684 lines
90 KiB
XML
684 lines
90 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e67569" ID-GBIF-Dataset="9f922048-e978-48d5-86c2-edeb0529c500" ID-GBIF-Taxon="183392463" ID-Pensoft-Pub="1864-8312-79-205" ID-Pensoft-UUID="74D6F43E57D75C658EB5DFB2040EA718" ID-ZooBank="5BE5330D71CA4F12BB2EC7B29A5C33B2" ModsDocID="1864-8312-79-205" checkinTime="1625591104043" checkinUser="pensoft" docAuthor="Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez & Klass, Klaus-Dieter" docDate="2021" docId="4A668C721AD95502974FA7553955584A" docLanguage="en" docName="ArthropodSystPhyl 79: 205-280" docOrigin="Arthropod Systematics & amp; Phylogeny 79" docPubDate="2021-07-05" docSource="http://dx.doi.org/10.3897/asp.79.e67569" docTitle="Attaphila bergi Bolivar 1901" docType="treatment" docVersion="3" id="74D6F43E57D75C658EB5DFB2040EA718" lastPageNumber="205" masterDocId="74D6F43E57D75C658EB5DFB2040EA718" masterDocTitle="Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants" masterLastPageNumber="280" masterPageNumber="205" pageId="0" pageNumber="205" updateTime="1632012750444" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bohn, Horst</mods:namePart>
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<mods:affiliation>Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, 81247 Muenchen, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">bohn@zi.biologie.uni-muenchen.de</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nehring, Volker</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0494-1428</mods:nameIdentifier>
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<mods:affiliation>Universitaet Freiburg, Biologie I, Hauptstrasse 1, 79104 Freiburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>G., Jonathan Rodriguez</mods:namePart>
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<mods:affiliation>Universidad del Valle, Grupo de Ecologia en Agroecosistemas y Habitats Naturales, Calle 13 No. 100 - 00, Ciudad Universitaria Melendez, Cali, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klass, Klaus-Dieter</mods:namePart>
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<mods:affiliation>Senckenberg Natural History Collections Dresden, Museum fuer Tierkunde, Koenigsbruecker Landstrasse 159, 01109 Dresden, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">klaus.klass@senckenberg.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-07-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>205</mods:start>
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<mods:end>280</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.79.e67569</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e67569</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-79-205</mods:identifier>
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<treatment ID-GBIF-Taxon="183392463" LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A" httpUri="http://treatment.plazi.org/id/4A668C721AD95502974FA7553955584A" lastPageNumber="205" pageId="0" pageNumber="205">
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<subSubSection pageId="0" pageNumber="205" type="nomenclature">
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<paragraph pageId="0" pageNumber="205">
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4.2.
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<taxonomicName LSID="4A668C72-1AD9-5502-974F-A7553955584A" authority="Bolivar, 1901" authorityName="Bolivar" authorityYear="1901" class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila bergi" order="Blattodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="bergi">
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Attaphila bergi
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<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
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, 1901
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="205" type="description">
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<paragraph pageId="0" pageNumber="205">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation." figureDoi="10.3897/asp.79.e67569.figure1" httpUri="https://binary.pensoft.net/fig/563270" pageId="0" pageNumber="205">Figs 1E, G</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago." figureDoi="10.3897/asp.79.e67569.figure2" httpUri="https://binary.pensoft.net/fig/563271" pageId="0" pageNumber="205">, 2G</figureCitation>
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<figureCitation captionStart="Figure 3 (A–J)" captionStartId="F3" captionText="Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text." figureDoi="10.3897/asp.79.e67569.figure3(a-f)" httpUri="https://binary.pensoft.net/fig/563272" pageId="0" pageNumber="205">, 3C, D</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct." figureDoi="10.3897/asp.79.e67569.figure5" httpUri="https://binary.pensoft.net/fig/563274" pageId="0" pageNumber="205">, 5C, D</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered)." figureDoi="10.3897/asp.79.e67569.figure7" httpUri="https://binary.pensoft.net/fig/563276" pageId="0" pageNumber="205">, 7A-E</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure12" httpUri="https://binary.pensoft.net/fig/563281" pageId="0" pageNumber="205">, 12B</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure13" httpUri="https://binary.pensoft.net/fig/563282" pageId="0" pageNumber="205">, 13D</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure14" httpUri="https://binary.pensoft.net/fig/563283" pageId="0" pageNumber="205">, 14B</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure15" httpUri="https://binary.pensoft.net/fig/563284" pageId="0" pageNumber="205">, 15A, I, J</figureCitation>
