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<document id="B66B4492C026BF5D8EB4CC7964D559A0" ID-CLB-Dataset="172793" ID-DOI="10.11646/zootaxa.5200.1.3" ID-GBIF-Dataset="8b762af5-ab4d-4bfd-b985-374e2ecc4cac" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251848" ID-ZooBank="C7E8D8A8-4DEF-4CC4-BBB6-5410DF6713DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666770397033" checkinUser="plazi" docAuthor="Flórez-V, Camilo &amp; Evangelista, Olivia" docDate="2022" docId="03AE60118033667284F1FA6D5A8BFDC8" docLanguage="en" docName="zootaxa.5199.1.3.pdf" docOrigin="Zootaxa 5200 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andinodarnis Flórez-V &amp; Evangelista 2022, gen. nov." docType="treatment" docVersion="6" lastPageNumber="40" masterDocId="FF971869803266718466FF845F17FFDE" masterDocTitle="Andinodarnis gen. nov., a new treehopper genus from the Andes (Hemiptera: Membracidae: Darninae)" masterLastPageNumber="50" masterPageNumber="37" pageNumber="38" updateTime="1699408245093" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFCFDAA515A3D04D599ECEA796363ADF">Andinodarnis gen. nov., a new treehopper genus from the Andes (Hemiptera: Membracidae: Darninae)</mods:title>
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<mods:name id="A07D42391379CAE396A9A6ED0325F03A" type="personal">
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<mods:roleTerm id="95968AA5AC15148145E958103E5DA284">Author</mods:roleTerm>
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<mods:namePart id="3BBD7AA54418C7B218E278109D3F20C0">Flórez-V, Camilo</mods:namePart>
<mods:affiliation id="0486DB7F4C3DE68AFB2002913A18D148">Grupo Biología CES, Colecciones Biológicas de la Universidad CES, Calle 10 A No. 22 - 04, Medellín-Antioquia, Colombia. Department of Entomology, The Pennsylvania State University, 501 ASI Building, University Park, PA, USA, 16802. CSIRO, AustralianNationalInsectCollection, GPOBox 1700, CanberraACT, Australia, 2601; https: // orcid. org / 0000 - 0001 - 8515 - 3548 Corresponding author. kmilofv @ gmail. com; https: // orcid. org / 0000 - 0002 - 6091 - 8150</mods:affiliation>
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<mods:namePart id="ECC8F4EB36D70C1950B6A1F361BCC8C5">Evangelista, Olivia</mods:namePart>
<mods:nameIdentifier id="53C728185A9DBB724DE78A1389898313" type="ORCID">0000-0001-8515-3548</mods:nameIdentifier>
<mods:nameIdentifier id="8CF2C48D001A2119ADEAAA3C00DC62EF" type="email">kmilofv@gmail.com</mods:nameIdentifier>
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<mods:title id="C6829744F724C9B14268007D2FD3F811">Zootaxa</mods:title>
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<mods:part id="1AB5976B554D32ECA6211A1B4D6F9F2B">
<mods:date id="67EC301ECD83790AADE97F2008C5377D">2022</mods:date>
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<mods:number id="B295EE56490DE252013C22E52117650E">2022-10-26</mods:number>
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<mods:detail id="7B43E6302A082C7718C11E67CCD8E9FF" type="volume">
<mods:number id="2987D39D9279C36EDE9C6CB548D42F8A">5200</mods:number>
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<mods:classification id="3DE59FAFE19C2B3E8100671AB0C42AD0">journal article</mods:classification>
<mods:identifier id="790F6D769BBAEA6AD5443A4B7D55A519" type="CLB-Dataset">172793</mods:identifier>
<mods:identifier id="8959F3B8D88D712FCAF8ABC0C1F20433" type="DOI">10.11646/zootaxa.5200.1.3</mods:identifier>
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<treatment id="03AE60118033667284F1FA6D5A8BFDC8" ID-DOI="http://doi.org/10.5281/zenodo.7254989" ID-GBIF-Taxon="203398737" ID-Zenodo-Dep="7254989" LSID="urn:lsid:plazi:treatment:03AE60118033667284F1FA6D5A8BFDC8" httpUri="http://treatment.plazi.org/id/03AE60118033667284F1FA6D5A8BFDC8" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38">
<subSubSection id="C31D828C8033667084F1FA6D5EA4F9DD" box="[151,435,1513,1540]" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="8BB8D1078033667084F1FA6D5EA4F9DD" blockId="1.[151,966,1513,1576]" box="[151,435,1513,1540]" pageId="1" pageNumber="38">
<heading id="D0F0666B8033667084F1FA6D5EA4F9DD" bold="true" box="[151,435,1513,1540]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="B9730D158033667084F1FA6D5EA4F9DD" bold="true" box="[151,435,1513,1540]" pageId="1" pageNumber="38">
