179 lines
19 KiB
XML
179 lines
19 KiB
XML
<document id="D54E794CE9462150A8343FD6E7E6939B" ID-DOI="10.5281/zenodo.176061" ID-GBIF-Dataset="9705d176-1598-469c-8614-33dc4a7e5af1" ID-ISSN="1175-5326" ID-Zenodo-Dep="176061" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459826834308" checkinUser="plazi" docAuthor="Wang, Tian-Tian, Liang, Jun-Hui & Ren, Dong" docDate="2007" docId="03BC87B1FF924B640680BF56EB6B8D9E" docLanguage="en" docName="zt01443p027.pdf" docOrigin="Zootaxa 1443" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Habroblattula Wang, Liang & Ren, 2007, gen. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FF85FFC9FF934B660617BB11E9058C6C" masterDocTitle="Variability of Habroblattula drepanoides gen. et. sp. nov. (Insecta: Blattaria: Blattulidae) from the Yixian Formation in Liaoning, China" masterLastPageNumber="27" masterPageNumber="17" pageNumber="18" updateTime="1698222403123" updateUser="plazi">
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<mods:title id="5F6A56EF6CC69DE4FFDC1CE57B7917C4">Variability of Habroblattula drepanoides gen. et. sp. nov. (Insecta: Blattaria: Blattulidae) from the Yixian Formation in Liaoning, China</mods:title>
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<mods:name id="F01F10BE4C0A7DF335C9429B9B882BB2" type="personal">
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<mods:namePart id="B8D42CBAA31A12D48D97C296614ACB9F">Wang, Tian-Tian</mods:namePart>
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<mods:namePart id="8D71B394B0447DC6E9C8E8D97578ABFD">Liang, Jun-Hui</mods:namePart>
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<mods:namePart id="DE743DF71BCD439F61B0372968FDAB9B">Ren, Dong</mods:namePart>
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<mods:title id="EDE8192F0B7562C7EE76CBF6F7FEAC11">Zootaxa</mods:title>
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<mods:part id="026AD12532A4F1B42B51E991784CE1DA">
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<mods:date id="F86942FC5C6AD1C04EFB1DC775A6D8A5">2007</mods:date>
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<mods:detail id="D00BEFA5F90226473892CC6479B4AC15" type="volume">
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<mods:number id="394D9CED10628A07FF61735A1F92E0DC">1443</mods:number>
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<mods:identifier id="15BA29D96E49B3213CE11A65B6B671EF" type="DOI">10.5281/zenodo.176061</mods:identifier>
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<treatment id="03BC87B1FF924B640680BF56EB6B8D9E" ID-DOI="http://doi.org/10.5281/zenodo.6245967" ID-GBIF-Taxon="119356989" ID-Zenodo-Dep="6245967" LSID="urn:lsid:plazi:treatment:03BC87B1FF924B640680BF56EB6B8D9E" httpUri="http://treatment.plazi.org/id/03BC87B1FF924B640680BF56EB6B8D9E" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
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<subSubSection id="C30F652CFF924B670680BF56EB0B880D" box="[151,526,1095,1121]" pageId="1" pageNumber="18" type="nomenclature">
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<paragraph id="8BAA36A7FF924B670680BF56EB0B880D" blockId="1.[151,526,1095,1121]" box="[151,526,1095,1121]" pageId="1" pageNumber="18">
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<heading id="D0E281CBFF924B670680BF56EB0B880D" bold="true" box="[151,526,1095,1121]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
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<emphasis id="B961EAB5FF924B670680BF56EB0B880D" bold="true" box="[151,526,1095,1121]" pageId="1" pageNumber="18">
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Genus
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<taxonomicName id="4C154D24FF924B6706FABF56E89D880D" box="[237,408,1095,1121]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B961EAB5FF924B6706FABF56E89D880D" bold="true" box="[237,408,1095,1121]" italics="true" pageId="1" pageNumber="18">Habroblattula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25257CEFF924B6707B7BF56EB0B880D" box="[416,526,1095,1121]" pageId="1" pageNumber="18" rank="genus">
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<taxonomicName id="4C154D24FF924B6707B7BF56E8CB880D" box="[416,462,1095,1121]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="gen">gen</taxonomicName>
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. nov.
