C30F655BFFB8FFE7E4E7F93AFBBF98D0" lastPageId="10" lastPageNumber="303" pageId="9" pageNumber="302" type="diagnosis">
<paragraph id="8BAA36D0FFB8FFE4E4E7F93AFEA99E1D" blockId="9.[151,1436,1688,2037]" pageId="9" pageNumber="302">
<emphasis id="B961EAC2FFB8FFE4E4E7F93AFEAE9FB1" bold="true" box="[199,321,1688,1713]" pageId="9" pageNumber="302">Diagnosis.</emphasis>
<taxonomicName id="4C154D53FFB8FFE4E568F93BFE409FB0" authority="Volynkin &amp; László, 2018" authorityName="Volynkin &amp; László" authorityYear="2018" box="[328,431,1689,1712]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="tsonga" status="sp. nov.">
<emphasis id="B961EAC2FFB8FFE4E568F93BFE409FB0" box="[328,431,1689,1712]" italics="true" pageId="9" pageNumber="302">T. tsonga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257B9FFB8FFE4E597F93BFDE19FB0" box="[439,526,1689,1712]" pageId="9" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="132E2A55FFB8FFE4E63DF93AFD6F9FB1" box="[541,640,1688,1713]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1885,1908]" captionTargetBox="[165,1435,196,1854]" captionTargetId="figure@5.[151,1436,195,1864]" captionTargetPageId="5" captionText="FIGURES 1–8. Tumicla spp.: adults. 1, T. arktos, holotype ♂, MoZambique (ANHRT); 2, T. arktos, paratype ♀, MoZambique (ANHRT); 3, T. tsonga, holotype ♂, MoZambique (ANHRT); 4, T. tsonga, paratype ♂, MoZambique (ANHRT); 5, T. tsonga, paratype ♀, MoZambique (ANHRT); 6, T. doa, holotype ♀, [TanZania] Kwedikundi (ZMB); 7, T. dimidiata, holotype ♀, Democratic Republic of the Congo (©NHMUK); 8, T. mutabilis, holotype ♀, [TanZania] D.O. Africa (ZMB)." httpUri="https://zenodo.org/record/1303151/files/figure.png" pageId="9" pageNumber="302">Figs 3–5</figureCitation>
) is externally similar to several other species of
<taxonomicName id="4C154D53FFB8FFE4E084F93AFB119FB1" authorityName="Wallengren" authorityYear="1863" box="[1188,1278,1688,1713]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC2FFB8FFE4E084F93AFB119FB1" box="[1188,1278,1688,1713]" italics="true" pageId="9" pageNumber="302">Tumicla</emphasis>
</taxonomicName>
like
<emphasis id="B961EAC2FFB8FFE4E117F93BFECD9FD4" italics="true" pageId="9" pageNumber="302">
<taxonomicName id="4C154D53FFB8FFE4E117F93BFA789FB1" authorityName="Durante" authorityYear="2008" box="[1335,1431,1688,1713]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="rubea">T. rubea</taxonomicName>
,
<taxonomicName id="4C154D53FFB8FFE4E4B7F91FFECD9FD4" authorityName="Wallengren" authorityYear="1860" box="[151,290,1725,1748]" class="Insecta" family="Erebidae" genus="Setina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="sagenaria">T. sagenaria</taxonomicName>
</emphasis>
,
<taxonomicName id="4C154D53FFB8FFE4E50FF91FFE429FD4" authorityName="Kuhne" authorityYear="2007" box="[303,429,1725,1748]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="B961EAC2FFB8FFE4E50FF91FFE429FD4" box="[303,429,1725,1748]" italics="true" pageId="9" pageNumber="302">T. gigantea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C154D53FFB8FFE4E59AF91FFE109FD5" authorityName="Kuhne" authorityYear="2007" box="[442,511,1724,1749]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="B961EAC2FFB8FFE4E59AF91FFE109FD5" box="[442,511,1724,1749]" italics="true" pageId="9" pageNumber="302">T. doa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C154D53FFB8FFE4E62DF91FFD7E9FD5" authorityName="Kuhne" authorityYear="2007" box="[525,657,1724,1749]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B961EAC2FFB8FFE4E62DF91FFD7E9FD5" box="[525,657,1724,1749]" italics="true" pageId="9" pageNumber="302">T. mutabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D53FFB8FFE4E6E5F91FFCF39FD4" authorityName="Durante" authorityYear="2008" box="[709,796,1725,1748]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B961EAC2FFB8FFE4E6E5F91FFCF39FD4" box="[709,796,1725,1748]" italics="true" pageId="9" pageNumber="302">T. rosea</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A55FFB8FFE4E70CF91EFC709FD5" box="[812,927,1724,1749]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURES 9–17" captionStartId-0="5.