<taxonomicName id="4C07AA848033667084F1FA6D5E2AF9DA" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[151,317,1513,1540]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9730D158033667084F1FA6D5E2AF9DA" bold="true" box="[151,317,1513,1540]" italics="true" pageId="1" pageNumber="38">Andinodarnis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240B06E803366708523FA6D5EA4F9DD" box="[325,435,1513,1539]" pageId="1" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31D828C8033667084F1F9895CD1F9F6" box="[151,966,1549,1576]" pageId="1" pageNumber="38" type="description">
<paragraph id="8BB8D1078033667084F1F9895CD1F9F6" blockId="1.[151,966,1513,1576]" box="[151,966,1549,1576]" pageId="1" pageNumber="38">
<uri id="FF96DD058033667084F1F9895CD1F9F6" box="[151,966,1549,1576]" pageId="1" pageNumber="38">
urn:lsid:zoobank.org:act:
<uuid id="FFA1EBD28033667085CCF9895CD1F9F6" box="[426,966,1549,1576]" pageId="1" pageNumber="38">264E9AEB-7003-4D71-A3B7-C85CD1973882</uuid>
</uri>
</paragraph>
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<subSubSection id="C31D828C8033667084F1F9C85D66F9BA" box="[151,625,1612,1636]" pageId="1" pageNumber="38" type="type_taxon">
<paragraph id="8BB8D1078033667084F1F9C85D66F9BA" blockId="1.[151,625,1612,1636]" box="[151,625,1612,1636]" pageId="1" pageNumber="38">
Type-species:
<taxonomicName id="4C07AA8480336670854EF9C85D66F9BA" ID-CoL="57DFH" authority="Stal, 1869" authorityName="Stal" authorityYear="1869" box="[296,625,1612,1636]" class="Insecta" family="Membracidae" genus="Tomogonia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="pectoralis">Tomogonia pectoralis Stål, 1869</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8033667084F1F9095C2BF885" pageId="1" pageNumber="38" type="diagnosis">
<paragraph id="8BB8D1078033667084F1F9095C2BF885" blockId="1.[151,1437,1677,2027]" pageId="1" pageNumber="38">
<emphasis id="B9730D158033667084F1F9095E03F979" bold="true" box="[151,276,1677,1703]" pageId="1" pageNumber="38">Diagnosis:</emphasis>
Head in anterior view subtriangular. Pronotum convex, finely punctate, smooth; suprahumeral horns acute, very close to humeral angles, slightly projected and curved backward in dorsal view; semicircular lateral impressions extending from behind humeral angles to apical third; posterior process extended slightly beyond M fork of forewing. Forewings entirely exposed, not concealed by pronotum. Pro- and mesothoracic femur bearing two rows of cucullate setae on ventral surface; metathoracic femur bearing one row of cucullate setae on ventral surface, fewer and comparatively smaller than other femora.
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8033667384A1F8E15E82FB7F" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="description">
<paragraph id="8BB8D1078033667084A1F8E15FF1F835" blockId="1.[151,1437,1677,2027]" pageId="1" pageNumber="38">
<emphasis id="B9730D158033667084A1F8E15E4CF8A1" bold="true" box="[199,347,1893,1919]" pageId="1" pageNumber="38">Description:</emphasis>
<emphasis id="B9730D15803366708503F8E15EB1F8A1" box="[357,422,1893,1919]" italics="true" pageId="1" pageNumber="38">Color</emphasis>
: Pale to ochraceous yellow, dark brown, or combination thereof; yellow patches bright green in live specimens. Labium, thoracic pleurites and sternites, coxae and base of metathoracic femur dark brown to black. Forewings amber to smoky hyaline with dark macula at apical third. Abdomen ochraceous, yellow to dark brown.
</paragraph>
<paragraph id="8BB8D1078030667384A1FF135BB3FF0B" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FF135E26FF6E" box="[199,305,151,176]" italics="true" pageId="2" pageNumber="39">Vestiture:</emphasis>
Head and pronotum smooth, shiny, and finely punctate. Body glabrous, except thoracic pleurites and sternites, coxae, ventral surface of femora and tibiae covered in golden or silver pubescence.
</paragraph>
<paragraph id="8BB8D1078030667384A1FF5B5EF3FEBB" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FF5B5E1AFF27" box="[199,269,223,249]" italics="true" pageId="2" pageNumber="39">Head:</emphasis>
Subtriangular; posterior margin of vertex, including supra-antennal ledges and frontoclypeus, forming slightly arcuate continuous outline; with feeble semicircular concavity between ocelli; lower half depressed, slightly directed posteriorly, frontoclypeus ovoid to subpentagonal with smooth round edges; dorsal margin arcuate; labium reaching mesothoracic coxae.
</paragraph>
<paragraph id="8BB8D1078030667384A1FEEB5BE3FD2F" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FEEB5E36FE57" box="[199,289,367,393]" italics="true" pageId="2" pageNumber="39">Thorax:</emphasis>
Pronotum convex, low, dorsally arched, moderately compressed laterally behind humeri; suprahumeral horns minute, conical, arising right above humeral angles, curved backward in dorsal view, slightly projected into acute tip; semicircular impressions, one at each side of posterior process, extending from behind humeral angles to apical third; dorsomedial carina inconspicuous anteriorly, gradually compressed and tectiform from mid-dorsum to pronotal apex; posterior process tapering into spine-like apex. Forewings entirely exposed, R initially divided into R
<subScript id="1783D3428030667384B8FDB15E1EFD9D" attach="left" box="[222,265,565,579]" fontSize="6" pageId="2" pageNumber="39">1+2+3</subScript>
and R
<subScript id="1783D342803066738530FDB15E67FD9D" attach="left" box="[342,368,565,579]" fontSize="6" pageId="2" pageNumber="39">4+5</subScript>
, with one
<emphasis id="B9730D1580306673858DFDA75EE1FDE2" box="[491,502,547,572]" italics="true" pageId="2" pageNumber="39">s</emphasis>
, one
<emphasis id="B9730D15803066738651FDA75D48FDE2" box="[567,607,547,572]" italics="true" pageId="2" pageNumber="39">r-m</emphasis>
, and two
<emphasis id="B9730D158030667386B4FDA75C1DFDE2" box="[722,778,547,572]" italics="true" pageId="2" pageNumber="39">m-cu</emphasis>
crossveins; crossvein
<emphasis id="B9730D1580306673806DFDA75B01FDE2" box="[1035,1046,547,572]" italics="true" pageId="2" pageNumber="39">s</emphasis>
slightly distad of
<emphasis id="B9730D15803066738081FDA75A18FDE2" box="[1255,1295,547,572]" italics="true" pageId="2" pageNumber="39">r-m</emphasis>
;
<emphasis id="B9730D15803066738179FDA75A3DFDE2" box="[1311,1322,547,572]" italics="true" pageId="2" pageNumber="39">s</emphasis>
and
<emphasis id="B9730D15803066738102FDA75A8BFDE2" box="[1380,1436,547,572]" italics="true" pageId="2" pageNumber="39">m-cu</emphasis>
delimiting two discoidal cells. Hind wing with one
<emphasis id="B9730D15803066738683FDC35C1AFDBE" box="[741,781,583,608]" italics="true" pageId="2" pageNumber="39">r-m</emphasis>
, and
<emphasis id="B9730D15803066738728FDC35C91FDBE" box="[846,902,583,608]" italics="true" pageId="2" pageNumber="39">m-cu</emphasis>
crossvein almost as long as M
<subScript id="1783D34280306673808FFDDD5A14FDB9" attach="left" box="[1257,1283,601,615]" fontSize="6" pageId="2" pageNumber="39">3+4</subScript>
, and R
<subScript id="1783D342803066738130FDDD5A67FDB9" attach="left" box="[1366,1392,601,615]" fontSize="6" pageId="2" pageNumber="39">4+5</subScript>
not fused with M
<subScript id="1783D34280306673854BFDF95E50FD55" attach="left" box="[301,327,637,651]" fontSize="6" pageId="2" pageNumber="39">1+2</subScript>
. Pro- and mesothoracic femur bearing two rows of cucullate setae on ventral surface (
<figureCitation id="133CCD82803066738175FDEF5A98FD5B" box="[1299,1423,618,645]" captionStart="FIGURE 5" captionStartId="9.[152,255,1790,1814]" captionTargetBox="[151,1436,181,1766]" captionTargetId="figure-20@9.[151,1436,181,1766]" captionTargetPageId="9" captionText="FIGURE 5. Holotype male of Andinodarnis atromeris sp. nov. (AB) Ventrolateral view and close-up of prothoracic femur, (C) Anteroventral view of pro- and mesothoracic femora, (D) Ventral view of pro- and mesothoracic femora, (EG) Metathoracic leg posterior view." figureDoi="http://doi.org/10.5281/zenodo.7251858" httpUri="https://zenodo.org/record/7251858/files/figure.png" pageId="2" pageNumber="39">Fig. 5AD</figureCitation>
), metathoracic femur bearing one row of cucullate setae on ventral surface, each row with few spines (7 or less), much less acute and pronounced in metathoracic femur; well-defined rows of cucullate setae on metathoracic tibia (three single rows: antero-dorsal, postero-dorsal and ventral = I, II and III
<emphasis id="B9730D158030667387E8FD535CDBFD2E" box="[910,972,727,752]" italics="true" pageId="2" pageNumber="39">sensu</emphasis>
<bibRefCitation id="EF96ACF68030667387B5FD535B41FD2F" author="Deitz, L. L." box="[979,1110,726,753]" pageId="2" pageNumber="39" pagination="1 - 177" refId="ref5507" refString="Deitz, L. L. (1975) Classification of the higher categories of the New World treehoppers (Homoptera: Membracidae). North Carolina Agricultural Experiment Station, Technical Bulletin, 225, 1 - 177." type="journal article" year="1975">Deitz, 1975</bibRefCitation>
) (
<figureCitation id="133CCD82803066738008FD535BFFFD2F" box="[1134,1256,726,753]" captionStart="FIGURE 5" captionStartId="9.[152,255,1790,1814]" captionTargetBox="[151,1436,181,1766]" captionTargetId="figure-20@9.[151,1436,181,1766]" captionTargetPageId="9" captionText="FIGURE 5. Holotype male of Andinodarnis atromeris sp. nov. (AB) Ventrolateral view and close-up of prothoracic femur, (C) Anteroventral view of pro- and mesothoracic femora, (D) Ventral view of pro- and mesothoracic femora, (EG) Metathoracic leg posterior view." figureDoi="http://doi.org/10.5281/zenodo.7251858" httpUri="https://zenodo.org/record/7251858/files/figure.png" pageId="2" pageNumber="39">Fig. 5EG</figureCitation>
).