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</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F652CFF924B670680BF83EBE688C0" box="[151,739,1170,1196]" pageId="1" pageNumber="18" type="materials_examined">
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<paragraph id="8BAA36A7FF924B670680BF83EBE688C0" blockId="1.[151,1437,1170,2036]" box="[151,739,1170,1196]" pageId="1" pageNumber="18">
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<emphasis id="B961EAB5FF924B670680BF83E83C88C0" bold="true" box="[151,313,1170,1196]" pageId="1" pageNumber="18">
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<typeStatus id="54AE8805FF924B670680BF83E9D188C0" box="[151,212,1170,1196]" pageId="1" pageNumber="18">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C154D24FF924B670757BF82EB7B88C0" box="[320,638,1171,1196]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="drepanoides" status="sp. nov.">
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<emphasis id="B961EAB5FF924B670757BF82EB7B88C0" box="[320,638,1171,1196]" italics="true" pageId="1" pageNumber="18">Habroblattula drepanoides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25257CEFF924B670492BF83EBE688C0" box="[645,739,1170,1196]" pageId="1" pageNumber="18" rank="species">
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<emphasis id="B961EAB5FF924B670492BF83EBE688C0" bold="true" box="[645,739,1170,1196]" pageId="1" pageNumber="18">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F652CFF924B6706D1BFABEC5F8948" pageId="1" pageNumber="18" type="diagnosis">
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<paragraph id="8BAA36A7FF924B6706D1BFABEC5F8948" blockId="1.[151,1437,1170,2036]" pageId="1" pageNumber="18">
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<emphasis id="B961EAB5FF924B6706D1BFABE83A88B8" bold="true" box="[198,319,1210,1236]" pageId="1" pageNumber="18">Diagnosis</emphasis>
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. Wings, pronotum and legs with bright and dark colour. Forewing Sc terminally branched, CuP strongly curved, all branches strong and dark, intercalaries and cross-veins distinct. Hindwing with pterostigma (dark macula in the R1), R1 and RS differentiated, CuA with abundant secondary branches.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F652CFF924B6706D2BE23EC228AE0" pageId="1" pageNumber="18" type="description">
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<paragraph id="8BAA36A7FF924B6706D2BE23EC228AE0" blockId="1.[151,1437,1170,2036]" pageId="1" pageNumber="18">
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<emphasis id="B961EAB5FF924B6706D2BE23E8588920" bold="true" box="[197,349,1330,1356]" pageId="1" pageNumber="18">Description.</emphasis>
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Forewings with more or less parallel margins and characteristic forewing coloration (with falciform dapple of the R and M area and with macula along with the basal part of the posterior margin). Costal space very short and narrow; Sc terminally branched; R slightly curved, multiple simple branches, ending before wing apex; apically descending M simplified; CuA with branches long, running parallel to margin; CuP strongly curved; Anal veins simple and with thick cross-veins (visible even in poorly preserved specimens). Hindwing with pterostigma (dark macula in the R1). Sc simple; R1 and RS differentiated; M without tertiary branchings; CuA with abundant secondary branches; CuP simple. Apex distinctly colored. Pronotum with anomalous dapple, wider than long. Legs long with bright dark macula, femur shorter than tibia; all tibiae very short, with numerous spurs. Body soft, terminal segments with multisegmented cerci and styli.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F652CFF924B6406D2BD8BEB548D3E" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="discussion">
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<paragraph id="8BAA36A7FF924B6706D2BD8BEC2D8AB0" blockId="1.[151,1437,1170,2036]" pageId="1" pageNumber="18">
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<emphasis id="B961EAB5FF924B6706D2BD8BE8328AD8" bold="true" box="[197,311,1690,1716]" pageId="1" pageNumber="18">Remarks</emphasis>
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.
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<taxonomicName id="4C154D24FF924B670751BD8AE8E88AD8" box="[326,493,1691,1716]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAB5FF924B670751BD8AE8E88AD8" box="[326,493,1691,1716]" italics="true" pageId="1" pageNumber="18">Habroblattula</emphasis>
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</taxonomicName>
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is assigned to the family
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<taxonomicName id="4C154D24FF924B67050ABD8BEAA48AD8" box="[797,929,1690,1716]" class="Insecta" family="Blattulidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="family">Blattulidae</taxonomicName>
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, because the forewing has long Sc, regular venation with distinct intercalaries and hindwing has simple CuP, branched A1. (Vishniakova 1983)
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</paragraph>
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<paragraph id="8BAA36A7FF924B6706D2BDFAEAF48B10" blockId="1.[151,1437,1170,2036]" pageId="1" pageNumber="18">
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<taxonomicName id="4C154D24FF924B6706D2BDFAE8698B68" box="[197,364,1771,1796]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B961EAB5FF924B6706D2BDFAE8698B68" box="[197,364,1771,1796]" italics="true" pageId="1" pageNumber="18">Habroblattula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25257CEFF924B670763BDFBE8E58B68" box="[372,480,1770,1796]" pageId="1" pageNumber="18" rank="genus">
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<taxonomicName id="4C154D24FF924B670763BDFBE8A78B68" box="[372,418,1770,1796]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="gen">gen</taxonomicName>
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. nov.