[151,264,1885,1908]" captionStartId-1="6.[151,264,1884,1907]" captionTargetBox-0="[165,1435,196,1854]" captionTargetBox-1="[170,1406,194,1862]" captionTargetId-0="figure@5.[151,1436,195,1864]" captionTargetId-1="figure@6.[170,1406,193,1865]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–8. Tumicla spp.: adults. 1, T. arktos, holotype ♂, MoZambique (ANHRT); 2, T. arktos, paratype ♀, MoZambique (ANHRT); 3, T. tsonga, holotype ♂, MoZambique (ANHRT); 4, T. tsonga, paratype ♂, MoZambique (ANHRT); 5, T. tsonga, paratype ♀, MoZambique (ANHRT); 6, T. doa, holotype ♀, [TanZania] Kwedikundi (ZMB); 7, T. dimidiata, holotype ♀, Democratic Republic of the Congo (©NHMUK); 8, T. mutabilis, holotype ♀, [TanZania] D.O. Africa (ZMB)." captionText-1="FIGURES 9–17. Tumicla spp.: adults. 9, T. gigantea, paratype ♂, Burundi (ZMB); 10, T. similis, holotype ♀, [Kenya] KibweZi (©NHMUK); 11, T. rosea, holotype ♀, Malawi, Nyassaland (©NHMUK); 12, T. rubea, paratype ♀, South Africa (©NHMUK); 13, T. sagenaria, lectotype ♀, [South Africa] Caffraria (NHRS, photo by T. Malm, ©2017NHRS); 14, T. sagenaria, paralectotype ♂, [South Africa] Caffraria (NHRS, photo by T. Malm, ©2017NHRS); 15, T. calimerae, holotype ♀, Senegal (©NHMUK); 16, T. distyi, ♂, Sierra Leone (ANHRT); 17, T. distyi, paratype ♀, Ghana (ZMB)." httpUri-0="https://zenodo.org/record/1303151/files/figure.png" httpUri-1="https://zenodo.org/record/1303153/files/figure.png" pageId="9" pageNumber="302">Figs 6–14</figureCitation>
,
<figureCitation id="132E2A55FFB8FFE4E78AF91EFC1C9FD5" box="[938,1011,1724,1749]" captionStart-0="FIGURES 18–21" captionStart-1="FIGURES 22–25" captionStartId-0="7.[151,264,1672,1695]" captionStartId-1="8.[151,264,1654,1677]" captionTargetBox-0="[153,1432,183,1639]" captionTargetBox-1="[187,1426,194,1623]" captionTargetId-0="figure@7.[151,1436,177,1652]" captionTargetId-1="figure@8.[151,1436,193,1633]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." captionText-1="FIGURES 22–25. Tumicla spp.: male genitalia. 22, T. mutabilis, paratype, Uganda, slide BMNH(E) Arct-6052 (©NHMUK); 23, T. gigantea, paratype, Urundi, slide AV2897 (ZMB); 24, T. sagenaria, paralectotype, [South Africa] Caffraria slide NHRS 11085 (photo by T. Malm, ©2017NHRS); 25, T. sagenaria, South Africa, slide BMNH(E) Arct-5861 (©NHMUK)." httpUri-0="https://zenodo.org/record/1303155/files/figure.png" httpUri-1="https://zenodo.org/record/1303158/files/figure.png" pageId="9" pageNumber="302">20–25</figureCitation>
,
<figureCitation id="132E2A55FFB8FFE4E7DEF91EFBA79FD5" box="[1022,1096,1724,1749]" captionStart-0="FIGURES 26–33" captionStart-1="FIGURES 34–36" captionStartId-0="9.[151,264,1473,1496]" captionStartId-1="10.[151,264,1662,1685]" captionTargetBox-0="[154,1436,194,1438]" captionTargetBox-1="[191,1391,519,1639]" captionTargetId-0="figure@9.[151,1436,193,1452]" captionTargetId-1="figure@10.[190,1395,517,1639]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 26–33. Tumicla spp.: female genitalia. 26, T. tsonga, paratype, MoZambique, slide AV3036 (ANHRT); 27, T. arktos, paratype, MoZambique, slide AV3044 (ANHRT); 28, T. doa, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30, T. similis, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31, T. mutabilis, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32, T. dimidiata, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK)." captionText-1="FIGURES 34–36. Tumicla spp.: female (34, 35) and male (36) genitalia. 34, T. gigantea, paratype, Urundi, slide AV2898 (ZMB); 35, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS); 36, T. distyi, Sierra Leone, slide AV2934 (ANHRT)." httpUri-0="https://zenodo.org/record/1303160/files/figure.png" httpUri-1="https://zenodo.org/record/1303162/files/figure.png" pageId="9" pageNumber="302">28–35</figureCitation>
), but clearly differs from them by its considerably darker hindwings. However, the clear distinctive characters are expressed in the configuration of the genitalia.