</paragraph>
<paragraph id="8BB8D1078030667384A1FD7F5B92FCCB" blockId="2.[151,1437,151,1365]" box="[199,1157,763,789]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FD7F5E2DFCCB" box="[199,314,763,789]" italics="true" pageId="2" pageNumber="39">Abdomen:</emphasis>
Terga smooth, lacking fenestrae, tuberosities or dorsal scars (
<figureCitation id="133CCD8280306673878BFD7F5B51FCCB" box="[1005,1094,763,789]" captionStart="FIGURE 3" captionStartId="6.[152,255,1677,1701]" captionTargetBox="[151,1436,181,1651]" captionTargetId="figure-22@6.[151,1436,181,1653]" captionTargetPageId="6" captionText="FIGURE 3. Female abdomen and genitalia of Andinodarnis pectoralis comb. nov. (Stål, 1869).Abdomen: (A) dorsal view, (B) ventral view, (C) abdominal sternites and pleurites. Female genitalia: (D) Gonoplac. (E) first valvulae, (F) close-up of apex of first valvulae, (G) second valvulae, (H) close-up of apex of second valvulae." figureDoi="http://doi.org/10.5281/zenodo.7251854" httpUri="https://zenodo.org/record/7251854/files/figure.png" pageId="2" pageNumber="39">Fig. 3A</figureCitation>
,
<figureCitation id="133CCD82803066738034FD7F5B60FCCB" box="[1106,1143,763,789]" captionStart="FIGURE 7" captionStartId="11.[152,255,1329,1353]" captionTargetBox="[151,1436,181,1305]" captionTargetId="figure-20@11.[151,1436,181,1305]" captionTargetPageId="11" captionText="FIGURE 7. Female abdomen and genitalia of Andinodarnis atromeris sp. nov. Abdomen: (A) lateral view. Female genitalia: (B) Gonoplac, (C) first valvulae, (D) close-up of apex of first valvulae, (E) second valvulae, (F) close-up of apex of second valvulae." figureDoi="http://doi.org/10.5281/zenodo.7251864" httpUri="https://zenodo.org/record/7251864/files/figure.png" pageId="2" pageNumber="39">7A</figureCitation>
).
</paragraph>
<paragraph id="8BB8D1078030667384A1FC9B5E44FC5F" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FC9B5E7CFCE7" box="[199,363,799,825]" italics="true" pageId="2" pageNumber="39">Male genitalia</emphasis>
: Subgenital plates somewhat rectangular, elongate, weakly curved in lateral view. Styles slender, hook-shaped, broader at middle. Aedeagus U-shaped, bearing several rows of minute teeth on upper anterior surface of posterior arm.
</paragraph>
<paragraph id="8BB8D1078030667384A1FC0F5B77FBA3" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FC0F5E9FFC7B" box="[199,392,907,933]" italics="true" pageId="2" pageNumber="39">Female genitalia</emphasis>
. Gonoplac round apically in lateral view, narrow basally and abruptly wider at apical half; ventral margin arched, sclerotized, covered in fine punctation and setae. First valvulae in lateral view weakly curved, apex triangular and acute; delicately ornate along dorsal margin, fine lines more or less perpendicular to longitudinal axis; ramus reaching apex of valvulae with pores along entire extension. Second valvulae blade shaped, distinctly more sclerotized than first valvulae, dorsally curved toward triangular apex, finely serrate; apical 1/5 covered in canals and pores; longitudinal ventral axis ornate basally with linear and oblique integumental processes, and apically with escamiform integumental processes; ramus extended almost to apex.
</paragraph>
<paragraph id="8BB8D1078030667384A1FB035E82FB7F" blockId="2.[151,1437,151,1365]" box="[199,405,1159,1185]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FB035E01FB7F" box="[199,278,1159,1185]" italics="true" pageId="2" pageNumber="39">Nymph</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8030667384A1FB2E5C76FAD3" pageId="2" pageNumber="39" type="distribution">
<paragraph id="8BB8D1078030667384A1FB2E5C76FAD3" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FB2E5E76FB1A" bold="true" box="[199,353,1194,1220]" pageId="2" pageNumber="39">Distribution.</emphasis>
<collectingCountry id="F310919780306673850DFB2F5D12FB1B" box="[363,517,1195,1221]" name="Colombia" pageId="2" pageNumber="39">COLOMBIA</collectingCountry>
:
<emphasis id="B9730D15803066738674FB2F5D97FB1B" box="[530,640,1195,1221]" italics="true" pageId="2" pageNumber="39">
<collectingRegion id="49C31FE5803066738674FB2F5D97FB1B" box="[530,640,1195,1221]" country="Colombia" name="Antioquia" pageId="2" pageNumber="39">Antioquia</collectingRegion>
</emphasis>
,
<emphasis id="B9730D158030667386F7FB2F5C14FB1A" box="[657,771,1195,1220]" italics="true" pageId="2" pageNumber="39">
<collectingRegion id="49C31FE58030667386F7FB2F5C14FB1A" box="[657,771,1195,1220]" country="Colombia" name="Putumayo" pageId="2" pageNumber="39">Putumayo</collectingRegion>
</emphasis>
,
<emphasis id="B9730D15803066738772FB2F5C95FB1B" box="[788,898,1195,1221]" italics="true" pageId="2" pageNumber="39">
<collectingRegion id="49C31FE5803066738772FB2F5C95FB1B" box="[788,898,1195,1221]" country="Colombia" name="Risaralda" pageId="2" pageNumber="39">Risaralda</collectingRegion>
</emphasis>
.
<collectingCountry id="F31091978030667387F2FB2F5B37FB1B" box="[916,1056,1195,1221]" name="Ecuador" pageId="2" pageNumber="39">ECUADOR</collectingCountry>
:
<emphasis id="B9730D1580306673804BFB2F5B88FB1B" box="[1069,1183,1195,1221]" italics="true" pageId="2" pageNumber="39">
<collectingRegion id="49C31FE580306673804BFB2F5B88FB1B" box="[1069,1183,1195,1221]" country="Ecuador" name="Pichincha" pageId="2" pageNumber="39">Pichincha</collectingRegion>
</emphasis>
(new record).
<collectingCountry id="F310919780306673812BFB2F5A8EFB1B" box="[1357,1433,1195,1221]" name="Peru" pageId="2" pageNumber="39">PERU</collectingCountry>
:
<emphasis id="B9730D158030667384F1FB4B5FCAFB37" box="[151,221,1231,1257]" italics="true" pageId="2" pageNumber="39">
<collectingRegion id="49C31FE58030667384F1FB4B5FCAFB37" box="[151,221,1231,1257]" country="Peru" name="Cusco" pageId="2" pageNumber="39">Cuzco</collectingRegion>
</emphasis>
. Members of
<taxonomicName id="4C07AA8480306673851FFB4B5D06FB37" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[377,529,1231,1257]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9730D1580306673851FFB4B5D06FB37" box="[377,529,1231,1257]" italics="true" pageId="2" pageNumber="39">Andinodarnis</emphasis>
</taxonomicName>
<emphasis id="B9730D1580306673867FFB4B5D95FB37" bold="true" box="[537,642,1231,1257]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A240B06E80306673867FFB4B5D95FB37" box="[537,642,1231,1257]" pageId="2" pageNumber="39" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are endemic to the Andean region. Recent specimens were collected in montane habitats ranging between 1800 and
<quantity id="4CFF7CE28030667386EAFB775C17FAD3" box="[652,768,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="39" unit="m" value="3000.0">3000 masl</quantity>
(
<figureCitation id="133CCD82803066738769FB775C43FAD3" box="[783,852,1267,1293]" captionStart="FIGURE 8" captionStartId="12.[152,255,1458,1482]" captionTargetBox="[165,1433,191,1426]" captionTargetId="figure-22@12.[155,1433,181,1434]" captionTargetPageId="12" captionText="FIGURE 8. Distribution map of Andinodarnis pectoralis comb. nov. (Stål, 1869) and A. atromeris sp. nov. (A) Belmira (Colombia: Antioquia), (B) PNN Tatamá (Colombia: Risaralda: Pueblo Rico), (C) San Rafael—Sibundoy (Colombia: Putumayo)." figureDoi="http://doi.org/10.5281/zenodo.7251866" httpUri="https://zenodo.org/record/7251866/files/figure.png" pageId="2" pageNumber="39">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8030667384A1FA935B68FA8B" pageId="2" pageNumber="39" type="etymology">
<paragraph id="8BB8D1078030667384A1FA935B68FA8B" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1FA935E59FAEF" bold="true" box="[199,334,1303,1329]" pageId="2" pageNumber="39">Etymology.</emphasis>
The generic name is composed of two words:
<emphasis id="B9730D15803066738707FA935CADFAEF" box="[865,954,1303,1329]" italics="true" pageId="2" pageNumber="39">andinus</emphasis>
, meaning from the Andes, and
<taxonomicName id="4C07AA84803066738158FA935A9BFAEE" authorityName="Fabricius" authorityYear="1803" box="[1342,1420,1303,1328]" class="Insecta" family="Membracidae" genus="Darnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9730D15803066738158FA935A9BFAEE" box="[1342,1420,1303,1328]" italics="true" pageId="2" pageNumber="39">Darnis</emphasis>
</taxonomicName>
, a genus in the same tribe that is somewhat similar in appearance. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8030667384F1FA065EBAFA42" box="[151,429,1410,1436]" pageId="2" pageNumber="39" type="multiple">
<paragraph id="8BB8D1078030667384F1FA065EBAFA42" blockId="2.[151,429,1410,1436]" box="[151,429,1410,1436]" pageId="2" pageNumber="39">
<heading id="D0F0666B8030667384F1FA065EBAFA42" bold="true" box="[151,429,1410,1436]" fontSize="11" level="1" pageId="2" pageNumber="39" reason="1">
<emphasis id="B9730D158030667384F1FA065EBAFA42" bold="true" box="[151,429,1410,1436]" pageId="2" pageNumber="39">Constituent species (2):</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8030667384F1FA415C3FF9DD" pageId="2" pageNumber="39" type="synonymic_list">
<paragraph id="8BB8D1078030667384F1FA415D08FA01" blockId="2.[151,808,1477,1539]" box="[151,543,1477,1504]" pageId="2" pageNumber="39">
<taxonomicName id="4C07AA848030667384F1FA415D08FA01" authority="Stal, 1869" authorityName="Stal" authorityYear="1869" box="[151,543,1477,1504]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="B9730D158030667384F1FA415EB0FA01" box="[151,423,1477,1503]" italics="true" pageId="2" pageNumber="39">Andinodarnis pectoralis</emphasis>
Stål, 1869
</taxonomicName>
</paragraph>
<paragraph id="8BB8D1078030667384F1FA6D5C3FF9DD" blockId="2.[151,808,1477,1539]" box="[151,808,1513,1539]" pageId="2" pageNumber="39">
<taxonomicName id="4C07AA848030667384F1FA6D5EB4F9DD" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[151,419,1513,1539]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="atromeris">
<emphasis id="B9730D158030667384F1FA6D5EB4F9DD" box="[151,419,1513,1539]" italics="true" pageId="2" pageNumber="39">Andinodarnis atromeris</emphasis>
</taxonomicName>
Flórez-V and Evangelista
<emphasis id="B9730D158030667386A8FA6D5C3FF9DD" bold="true" box="[718,808,1513,1539]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A240B06E8030667386A8FA6D5C3FF9DD" box="[718,808,1513,1539]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8030667284A1F9A85A99FEC3" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39" type="discussion">
<paragraph id="8BB8D1078030667384A1F9A85E7CF908" blockId="2.[151,1437,1580,2039]" pageId="2" pageNumber="39">
<emphasis id="B9730D158030667384A1F9A85E2CF998" bold="true" box="[199,315,1580,1606]" pageId="2" pageNumber="39">Remarks.</emphasis>
Representatives of
<taxonomicName id="4C07AA84803066738678F9A85DA1F998" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[542,694,1580,1606]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9730D15803066738678F9A85DA1F998" box="[542,694,1580,1606]" italics="true" pageId="2" pageNumber="39">Andinodarnis</emphasis>
</taxonomicName>
<emphasis id="B9730D158030667386D9F9A85C3DF998" bold="true" box="[703,810,1580,1606]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A240B06E8030667386D9F9A85C3DF998" box="[703,810,1580,1606]" pageId="2" pageNumber="39" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are infrequently collected and scarce in entomological collections. Several collecting events targeting this group yielded only a few specimens (six new exemplars in four collecting sites). Current records indicate that
<taxonomicName id="4C07AA848030667386F8F9F05C39F950" authorityName="Stal" authorityYear="1869" box="[670,814,1652,1678]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="B9730D158030667386F8F9F05C39F950" box="[670,814,1652,1678]" italics="true" pageId="2" pageNumber="39">A. pectoralis</emphasis>
</taxonomicName>
occurs at mid to high elevation along the Northern and Central Andes (
<quantity id="4CFF7CE2803066738523F91C5D12F96C" box="[325,517,1688,1714]" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="3.0" metricValueMin="1.8" pageId="2" pageNumber="39" unit="m" value="2400.0" valueMax="3000.0" valueMin="1800.0">18003000 masl</quantity>
), while
<taxonomicName id="4C07AA84803066738638F91C5DFEF96C" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[606,745,1688,1714]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="atromeris" status="sp. nov.">
<emphasis id="B9730D15803066738638F91C5DFEF96C" box="[606,745,1688,1714]" italics="true" pageId="2" pageNumber="39">A. atromeris</emphasis>
</taxonomicName>
<emphasis id="B9730D15803066738696F91C5C5DF96C" bold="true" box="[752,842,1688,1714]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A240B06E803066738696F91C5C5DF96C" box="[752,842,1688,1714]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is restricted to the western area of Colombian Andes (
<quantity id="4CFF7CE28030667384F9F9385E77F908" box="[159,352,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="2.125" metricValueMax="2.45" metricValueMin="1.8" pageId="2" pageNumber="39" unit="m" value="2125.0" valueMax="2450.0" valueMin="1800.0">18002450 masl</quantity>
).
</paragraph>
<paragraph id="8BB8D1078030667284A1F9645A99FEC3" blockId="2.[151,1437,1580,2039]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
The genus can be unambiguously ascribed to
<taxonomicName id="4C07AA848030667386B9F9645C22F924" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[735,821,1760,1786]" class="Insecta" family="Membracidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="tribe" tribe="Darnini">Darnini</taxonomicName>
based on the femoral cucullate setae (
<figureCitation id="133CCD8280306673809EF9645A57F925" box="[1272,1344,1760,1787]" captionStart="FIGURE 5" captionStartId="9.[152,255,1790,1814]" captionTargetBox="[151,1436,181,1766]" captionTargetId="figure-20@9.[151,1436,181,1766]" captionTargetPageId="9" captionText="FIGURE 5. Holotype male of Andinodarnis atromeris sp. nov. (AB) Ventrolateral view and close-up of prothoracic femur, (C) Anteroventral view of pro- and mesothoracic femora, (D) Ventral view of pro- and mesothoracic femora, (EG) Metathoracic leg posterior view." figureDoi="http://doi.org/10.5281/zenodo.7251858" httpUri="https://zenodo.org/record/7251858/files/figure.png" pageId="2" pageNumber="39">Fig. 5</figureCitation>
). Tribal constituents in the clade with the same pronotal syndrome lack suprahumeral horns and have forewings partially to entirely concealed by the posterior process, in addition to more pronounced and numerous femoral cucullate setae.