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</taxonomicNameLabel>
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is closely related to
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<emphasis id="B961EAB5FF924B6704CFBDFAEA338B68" box="[728,822,1771,1796]" italics="true" pageId="1" pageNumber="18">Svabula</emphasis>
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<bibRefCitation id="EF844B56FF924B670528BDFBED008B68" author="Vrsansky" box="[831,1029,1770,1796]" pageId="1" pageNumber="27" refString="VrSansky, P. (2005 b) Lower Cretaceous cockroaches and mantids (Insecta: Blattaria, Mantodea) from the Sharin-Gol in Mongolia. Entomological Problems, 35, 163 - 167." type="journal article" year="2005" yearSuffix="b">Vršanský, 2005b</bibRefCitation>
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from the Berriasian Lower Cretaceous sediments of Sharin-gol in
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<collectingCountry id="F3027637FF924B67043FBC03EB9B8B40" box="[552,670,1810,1836]" name="Mongolia" pageId="1" pageNumber="18">Mongolia</collectingCountry>
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. Synapomorphies include wider forewing with margins parallel, branches strong, straight and dark, clavus with cross-veins.
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<emphasis id="B961EAB5FF924B670576BC2AEABA8B38" box="[865,959,1851,1876]" italics="true" pageId="1" pageNumber="18">Svabula</emphasis>
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differs in small forewing size, different coloured pattern, having less apparent intercalaries and thick cross-veins.
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</paragraph>
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<paragraph id="8BAA36A7FF924B6706D2BC9BEC988B98" blockId="1.[151,1437,1170,2036]" pageId="1" pageNumber="18">
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Hindwing of the new genus resembles
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<emphasis id="B961EAB5FF924B670489BC9AEBEF8BC8" box="[670,746,1931,1956]" italics="true" pageId="1" pageNumber="18">Kridla</emphasis>
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<bibRefCitation id="EF844B56FF924B6704E2BC9BEAB98BC8" author="Vrsansky" box="[757,956,1930,1956]" pageId="1" pageNumber="27" refString="VrSansky, P. (2005 a) A fossil insect in a drilling core sample-cockroach Kridla stastia gen. et sp. nov. (Blattulidae) from the cretaceous of the Verkhne-Bureinskaya Depression in Eastern Russia. Entomological Problems, 35, 115 - 116." type="journal article" year="2005" yearSuffix="a">Vršanský, 2005a</bibRefCitation>
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from the Albian or Cenomanian Upper Cretaceous sediments of the Kyndalskaya Formation in having distinct pterostigma and distinct apex coloration, but differs from the latter by secondarily branched R1, and A2, simple M and more abundant cross-veins.
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</paragraph>
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<paragraph id="8BAA36A7FF914B6406D1BB89ED5A8CB6" blockId="2.[151,1436,152,498]" pageId="2" pageNumber="19">
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The new genus differs from genus
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<taxonomicName id="4C154D24FF914B640477BB89EAA18CDE" authority="Vrsansky, 2003" authorityName="Vrsansky" authorityYear="2003" box="[608,932,152,178]" class="Insecta" family="Blattulidae" genus="Tarakanula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAB5FF914B640477BB89EBE38CDD" box="[608,742,152,177]" italics="true" pageId="2" pageNumber="19">Tarakanula</emphasis>
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<bibRefCitation id="EF844B56FF914B6404FABB89EAA18CDE" author="Vrsansky" box="[749,932,152,178]" pageId="2" pageNumber="27" refString="VrSansky, P. (2003) Unique assemblage of Dictyoptera (Insecta-Blattaria, Mantodea, Isoptera) from the Lower Cretaceous of Bon Tsagaan Nuur in Mongolia. Entomological Problems, 1 - 2, 119 - 151." type="book chapter" year="2003">Vršanský, 2003</bibRefCitation>
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</taxonomicName>
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by prominently coloured wings, pronotum and legs of new genus, with more dense veins and with apparent thick cross-veins.