</paragraph>
<paragraph id="8BAA36D0FFB8FFE7E4E7F88AFA6C99D5" blockId="9.[151,1436,1688,2037]" lastBlockId="10.[151,1436,151,464]" lastPageId="10" lastPageNumber="303" pageId="9" pageNumber="302">
The male genitalia (
<figureCitation id="132E2A55FFB8FFE4E587F88AFE169E40" box="[423,505,1832,1857]" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="9" pageNumber="302">Fig. 18</figureCitation>
) is similar to those of
<taxonomicName id="4C154D53FFB8FFE4E6D4F88BFCA09E41" authorityName="Durante" authorityYear="2008" box="[756,847,1832,1857]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="B961EAC2FFB8FFE4E6D4F88BFCA09E41" box="[756,847,1832,1857]" italics="true" pageId="9" pageNumber="302">T. rubea</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A55FFB8FFE4E741F88AFC5B9E40" box="[865,948,1832,1857]" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="9" pageNumber="302">Fig. 20</figureCitation>
),
<taxonomicName id="4C154D53FFB8FFE4E7E9F88BFBBA9E40" authorityName="Wallengren" authorityYear="1860" box="[969,1109,1833,1856]" class="Insecta" family="Erebidae" genus="Setina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="B961EAC2FFB8FFE4E7E9F88BFBBA9E40" box="[969,1109,1833,1856]" italics="true" pageId="9" pageNumber="302">T. sagenaria</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A55FFB8FFE4E043F88AFB0A9E41" box="[1123,1253,1832,1857]" captionStart="FIGURES 22–25" captionStartId="8.[151,264,1654,1677]" captionTargetBox="[187,1426,194,1623]" captionTargetId="figure@8.[151,1436,193,1633]" captionTargetPageId="8" captionText="FIGURES 22–25. Tumicla spp.: male genitalia. 22, T. mutabilis, paratype, Uganda, slide BMNH(E) Arct-6052 (©NHMUK); 23, T. gigantea, paratype, Urundi, slide AV2897 (ZMB); 24, T. sagenaria, paralectotype, [South Africa] Caffraria slide NHRS 11085 (photo by T. Malm, ©2017NHRS); 25, T. sagenaria, South Africa, slide BMNH(E) Arct-5861 (©NHMUK)." httpUri="https://zenodo.org/record/1303158/files/figure.png" pageId="9" pageNumber="302">Figs 24, 25</figureCitation>
),
<taxonomicName id="4C154D53FFB8FFE4E0DAF88BFAB09E41" authorityName="Durante" authorityYear="2008" box="[1274,1375,1832,1857]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B961EAC2FFB8FFE4E0DAF88BFAB09E41" box="[1274,1375,1832,1857]" italics="true" pageId="9" pageNumber="302">T. similis</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A55FFB8FFE4E14EF88AFF5B9E65" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="9" pageNumber="302">Fig. 21</figureCitation>
) and
<taxonomicName id="4C154D53FFB8FFE4E4DAF8EFFEB09E65" authority="Volynkin &amp; László, 2018" authorityName="Volynkin &amp; László" authorityYear="2018" box="[250,351,1868,1893]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="arktos" status="sp. nov.">
<emphasis id="B961EAC2FFB8FFE4E4DAF8EFFEB09E65" box="[250,351,1868,1893]" italics="true" pageId="9" pageNumber="302">T. arktos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257B9FFB8FFE4E549F8EEFE299E64" box="[361,454,1868,1893]" pageId="9" pageNumber="302" rank="species">
<emphasis id="B961EAC2FFB8FFE4E549F8EEFE509E65" bold="true" box="[361,447,1868,1893]" pageId="9" pageNumber="302">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="132E2A55FFB8FFE4E5F9F8EEFDC19E65" box="[473,558,1868,1893]" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="9" pageNumber="302">Fig. 19</figureCitation>
) due to the absence of thorn-like carinal process, but differs clearly by the configuration of vesica which has three diverticula basally and subbasally. In addition there is a distal row of robust, slightly curved, thorn-like cornuti in
<taxonomicName id="4C154D53FFB8FFE4E6A5F837FD049EAC" authority="Volynkin &amp; László, 2018" authorityName="Volynkin &amp; László" authorityYear="2018" box="[645,747,1941,1964]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="tsonga" status="sp. nov.">
<emphasis id="B961EAC2FFB8FFE4E6A5F837FD049EAC" box="[645,747,1941,1964]" italics="true" pageId="9" pageNumber="302">T. tsonga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257B9FFB8FFE4E6D1F836FCA59EAD" box="[753,842,1940,1965]" pageId="9" pageNumber="302" rank="species">
<emphasis id="B961EAC2FFB8FFE4E6D1F836FCA59EAD" bold="true" box="[753,842,1940,1965]" pageId="9" pageNumber="302">sp. nov.</emphasis>
</taxonomicNameLabel>
, whereas there are two bunches of considerably finer cornuti of vesica in
<taxonomicName id="4C154D53FFB8FFE4E556F81BFDEE9ED0" authorityName="Wallengren" authorityYear="1860" box="[374,513,1977,2000]" class="Insecta" family="Erebidae" genus="Setina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="B961EAC2FFB8FFE4E556F81BFDEE9ED0" box="[374,513,1977,2000]" italics="true" pageId="9" pageNumber="302">T. sagenaria</emphasis>
</taxonomicName>
, much fewer, straight cornuti in
<taxonomicName id="4C154D53FFB8FFE4E750F81BFC3D9ED1" authorityName="Volynkin &amp; László" authorityYear="2018" box="[880,978,1976,2001]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="arktos">
<emphasis id="B961EAC2FFB8FFE4E750F81BFC3D9ED1" box="[880,978,1976,2001]" italics="true" pageId="9" pageNumber="302">T. arktos</emphasis>
</taxonomicName>
, much finer, and more numerous cornuti in
<taxonomicName id="4C154D53FFB8FFE4E495F87FFEFD9EF5" authorityName="Durante" authorityYear="2008" box="[181,274,2012,2037]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="B961EAC2FFB8FFE4E495F87FFEFD9EF5" box="[181,274,2012,2037]" italics="true" pageId="9" pageNumber="302">T. rubea</emphasis>
</taxonomicName>
and no cornuti on the vesica of
<emphasis id="B961EAC2FFB8FFE4E6A5F87FFD1D9EF5" box="[645,754,2012,2037]" italics="true" pageId="9" pageNumber="302">
<taxonomicName id="4C154D53FFB8FFE4E6A5F87FFD019EF5" authorityName="Durante" authorityYear="2008" box="[645,750,2012,2037]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="similis">T. similis</taxonomicName>
.
</emphasis>
Due to the similar basic configuration of uncus, valvae and cornuti of vesica, the new species’ closest relative is most probably
<taxonomicName id="4C154D53FFBBFFE7E783FF3BFBEF99B0" authorityName="Durante" authorityYear="2008" box="[931,1024,151,176]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="B961EAC2FFBBFFE7E783FF3BFBEF99B0" box="[931,1024,151,176]" italics="true" pageId="10" pageNumber="303">T. rubea</emphasis>
</taxonomicName>
, although the relationships between species within the entire genus could be clarified most reliably by application of molecular / biological methods.
</paragraph>
<paragraph id="8BAA36D0FFBBFFE7E4E7FF7DFBBF98D0" blockId="10.[151,1436,151,464]" pageId="10" pageNumber="303">
The female genitalia of
<taxonomicName id="4C154D53FFBBFFE7E5F1FF43FDD899F8" authority="Volynkin &amp; László, 2018" authorityName="Volynkin &amp; László" authorityYear="2018" box="[465,567,225,248]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="tsonga" status="sp. nov.">
<emphasis id="B961EAC2FFBBFFE7E5F1FF43FDD899F8" box="[465,567,225,248]" italics="true" pageId="10" pageNumber="303">T. tsonga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257B9FFBBFFE7E61EFF7DFD7899F8" box="[574,663,223,248]" pageId="10" pageNumber="303" rank="species">
<emphasis id="B961EAC2FFBBFFE7E61EFF7DFD7899F8" bold="true" box="[574,663,223,248]" pageId="10" pageNumber="303">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="132E2A55FFBBFFE7E686FF7DFD1899F8" box="[678,759,223,248]" captionStart="FIGURES 26–33" captionStartId="9.[151,264,1473,1496]" captionTargetBox="[154,1436,194,1438]" captionTargetId="figure@9.[151,1436,193,1452]" captionTargetPageId="9" captionText="FIGURES 26–33. Tumicla spp.: female genitalia. 26, T. tsonga, paratype, MoZambique, slide AV3036 (ANHRT); 27, T. arktos, paratype, MoZambique, slide AV3044 (ANHRT); 28, T. doa, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30, T. similis, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31, T. mutabilis, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32, T. dimidiata, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK)." httpUri="https://zenodo.org/record/1303160/files/figure.png" pageId="10" pageNumber="303">Fig. 26</figureCitation>
) is similar to those of
<taxonomicName id="4C154D53FFBBFFE7E7D0FF43FBDB99F8" authorityName="Kuhne" authorityYear="2007" box="[1008,1076,223,248]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="B961EAC2FFBBFFE7E7D0FF43FBDB99F8" box="[1008,1076,223,248]" italics="true" pageId="10" pageNumber="303">T. doa</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A55FFBBFFE7E065FF7DFB7999F8" box="[1093,1174,223,248]" captionStart="FIGURES 26–33" captionStartId="9.[151,264,1473,1496]" captionTargetBox="[154,1436,194,1438]" captionTargetId="figure@9.[151,1436,193,1452]" captionTargetPageId="9" captionText="FIGURES 26–33. Tumicla spp.: female genitalia. 26, T. tsonga, paratype, MoZambique, slide AV3036 (ANHRT); 27, T. arktos, paratype, MoZambique, slide AV3044 (ANHRT); 28, T. doa, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30, T. similis, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31, T. mutabilis, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32, T. dimidiata, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK)." httpUri="https://zenodo.org/record/1303160/files/figure.png" pageId="10" pageNumber="303">Fig. 28</figureCitation>
) and
<taxonomicName id="4C154D53FFBBFFE7E0F4FF43FAC599F8" authorityName="Durante" authorityYear="2008" box="[1236,1322,225,248]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B961EAC2FFBBFFE7E0F4FF43FAC599F8" box="[1236,1322,225,248]" italics="true" pageId="10" pageNumber="303">T. rosea</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A55FFBBFFE7E11CFF7DFA6099F8" box="[1340,1423,223,248]" captionStart="FIGURES 26–33" captionStartId="9.[151,264,1473,1496]" captionTargetBox="[154,1436,194,1438]" captionTargetId="figure@9.[151,1436,193,1452]" captionTargetPageId="9" captionText="FIGURES 26–33. Tumicla spp.: female genitalia. 26, T. tsonga, paratype, MoZambique, slide AV3036 (ANHRT); 27, T. arktos, paratype, MoZambique, slide AV3044 (ANHRT); 28, T. doa, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30, T. similis, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31, T. mutabilis, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32, T. dimidiata, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK)." httpUri="https://zenodo.org/record/1303160/files/figure.png" pageId="10" pageNumber="303">Fig. 29</figureCitation>
), but differs by its more strongly sclerotized, anteriorly and posteriorly broadened, more or less hour-glass shaped ductus bursae, which is rather funnel-like in the related species. The laterally situated, globular shaped corpus bursae of the new species is considerably smaller than that of
<taxonomicName id="4C154D53FFBBFFE7E754FEEFFC539865" authorityName="Kuhne" authorityYear="2007" box="[884,956,332,357]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="B961EAC2FFBBFFE7E754FEEFFC539865" box="[884,956,332,357]" italics="true" pageId="10" pageNumber="303">T. doa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D53FFBBFFE7E7DBFEEFFBB99864" authorityName="Durante" authorityYear="2008" box="[1019,1110,333,356]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B961EAC2FFBBFFE7E7DBFEEFFBB99864" box="[1019,1110,333,356]" italics="true" pageId="10" pageNumber="303">T. rosea</emphasis>
</taxonomicName>
situated proximally in both species. In addition the new species has a rounded, scobinated signum bursae, whereas signum bursae is absent in both related species; the appendix bursae of the new species is the largest in the group, with conspicuous lateral sclerotization, while the appendix bursae of
<taxonomicName id="4C154D53FFBBFFE7E6A6FE1BFD2598D0" authorityName="Kuhne" authorityYear="2007" box="[646,714,439,464]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="B961EAC2FFBBFFE7E6A6FE1BFD2598D0" box="[646,714,439,464]" italics="true" pageId="10" pageNumber="303">T. doa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D53FFBBFFE7E722FE1BFCB798D0" authorityName="Durante" authorityYear="2008" box="[770,856,441,464]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B961EAC2FFBBFFE7E722FE1BFCB798D0" box="[770,856,441,464]" italics="true" pageId="10" pageNumber="303">T. rosea</emphasis>
</taxonomicName>
is fully membranous.
</paragraph>
</subSubSection>
<caption id="DF6A6658FFBBFFE7E4B7F9DCFD889FD2" httpUri="https://zenodo.org/record/1303162/files/figure.png" pageId="10" pageNumber="303" startId="10.[151,264,1662,1685]" targetBox="[191,1391,519,1639]" targetPageId="10">
<paragraph id="8BAA36D0FFBBFFE7E4B7F9DCFD889FD2" blockId="10.[151,1437,1662,1746]" pageId="10" pageNumber="303">
<emphasis id="B961EAC2FFBBFFE7E4B7F9DCFEBF9F94" bold="true" box="[151,336,1662,1685]" pageId="10" pageNumber="303">FIGURES 34–36</emphasis>
.
<taxonomicName id="4C154D53FFBBFFE7E540F9DDFE5E9F95" authorityName="Wallengren" authorityYear="1863" box="[352,433,1663,1685]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC2FFBBFFE7E540F9DDFE5E9F95" box="[352,433,1663,1685]" italics="true" pageId="10" pageNumber="303">Tumicla</emphasis>
</taxonomicName>
spp.: female (34, 35) and male (36) genitalia. 34,
<taxonomicName id="4C154D53FFBBFFE7E7ECF922FBAE9F95" authorityName="Kuhne" authorityYear="2007" box="[972,1089,1664,1685]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="B961EAC2FFBBFFE7E7ECF922FBAE9F95" box="[972,1089,1664,1685]" italics="true" pageId="10" pageNumber="303">T. gigantea</emphasis>
</taxonomicName>
, paratype, Urundi, slide AV2898 (ZMB); 35,
<taxonomicName id="4C154D53FFBBFFE7E531F93DFE609FB4" authorityName="Wallengren" authorityYear="1860" box="[273,399,1695,1716]" class="Insecta" family="Erebidae" genus="Setina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="B961EAC2FFBBFFE7E531F93DFE609FB4" box="[273,399,1695,1716]" italics="true" pageId="10" pageNumber="303">T. sagenaria</emphasis>
</taxonomicName>
, lectotype, [South Africa] Caffraria, slide NHRS 0 0 0 0 11009 (photo by T. Malm, ©2017NHRS); 36,
<taxonomicName id="4C154D53FFBBFFE7E1A9F93DFF219FD2" authorityName="Kuhne" authorityYear="2007" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="distyi">
<emphasis id="B961EAC2FFBBFFE7E1A9F93DFF219FD2" italics="true" pageId="10" pageNumber="303">T. distyi</emphasis>
</taxonomicName>
, Sierra Leone, slide AV2934 (ANHRT).
</paragraph>
</caption>
<subSubSection id="C30F655BFFBBFFE6E4E7F958FB469E00" lastPageId="11" lastPageNumber="304" pageId="10" pageNumber="303" type="description">
<paragraph id="8BAA36D0FFBBFFE7E4E7F958FBD79E5B" blockId="10.[151,1437,1786,2027]" pageId="10" pageNumber="303">
<emphasis id="B961EAC2FFBBFFE7E4E7F958FEBE9E13" bold="true" box="[199,337,1786,1811]" pageId="10" pageNumber="303">Description</emphasis>
.
<emphasis id="B961EAC2FFBBFFE7E57FF958FE4C9E13" bold="true" box="[351,419,1786,1811]" pageId="10" pageNumber="303">Adult</emphasis>
(
<figureCitation id="132E2A55FFBBFFE7E591F958FDF89E13" box="[433,535,1786,1811]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1885,1908]" captionTargetBox="[165,1435,196,1854]" captionTargetId="figure@5.[151,1436,195,1864]" captionTargetPageId="5" captionText="FIGURES 1–8. Tumicla spp.: adults. 1, T. arktos, holotype ♂, MoZambique (ANHRT); 2, T. arktos, paratype ♀, MoZambique (ANHRT); 3, T. tsonga, holotype ♂, MoZambique (ANHRT); 4, T. tsonga, paratype ♂, MoZambique (ANHRT); 5, T. tsonga, paratype ♀, MoZambique (ANHRT); 6, T. doa, holotype ♀, [TanZania] Kwedikundi (ZMB); 7, T. dimidiata, holotype ♀, Democratic Republic of the Congo (©NHMUK); 8, T. mutabilis, holotype ♀, [TanZania] D.O. Africa (ZMB)." httpUri="https://zenodo.org/record/1303151/files/figure.png" pageId="10" pageNumber="303">Figs 3–5</figureCitation>
). Forewing length
<quantity id="4CED9B35FFBBFFE7E6C9F958FC8F9E12" box="[745,864,1786,1810]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="10" pageNumber="303" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
in males and
<quantity id="4CED9B35FFBBFFE7E7D9F958FB9D9E12" box="[1017,1138,1786,1811]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="10" pageNumber="303" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
in females. Male antennae ciliate, female antennae filiform. Intraspecific variability rather low, expressed only by the different thickness of the elements of forewing pattern, especially the ante-, postmedial and medial lines.
</paragraph>
<paragraph id="8BAA36D0FFBBFFE7E4E7F8C4FDC29E7F" blockId="10.[151,1437,1786,2027]" box="[199,557,1894,1919]" pageId="10" pageNumber="303">
<emphasis id="B961EAC2FFBBFFE7E4E7F8C4FEEB9E7F" box="[199,260,1894,1919]" italics="true" pageId="10" pageNumber="303">Head</emphasis>
: brick-red, frons blackish.
</paragraph>
<paragraph id="8BAA36D0FFBBFFE6E4E7F828FA6899D5" blockId="10.[151,1437,1786,2027]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="304" pageId="10" pageNumber="303">
<emphasis id="B961EAC2FFBBFFE7E4E7F828FEF79EA3" box="[199,280,1930,1955]" italics="true" pageId="10" pageNumber="303">Thorax</emphasis>
: brick-red, with four black spots mediolaterally; Forewing moderately broad, broader in female, with rounded apex and oblique tornus; forewing ground colour brick-red in males, pale brick-red in females; wing pattern blackish, consists of two large subbasal spots, zigzagged ante-, postmedial and medial lines connecting each other, semilunar discal spot, and interrupted subterminal line consisting of dashes of various length; cilia black. Hindwing dark brick-red in males, paler in females, cilia blackish around apex and orange-red posteriorly.
</paragraph>
<caption id="DF6A6658FFBAFFE6E4B7FB30FECD9DE6" httpUri="https://zenodo.org/record/1303164/files/figure.png" pageId="11" pageNumber="304" startId="11.[151,264,1170,1193]" targetBox="[195,1378,263,1143]" targetPageId="11">
<paragraph id="8BAA36D0FFBAFFE6E4B7FB30FECD9DE6" blockId="11.[151,1436,1170,1254]" pageId="11" pageNumber="304">
<emphasis id="B961EAC2FFBAFFE6E4B7FB30FEA19DA8" bold="true" box="[151,334,1170,1193]" pageId="11" pageNumber="304">FIGURES 37–39</emphasis>
.
<taxonomicName id="4C154D53FFBAFFE6E57CFB31FE419DA9" authorityName="Wallengren" authorityYear="1863" box="[348,430,1171,1193]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC2FFBAFFE6E57CFB31FE419DA9" box="[348,430,1171,1193]" italics="true" pageId="11" pageNumber="304">Tumicla</emphasis>
</taxonomicName>
spp.: female genitalia. 37,
<taxonomicName id="4C154D53FFBAFFE6E6E7FB36FCF89DA9" authorityName="Kuhne" authorityYear="2007" box="[711,791,1171,1193]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="distyi">
<emphasis id="B961EAC2FFBAFFE6E6E7FB36FCF89DA9" box="[711,791,1171,1193]" italics="true" pageId="11" pageNumber="304">T. distyi</emphasis>
</taxonomicName>
, paratype, Ghana, slide 170/2004 Kühne (ZMB); 38,
<taxonomicName id="4C154D53FFBAFFE6E164FB36FA7B9DA9" authorityName="Kuhne" authorityYear="2007" box="[1348,1428,1171,1193]" class="Insecta" family="Erebidae" genus="Asura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="distyi">
<emphasis id="B961EAC2FFBAFFE6E164FB36FA7B9DA9" box="[1348,1428,1171,1193]" italics="true" pageId="11" pageNumber="304">T. distyi</emphasis>
</taxonomicName>
, Ghana, slide MwM 33770 Volynkin (MwM/ZSM); 39,
<emphasis id="B961EAC2FFBAFFE6E735FB11FC4E9DC8" box="[789,929,1202,1224]" italics="true" pageId="11" pageNumber="304">
K.
<taxonomicName id="4C154D53FFBAFFE6E71CFB10FC4E9DC8" authorityName="Durante" authorityYear="2008" box="[828,929,1202,1224]" class="Insecta" family="Arctiidae" genus="Asuroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="calimerae">calimerae</taxonomicName>
</emphasis>
, paratype, Senegal, slide NHMUK010314048 (©NHMUK).
</paragraph>
</caption>
<paragraph id="8BAA36D0FFBAFFE6E4E7FAB1FC509C2C" blockId="11.[151,1437,1299,2008]" box="[199,959,1299,1324]" pageId="11" pageNumber="304">
<emphasis id="B961EAC2FFBAFFE6E4E7FAB1FEDE9C2C" box="[199,305,1299,1324]" italics="true" pageId="11" pageNumber="304">Abdomen</emphasis>
: brick-red, in male with admixture of blackish hair scales.
</paragraph>
<paragraph id="8BAA36D0FFBAFFE6E4E7FA95FBD59F28" blockId="11.[151,1437,1299,2008]" pageId="11" pageNumber="304">
<emphasis id="B961EAC2FFBAFFE6E4E7FA95FE809C50" box="[199,367,1335,1360]" italics="true" pageId="11" pageNumber="304">Male genitalia</emphasis>
(
<figureCitation id="132E2A55FFBAFFE6E5A1FA95FE379C4F" box="[385,472,1335,1360]" captionStart="FIGURES 18–21" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[153,1432,183,1639]" captionTargetId="figure@7.[151,1436,177,1652]" captionTargetPageId="7" captionText="FIGURES 18–21. Tumicla spp.: male genitalia. 18, T. tsonga, holotype, MoZambique, slide AV3035 (ANHRT); 19, T. arktos, holotype, MoZambique, slide AV3042 (ANHRT); 20, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 21, T. similis, paratype, Kenya, KibweZi, slide BMNH(E) Arct-4656 (©NHMUK)." httpUri="https://zenodo.org/record/1303155/files/figure.png" pageId="11" pageNumber="304">Fig. 18</figureCitation>
). Uncus moderately long, narrow, distally curved, hook-like, apically pointed; tuba analis broad, scaphium thin, weakly sclerotized, subscaphium represented by a field of fine spinulose scobination; tegumen moderately long and broad; juxta broad, shield-like; vinculum short, V-like, apically rounded; valva elongated, medially broadened, its distal half slightly curved ventrally, apically strongly tapered, with rounded tip; costal margin with short irregular dentation at apex; sacculus short, narrow, weakly setose, without processes. Aedeagus short, straight; vesica broad and long, directed ventrally, with one basal and two lateral subbasal diverticula and a distal row of robust, curved, thorn-like cornuti of different length.