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow)." figureDoi="10.3897/asp.79.e67569.figure16" httpUri="https://binary.pensoft.net/fig/563285" pageId="0" pageNumber="205">, 16A, B</figureCitation>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F." figureDoi="10.3897/asp.79.e67569.figure17" httpUri="https://binary.pensoft.net/fig/563286" pageId="0" pageNumber="205">, 17H</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars." figureDoi="10.3897/asp.79.e67569.figure18" httpUri="https://binary.pensoft.net/fig/563287" pageId="0" pageNumber="205">, 18C, D</figureCitation>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)" figureDoi="10.3897/asp.79.e67569.figure19" httpUri="https://binary.pensoft.net/fig/563288" pageId="0" pageNumber="205">, 19B</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G)." figureDoi="10.3897/asp.79.e67569.figure22" httpUri="https://binary.pensoft.net/fig/563291" pageId="0" pageNumber="205">, 22C, D</figureCitation>
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||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga." figureDoi="10.3897/asp.79.e67569.figure24" httpUri="https://binary.pensoft.net/fig/563293" pageId="0" pageNumber="205">, 24C-G</figureCitation>
|
||
<figureCitation captionStartId="S1" pageId="0" pageNumber="205">, S1C</figureCitation>
|
||
<figureCitation captionStartId="S3" pageId="0" pageNumber="205">, S3B-D</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<taxonomicName authorityName="Bolivar" authorityYear="1901" class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila bergi" order="Blattodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="bergi">Attaphila bergi</taxonomicName>
|
||
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
|
||
, 1901: 335, pl. 6;
|
||
<bibRefCitation author="Princis, K" journalOrPublisher="Insectes Sociaux" pageId="0" pageNumber="205" refId="B67" refString="Princis, K, 1963. Blattariae: Subordo Polyphagoidea: Fam.: Homoeogamiidae, Euthyrrhaphidae, Latindiidae, Anacompsidae, Atticolidae, Attaphilidae. Subordo Blaberoidea: Fam. Blaberidae. In: Beier M (Ed.), Orthopterorum Catalogus Pars 4: 75-172. Junk, s'Gravenhage." title="Blattariae: Subordo Polyphagoidea: Fam.: Homoeogamiidae, Euthyrrhaphidae, Latindiidae, Anacompsidae, Atticolidae, Attaphilidae. Subordo Blaberoidea: Fam. Blaberidae. In: Beier M (Ed.), Orthopterorum Catalogus Pars 4: 75 - 172. Junk, s'Gravenhage." year="1963">Princis 1963</bibRefCitation>
|
||
: 111.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<taxonomicName class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila bergi subsp. var. var. minor" order="Blattodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="bergi" subSpecies="var." variety="minor">Attaphila bergi var. minor</taxonomicName>
|
||
Bruch, 1916: 329, fig. 17A-F.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="205">Material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Type material</emphasis>
|
||
(presumably lost).
|
||
<materialsCitation ID-GBIF-Occurrence="3314641315" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Argentina" location="Material" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Buenos Aires" typeStatus="Syntypes">
|
||
<typeStatus>Syntypes</typeStatus>
|
||
,
|
||
<specimenCount type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount type="female">6♀</specimenCount>
|
||
,
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
(
|
||
<collectingRegion country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
|
||
?),
|
||
<collectingCountry name="Uruguay">Uruguay</collectingCountry>
|
||
. -
|
||
<emphasis bold="true" pageId="0" pageNumber="205">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:B0B63A0E38D34954E8BC1056540C9E2E" country="Argentina" name="Material" stateProvince="Buenos Aires">Material</location>
|
||
studied
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3314641328" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Argentina" location="Bo" municipality="Bo" specimenCount="2" specimenCount-female="2" typeStatus="syntype">
|
||
<specimenCount type="female">2♀</specimenCount>
|
||
, [
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
], B[ueno]s. A[ire]s (abdomen of each on one slide:
|
||
<collectingMunicipality>Bo</collectingMunicipality>
|
||
1284,
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:03E66AAFB474BE18BAC4BBF83772DD7C" country="Argentina" municipality="Bo" name="Bo">Bo</location>
|
||
1440); 4L
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3314641334" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Prov., Aires, Castelar, M, Viana, R. Maniglia" country="Argentina" location="Argentina" specimenCount="♀" typeStatus="syntype">
|
||
<specimenCount>♀</specimenCount>
|
||
,
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
, [
|
||
<collectorName>Prov.</collectorName>
|
||
] B[ueno]s
|
||
<collectorName>Aires</collectorName>
|
||
,
|
||
<collectorName>Castelar, M</collectorName>
|
||
.