<taxonomicName id="4C07AA848030667384F1F8C85E38F8B8" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[151,303,1868,1894]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9730D158030667384F1F8C85E38F8B8" box="[151,303,1868,1894]" italics="true" pageId="2" pageNumber="39">Andinodarnis</emphasis>
</taxonomicName>
<emphasis id="B9730D15803066738551F8C85EB7F8B8" bold="true" box="[311,416,1868,1894]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A240B06E803066738551F8C85EB7F8B8" box="[311,416,1868,1894]" pageId="2" pageNumber="39" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is, therefore, the only genus in the raindrop group (dewdrop
<emphasis id="B9730D15803066738006F8C95B89F8B8" box="[1120,1182,1869,1894]" italics="true" pageId="2" pageNumber="39">sensu</emphasis>
<bibRefCitation id="EF96ACF68030667380C3F8C95A4BF8B8" author="Roy, L. &amp; Guilbert, E. &amp; Bourgoin, T." box="[1189,1372,1868,1895]" pageId="2" pageNumber="39" pagination="273 - 288" refId="ref6211" refString="Roy, L., Guilbert, E. &amp; Bourgoin, T. (2007) Phylogenetic patterns of mimicry strategies in Darnini (Hemiptera: Membracidae). Annales de La Societe Entomologique de France, 43 (3), 273 - 288. https: // doi. org / 10.1080 / 00379271.2007.10697523" type="journal article" year="2007">
Roy
<emphasis id="B9730D158030667380BDF8C95A03F8B8" box="[1243,1300,1868,1894]" italics="true" pageId="2" pageNumber="39">et al.</emphasis>
, 2007
</bibRefCitation>
) with fully exposed wings and suprahumeral horns (
<figureCitation id="133CCD828030667386F9F8F45DF8F854" box="[671,751,1904,1930]" captionStart="FIGURE 2" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[163,1424,181,1919]" captionTargetId="figure-20@5.[163,1424,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Andinodarnis pectoralis comb. nov. (Stål, 1869). (AD) Female from Colombia in frontal, dorsal, lateral, and frontolateral view, respectively. (EG) Female from Peru in frontal, dorsal and lateral view, respectively." figureDoi="http://doi.org/10.5281/zenodo.7251852" httpUri="https://zenodo.org/record/7251852/files/figure.png" pageId="2" pageNumber="39">Figs. 2</figureCitation>
,
<figureCitation id="133CCD8280306673869DF8F45C1BF854" box="[763,780,1904,1930]" captionStart="FIGURE 4" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[153,1434,181,1919]" captionTargetId="figure-22@8.[153,1434,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Andinodarnis atromeris sp. nov. (AD) Holotype male before DNA extraction in frontal, dorsal, lateral, and frontolateral view, respectively. (EH) Holotype male after DNA extraction in frontal, dorsal, lateral, and frontolateral view, respectively." figureDoi="http://doi.org/10.5281/zenodo.7251856" httpUri="https://zenodo.org/record/7251856/files/figure.png" pageId="2" pageNumber="39">4</figureCitation>
). These two traits are also observed in
<taxonomicName id="4C07AA848030667380A2F8F45A2EF854" authorityName="Kirkaldy" authorityYear="1904" box="[1220,1337,1904,1930]" class="Insecta" family="Membracidae" genus="Sundarion" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9730D158030667380A2F8F45A2EF854" box="[1220,1337,1904,1930]" italics="true" pageId="2" pageNumber="39">Sundarion</emphasis>
</taxonomicName>
, a genus in the tribe outside the raindrop group, which differs in having much larger, strongly projected suprahumeral horns, which are prismatic and often truncate apically. A phylogenetic framework is required to verify whether these topologically similar structures are homologous, considering stark differences in pronotal shape and vestiture. In
<taxonomicName id="4C07AA848031667284F1FF135E38FF6F" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[151,303,151,177]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9730D158031667284F1FF135E38FF6F" box="[151,303,151,177]" italics="true" pageId="3" pageNumber="40">Andinodarnis</emphasis>
</taxonomicName>
<emphasis id="B9730D15803166728551FF135EB6FF6F" bold="true" box="[311,417,151,177]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A240B06E803166728551FF135EB6FF6F" box="[311,417,151,177]" pageId="3" pageNumber="40" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, the suprahumeral horns are conical, minute, lacking a longitudinal carina, and the surface of the pronotum is ultra-smooth, shiny, and finely punctured. Additional distinguishing features of
<taxonomicName id="4C07AA84803166728162FF3F5A8BFF0B" authority="Flórez-V &amp; Evangelista, 2022" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[1284,1436,187,213]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9730D15803166728162FF3F5A8BFF0B" box="[1284,1436,187,213]" italics="true" pageId="3" pageNumber="40">Andinodarnis</emphasis>
</taxonomicName>
<emphasis id="B9730D158031667284F1FF5B5E17FF27" bold="true" box="[151,256,223,249]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A240B06E8031667284F1FF5B5E17FF27" box="[151,256,223,249]" pageId="3" pageNumber="40" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
include a triangular head, femora with few cucullate setae on the ventral surface (rows with seven spines or less), and a widely curved upward female second valvulae with one or two prominent teeth apically (
<figureCitation id="133CCD82803166728146FE875A74FEC3" box="[1312,1379,259,285]" captionStart="FIGURE 3" captionStartId="6.[152,255,1677,1701]" captionTargetBox="[151,1436,181,1651]" captionTargetId="figure-22@6.[151,1436,181,1653]" captionTargetPageId="6" captionText="FIGURE 3. Female abdomen and genitalia of Andinodarnis pectoralis comb. nov. (Stål, 1869).Abdomen: (A) dorsal view, (B) ventral view, (C) abdominal sternites and pleurites. Female genitalia: (D) Gonoplac. (E) first valvulae, (F) close-up of apex of first valvulae, (G) second valvulae, (H) close-up of apex of second valvulae." figureDoi="http://doi.org/10.5281/zenodo.7251854" httpUri="https://zenodo.org/record/7251854/files/figure.png" pageId="3" pageNumber="40">Fig. 3</figureCitation>