</paragraph>
<paragraph id="8BAA36D0FFBAFFE6E4E7F991FB469E00" blockId="11.[151,1437,1299,2008]" pageId="11" pageNumber="304">
<emphasis id="B961EAC2FFBAFFE6E4E7F991FE649F4C" box="[199,395,1587,1612]" italics="true" pageId="11" pageNumber="304">Female genitalia</emphasis>
(
<figureCitation id="132E2A55FFBAFFE6E5BFF991FE179F4C" box="[415,504,1587,1612]" captionStart="FIGURES 26–33" captionStartId="9.[151,264,1473,1496]" captionTargetBox="[154,1436,194,1438]" captionTargetId="figure@9.[151,1436,193,1452]" captionTargetPageId="9" captionText="FIGURES 26–33. Tumicla spp.: female genitalia. 26, T. tsonga, paratype, MoZambique, slide AV3036 (ANHRT); 27, T. arktos, paratype, MoZambique, slide AV3044 (ANHRT); 28, T. doa, holotype, [TanZania] Kwedikundi, slide 388/2004 Kühne (ZMB); 29, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 30, T. similis, paratype, Kenya, KibweZi, slide NHMUK010314047 (©NHMUK); 31, T. mutabilis, holotype, [TanZania] D.O. Africa, slide 387/2004 Kühne, re-mounted by A.V. Volynkin (ZMB); 32, T. dimidiata, holotype, Democratic Republic of the Congo, slide BMNH(E) Arct-5865 (©NHMUK); 33, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NHMUK)." httpUri="https://zenodo.org/record/1303160/files/figure.png" pageId="11" pageNumber="304">Fig. 26</figureCitation>
). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses anteriores and posteriores moderately long, thin, and equal in length; ostium bursae very broad with slightly arched margin; ductus bursae broad, dorso-ventrally flattened, medially laced, heavily sclerotized, rugose at connection with corpus bursae; appendix bursae larger than corpus bursae, directed anteriorly, moderately sclerotized laterally, with ductus seminalis originating apically; corpus bursae small, globular, situated laterally, membranous, with a medium large, more or less rounded weakly scobinated plate of signum.
</paragraph>
</subSubSection>
<subSubSection id="C30F655BFFBAFFE6E4E7F8A9FB049EB4" pageId="11" pageNumber="304" type="distribution">
<paragraph id="8BAA36D0FFBAFFE6E4E7F8A9FB049EB4" blockId="11.[151,1437,1299,2008]" pageId="11" pageNumber="304">
<emphasis id="B961EAC2FFBAFFE6E4E7F8A9FDF19E24" bold="true" box="[199,542,1803,1828]" pageId="11" pageNumber="304">Distribution and bionomics</emphasis>
.
<taxonomicName id="4C154D53FFBAFFE6E615F8A9FD069E23" authorityName="Volynkin &amp; László" authorityYear="2018" box="[565,745,1803,1828]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="tsonga">
<emphasis id="B961EAC2FFBAFFE6E615F8A9FD069E23" box="[565,745,1803,1828]" italics="true" pageId="11" pageNumber="304">Tumicla tsonga</emphasis>
</taxonomicName>
is currently only known from the southernmost part of
<collectingCountry id="F3027640FFBAFFE6E4B7F88DFEC09E48" box="[151,303,1839,1864]" name="Mozambique" pageId="11" pageNumber="304">Mozambique</collectingCountry>
. All specimens of the
<typeStatus id="54AE8872FFBAFFE6E60EF892FDB19E48" box="[558,606,1840,1864]" pageId="11" pageNumber="304">type</typeStatus>
series were collected at mercury vapour light and by actinic light trap in various habitats of the
<collectingRegion id="49D1F832FFBAFFE6E5BDF8F6FE1B9E6B" box="[413,500,1876,1899]" country="Mozambique" name="Maputo" pageId="11" pageNumber="304">Maputo</collectingRegion>
Special Reserve, i.e. in dune forest (
<typeStatus id="54AE8872FFBAFFE6E7BAF8F6FC239E6C" box="[922,972,1876,1900]" pageId="11" pageNumber="304">type</typeStatus>
locality, see
<figureCitation id="132E2A55FFBAFFE6E043F8F1FB569E6C" box="[1123,1209,1875,1900]" captionStart="FIGURES 40–41" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[151,1435,203,1926]" captionTargetId="figure@12.[151,1436,187,1926]" captionTargetPageId="12" captionText="FIGURES 40–41. MoZambique, Maputo Special Reserve, habitats of the new species of Tumicla. 40, Ponta Milibangalala, dune forest, 26°26'58.6''S, 32°55'29.8''E, 24.XI.2016, the type locality of T. tsonga (Photo by Gy. M. LásZló); 41, west Gate, sand thicket, 26°30'14.2''S, 32°42'59.6''E, 26.XI.2016, the type locality of T. arktos (Photo by Gy. M. LásZló)." httpUri="https://zenodo.org/record/1303166/files/figure.png" pageId="11" pageNumber="304">Fig. 40</figureCitation>
), sand thicket, open woodland and sand forest. The specimens have been collected during the beginning of the wet season and in the dry season as well, the species is most probably polyvoltine. Food plant and early stages unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30F655BFFBAFFE6E4E7F81DFB8F9ED8" box="[199,1120,1983,2008]" pageId="11" pageNumber="304" type="etymology">
<paragraph id="8BAA36D0FFBAFFE6E4E7F81DFB8F9ED8" blockId="11.[151,1437,1299,2008]" box="[199,1120,1983,2008]" pageId="11" pageNumber="304">
<emphasis id="B961EAC2FFBAFFE6E4E7F81DFEA99ED8" bold="true" box="[199,326,1983,2008]" pageId="11" pageNumber="304">Etymology</emphasis>
. Tsonga is the name of a major ethnic group in southern
<collectingCountry id="F3027640FFBAFFE6E7E4F81DFBB49ED8" box="[964,1115,1983,2008]" name="Mozambique" pageId="11" pageNumber="304">Mozambique</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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