|
||
<collectorName>Viana</collectorName>
|
||
and
|
||
<collectorName>R.Maniglia</collectorName>
|
||
, # 41175 (together with a worker of
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
on one pin); 4L
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3314641313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Prov. & Aires & Castelar, M & Viana & R. Maniglia" country="Argentina" location="Bo" specimenCount="♀" typeStatus="syntype">
|
||
<specimenCount>♀</specimenCount>
|
||
, same data as preceding (together with a worker of
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
on one pin) (abdomen of 1L
|
||
<specimenCount>♀</specimenCount>
|
||
on one slide:
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:BD2414A41BC59E0FCFD8F3A6C396F1AB" country="Argentina" name="Bo">Bo</location>
|
||
1285)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3314641323" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Argentina" location="Prov." specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Entre Rios" typeStatus="syntype">
|
||
<specimenCount type="male">3♂</specimenCount>
|
||
,
|
||
<specimenCount type="female">1♀</specimenCount>
|
||
, [
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
], [
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:34C0CA5DBCA3FB1CAA72138583675B50" country="Argentina" name="Prov." stateProvince="Entre Rios">Prov.</location>
|
||
]
|
||
<collectingRegion country="Argentina" name="Entre Rios">
|
||
Entre
|
||
<normalizedToken originalValue="Ríos">Rios</normalizedToken>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation ID-GBIF-Occurrence="3314641306" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Paranacito, Daguerre" country="Argentina" location="Bo" municipality="Bo" specimenCount="2" specimenCount-male="2" stateProvince="Buenos Aires" typeStatus="syntype">
|
||
<collectorName>Paranacito</collectorName>
|
||
, [1931, acc. to loan form],
|
||
<collectorName>Daguerre</collectorName>
|
||
, # 28 994 (together with a worker of
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
on one pin) (
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
completely on two slides: Bo 1286; abdomen and tegmina of
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
on two slides:
|
||
<collectingMunicipality>Bo</collectingMunicipality>
|
||
1443; abdomen of
|
||
<specimenCount>♀</specimenCount>
|
||
on one slide:
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:37D237241AAC7811C7309E7C819C3E05" country="Argentina" municipality="Bo" name="Bo" stateProvince="Buenos Aires">Bo</location>
|
||
1275) (M.
|
||
<collectingRegion country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
. -
|
||
<materialsCitation ID-GBIF-Occurrence="3314641317" collectingDate="1918-07" collectorName="Moreno BA, M. D. Jurado, M. Maastricht, La Plata, C. Bruch" country="Argentina" county="Bo" location="La Plata" municipality="Bo" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Buenos Aires" typeStatus="syntype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount type="female">1♀</specimenCount>
|
||
, [
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
, Prov.
|
||
<collectingRegion country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
|
||
],
|
||
<collectorName>Moreno BA</collectorName>
|
||
, 30.XI.[19]38,
|
||
<collectorName>M.D.Jurado</collectorName>
|
||
, # MACN-En 7781/7780 (
|
||
<specimenCount>♂</specimenCount>
|
||
completely on two slides:
|
||
<collectingCounty>Bo</collectingCounty>
|
||
1283; abdomen and part of legs of the
|
||
<specimenCount>♀</specimenCount>
|
||
on one slide:
|
||
<collectingMunicipality>Bo</collectingMunicipality>
|
||
1282) (
|
||
<collectorName>M. Maastricht</collectorName>
|
||
, NHMM 007, 008); 10L,
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
, [Prov.