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</paragraph>
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<paragraph id="8BAA36A7FF914B6406D1BBF9EB548D3E" blockId="2.[151,1436,152,498]" pageId="2" pageNumber="19">
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<taxonomicName id="4C154D24FF914B6406D1BBF9E8688D6D" box="[198,365,232,257]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B961EAB5FF914B6406D1BBF9E8688D6D" box="[198,365,232,257]" italics="true" pageId="2" pageNumber="19">Habroblattula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25257CEFF914B64076DBBF9E8EE8D6E" box="[378,491,232,258]" pageId="2" pageNumber="19" rank="genus">
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<taxonomicName id="4C154D24FF914B64076DBBF9E8AD8D6E" box="[378,424,232,258]" class="Insecta" family="Blattulidae" genus="Habroblattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="gen">gen</taxonomicName>
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. nov.
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</taxonomicNameLabel>
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is easily differentiated from
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<taxonomicName id="4C154D24FF914B64054DBBF9ED618D6E" authority="Giebel, 1856" authorityName="Giebel" authorityYear="1856" box="[858,1124,232,258]" class="Insecta" family="Blattulidae" genus="Elisama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAB5FF914B64054DBBF9EABF8D6D" box="[858,954,232,257]" italics="true" pageId="2" pageNumber="19">Elisama</emphasis>
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Giebel, 1856
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</taxonomicName>
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and
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<taxonomicName id="4C154D24FF914B6402BFBBF9E9D48D46" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" class="Insecta" family="Blattulidae" genus="Blattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAB5FF914B6402BFBBF9EC028D6D" box="[1192,1287,232,257]" italics="true" pageId="2" pageNumber="19">Blattula</emphasis>
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Handlirsch, 1906
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</taxonomicName>
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–1908 by having coloured wings, pronotum and legs, the forewing with strong, dark branches and apparent cross-veins in clavus and Cu area.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F652CFF914B6406D1BA71EB7F8DCE" pageId="2" pageNumber="19" type="description">
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<paragraph id="8BAA36A7FF914B6406D1BA71EB7F8DCE" blockId="2.[151,1436,152,498]" pageId="2" pageNumber="19">
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<emphasis id="B961EAB5FF914B6406D1BA71E8498D15" box="[198,332,352,377]" italics="true" pageId="2" pageNumber="19">Xonpepetla</emphasis>
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Cifuentes-Ruiz et
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<bibRefCitation id="EF844B56FF914B64043EBA71EBE58D16" author="Vrsansky" box="[553,736,352,378]" pageId="2" pageNumber="27" refString="VrSansky, P. & Ansorge, J. (2006) Lower Jurassic cockroaches (Insecta, Blattaria) from Germany and England. African Invertebrates, 47, 27 pp." type="book" year="2006">Vršanský, 2006</bibRefCitation>
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from the Campanian of
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<collectingCountry id="F3027637FF914B640217BA71ED5F8D16" box="[1024,1114,352,378]" name="Mexico" pageId="2" pageNumber="19">Mexico</collectingCountry>
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differ in being robust body with very short and very wide forewings.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C30F652CFF914B6406D1BABEEB6B8D9E" pageId="2" pageNumber="19" type="etymology">
|
||
<paragraph id="8BAA36A7FF914B6406D1BABEEB6B8D9E" blockId="2.[151,1436,152,498]" pageId="2" pageNumber="19">
|
||
<emphasis id="B961EAB5FF914B6406D1BABEE8488DA5" bold="true" box="[198,333,431,457]" pageId="2" pageNumber="19">Etymology</emphasis>
|
||
: The name is derived from the Greek prefix
|
||
<emphasis id="B961EAB5FF914B64057ABAA1EAA18DA5" box="[877,932,432,457]" italics="true" pageId="2" pageNumber="19">habr</emphasis>
|
||
(meaning “elegancy”) and the
|
||
<typeStatus id="54AE8805FF914B64030BBAA1EC4B8DA6" box="[1308,1358,432,458]" pageId="2" pageNumber="19">type</typeStatus>
|
||
genus
|
||
<taxonomicName id="4C154D24FF914B640680BAC9E9F08D9D" box="[151,245,472,497]" class="Insecta" family="Blattulidae" genus="Blattula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961EAB5FF914B640680BAC9E9F08D9D" box="[151,245,472,497]" italics="true" pageId="2" pageNumber="19">Blattula</emphasis>
|
||
</taxonomicName>
|
||
(the
|
||
<typeStatus id="54AE8805FF914B64072EBAC9E8698D9E" box="[313,364,472,498]" pageId="2" pageNumber="19">type</typeStatus>
|
||
genus of this family).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |