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<document id="4CBBAD178348F635EA2C9AE0904F1139" ID-DOI="10.5281/zenodo.195391" ID-GBIF-Dataset="369a933b-24a3-49ce-a98c-7c1bb931a178" ID-ISSN="1175-5326" ID-Zenodo-Dep="195391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226407764" checkinUser="plazi" docAuthor="Lindquist, Evert E. &amp; Moraza, Maria L." docDate="2010" docId="03ACF95E270D3D4EFF6DFF74FC949705" docLanguage="en" docName="zt02479p021.pdf" docOrigin="Zootaxa 2479" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Opilioseius grallator Lindquist &amp; Moraza, 2010, new species" docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="FF95812627033D5FFFFAFFE3FFF69522" masterDocTitle="Revised diagnosis of the family Blattisociidae (Acari: Mesostigmata: Phytoseioidea), with a key to its genera and description of a new fungus- inhabiting genus from Costa Rica" masterLastPageNumber="21" masterPageNumber="1" pageNumber="15" updateTime="1698238420118" updateUser="plazi">
<mods:mods id="B653C630E098DD7ABEB477F0FEA7C66D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="643E1C7F99F5BB3C2BE5CCE665A48A2E">
<mods:title id="9D329BE0928BAE43D6B659C9166ACDCE">Revised diagnosis of the family Blattisociidae (Acari: Mesostigmata: Phytoseioidea), with a key to its genera and description of a new fungus- inhabiting genus from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="535E5EF9C584239619465B0AE23C2F67" type="personal">
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<mods:namePart id="2F0E0B347755A5A81D272B60A8FA90DD">Lindquist, Evert E.</mods:namePart>
</mods:name>
<mods:name id="F809E431F84E85E3CA8039EB47B1C33B" type="personal">
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<mods:namePart id="C87CC0EAACA30508DE48328C768921E8">Moraza, Maria L.</mods:namePart>
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<mods:typeOfResource id="BE20B60D39B5A13EE5A6BEE1C7BE1C7A">text</mods:typeOfResource>
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<mods:part id="36DF4AA132023492817866C28C77EBBD">
<mods:date id="49A2361CCDE4CA5FB098E772CE664E0A">2010</mods:date>
<mods:detail id="9AC2077E910E9F09022038FC54F387A6" type="volume">
<mods:number id="EE4BEB833F4230D486D4F558ED849457">2479</mods:number>
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<mods:classification id="96230035E64E848319577FAB03D880DF">journal article</mods:classification>
<mods:identifier id="19C2729AB2B9DC849F66C64A2A1E476B" type="DOI">10.5281/zenodo.195391</mods:identifier>
<mods:identifier id="6EFE370BE277A9BE02684DF8C2A1C5C9" type="GBIF-Dataset">369a933b-24a3-49ce-a98c-7c1bb931a178</mods:identifier>
<mods:identifier id="FE9A112EFA5D0DC088C1260DA715ABFF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="952BD78F9B2895F2ECAEBFE2A212492D" type="Zenodo-Dep">195391</mods:identifier>
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<treatment id="03ACF95E270D3D4EFF6DFF74FC949705" ID-DOI="http://doi.org/10.5281/zenodo.6198579" ID-GBIF-Taxon="119386428" ID-Zenodo-Dep="6198579" LSID="urn:lsid:plazi:treatment:03ACF95E270D3D4EFF6DFF74FC949705" httpUri="http://treatment.plazi.org/id/03ACF95E270D3D4EFF6DFF74FC949705" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="C31F1BC3270D3D51FF6DFF74FE3895F5" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BBA4848270D3D51FF6DFF74FDD09593" blockId="14.[151,550,151,215]" box="[151,550,151,177]" pageId="14" pageNumber="15">
<heading id="D0F2FF24270D3D51FF6DFF74FDD09593" bold="true" box="[151,550,151,177]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B971945A270D3D51FF6DFF74FDD09593" bold="true" box="[151,550,151,177]" pageId="14" pageNumber="15">
<taxonomicName id="4C0533CB270D3D51FF6DFF74FE799593" ID-CoL="9BDJL" box="[151,399,151,177]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
<emphasis id="B971945A270D3D51FF6DFF74FE799593" bold="true" box="[151,399,151,177]" italics="true" pageId="14" pageNumber="15">Opilioseius grallator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2422921270D3D51FE6CFF74FDD09593" box="[406,550,151,177]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA4848270D3D51FF6DFF5EFE3895F5" blockId="14.[151,550,151,215]" box="[151,462,189,215]" pageId="14" pageNumber="15">
(
<figureCitation id="133E54CD270D3D51FF65FF5EFEE195F5" box="[159,279,189,215]" captionStart-0="FIGURES 1 – 11" captionStart-1="FIGURES 12 – 13" captionStart-2="FIGURES 14 – 16" captionStart-3="FIGURES 17 – 22" captionStart-4="FIGURES 23 – 31. 23 – 29" captionStartId-0="4.[151,269,1877,1901]" captionStartId-1="5.[151,272,1979,2003]" captionStartId-2="6.[151,270,1764,1788]" captionStartId-3="8.[151,269,1841,1865]" captionStartId-4="9.[151,271,1803,1827]" captionTargetBox-0="[157,1435,197,1824]" captionTargetBox-1="[157,1436,500,1919]" captionTargetBox-2="[154,1435,194,1763]" captionTargetBox-3="[150,1436,197,1772]" captionTargetBox-4="[163,1436,194,1764]" captionTargetId-0="figure@4.[157,1442,197,1853]" captionTargetId-1="figure@5.[153,1438,494,1958]" captionTargetId-2="figure@6.[154,1439,194,1789]" captionTargetId-3="figure@8.[132,1455,197,1935]" captionTargetId-4="figure@9.[163,1448,194,1782]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." captionText-1="FIGURES 12 – 13. Opilioseius grallator n. sp., adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted." captionText-2="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." captionText-3="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." captionText-4="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri-0="https://zenodo.org/record/195392/files/figure.png" httpUri-1="https://zenodo.org/record/195393/files/figure.png" httpUri-2="https://zenodo.org/record/195394/files/figure.png" httpUri-3="https://zenodo.org/record/195395/files/figure.png" httpUri-4="https://zenodo.org/record/195396/files/figure.png" pageId="14" pageNumber="15">Figs 1–29</figureCitation>
,
<figureCitation id="133E54CD270D3D51FED9FF5EFE9995F5" box="[291,367,189,215]" captionStart-0="FIGURES 32 – 33" captionStart-1="FIGURES 34 – 38. 34 – 35" captionStartId-0="10.[151,269,1980,2004]" captionStartId-1="11.[151,270,1817,1841]" captionTargetBox-0="[154,1436,642,1930]" captionTargetBox-1="[157,1436,204,1770]" captionTargetId-0="figure@10.[154,1439,642,1958]" captionTargetId-1="figure@11.[156,1441,197,1794]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 32 – 33. Opilioseius grallator n. sp., adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view." captionText-1="FIGURES 34 – 38. 34 – 35, Opilioseius grallator n. sp., deutonymph: 34, idiosoma, dorsal aspect; 35, idiosoma, ventral aspect. 36 – 38, leg III, tarsal apex and pretarsus, showing different forms of ventroapical tarsal processes and conjectured acrotarsal demarcation: 36, Adenoseius sp. (Sejidae), anterolateral view; 37, Euzercon sp. (Euzerconidae), ventral view; 38, Arctacarus rostratus Evans (Arctacaridae), ventral view. Abbreviation: acr mar, acrotarsal margin." httpUri-0="https://zenodo.org/record/195397/files/figure.png" httpUri-1="https://zenodo.org/record/195398/files/figure.png" pageId="14" pageNumber="15">32–35</figureCitation>
,
<figureCitation id="133E54CD270D3D51FE81FF5EFE3095F5" box="[379,454,189,215]" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="14" pageNumber="15">39, 40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31F1BC3270D3D51FF6DFEE4FADF94B0" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BBA4848270D3D51FF6DFEE4FADF94B0" blockId="14.[151,1437,263,2007]" pageId="14" pageNumber="15">
<emphasis id="B971945A270D3D51FF6DFEE4FEEE9403" bold="true" box="[151,280,263,289]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
With the attributes of the genus. In addition, dorsal shield with setae
<emphasis id="B971945A270D3D51FBA3FEEBFB859403" box="[1113,1139,264,289]" italics="true" pageId="14" pageNumber="15">z2</emphasis>
,
<emphasis id="B971945A270D3D51FB79FEEBFB689403" box="[1155,1182,264,289]" italics="true" pageId="14" pageNumber="15">z5</emphasis>
,
<emphasis id="B971945A270D3D51FB57FEEBFB339403" box="[1197,1221,264,289]" italics="true" pageId="14" pageNumber="15">j6</emphasis>
nearly as short as clunal setae
<emphasis id="B971945A270D3D51FED6FECEFEBF9464" box="[300,329,301,326]" italics="true" pageId="14" pageNumber="15">J5</emphasis>
, with
<emphasis id="B971945A270D3D51FE6CFECEFE479464" box="[406,433,301,326]" italics="true" pageId="14" pageNumber="15">s5</emphasis>
the longest, over half as long as dorsal shield, and with
<emphasis id="B971945A270D3D51FBA6FECEFB8D9464" box="[1116,1147,301,326]" italics="true" pageId="14" pageNumber="15">Z3</emphasis>
and
<emphasis id="B971945A270D3D51FB4DFECEFB219464" box="[1207,1239,301,326]" italics="true" pageId="14" pageNumber="15">Z4</emphasis>
subequally next longest, slightly longer than
<emphasis id="B971945A270D3D51FE1CFEB1FDF39449" box="[486,517,338,363]" italics="true" pageId="14" pageNumber="15">Z5</emphasis>
; peritremes reaching only to level of setae
<emphasis id="B971945A270D3D51FBFAFEB1FBEC9449" box="[1024,1050,338,363]" italics="true" pageId="14" pageNumber="15">z4</emphasis>
; coxa II with seta
<emphasis id="B971945A270D3D51FB0EFEB1FAE69449" box="[1268,1296,338,363]" italics="true" pageId="14" pageNumber="15">av</emphasis>
minute; leg IV with greatly elongated setae on genu (
<emphasis id="B971945A270D3D51FD81FE9BFD4794B3" box="[635,689,376,402]" italics="true" pageId="14" pageNumber="15">pd -1</emphasis>
), tibia (
<emphasis id="B971945A270D3D51FCF4FE9BFCB294B3" box="[782,836,376,402]" italics="true" pageId="14" pageNumber="15">ad -1</emphasis>
), basitarsus (
<emphasis id="B971945A270D3D51FC1BFE9BFBE194B3" box="[993,1047,376,402]" italics="true" pageId="14" pageNumber="15">ad -4</emphasis>
), and telotarsus (
<emphasis id="B971945A270D3D51FB18FE9BFAEE94B3" box="[1250,1304,376,402]" italics="true" pageId="14" pageNumber="15">ad -3</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F1BC3270D3D4FFF3CFE7EFB539705" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="8BBA4848270D3D51FF3CFE7EFC97979E" blockId="14.[151,1437,263,2007]" pageId="14" pageNumber="15">
<emphasis id="B971945A270D3D51FF3CFE7EFDD99495" bold="true" box="[198,559,413,439]" pageId="14" pageNumber="15">Description. ADULT FEMALE.</emphasis>
Dorsal shield 290-350 long, 165–200 at its greatest width at level of setae
<emphasis id="B971945A270D3D51FF6DFE21FF4494F9" box="[151,178,450,475]" italics="true" pageId="14" pageNumber="15">r3</emphasis>
; shield smooth, unornamented over entire surface (
<figureCitation id="133E54CD270D3D51FCD9FE21FC8B94FE" box="[803,893,450,476]" captionStart="FIGURES 12 – 13" captionStartId="5.[151,272,1979,2003]" captionTargetBox="[157,1436,500,1919]" captionTargetId="figure@5.[153,1438,494,1958]" captionTargetPageId="5" captionText="FIGURES 12 – 13. Opilioseius grallator n. sp., adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/195393/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
); with 23 pairs of attenuated, mostly smooth setae, except
<emphasis id="B971945A270D3D51FEC9FE0BFEB99723" box="[307,335,488,513]" italics="true" pageId="14" pageNumber="15">J5</emphasis>
and few of longest setae sparsely, almost imperceptibly barbed basally. Dorsal setae of greatly different lengths,
<emphasis id="B971945A270D3D51FE93FDEEFE739704" box="[361,389,525,550]" italics="true" pageId="14" pageNumber="15">J5</emphasis>
shortest (4–6),
<emphasis id="B971945A270D3D51FDBBFDEEFDAD9704" box="[577,603,525,550]" italics="true" pageId="14" pageNumber="15">z2</emphasis>
,
<emphasis id="B971945A270D3D51FD91FDEEFD739704" box="[619,645,525,550]" italics="true" pageId="14" pageNumber="15">z5</emphasis>
,
<emphasis id="B971945A270D3D51FD6EFDEEFD5A9704" box="[660,684,525,550]" italics="true" pageId="14" pageNumber="15">j6</emphasis>
nearly as short (7–12),
<emphasis id="B971945A270D3D51FC32FDEEFC299704" box="[968,991,525,550]" italics="true" pageId="14" pageNumber="15">j1</emphasis>
moderately short (25–30),
<emphasis id="B971945A270D3D51FADEFDEEFAC89704" box="[1316,1342,525,550]" italics="true" pageId="14" pageNumber="15">s5</emphasis>
longest (193–220),
<emphasis id="B971945A270D3D51FEDAFDD1FEC99769" box="[288,319,562,587]" italics="true" pageId="14" pageNumber="15">Z3</emphasis>
(125–145) and
<emphasis id="B971945A270D3D51FE03FDD1FDEF9769" box="[505,537,562,587]" italics="true" pageId="14" pageNumber="15">Z4</emphasis>
(125–138) subequally next longest, followed by
<emphasis id="B971945A270D3D51FBA3FDD1FB819769" box="[1113,1143,562,587]" italics="true" pageId="14" pageNumber="15">Z5</emphasis>
(105–115),
<emphasis id="B971945A270D3D51FAF2FDD1FAD79769" box="[1288,1313,562,587]" italics="true" pageId="14" pageNumber="15">r3</emphasis>
(88–100),
<emphasis id="B971945A270D3D51FF6DFDBBFF589753" box="[151,174,600,625]" italics="true" pageId="14" pageNumber="15">j5</emphasis>
(55–68); other setae of different moderate lengths (27–58) (
<figureCitation id="133E54CD270D3D51FC89FDBBFC3D9750" box="[883,971,600,626]" captionStart="FIGURES 12 – 13" captionStartId="5.[151,272,1979,2003]" captionTargetBox="[157,1436,500,1919]" captionTargetId="figure@5.[153,1438,494,1958]" captionTargetPageId="5" captionText="FIGURES 12 – 13. Opilioseius grallator n. sp., adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/195393/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
). Lateral soft cuticle with three pairs of simple
<emphasis id="B971945A270D3D51FF16FD9EFEE497B4" box="[236,274,637,663]" italics="true" pageId="14" pageNumber="15">r -R</emphasis>
marginal setae (30–55). Peritrematal plates uniting with dorsal shield anteriorly at level of setae
<emphasis id="B971945A270D3D51FA80FD9EFA6597B4" box="[1402,1427,637,662]" italics="true" pageId="14" pageNumber="15">s1</emphasis>
; peritremes shortened, reaching anteriorly to level of setae
<emphasis id="B971945A270D3D51FCBAFD41FCAC9799" box="[832,858,674,699]" italics="true" pageId="14" pageNumber="15">z4</emphasis>
.
</paragraph>
<paragraph id="8BBA4848270D3D51FF3CFD2BFF409085" blockId="14.[151,1437,263,2007]" pageId="14" pageNumber="15">
Tritosternum (
<figureCitation id="133E54CD270D3D51FE8BFD2BFE4F97C0" box="[369,441,712,738]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
) with base twice as long (23–28) as wide, with laciniae free for about 0.8 of total length (70–79 excluding base), their fused area with a chevron of denticles. Presternal area with pair of weakly sclerotized plates consolidated with sternal shield (
<figureCitation id="133E54CD270D3D51FC98FCF1FC4B960E" box="[866,957,786,812]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="14" pageNumber="15">Fig. 16</figureCitation>
). Sternal shield mid-lateral length from anterior edge of sternal/presternal margin to posterior margin 95–105, narrowest width 70–80, unornamented, weakly sclerotized; anterior margin deeply emarginate medially; lateral margins indented at level between lyrifissures
<date id="FFBB6E88270D3D51FEDEFC61FEBF96B9" box="[292,329,898,923]" pageId="14" pageNumber="15">
<emphasis id="B971945A270D3D51FEDEFC61FEBF96B9" box="[292,329,898,923]" italics="true" pageId="14" pageNumber="15">iv1</emphasis>
</date>
and setae
<emphasis id="B971945A270D3D51FE32FC61FE1D96B9" box="[456,491,898,923]" italics="true" pageId="14" pageNumber="15">st2</emphasis>
, and with weak endopodal extensions between coxae I–II and II–III; posterior margin slightly concave. Sternal setae
<emphasis id="B971945A270D3D51FDA0FC4BFD5E96E3" box="[602,680,936,962]" italics="true" pageId="14" pageNumber="15">st1-st4</emphasis>
of similar length (50–60),
<emphasis id="B971945A270D3D51FC1EFC4BFBF096E3" box="[996,1030,936,961]" italics="true" pageId="14" pageNumber="15">st4</emphasis>
on weakly sclerotized metasternal plates. Endopodal strips weakly developed between coxae III and IV. Epigynal shield 96-120 long, 80-95 wide at level of setae
<emphasis id="B971945A270D3D51FE5EFC11FE319129" box="[420,455,1010,1035]" italics="true" pageId="14" pageNumber="15">st5</emphasis>
, unornamented, axe-shaped with truncate posterior margin and tapered, slightly concave lateral margins, with genital setae
<emphasis id="B971945A270D3D51FD6DFBFBFD4C9113" box="[663,698,1048,1073]" italics="true" pageId="14" pageNumber="15">st5</emphasis>
(51–55) subequally as long as
<emphasis id="B971945A270D3D51FBD6FBFBFB8D9113" box="[1068,1147,1048,1074]" italics="true" pageId="14" pageNumber="15">st1-st4</emphasis>
. Paragenital poroids
<date id="FFBB6E88270D3D51FA82FBFBFA6A9113" box="[1400,1436,1048,1073]" pageId="14" pageNumber="15">
<emphasis id="B971945A270D3D51FA82FBFBFA6A9113" box="[1400,1436,1048,1073]" italics="true" pageId="14" pageNumber="15">iv5</emphasis>
</date>
well removed on soft cuticle from posterolateral corners of epigynal shield. Ventrianal shield unornamented, smaller than epigynal shield, midlength 66–75, greatest width near convex anterior margin 55–65, with two pairs of opisthogastric setae
<emphasis id="B971945A270D3D51FE16FB6BFDEC9183" box="[492,538,1160,1185]" italics="true" pageId="14" pageNumber="15">JV2</emphasis>
(45-60),
<emphasis id="B971945A270D3D51FD70FB6BFD4A9183" box="[650,700,1160,1185]" italics="true" pageId="14" pageNumber="15">ZV2</emphasis>
(35–52), subequally short paranal (21–24) and postanal setae (18–20), and gland pores
<emphasis id="B971945A270D3D51FE31FB4EFE0091E4" box="[459,502,1197,1222]" italics="true" pageId="14" pageNumber="15">gv3</emphasis>
(
<figureCitation id="133E54CD270D3D51FDFDFB4EFD9691E5" box="[519,608,1197,1223]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="14" pageNumber="15">Fig. 16</figureCitation>
). Soft cuticle with six pairs of opisthogastric setae
<emphasis id="B971945A270D3D51FB3CFB4EFB0391E4" box="[1222,1269,1197,1222]" italics="true" pageId="14" pageNumber="15">JV1</emphasis>
(40–43),
<emphasis id="B971945A270D3D51FA91FB4EFA6A91E4" box="[1387,1436,1197,1222]" italics="true" pageId="14" pageNumber="15">ZV1</emphasis>
(40–45),
<emphasis id="B971945A270D3D51FEF9FB30FEC391CE" box="[259,309,1235,1260]" italics="true" pageId="14" pageNumber="15">ZV3</emphasis>
(24–30),
<emphasis id="B971945A270D3D51FE53FB30FE2091CE" box="[425,470,1235,1260]" italics="true" pageId="14" pageNumber="15">JV4</emphasis>
(65–72),
<emphasis id="B971945A270D3D51FDB6FB30FD8A91CE" box="[588,636,1235,1260]" italics="true" pageId="14" pageNumber="15">ZV4</emphasis>
(38–42),
<emphasis id="B971945A270D3D51FD0BFB30FCE991CE" box="[753,799,1235,1260]" italics="true" pageId="14" pageNumber="15">JV5</emphasis>
(80–87), flanked by posteriormost pair of marginal
<emphasis id="B971945A270D3D51FA70FB30FA6A91CE" box="[1418,1436,1235,1260]" italics="true" pageId="14" pageNumber="15">R</emphasis>
setae. Peritrematal shield slightly wider than stigma at that level, bearing three poroids, three gland pores, and consolidated with slender exopodal strip curving behind coxa IV (
<figureCitation id="133E54CD270D3D51FC5CFAFEFBC79015" box="[934,1073,1309,1335]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="14" pageNumber="15">Figs 15, 16</figureCitation>
). Spermathecal apparatus with long (40–42), slender, thinly sclerotized calyx, and a minor duct arising from embolus near base of calyx, seeming to adhere to calyx wall and ending amidst folds of highly plicate and expandable vesicle (
<figureCitation id="133E54CD270D3D51FADDFA8BFA8990A0" box="[1319,1407,1384,1410]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="14" pageNumber="15">Figs 14</figureCitation>
a– b).
</paragraph>
<paragraph id="8BBA4848270D3D51FF3EFA51FB7492F5" blockId="14.[151,1437,263,2007]" pageId="14" pageNumber="15">
Gnathotectum with anterior margin irregularly convex to nearly truncate, denticulate (
<figureCitation id="133E54CD270D3D51FB4DFA51FAFB90EE" box="[1207,1293,1458,1484]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
). Cheliceral shaft, excluding basal section, 85–87 long, with stout digits; dorsal face of fixed digit with blunt protuberance at base of short, basally thick, dorsal seta (
<figureCitation id="133E54CD270D3D51FD75FA1EFD119335" box="[655,743,1533,1559]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
); fixed digit with minute (1.5–2.0) pilus dentilis, with row of 9 or 10 coarse teeth along entire masticatory surface and offset subapical tooth (
<figureCitation id="133E54CD270D3D51FB92F9C1FB15931E" box="[1128,1251,1570,1596]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Figs 8, 10</figureCitation>
); movable digit (30–35) with row of 5 or 6 similarly coarse teeth along apical half of masticatory surface, followed basally by series of several indentations that accommodate more basal teeth of fixed digit; movable digit with two paraxial grooves running proximally and ventrally, one on apical third, leading from base of subapical tooth, one on mid-third, leading from base of second tooth (
<figureCitation id="133E54CD270D3D51FCDFF95BFC6393F0" box="[805,917,1720,1746]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Figs 8, 9</figureCitation>
). Corniculi with medial edges and pointed apices well separated but parallel; internal malae slightly longer than corniculi (
<figureCitation id="133E54CD270D3D51FBAFF93EFB6A93D5" box="[1109,1180,1757,1783]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
). Deutosternum with anterior five rows of denticles each with 5-7 fine teeth, the first, third and fifth rows flanked by arched denticulate extensions (
<figureCitation id="133E54CD270D3D51FE54F8CBFE059260" box="[430,499,1832,1858]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
), sixth and seventh rows widened, multidenticulate. Subcapitulum with lateral hypostomatic setae
<emphasis id="B971945A270D3D51FE87F8AEFE5F9244" box="[381,425,1869,1894]" italics="true" pageId="14" pageNumber="15">hp2</emphasis>
subequally as long as anterior pair
<emphasis id="B971945A270D3D51FCBCF8AEFC849244" box="[838,882,1869,1894]" italics="true" pageId="14" pageNumber="15">hp1</emphasis>
(25–30); medial pair
<emphasis id="B971945A270D3D51FB8AF8AEFB6A9244" box="[1136,1180,1869,1894]" italics="true" pageId="14" pageNumber="15">hp3</emphasis>
longer (30–35), basal pair
<emphasis id="B971945A270D3D51FF36F890FF1E92AE" box="[204,232,1907,1932]" italics="true" pageId="14" pageNumber="15">pc</emphasis>
longest (39–42). Palpal length 105–110, not elongated. Palptrochanter setae slender, inner seta slightly longer than outer one; palpfemur with seta
<emphasis id="B971945A270D3D51FD61F87BFD459293" box="[667,691,1944,1969]" italics="true" pageId="14" pageNumber="15">pl</emphasis>
(30–35) the longest of palpal setae, and with seta
<emphasis id="B971945A270D3D51FAF6F87BFAD59293" box="[1292,1315,1944,1969]" italics="true" pageId="14" pageNumber="15">al</emphasis>
spatulate, with flat tip; palpgenu with setae
<emphasis id="B971945A270D3D51FDE5F85EFDB892F4" box="[543,590,1981,2007]" italics="true" pageId="14" pageNumber="15">al -1</emphasis>
and
<emphasis id="B971945A270D3D51FD7AF85EFD5992F4" box="[640,687,1981,2007]" italics="true" pageId="14" pageNumber="15">al -2</emphasis>
spatulate, with rounded tips (
<figureCitation id="133E54CD270D3D51FBF3F85EFB8592F5" box="[1033,1139,1981,2007]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="14" pageNumber="15">Figs 6, 7</figureCitation>
).
</paragraph>
<paragraph id="8BBA4848270C3D50FF3EFF7BFBF3979E" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
Legs IV the longest (810–875, excluding pretarsus) of legs, about 2.4-2.7 as long as dorsal shield (
<figureCitation id="133E54CD270C3D50FA96FF7BFF4E95F5" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="15" pageNumber="16">Fig. 39</figureCitation>
); other leg lengths: I 705–730, II 458–490, III 498–540. Legs II, III, IV with tarsus 3.2–3.7, 3.9–4.1, 3.6– 3.7 times, respectively, as long as tibia. Tarsi II–IV with pair of apical setal processes
<emphasis id="B971945A270C3D50FB6DFF01FB3B95D9" box="[1175,1229,226,252]" italics="true" pageId="15" pageNumber="16">ad -1</emphasis>
,
<emphasis id="B971945A270C3D50FB26FF01FAE595D9" box="[1244,1299,226,252]" italics="true" pageId="15" pageNumber="16">pd -1</emphasis>
minute (8– 10), less than 0.2 the length of pretarsus to base of claws (50–55 for II-III, 80–85 for IV) (
<figureCitation id="133E54CD270C3D50FB13FEEBFAB29400" box="[1257,1348,264,290]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="15" pageNumber="16">Fig. 21</figureCitation>
). Leg I segment length ratios, femur: genu: tibia: tarsus, about 1.0: 0.8: 0.95–1.0: 2.0–2.1; tarsus (265–268) longer than femur (125–139), genu (100–111), and tibia (123–130); length of pretarsus I to base of claws 20–26. Coxae II–IV each with a raised boss on ventral surface (
<figureCitation id="133E54CD270C3D50FCD9FE9BFC8F94B0" box="[803,889,376,402]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="15" pageNumber="16">Fig. 16</figureCitation>
); coxal setae simple, moderately long (30–40), except coxa II
<emphasis id="B971945A270C3D50FEB1FE7EFE919494" box="[331,359,413,438]" italics="true" pageId="15" pageNumber="16">av</emphasis>
short (5–6). Legs I to IV with chaetotactic formulae of segments as described for genus. Legs I and II (
<figureCitation id="133E54CD270C3D50FEC5FE21FE3394FE" box="[319,453,450,476]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="15" pageNumber="16">Figs 17, 18</figureCitation>
) without notably long setae, except tibia I
<emphasis id="B971945A270C3D50FC44FE21FC0294F9" box="[958,1012,450,476]" italics="true" pageId="15" pageNumber="16">ad -3</emphasis>
(52–60) and basitarsus II
<emphasis id="B971945A270C3D50FAD3FE21FAA994F9" box="[1321,1375,450,476]" italics="true" pageId="15" pageNumber="16">ad -4</emphasis>
(38– 45) slightly elongated; genu I setae
<emphasis id="B971945A270C3D50FD9FFE0BFD689723" box="[613,670,488,514]" italics="true" pageId="15" pageNumber="16">pd -2</emphasis>
,
<emphasis id="B971945A270C3D50FD4BFE0BFD1C9723" box="[689,746,488,514]" italics="true" pageId="15" pageNumber="16">pd -3</emphasis>
with blunt bent tips, stouter and shorter (17–20) than neighboring
<emphasis id="B971945A270C3D50FED5FDEEFEBA9704" box="[303,332,525,550]" italics="true" pageId="15" pageNumber="16">ad</emphasis>
setae. Leg III with tibial
<emphasis id="B971945A270C3D50FD85FDEEFD439704" box="[639,693,525,551]" italics="true" pageId="15" pageNumber="16">ad -1</emphasis>
(42–50) and
<emphasis id="B971945A270C3D50FCADFDEEFC789704" box="[855,910,525,551]" italics="true" pageId="15" pageNumber="16">pd -2</emphasis>
(55–72) somewhat elongated (
<figureCitation id="133E54CD270C3D50FAFBFDEEFAAC9705" box="[1281,1370,525,551]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="15" pageNumber="16">Fig. 19</figureCitation>
), and basitarsal
<emphasis id="B971945A270C3D50FEF7FDD1FEB59769" box="[269,323,562,588]" italics="true" pageId="15" pageNumber="16">ad -4</emphasis>
slightly so. Leg IV with femoral setae
<emphasis id="B971945A270C3D50FCEBFDD1FCE89769" box="[785,798,562,587]" italics="true" pageId="15" pageNumber="16">v</emphasis>
and
<emphasis id="B971945A270C3D50FCA2FDD1FC719769" box="[856,903,562,588]" italics="true" pageId="15" pageNumber="16">al -1</emphasis>
somewhat elongated (60–70),
<emphasis id="B971945A270C3D50FB09FDD1FAF69769" box="[1267,1280,562,587]" italics="true" pageId="15" pageNumber="16">v</emphasis>
stiff, aligned proximal to
<emphasis id="B971945A270C3D50FED2FDBBFEAE9753" box="[296,344,600,626]" italics="true" pageId="15" pageNumber="16">
al -
<quantity id="4CFDE5AD270C3D50FEB3FDBBFEAE9753" box="[329,344,600,625]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="16" unit="in" value="1.0">1</quantity>
</emphasis>
in an anterolateral position (
<figureCitation id="133E54CD270C3D50FD54FDBBFCF19750" box="[686,775,600,626]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="15" pageNumber="16">Fig. 20</figureCitation>
); genual seta
<emphasis id="B971945A270C3D50FC51FDBBFC179753" box="[939,993,600,626]" italics="true" pageId="15" pageNumber="16">pd -1</emphasis>
(235–260) and tibial seta
<emphasis id="B971945A270C3D50FAE3FDBBFAB99753" box="[1305,1359,600,626]" italics="true" pageId="15" pageNumber="16">ad -1</emphasis>
(220– 280) greatly elongated, tibial
<emphasis id="B971945A270C3D50FE09FD9EFDDF97B4" box="[499,553,637,663]" italics="true" pageId="15" pageNumber="16">pd -1</emphasis>
(80–85) slightly so; basitarsal seta
<emphasis id="B971945A270C3D50FC36FD9EFBF497B4" box="[972,1026,637,663]" italics="true" pageId="15" pageNumber="16">ad -4</emphasis>
(212–242) and telotarsal seta
<emphasis id="B971945A270C3D50FA9CFD9EFA6A97B4" box="[1382,1436,637,663]" italics="true" pageId="15" pageNumber="16">ad -3</emphasis>
(202–225) greatly elongated, basitarsal
<emphasis id="B971945A270C3D50FD9DFD41FD6B9799" box="[615,669,674,700]" italics="true" pageId="15" pageNumber="16">pd -4</emphasis>
(95–100) slightly so (
<figureCitation id="133E54CD270C3D50FC5AFD41FC0E979E" box="[928,1016,674,700]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="15" pageNumber="16">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BBA4848270C3D50FF3CFD24FC5496BE" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
<emphasis id="B971945A270C3D50FF3CFD24FE8797C3" bold="true" box="[198,369,711,737]" pageId="15" pageNumber="16">ADULT MALE.</emphasis>
Dorsal shield 224–290 long, 185-200 at its greatest width at level of setae
<emphasis id="B971945A270C3D50FB1EFD2BFB0897C3" box="[1252,1278,712,737]" italics="true" pageId="15" pageNumber="16">r5</emphasis>
; shield form, ornamentation, and setation as in female except more expanded anterolaterally so as to bear setae
<emphasis id="B971945A270C3D50FAEAFD0EFADF9624" box="[1296,1321,749,774]" italics="true" pageId="15" pageNumber="16">r5</emphasis>
and to be more broadly united to peritrematal shields at level of setae
<emphasis id="B971945A270C3D50FCAAFCF1FC9C9609" box="[848,874,786,811]" italics="true" pageId="15" pageNumber="16">r3</emphasis>
(
<figureCitation id="133E54CD270C3D50FC83FCF1FC26960E" box="[889,976,786,812]" captionStart="FIGURES 32 – 33" captionStartId="10.[151,269,1980,2004]" captionTargetBox="[154,1436,642,1930]" captionTargetId="figure@10.[154,1439,642,1958]" captionTargetPageId="10" captionText="FIGURES 32 – 33. Opilioseius grallator n. sp., adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view." httpUri="https://zenodo.org/record/195397/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
). Form and proportionate size of dorsal shield setae as in female, except setae collectively about 15-20 percent shorter,
<emphasis id="B971945A270C3D50FBCEFCDBFBB79673" box="[1076,1089,824,849]" italics="true" pageId="15" pageNumber="16">e</emphasis>
.
<emphasis id="B971945A270C3D50FBB2FCDBFBA19673" box="[1096,1111,824,849]" italics="true" pageId="15" pageNumber="16">g</emphasis>
.,
<emphasis id="B971945A270C3D50FB96FCDBFB709673" box="[1132,1158,824,849]" italics="true" pageId="15" pageNumber="16">z2</emphasis>
,
<emphasis id="B971945A270C3D50FB6EFCDBFB589673" box="[1172,1198,824,849]" italics="true" pageId="15" pageNumber="16">z5</emphasis>
,
<emphasis id="B971945A270C3D50FB46FCDBFB229673" box="[1212,1236,824,849]" italics="true" pageId="15" pageNumber="16">j6</emphasis>
(5–10),
<emphasis id="B971945A270C3D50FACCFCDBFAA69673" box="[1334,1360,824,849]" italics="true" pageId="15" pageNumber="16">s5</emphasis>
(153– 162),
<emphasis id="B971945A270C3D50FF21FCBEFF0C9654" box="[219,250,861,886]" italics="true" pageId="15" pageNumber="16">Z3</emphasis>
,
<emphasis id="B971945A270C3D50FEF2FCBEFED19654" box="[264,295,861,886]" italics="true" pageId="15" pageNumber="16">Z4</emphasis>
(100–105),
<emphasis id="B971945A270C3D50FE4CFCBEFE229654" box="[438,468,861,886]" italics="true" pageId="15" pageNumber="16">Z5</emphasis>
(68–70),
<emphasis id="B971945A270C3D50FDBDFCBEFD979654" box="[583,609,861,886]" italics="true" pageId="15" pageNumber="16">r3</emphasis>
(73–80),
<emphasis id="B971945A270C3D50FD29FCBEFD1D9654" box="[723,747,861,886]" italics="true" pageId="15" pageNumber="16">j5</emphasis>
(40–52). Lateral soft cuticle without any
<emphasis id="B971945A270C3D50FB2AFCBEFB149654" box="[1232,1250,861,886]" italics="true" pageId="15" pageNumber="16">R</emphasis>
-marginal setae. Peritremes as in female, reaching anteriorly to level of setae
<emphasis id="B971945A270C3D50FCA7FC61FC6D96B9" box="[861,923,898,924]" italics="true" pageId="15" pageNumber="16">z4-s4</emphasis>
.
</paragraph>
<paragraph id="8BBA4848270C3D50FF3EFC4BFAFB91CE" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
Tritosternum as in female, including chevron of denticles on fused basal portion of laciniae. Ventral shields unornamented, as in female. Sternitigenital shield mid-lateral length 132–140, width 80 at level between coxae II, with five pairs of setae of similar lengths (40–47) and two pairs of poroids; lateral margins slightly indented between coxae II, deeply incised between bases of setae
<emphasis id="B971945A270C3D50FBF7FBFBFBC69113" box="[1037,1072,1048,1073]" italics="true" pageId="15" pageNumber="16">st3</emphasis>
and
<emphasis id="B971945A270C3D50FB91FBFBFB789113" box="[1131,1166,1048,1073]" italics="true" pageId="15" pageNumber="16">st4</emphasis>
, and again behind
<emphasis id="B971945A270C3D50FA88FBFBFA639113" box="[1394,1429,1048,1073]" italics="true" pageId="15" pageNumber="16">st4</emphasis>
, partly delineating metasternal and genital portions of shield (
<figureCitation id="133E54CD270C3D50FC9AFBDEFC4E9175" box="[864,952,1085,1111]" captionStart="FIGURES 32 – 33" captionStartId="10.[151,269,1980,2004]" captionTargetBox="[154,1436,642,1930]" captionTargetId="figure@10.[154,1439,642,1958]" captionTargetPageId="10" captionText="FIGURES 32 – 33. Opilioseius grallator n. sp., adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view." httpUri="https://zenodo.org/record/195397/files/figure.png" pageId="15" pageNumber="16">Fig. 33</figureCitation>
). Endopodal strips alongside coxae II–III and III–IV weakly developed, free from sternitigenital shield. Ventrianal shield mid-length (70–75), greatest midlateral width (83–94) at level of setae
<emphasis id="B971945A270C3D50FD72FB6BFD419183" box="[648,695,1160,1185]" italics="true" pageId="15" pageNumber="16">JV2</emphasis>
, with irregularly convex lateral margins, bearing the only four pairs of opisthogastric setae present,
<emphasis id="B971945A270C3D50FDB7FB4EFD8D91E4" box="[589,635,1197,1222]" italics="true" pageId="15" pageNumber="16">JV1</emphasis>
,
<emphasis id="B971945A270C3D50FD71FB4EFD4D91E4" box="[651,699,1197,1222]" italics="true" pageId="15" pageNumber="16">ZV2</emphasis>
,
<emphasis id="B971945A270C3D50FD31FB4EFD0E91E4" box="[715,760,1197,1222]" italics="true" pageId="15" pageNumber="16">JV2</emphasis>
,
<emphasis id="B971945A270C3D50FCF2FB4EFCC091E4" box="[776,822,1197,1222]" italics="true" pageId="15" pageNumber="16">JV5</emphasis>
plus the circumanal setae; form and relative size of these setae as in female, but collectively about 25 percent shorter. Metapodal plates indiscernible.
</paragraph>
<paragraph id="8BBA4848270C3D50FF3EFB1BFC869335" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
Gnathotectum (
<figureCitation id="133E54CD270C3D50FE80FB1BFE269030" box="[378,464,1272,1298]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="15" pageNumber="16">Fig. 28</figureCitation>
) similar to that of female. Cheliceral fixed digit with minute pilus dentilis and row of 6-8 coarse teeth (
<figureCitation id="133E54CD270C3D50FE6AFAFEFE189015" box="[400,494,1309,1335]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="15" pageNumber="16">Fig. 25</figureCitation>
); movable digit (27–28) with two or three teeth, or these are indentations to receive some of apposed teeth of fixed digit; spermatodactyl a complex structure, which in dorsoventral view seems to have a pointed spearlike process (21) covering the twisted apex of the main dactyl shaft (length 16 to concave apex) that bears the sperm transfer tube (
<figureCitation id="133E54CD270C3D50FD0AFA6EFC8B9085" box="[752,893,1421,1447]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="15" pageNumber="16">Figs 25–27</figureCitation>
). Corniculi (28–29) more slender and widely spaced basally, with their apices more flared than in female; internal malae slightly longer and more acute apically than in female (
<figureCitation id="133E54CD270C3D50FE47FA3BFDEE90D0" box="[445,536,1496,1522]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="15" pageNumber="16">Fig. 29</figureCitation>
). Subcapitulum with rows of deutosternal denticles and relative lengths of setae as in female. Palpal structures similar to those in female.
</paragraph>
<paragraph id="8BBA4848270C3D50FF3EF9C1FD83938E" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
Relative leg lengths, excluding pretarsi, similar to those of female, I (607–616), II (397–410), III (434– 446), IV (695–744). Leg I segment length ratios, leg coxal structures and setae (including short
<emphasis id="B971945A270C3D50FAFBF9ABFAEB9343" box="[1281,1309,1608,1633]" italics="true" pageId="15" pageNumber="16">av</emphasis>
on coxa II and slightly spinelike
<emphasis id="B971945A270C3D50FE56F98EFE1293A4" box="[428,484,1645,1670]" italics="true" pageId="15" pageNumber="16">pd-2</emphasis>
and
<emphasis id="B971945A270C3D50FDD8F98EFDAD93A4" box="[546,603,1645,1670]" italics="true" pageId="15" pageNumber="16">pd-3</emphasis>
on genu I), and relative leg setal lengths as in female except setae collectively about 15–25 percent shorter.
</paragraph>
<paragraph id="8BBA4848270C3D50FF3CF954FBC19290" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
<emphasis id="B971945A270C3D50FF3CF954FE8B93F3" bold="true" box="[198,381,1719,1745]" pageId="15" pageNumber="16">DEUTONYMPH.</emphasis>
Dorsal shield weakly sclerotized, 261 long, 148 at its greatest width at level of setae
<emphasis id="B971945A270C3D50FA80F95BFA6293F3" box="[1402,1428,1720,1745]" italics="true" pageId="15" pageNumber="16">s5</emphasis>
; shield without discernible lateral incisions, its form and setation as in female except less expanded anterolaterally so that setae
<emphasis id="B971945A270C3D50FE1AF8E0FE0C923E" box="[480,506,1795,1820]" italics="true" pageId="15" pageNumber="16">s1</emphasis>
and
<emphasis id="B971945A270C3D50FDCEF8E0FDBB923E" box="[564,589,1795,1820]" italics="true" pageId="15" pageNumber="16">r3</emphasis>
on soft cuticle, and anterior apices of peritrematal shields ending freely (
<figureCitation id="133E54CD270C3D50FF65F8CBFF0E9260" box="[159,248,1832,1858]" captionStart="FIGURES 34 – 38. 34 – 35" captionStartId="11.[151,270,1817,1841]" captionTargetBox="[157,1436,204,1770]" captionTargetId="figure@11.[156,1441,197,1794]" captionTargetPageId="11" captionText="FIGURES 34 – 38. 34 – 35, Opilioseius grallator n. sp., deutonymph: 34, idiosoma, dorsal aspect; 35, idiosoma, ventral aspect. 36 – 38, leg III, tarsal apex and pretarsus, showing different forms of ventroapical tarsal processes and conjectured acrotarsal demarcation: 36, Adenoseius sp. (Sejidae), anterolateral view; 37, Euzercon sp. (Euzerconidae), ventral view; 38, Arctacarus rostratus Evans (Arctacaridae), ventral view. Abbreviation: acr mar, acrotarsal margin." httpUri="https://zenodo.org/record/195398/files/figure.png" pageId="15" pageNumber="16">Fig. 34</figureCitation>
). Form and proportionate size of dorsal shield setae as in female, except setae collectively about 15- 20 percent shorter. Lateral soft cuticle with four pairs of marginal setae,
<emphasis id="B971945A270C3D50FC0EF8AEFBF89244" box="[1012,1038,1869,1894]" italics="true" pageId="15" pageNumber="16">r3</emphasis>
,
<emphasis id="B971945A270C3D50FBE4F8AEFBCE9244" box="[1054,1080,1869,1894]" italics="true" pageId="15" pageNumber="16">r5</emphasis>
,
<emphasis id="B971945A270C3D50FBB2F8AEFB9E9244" box="[1096,1128,1869,1894]" italics="true" pageId="15" pageNumber="16">R4</emphasis>
,
<emphasis id="B971945A270C3D50FB82F8AEFB6E9244" box="[1144,1176,1869,1894]" italics="true" pageId="15" pageNumber="16">R5</emphasis>
. Peritrematal shield a thin strip, reaching anteriorly to level of setae
<emphasis id="B971945A270C3D50FD41F890FD2B92AE" box="[699,733,1907,1932]" italics="true" pageId="15" pageNumber="16">st1</emphasis>
, and extending posteriorly from stigma only to bear poroid
<emphasis id="B971945A270C3D50FF6DF87BFF4A9293" box="[151,188,1944,1969]" italics="true" pageId="15" pageNumber="16">ip3</emphasis>
and gland pore
<emphasis id="B971945A270C3D50FE83F87BFE539293" box="[377,421,1944,1969]" italics="true" pageId="15" pageNumber="16">gp3</emphasis>
; peritreme reaching anteriorly to mid-level of coxae III.
</paragraph>
<paragraph id="8BBA484827133D4FFF3EFF7BFCFE94B0" blockId="16.[151,1437,152,2007]" pageId="16" pageNumber="17">
Tritosternum as in female, including chevron of denticles on fused basal portion of laciniae. Intercoxal shield so poorly sclerotized that its delineation hardly perceptible, with four pairs of setae and two pairs of poroids,
<emphasis id="B971945A27133D4FFF05FF01FEB895D9" box="[255,334,226,252]" italics="true" pageId="16" pageNumber="17">st1-st3</emphasis>
of similar lengths (38–44),
<emphasis id="B971945A27133D4FFD66FF01FD4995D9" box="[668,703,226,251]" italics="true" pageId="16" pageNumber="17">st4</emphasis>
shorter (26), poroids
<date id="FFBB6E8827133D4FFC39FF01FC1195D9" box="[963,999,226,251]" pageId="16" pageNumber="17">
<emphasis id="B971945A27133D4FFC39FF01FC1195D9" box="[963,999,226,251]" italics="true" pageId="16" pageNumber="17">iv3</emphasis>
</date>
absent; setae
<emphasis id="B971945A27133D4FFB75FF01FB4795D9" box="[1167,1201,226,251]" italics="true" pageId="16" pageNumber="17">st5</emphasis>
(35) on soft cuticle (
<figureCitation id="133E54CD27133D4FFF5AFEEBFF0C9400" box="[160,250,264,290]" captionStart="FIGURES 34 – 38. 34 – 35" captionStartId="11.[151,270,1817,1841]" captionTargetBox="[157,1436,204,1770]" captionTargetId="figure@11.[156,1441,197,1794]" captionTargetPageId="11" captionText="FIGURES 34 – 38. 34 – 35, Opilioseius grallator n. sp., deutonymph: 34, idiosoma, dorsal aspect; 35, idiosoma, ventral aspect. 36 – 38, leg III, tarsal apex and pretarsus, showing different forms of ventroapical tarsal processes and conjectured acrotarsal demarcation: 36, Adenoseius sp. (Sejidae), anterolateral view; 37, Euzercon sp. (Euzerconidae), ventral view; 38, Arctacarus rostratus Evans (Arctacaridae), ventral view. Abbreviation: acr mar, acrotarsal margin." httpUri="https://zenodo.org/record/195398/files/figure.png" pageId="16" pageNumber="17">Fig. 35</figureCitation>
). Endopodal strips alongside coxae II-III developed, fused to intercoxal shield. Anal shield weakly sclerotized, oval (its anterior margin folded on specimen at hand), midlateral width (52) at level of setae
<emphasis id="B971945A27133D4FFA99FECEFA659464" box="[1379,1427,301,326]" italics="true" pageId="16" pageNumber="17">ZV2</emphasis>
; paranal setae slightly longer than postanal seta. Eight pairs of opisthogastric setae on soft cuticle,
<emphasis id="B971945A27133D4FFAD9FEB1FAA29449" box="[1315,1364,338,363]" italics="true" pageId="16" pageNumber="17">ZV3</emphasis>
,
<emphasis id="B971945A27133D4FFA9EFEB1FA629449" box="[1380,1428,338,363]" italics="true" pageId="16" pageNumber="17">ZV4</emphasis>
, shortest,
<emphasis id="B971945A27133D4FFEFBFE9BFED894B3" box="[257,302,376,401]" italics="true" pageId="16" pageNumber="17">JV5</emphasis>
longest. Metapodal plates indiscernible.
</paragraph>
<paragraph id="8BBA484827133D4FFF3EFE7EFB8D9495" blockId="16.[151,1437,152,2007]" box="[196,1147,413,439]" pageId="16" pageNumber="17">Gnathotectum, chelicerae, subcapitulum, and palpal structures as in adult female.</paragraph>
<paragraph id="8BBA484827133D4FFF3EFE21FB539705" blockId="16.[151,1437,152,2007]" pageId="16" pageNumber="17">
Relative leg lengths, excluding pretarsi, I (527), II (377), III (453), IV (615); pretarsi, I (28), II-III (41), IV (70). Leg I segment length ratios, leg coxal structures and setae (including short
<emphasis id="B971945A27133D4FFBA3FE0BFB839723" box="[1113,1141,488,513]" italics="true" pageId="16" pageNumber="17">av</emphasis>
on coxa II), and relative leg setal lengths as in adult female except setae collectively about 25–30 percent shorter.
</paragraph>
</subSubSection>
<subSubSection id="C31F1BC327133D4FFF3CFDD1FBCE9655" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BBA484827133D4FFF3CFDD1FBCE9655" blockId="16.[151,1437,152,2007]" pageId="16" pageNumber="17">
<emphasis id="B971945A27133D4FFF3CFDD1FE8B976E" bold="true" box="[198,381,562,588]" pageId="16" pageNumber="17">
<typeStatus id="54BEF6EA27133D4FFF3CFDD1FEF7976E" box="[198,257,562,588]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
All specimens from samples collected on the area of the La Selva Biological Station, Heredia Province,
<collectingCountry id="F31208D827133D4FFE8EFDBBFDE99750" box="[372,543,600,626]" name="Costa Rica" pageId="16" pageNumber="17">COSTA RICA</collectingCountry>
(
<geoCoordinate id="EE312E8F27133D4FFDCBFDBBFD3E9750" box="[561,712,600,626]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="10.433611">10° 26' 1&quot; N</geoCoordinate>
,
<geoCoordinate id="EE312E8F27133D4FFD2FFDBBFC979750" box="[725,865,600,626]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="15" value="-84.01723">84° 1' 2&quot; W</geoCoordinate>
, elevation
<quantity id="4CFDE5AD27133D4FFC1BFDBBFBAB9750" box="[993,1117,600,626]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="16" pageNumber="17" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</quantity>
):
<typeStatus id="54BEF6EA27133D4FFB88FDBBFB049750" box="[1138,1266,600,626]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
: adult female,
<quantity id="4CFDE5AD27133D4FFF6DFD9EFF1597B5" box="[151,227,637,663]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="150.0">150 m</quantity>
along Surá trail, ex “Hongo” (fungus),
<date id="FFBB6E8827133D4FFD38FD9EFC6F97B5" box="[706,921,637,663]" pageId="16" pageNumber="17" value="1995-02-21">21 February 1995</date>
, coll.
<collectingCountry id="F31208D827133D4FFC19FD9EFBC297B5" box="[995,1076,637,663]" name="Kazakhstan" pageId="16" pageNumber="17">ALAS</collectingCountry>
.
<typeStatus id="54BEF6EA27133D4FFBBAFD9EFB3D97B5" box="[1088,1227,637,663]" pageId="16" pageNumber="17" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D0383C127133D4FFB26FD9EFAC697B5" box="[1244,1328,637,663]" pageId="16" pageNumber="17" type="adult">9 adult</specimenCount>
females, with same data as
<typeStatus id="54BEF6EA27133D4FFE7AFD41FE04979E" box="[384,498,674,700]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D0383C127133D4FFDFAFD41FDA1979E" box="[512,599,674,700]" pageId="16" pageNumber="17" type="adult">8 adult</specimenCount>
females,
<specimenCount id="9D0383C127133D4FFD28FD41FCDC979E" box="[722,810,674,700]" pageId="16" pageNumber="17" type="adult">4 adult</specimenCount>
males, with same data as
<typeStatus id="54BEF6EA27133D4FFB8CFD41FB12979E" box="[1142,1252,674,700]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
except fungus determined as
<taxonomicName id="4C0533CB27133D4FFEB5FD2BFE1797C3" box="[335,481,712,737]" class="Agaricomycetes" family="Polyporaceae" genus="Coriolopsis" kingdom="Fungi" order="Polyporales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="genus">
<emphasis id="B971945A27133D4FFEB5FD2BFE1797C3" box="[335,481,712,737]" italics="true" pageId="16" pageNumber="17">Coriolopsis</emphasis>
</taxonomicName>
, and coll. E.E. Lindquist;
<specimenCount id="9D0383C127133D4FFCC8FD2BFC7F97C0" box="[818,905,712,738]" pageId="16" pageNumber="17" type="adult">1 adult</specimenCount>
female, 1 deutonymph, with same data as
<typeStatus id="54BEF6EA27133D4FFF6DFD0EFF089625" box="[151,254,749,775]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
except
<date id="FFBB6E8827133D4FFEA2FD0EFE099625" box="[344,511,749,775]" pageId="16" pageNumber="17" value="1995-03-09">9 March 1995</date>
;
<specimenCount id="9D0383C127133D4FFDF1FD0EFDAA9625" box="[523,604,749,775]" pageId="16" pageNumber="17" type="adult">1 adult</specimenCount>
female, Arboleda area, ex Hongo,
<date id="FFBB6E8827133D4FFC0EFD0EFB5F9625" box="[1012,1193,749,775]" pageId="16" pageNumber="17" value="1995-03-28">28 March 1995</date>
, coll.
<collectingCountry id="F31208D827133D4FFB0AFD0EFAB69625" box="[1264,1344,749,775]" name="Kazakhstan" pageId="16" pageNumber="17">ALAS</collectingCountry>
;
<specimenCount id="9D0383C127133D4FFAB6FD0EFA6A9625" box="[1356,1436,749,775]" pageId="16" pageNumber="17" type="adult">2 adult</specimenCount>
females,
<quantity id="4CFDE5AD27133D4FFEFCFCF1FEA4960E" box="[262,338,786,812]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="16" pageNumber="17" unit="m" value="645.0">645 m</quantity>
along Suampo trail, ex fungus,
<date id="FFBB6E8827133D4FFD27FCF1FC7D960E" box="[733,907,786,812]" pageId="16" pageNumber="17" value="1995-04-18">18 April 1995</date>
, coll.
<collectingCountry id="F31208D827133D4FFC22FCF1FBDD960E" box="[984,1067,786,812]" name="Kazakhstan" pageId="16" pageNumber="17">ALAS</collectingCountry>
;
<specimenCount id="9D0383C127133D4FFBCDFCF1FB7A960E" box="[1079,1164,786,812]" pageId="16" pageNumber="17" type="adult">2 adult</specimenCount>
females,
<quantity id="4CFDE5AD27133D4FFAF9FCF1FAA6960E" box="[1283,1360,786,812]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="16" pageNumber="17" unit="m" value="345.0">345 m</quantity>
along Holdridge trail, ex fungus,
<date id="FFBB6E8827133D4FFE26FCDBFD719670" box="[476,647,824,850]" pageId="16" pageNumber="17" value="1995-04-18">18 April 1995</date>
, coll.
<collectingCountry id="F31208D827133D4FFD2BFCDBFCD49670" box="[721,802,824,850]" name="Kazakhstan" pageId="16" pageNumber="17">ALAS</collectingCountry>
;
<specimenCount id="9D0383C127133D4FFCD5FCDBFC779670" box="[815,897,824,850]" pageId="16" pageNumber="17" type="adult">5 adult</specimenCount>
females, ca
<quantity id="4CFDE5AD27133D4FFBE2FCDBFB929670" box="[1048,1124,824,850]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="17" unit="m" value="300.0">300 m</quantity>
along Occidental trail, ex polypore fungi on canopy-shaded log,
<date id="FFBB6E8827133D4FFDA1FCBEFCB69655" box="[603,832,861,887]" pageId="16" pageNumber="17" value="1992-11-13">13 November 1992</date>
, coll. E.E. Lindquist.
</paragraph>
</subSubSection>
<subSubSection id="C31F1BC327133D4FFF3CFC61FD9096E0" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BBA484827133D4FFF3CFC61FD9096E0" blockId="16.[151,1437,152,2007]" pageId="16" pageNumber="17">
<emphasis id="B971945A27133D4FFF3CFC61FEA596BE" bold="true" box="[198,339,898,924]" pageId="16" pageNumber="17">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C0533CB27133D4FFDAAFC61FD4F96B9" box="[592,697,898,923]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="grallator">
<emphasis id="B971945A27133D4FFDAAFC61FD4F96B9" box="[592,697,898,923]" italics="true" pageId="16" pageNumber="17">grallator</emphasis>
</taxonomicName>
, is masculine and based on the Latin “
<emphasis id="B971945A27133D4FFB85FC61FB3096B9" box="[1151,1222,898,923]" italics="true" pageId="16" pageNumber="17">gralla</emphasis>
”, meaning “stilt”, and indicates “one that walks on stilts”.
</paragraph>
</subSubSection>
<subSubSection id="C31F1BC327133D4EFF3CFC2EFC949705" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BBA484827133D4FFF3CFC2EFCEA91E5" blockId="16.[151,1437,152,2007]" pageId="16" pageNumber="17">
<emphasis id="B971945A27133D4FFF3CFC2EFEB696C5" bold="true" box="[198,320,973,999]" pageId="16" pageNumber="17">Remarks.</emphasis>
Resembling minute opilionid-like creatures (
<figureCitation id="133E54CD27133D4FFCACFC2EFC5B96C5" box="[854,941,973,999]" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="16" pageNumber="17">Fig. 39</figureCitation>
), these spindly-legged mites move slowly along the undersurfaces of the fungal fruiting bodies (sporocarps), which consist of a series of parallel laminae. The apices of all six of legs II to IV invariably rest on the tips of the laminae, like tiny feet on the joists of a subflooring. In feeding, they protrude their gnathosoma down between the laminae, without taking legs III–IV off the tips of the laminae. Whether they feed on fungal matter or on some of the various microarthropods (other mites, springtails, tiny beetles) that move between the laminae, was not ascertained (R.K. Colwell, personal observations,
<date id="FFBB6E8827133D4FFDA2FB4EFCF991E5" box="[600,783,1197,1223]" pageId="16" pageNumber="17" value="1995-03-13">13 March 1995</date>
).
</paragraph>
<paragraph id="8BBA484827133D4EFF3CFB31FB1495F5" blockId="16.[151,1437,152,2007]" lastBlockId="17.[151,1437,152,551]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Full-sized eggs within adult female
<taxonomicName id="4C0533CB27133D4FFD86FB30FC8191CE" box="[636,887,1235,1260]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="grallator">
<emphasis id="B971945A27133D4FFD86FB30FC8191CE" box="[636,887,1235,1260]" italics="true" pageId="16" pageNumber="17">Opilioseius grallator</emphasis>
</taxonomicName>
are remarkably large relative to the female’s idiosoma (
<figureCitation id="133E54CD27133D4FFEECFB1BFE859030" box="[278,371,1272,1298]" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="16" pageNumber="17">Fig. 40</figureCitation>
). Maximum length of eight eggs ranged from 250 to 300 µm, and were 0.74 to 0.85 the maternal females’ dorsal shield lengths of 312 to 337 µm, bearing in mind that the dorsal shield is fully developed, extending from just anterior of the vertical setae
<emphasis id="B971945A27133D4FFCAEFAA0FC9D907E" box="[852,875,1347,1372]" italics="true" pageId="16" pageNumber="17">j1</emphasis>
to just posterior to the clunals
<emphasis id="B971945A27133D4FFB28FAA0FB1B907E" box="[1234,1261,1347,1372]" italics="true" pageId="16" pageNumber="17">J5</emphasis>
. Females have an abundance of soft pliable cuticle between the dorsal and ventral shields, and a small ventrianal shield, which permit considerable idiosomatic expansion to accommodate such large eggs. One slide preparation of a female in lateral view shows a tremendous dorsoventral opisthosomatic distension that accommodates the greatest length of the egg at vertical right angles to that of her idiosoma, thus orienting the smallest egg dimension (width) for issue from the genital opening. Subsequent oviposition of such large eggs, whose greatest widths range from 192 to 210 µm, through a genital orifice whose width between the coxae of legs IV appears to be 130–150 µm, must depend on considerable egg flexibility and resilience (see
<bibRefCitation id="EF9435B927133D4FFB27F9ABFA9B9340" author="Evans" box="[1245,1389,1608,1634]" pageId="16" pageNumber="20" refString="Evans, G. O. (1992) Principles of Acarology. C. A. B. International, Wallingford, xviii + 563 pp." type="book" year="1992">Evans 1992</bibRefCitation>
), as there is no evidence of maternal idiosomatic rupture. Few data are available in the literature concerning this aspect of egg size among free-living mesostigmatic mites. Fully adequate metric data for egg size, i.e. egg mass or volume compared to adult female mass or volume, are not available. However, egg length/dorsal shield length ratios of 0.48 to 0.57 are evident in the phytoseiids
<taxonomicName id="4C0533CB27133D4FFC4BF93EFA9E93D5" authority="Muma" authorityName="Muma" box="[945,1384,1757,1783]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromalus" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B971945A27133D4FFC4BF93EFB0893D4" box="[945,1278,1757,1782]" italics="true" pageId="16" pageNumber="17">Typhlodromalus peregrinus</emphasis>
(Muma)
</taxonomicName>
and
<taxonomicName id="4C0533CB27133D4FFF6DF8E0FDF6923E" authority="Prasad" authorityName="Prasad" box="[151,512,1794,1820]" class="Arachnida" family="Phytoseiidae" genus="Phytoseius" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="B971945A27133D4FFF6DF8E0FE5E923E" box="[151,424,1795,1820]" italics="true" pageId="16" pageNumber="17">Phytoseius hawaiiensis</emphasis>
Prasad
</taxonomicName>
, a family with relatively large egg sizes (
<bibRefCitation id="EF9435B927133D4FFC1BF8E1FB83923E" author="Muma" box="[993,1141,1794,1820]" pageId="16" pageNumber="20" refString="Muma, M. H. (1969) Typhlodromalus peregrinus (Muma) (Acarina: Phytoseiidae) on Florida citrus. In: Evans, G. O. (Ed.), Proceedings of the 2 nd International Congress of Acarology, Sutton Bonington, England (1967), Akademia Kiado, Budapest, pp. 135 - 148." type="journal article" year="1969">Muma 1969</bibRefCitation>
;
<bibRefCitation id="EF9435B927133D4FFB78F8E1FF219260" author="Sanderson" pageId="16" pageNumber="21" refString="Sanderson, J. P. &amp; McMurtry, J. A. (1984) Life history studies of the predaceous mite Phytoseius hawaiiensis. Entomologia experimentalis et applicata., 35, 227 - 234." type="journal article" year="1984">Sanderson &amp; McMurtry 1984</bibRefCitation>
). A similar ratio of 0.45 to 0.48 is evident from the mature, newly-laid egg length versus the maternal dorsal shield length of
<taxonomicName id="4C0533CB27133D4FFE5EF8AEFCF59245" authority="Treat" authorityName="Treat" box="[420,771,1869,1895]" class="Arachnida" family="Ascidae" genus="Blattisocius" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="patagiorum">
<emphasis id="B971945A27133D4FFE5EF8AEFD499244" box="[420,703,1869,1894]" italics="true" pageId="16" pageNumber="17">Blattisocius patagiorum</emphasis>
Treat
</taxonomicName>
given by
<bibRefCitation id="EF9435B927133D4FFC83F8AEFBF99245" author="Treat" box="[889,1039,1869,1895]" pageId="16" pageNumber="21" refString="Treat, A. E. (1966) A new Blattisocius (Acarina: Mesostigmata) from noctuid moths. Journal of the New York Entomological Society, 74, 143 - 159." type="journal article" year="1966">Treat (1966)</bibRefCitation>
. Other than
<taxonomicName id="4C0533CB27133D4FFB65F8AEFA639244" box="[1183,1429,1869,1894]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="grallator">
<emphasis id="B971945A27133D4FFB65F8AEFA639244" box="[1183,1429,1869,1894]" italics="true" pageId="16" pageNumber="17">Opilioseius grallator</emphasis>
</taxonomicName>
, the relatively largest eggs we've encountered are one in a female of
<taxonomicName id="4C0533CB27133D4FFC53F890FAE792AE" authority="Berlese" authorityName="Berlese" box="[937,1297,1906,1932]" class="Arachnida" family="Ascidae" genus="Zercoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="spathuliger">
<emphasis id="B971945A27133D4FFC53F890FB4592AE" box="[937,1203,1907,1932]" italics="true" pageId="16" pageNumber="17">Zercoseius spathuliger</emphasis>
Berlese
</taxonomicName>
with length ratio of 0.67, and those of two undescribed species of
<taxonomicName id="4C0533CB27133D4FFCDDF87BFC4D9293" box="[807,955,1944,1969]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27133D4FFCDDF87BFC4D9293" box="[807,955,1944,1969]" italics="true" pageId="16" pageNumber="17">Aceodromus</emphasis>
</taxonomicName>
, with length ratios of 0.51 to 0.63 from
<collectingCountry id="F31208D827133D4FFF6DF85EFF2992F5" box="[151,223,1981,2007]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
and 0.60 to 0.70 from the La Selva area in
<collectingCountry id="F31208D827133D4FFD2CF85EFCAE92F5" box="[726,856,1981,2007]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
, the highest ratio being from an obviously mature egg containing a pharate larva. We are unaware of any other non-parasitic mesostigmatans having a fully developed dorsal shield (i.e., bearing setae
<emphasis id="B971945A27123D4EFD6AFF5EFD5195F4" box="[656,679,189,214]" italics="true" pageId="17" pageNumber="18">j1</emphasis>
to
<emphasis id="B971945A27123D4EFD36FF5EFD1E95F4" box="[716,744,189,214]" italics="true" pageId="17" pageNumber="18">J5</emphasis>
) that have such proportionately large eggs.
</paragraph>
<paragraph id="8BBA484827123D4EFF3EFF01FC949705" blockId="17.[151,1437,152,551]" pageId="17" pageNumber="18">
These stilt-legged mites coexist with a remarkable variety of other blattisociid mites occupying the same fungi. Although slide preparations and identifications were not made of any of the trombidiform or sarcoptiform (oribatid and astigmatan) mites that may occupy the same fungi, we have determined material of the following mesostigmatic taxa from just one of the several samples from which
<taxonomicName id="4C0533CB27123D4EFB94FEB1FA909449" box="[1134,1382,338,363]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="grallator">
<emphasis id="B971945A27123D4EFB94FEB1FA909449" box="[1134,1382,338,363]" italics="true" pageId="17" pageNumber="18">Opilioseius grallator</emphasis>
</taxonomicName>
was recovered: an undescribed blattisociid genus with one species (which we plan to describe next), 11 species of
<taxonomicName id="4C0533CB27123D4EFF6DFE7EFEE39494" box="[151,277,413,438]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFF6DFE7EFEE39494" box="[151,277,413,438]" italics="true" pageId="17" pageNumber="18">Lasioseius</emphasis>
</taxonomicName>
, four species of
<taxonomicName id="4C0533CB27123D4EFE24FE7EFD9D9494" box="[478,619,413,438]" class="Arachnida" family="Ascidae" genus="Cheiroseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFE24FE7EFD9D9494" box="[478,619,413,438]" italics="true" pageId="17" pageNumber="18">Cheiroseius</emphasis>
</taxonomicName>
, five species of
<taxonomicName id="4C0533CB27123D4EFCCBFE7EFC039494" box="[817,1013,413,438]" class="Arachnida" family="Digamasellidae" genus="Dendrolaelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFCCBFE7EFC039494" box="[817,1013,413,438]" italics="true" pageId="17" pageNumber="18">Dendrolaelaspis</emphasis>
</taxonomicName>
, two species of
<taxonomicName id="4C0533CB27123D4EFB43FE7EFA949494" box="[1209,1378,413,438]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFB43FE7EFA949494" box="[1209,1378,413,438]" italics="true" pageId="17" pageNumber="18">Cosmolaelaps</emphasis>
</taxonomicName>
, and one each of
<taxonomicName id="4C0533CB27123D4EFEDDFE21FE9694F9" box="[295,352,450,475]" class="Arachnida" family="Ascidae" genus="Asca" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFEDDFE21FE9694F9" box="[295,352,450,475]" italics="true" pageId="17" pageNumber="18">Asca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0533CB27123D4EFE95FE21FE1D94F9" box="[367,491,450,475]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFE95FE21FE1D94F9" box="[367,491,450,475]" italics="true" pageId="17" pageNumber="18">Hypoaspis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0533CB27123D4EFE00FE21FD9C94F9" box="[506,618,450,475]" class="Arachnida" family="Parasitidae" genus="Parasitus" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFE00FE21FD9C94F9" box="[506,618,450,475]" italics="true" pageId="17" pageNumber="18">Parasitus</emphasis>
</taxonomicName>
, and an undetermined (perhaps undescribed) genus of
<taxonomicName id="4C0533CB27123D4EFB0BFE21FA8794FE" box="[1265,1393,450,476]" class="Arachnida" family="Veigaiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Veigaiidae</taxonomicName>
. Of these 27 species, 16 were abundant (nine
<taxonomicName id="4C0533CB27123D4EFD83FE0BFD039723" box="[633,757,488,513]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFD83FE0BFD039723" box="[633,757,488,513]" italics="true" pageId="17" pageNumber="18">Lasioseius</emphasis>
</taxonomicName>
, three
<taxonomicName id="4C0533CB27123D4EFCBEFE0BFC399723" box="[836,975,488,513]" class="Arachnida" family="Ascidae" genus="Cheiroseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFCBEFE0BFC399723" box="[836,975,488,513]" italics="true" pageId="17" pageNumber="18">Cheiroseius</emphasis>
</taxonomicName>
, four
<taxonomicName id="4C0533CB27123D4EFBEFFE0BFB239723" box="[1045,1237,488,513]" class="Arachnida" family="Digamasellidae" genus="Dendrolaelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B971945A27123D4EFBEFFE0BFB239723" box="[1045,1237,488,513]" italics="true" pageId="17" pageNumber="18">Dendrolaelaspis</emphasis>
</taxonomicName>
), indicating they were regular members of that particular bracket fungal guild.
</paragraph>
</subSubSection>
</treatment>
</document>
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