|
||
<collectingRegion country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
|
||
],
|
||
<collectorName>
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:9523A4D2EAA5EE8D73A01E2754D283F1" country="Argentina" county="Bo" municipality="Bo" name="La Plata" stateProvince="Buenos Aires">La Plata</location>
|
||
</collectorName>
|
||
,
|
||
<collectingDate value="1918-07">VII.1918</collectingDate>
|
||
,
|
||
<collectorName>C.Bruch</collectorName>
|
||
(on 4 pins, each with a worker of
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
) (1L
|
||
<specimenCount>♂</specimenCount>
|
||
completely on two slides: Bo 1230; 1L
|
||
<specimenCount>♀</specimenCount>
|
||
completely on two slides:
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:F026E3D9ADD7A44DF0E6B0E299D0431A" country="Argentina" county="Bo" municipality="Bo" name="Bo" stateProvince="Buenos Aires">Bo</location>
|
||
1231) (
|
||
<collectorName>M. Maastricht</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
. -
|
||
<materialsCitation ID-GBIF-Occurrence="3314641304" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Argentina" location="Prov." specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios" typeStatus="syntype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
[
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
], [
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:360AF9625C540BFF620C663D33C95AF6" country="Argentina" name="Prov." stateProvince="Entre Rios">Prov.</location>
|
||
]
|
||
<collectingRegion country="Argentina" name="Entre Rios">
|
||
Entre
|
||
<normalizedToken originalValue="Ríos">Rios</normalizedToken>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation ID-GBIF-Occurrence="3314641347" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Paranacito, Daguerre" country="Argentina" location="Bo" specimenCount="1" stateProvince="Entre Rios" typeStatus="syntype">
|
||
<collectorName>Paranacito</collectorName>
|
||
, [1931, acc. to loan form],
|
||
<collectorName>Daguerre</collectorName>
|
||
, # 28 944 (from a pin with a worker of
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
) (completely on two slides:
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:9EE81FB9CCC8F05EEFEB078068F5E027" country="Argentina" name="Bo" stateProvince="Entre Rios">Bo</location>
|
||
1274)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3314641307" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Prov., Rosario, Coll. Hubrich" country="Argentina" location="Bo" specimenCount="1" specimenCount-female="1" stateProvince="Santa Fe" typeStatus="syntype">
|
||
<specimenCount type="female">1♀</specimenCount>
|
||
, [
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
,
|
||
<collectorName>Prov.</collectorName>
|
||
<collectingRegion country="Argentina" name="Santa Fe">
|
||
Santa
|
||
<normalizedToken originalValue="Fé">Fe</normalizedToken>
|
||
</collectingRegion>
|
||
],
|
||
<collectorName>Rosario</collectorName>
|
||
,
|
||
<collectorName>Coll. Hubrich</collectorName>
|
||
, # 15 A 16 (completely on two slides:
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:AA3AE002B94EFF54C9594327C18D872B" country="Argentina" name="Bo" stateProvince="Santa Fe">Bo</location>
|
||
1239); 6L
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3314641324" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Rosario, Coll. Hubrich, ZS Munich" country="Argentina" location="Bo" specimenCount="♀" typeStatus="syntype">
|
||
<specimenCount>♀</specimenCount>
|
||
, [
|
||
<collectingCountry name="Argentina">Argentina</collectingCountry>
|
||
],
|
||
<collectorName>Rosario</collectorName>
|
||
(?),
|
||
<collectorName>Coll. Hubrich</collectorName>
|
||
, # R.66, B.p.77 (together with a worker of
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
on one cardboard) (1L
|
||
<specimenCount>♀</specimenCount>
|
||
completely on one slide:
|
||
<location LSID="urn:lsid:plazi:treatment:4A668C721AD95502974FA7553955584A:C142DBC422302E76C1D9658477EE4405" country="Argentina" name="Bo">Bo</location>
|
||
1238) (
|
||
<collectorName>ZS Munich</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="205">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Male</emphasis>
|
||
Well characterised by several features unique in
|
||
<taxonomicName class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila" order="Isoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Attaphila</emphasis>
|
||
</taxonomicName>
|
||
: the rather inconspicuous specialisation anteromedially on
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T1</emphasis>
|
||
with a pair of areas showing strongly developed microreticulation (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">msa1</emphasis>
|
||
), the bristle distribution on
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T6,7</emphasis>
|
||
with some bristles on the surface of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T6</emphasis>
|
||
and long bristles along the posterior borders, and the specially shaped virga (with a very strong sinusoidal curvature). With
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
it shares the absence of long bristles on surface and borders of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T1</emphasis>
|
||
.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Female</emphasis>
|
||
: Distinguished from all other species by
|
||
<emphasis bold="true" pageId="0" pageNumber="205">S7</emphasis>
|
||
having a continuous, uninterrupted transversal ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="description">
|
||
<paragraph pageId="0" pageNumber="205">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Size</emphasis>
|
||
Length of body (dried, type specimens after
|
||
<bibRefCitation author="Bolivar, I" journalOrPublisher="Comunicaciones del Museo Nacional de Buenos Aires" pageId="0" pageNumber="205" pagination="331 - 336" refId="B11" refString="Bolivar, I, 1901. Un nuevo ortoptero mirmecofilo Attaphila Bergi. Comunicaciones del Museo Nacional de Buenos Aires 1 (10): 331 - 336" title="Un nuevo ortoptero mirmecofilo Attaphila Bergi." volume="1" year="1901">
|
||
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
|
||
1901
|
||
</bibRefCitation>
|
||
): male 2.8 mm, female 2.8 mm.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Surface bristles of tergites 2-5</emphasis>
|
||
arranged in about two irregular transversal rows (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered)." figureDoi="10.3897/asp.79.e67569.figure7" httpUri="https://binary.pensoft.net/fig/563276" pageId="0" pageNumber="205">7A, D</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="205">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Transversal ridges</emphasis>
|
||
tr2-5
|
||
</emphasis>
|
||
without distinct excurvations (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered)." figureDoi="10.3897/asp.79.e67569.figure7" httpUri="https://binary.pensoft.net/fig/563276" pageId="0" pageNumber="205">7A, D</figureCitation>
|
||
, compare grey arrows in Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior." figureDoi="10.3897/asp.79.e67569.figure11" httpUri="https://binary.pensoft.net/fig/563280" pageId="0" pageNumber="205">11A</figureCitation>
|
||
, and in Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2." figureDoi="10.3897/asp.79.e67569.figure9" httpUri="https://binary.pensoft.net/fig/563278" pageId="0" pageNumber="205">9B</figureCitation>
|
||
for male
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr2</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Male</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Tegmina</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 3 (A–J)" captionStartId="F3" captionText="Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text." figureDoi="10.3897/asp.79.e67569.figure3(a-f)" httpUri="https://binary.pensoft.net/fig/563272" pageId="0" pageNumber="205">3C, D</figureCitation>
|
||
) widest at about 2/3 of length; overall shape more rectangular than triangular; posterior border straight or slightly concave near tegmen base (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">fwar</emphasis>
|
||
), slightly converging with the apical border, soon bending into a rather tight curvation (as compared to
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
), finally joining the apical border in a still tighter curvation (though less tight than in
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
); apical border transversal, fairly straight; surface bristles strong (stronger than in
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Hindwings</emphasis>
|
||
lanceolate, with narrowly rounded apex (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation." figureDoi="10.3897/asp.79.e67569.figure1" httpUri="https://binary.pensoft.net/fig/563270" pageId="0" pageNumber="205">1G</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Glandular pores</emphasis>
|
||
only on
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T1</emphasis>
|
||
, in the area of the specialisation and some more around (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct." figureDoi="10.3897/asp.79.e67569.figure5" httpUri="https://binary.pensoft.net/fig/563274" pageId="0" pageNumber="205">5C, D</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Tergite 1</emphasis>
|
||
without long bristles on surface and borders (in contrast to long bristles on
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T2-5</emphasis>
|
||
); medially, about at the level of the medially obsolete ridge
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr1</emphasis>
|
||
(far anteriorly on
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T1-a</emphasis>
|
||
), with an inconspicuous specialisation (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">msa1</emphasis>
|
||
) consisting of a pair of small areas showing a net-like pattern produced by delicate furrows or ridges along which tiny glandular pores are sporadically arranged; area in between either unstructured or with dispersed small pores, occasionally also with few small bristles (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct." figureDoi="10.3897/asp.79.e67569.figure5" httpUri="https://binary.pensoft.net/fig/563274" pageId="0" pageNumber="205">5C, D</figureCitation>
|
||
). Ridge
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr1</emphasis>
|
||
interrupted by the specialisation.
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Tergite 2</emphasis>
|
||
without specialisations.
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Tergites 6,7</emphasis>
|
||
: (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered)." figureDoi="10.3897/asp.79.e67569.figure7" httpUri="https://binary.pensoft.net/fig/563276" pageId="0" pageNumber="205">7B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure13" httpUri="https://binary.pensoft.net/fig/563282" pageId="0" pageNumber="205">13D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure15" httpUri="https://binary.pensoft.net/fig/563284" pageId="0" pageNumber="205">15A, I</figureCitation>
|
||
(larval)) Median lobe of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T7</emphasis>
|
||
very short, but distinct; transversal ridge
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr6</emphasis>
|
||
well developed,
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr7</emphasis>
|
||
rather weakly developed, perhaps with gaps in lateral parts; posterior border of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T6</emphasis>
|
||
and
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T7</emphasis>
|
||
with a row of rather long and strong bristles, almost reaching the size of those of the lateral borders; surface of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T6</emphasis>
|
||
in the posterior half with dispersed bristles of slightly smaller size, sometimes distributed in two groups, of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T7</emphasis>
|
||
only with few isolated and usually much smaller bristles.