,
<figureCitation id="133CCD82803166728116FE875A97FEC3" box="[1392,1408,259,285]" captionStart="FIGURE 7" captionStartId="11.[152,255,1329,1353]" captionTargetBox="[151,1436,181,1305]" captionTargetId="figure-20@11.[151,1436,181,1305]" captionTargetPageId="11" captionText="FIGURE 7. Female abdomen and genitalia of Andinodarnis atromeris sp. nov. Abdomen: (A) lateral view. Female genitalia: (B) Gonoplac, (C) first valvulae, (D) close-up of apex of first valvulae, (E) second valvulae, (F) close-up of apex of second valvulae." figureDoi="http://doi.org/10.5281/zenodo.7251864" httpUri="https://zenodo.org/record/7251864/files/figure.png" pageId="3" pageNumber="40">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8031667284F1FEE05EE3FEA0" box="[151,500,355,382]" pageId="3" pageNumber="40" type="multiple">
<paragraph id="8BB8D1078031667284F1FEE05EE3FEA0" blockId="3.[151,500,355,382]" box="[151,500,355,382]" pageId="3" pageNumber="40">
<heading id="D0F0666B8031667284F1FEE05EE3FEA0" bold="true" box="[151,500,355,382]" fontSize="11" level="1" pageId="3" pageNumber="40" reason="1">
<emphasis id="B9730D158031667284F1FEE05EE3FEA0" bold="true" box="[151,500,355,382]" pageId="3" pageNumber="40">
Key to
<taxonomicName id="4C07AA84803166728497FEE75EE3FEA0" box="[241,500,355,382]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B9730D15803166728497FEE75E80FEA0" bold="true" box="[241,407,355,382]" italics="true" pageId="3" pageNumber="40">Andinodarnis</emphasis>
species
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D828C8031667284F1FE2D5A8BFDC8" pageId="3" pageNumber="40" type="key">
<keyStep id="30F3C9A28031667284F1FE2D5A8BFDC8" pageId="3" pageNumber="40">
<key id="FF9686368031667284F1FE2D5A8BFDC8" pageId="3" pageNumber="40">
<paragraph id="8BB8D1078031667284F1FE2D5A8BFE02" blockId="3.[151,1436,425,535]" pageId="3" pageNumber="40">
<keyLead id="30F672328031667284F1FE2D5A8BFE02" pageId="3" pageNumber="40">
1. Head, pronotum and legs pale to ochraceous yellow, forewings amber hyaline, highest point of dorsal outline past mid-dorsum (
<figureCitation id="133CCD8280316672848FFE425E34FE05" box="[233,291,454,476]" captionStart="FIGURE 2" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[163,1424,181,1919]" captionTargetId="figure-20@5.[163,1424,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Andinodarnis pectoralis comb. nov. (Stål, 1869). (AD) Female from Colombia in frontal, dorsal, lateral, and frontolateral view, respectively. (EG) Female from Peru in frontal, dorsal and lateral view, respectively." figureDoi="http://doi.org/10.5281/zenodo.7251852" httpUri="https://zenodo.org/record/7251852/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
)..........................................
<taxonomicName id="4C07AA84803166728740FE435CDBFE02" authority="Stal" authorityName="Stal" authorityYear="2022" box="[806,972,454,476]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="pectoralis" status="comb. nov.">
<emphasis id="B9730D15803166728740FE435CB6FE05" box="[806,929,454,476]" italics="true" pageId="3" pageNumber="40">A. pectoralis</emphasis>
Stål
</taxonomicName>
<emphasis id="B9730D158031667287B4FE425B57FE05" bold="true" box="[978,1088,454,475]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A240B06E8031667287B4FE425B57FE05" box="[978,1088,454,475]" pageId="3" pageNumber="40" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(=
<taxonomicName id="4C07AA8480316672803AFE435A82FE02" ID-CoL="57DFH" authority="Stal, 1869" authorityName="Stal" authorityYear="1869" box="[1116,1429,454,476]" class="Insecta" family="Membracidae" genus="Tomogonia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="B9730D1580316672803AFE435A3AFE05" box="[1116,1325,454,476]" italics="true" pageId="3" pageNumber="40">Tomogonia pectoralis</emphasis>
Stål, 1869
</taxonomicName>
)
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</paragraph>
<paragraph id="8BB8D1078031667284F1FE605A8BFDC8" blockId="3.[151,1436,425,535]" pageId="3" pageNumber="40">
<keyLead id="30F672328031667284F1FE605A8BFDC8" pageId="3" pageNumber="40">
1. Head and metopidium dark brown, posterior process yellow from humeral angles to apex, forewings smoky hyaline, highest point of dorsal outline approximately at mid-dorsum (
<figureCitation id="133CCD82803166728681FD855C36FDC8" box="[743,801,513,535]" captionStart="FIGURE 4" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[153,1434,181,1919]" captionTargetId="figure-22@8.[153,1434,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Andinodarnis atromeris sp. nov. (AD) Holotype male before DNA extraction in frontal, dorsal, lateral, and frontolateral view, respectively. (EH) Holotype male after DNA extraction in frontal, dorsal, lateral, and frontolateral view, respectively." figureDoi="http://doi.org/10.5281/zenodo.7251856" httpUri="https://zenodo.org/record/7251856/files/figure.png" pageId="3" pageNumber="40">Fig. 4</figureCitation>
)..............
<taxonomicName id="4C07AA848031667287BEFD855B59FDC9" authorityName="Flórez-V &amp; Evangelista" authorityYear="2022" box="[984,1102,513,535]" class="Insecta" family="Membracidae" genus="Andinodarnis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="atromeris">
<emphasis id="B9730D158031667287BEFD855B59FDC9" box="[984,1102,513,535]" italics="true" pageId="3" pageNumber="40">A. atromeris</emphasis>
</taxonomicName>
Flórez-V and Evangelista
<emphasis id="B9730D15803166728136FD855A8BFDC8" bold="true" box="[1360,1436,513,534]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A240B06E803166728136FD855A8BFDC8" box="[1360,1436,513,534]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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