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Subgenital lobe</emphasis>
|
||
: (Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga." figureDoi="10.3897/asp.79.e67569.figure24" httpUri="https://binary.pensoft.net/fig/563293" pageId="0" pageNumber="205">24C, D</figureCitation>
|
||
) distal part with a deep excavation only along left side; lobe posterior to level of excavation widely tongue-shaped; with two styli, the larger left one (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">sll</emphasis>
|
||
) conical, inserted at base of excavation, not reaching tip of lobe, the much smaller right one (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">slr</emphasis>
|
||
, relatively larger than in
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
) knob-like, situated subterminally on right flank of tip; stylus and borders of distal lobe with numerous bristles, most of them large (on average longer and stronger than in
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
), some rather long and strong.
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Phallomeres</emphasis>
|
||
: (Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga." figureDoi="10.3897/asp.79.e67569.figure24" httpUri="https://binary.pensoft.net/fig/563293" pageId="0" pageNumber="205">24D-G</figureCitation>
|
||
) Sclerotised part of hook (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">h</emphasis>
|
||
) from the long, wide base (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">b</emphasis>
|
||
) very gradually narrowing into a slender, very short neck (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">n</emphasis>
|
||
; shorter than in
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
) which is evenly curved into the slightly wider claw part (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">cl</emphasis>
|
||
; neck thus hardly set off from both the base and the claw part). Endophallic apodeme (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">ea</emphasis>
|
||
) not narrowed at base (forking site of sclerite). Relative to the axis of the endophallic apodeme (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">ea</emphasis>
|
||
), the virga (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">vi</emphasis>
|
||
) shows a distinct, deep sinusoidal curvature to the left, the apical part being very strongly and tightly curved (back into the
|
||
<emphasis bold="true" pageId="0" pageNumber="205">ea</emphasis>
|
||
-axis), and gradually narrowed to a rather acute tip ending shortly after the termination of the recurvation, hence hook-like (presence of a groove on the virga unclear).
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Paraprocts</emphasis>
|
||
: Both lacking a sclerotised projection (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered)." figureDoi="10.3897/asp.79.e67569.figure7" httpUri="https://binary.pensoft.net/fig/563276" pageId="0" pageNumber="205">7C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="205">Female</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Tergites 6,7</emphasis>
|
||
: (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered)." figureDoi="10.3897/asp.79.e67569.figure7" httpUri="https://binary.pensoft.net/fig/563276" pageId="0" pageNumber="205">7E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure12" httpUri="https://binary.pensoft.net/fig/563281" pageId="0" pageNumber="205">12B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure14" httpUri="https://binary.pensoft.net/fig/563283" pageId="0" pageNumber="205">14B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7." figureDoi="10.3897/asp.79.e67569.figure15" httpUri="https://binary.pensoft.net/fig/563284" pageId="0" pageNumber="205">15J</figureCitation>
|
||
(larval)) Median lobe of
|
||
<emphasis bold="true" pageId="0" pageNumber="205">T7</emphasis>
|
||
rather long; transversal ridges
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr6</emphasis>
|
||
and
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr7</emphasis>
|
||
complete,
|
||
<emphasis bold="true" pageId="0" pageNumber="205">tr6</emphasis>
|
||
sublaterally with a distinct bend; surface bristles numerous and rather long and strong (in length comparable to those of
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
, but less strong), strongly focused to central part of surface area.
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Subgenital plate</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow)." figureDoi="10.3897/asp.79.e67569.figure16" httpUri="https://binary.pensoft.net/fig/563285" pageId="0" pageNumber="205">16A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars." figureDoi="10.3897/asp.79.e67569.figure18" httpUri="https://binary.pensoft.net/fig/563287" pageId="0" pageNumber="205">18C, D</figureCitation>
|
||
) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">sr7-l</emphasis>
|
||
) almost straight; median part (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">sr7-m</emphasis>
|
||
) complete, slightly (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow)." figureDoi="10.3897/asp.79.e67569.figure16" httpUri="https://binary.pensoft.net/fig/563285" pageId="0" pageNumber="205">16A</figureCitation>
|
||
) to strongly (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow)." figureDoi="10.3897/asp.79.e67569.figure16" httpUri="https://binary.pensoft.net/fig/563285" pageId="0" pageNumber="205">16B</figureCitation>
|
||
) bisinuate.
|
||
<emphasis bold="true" pageId="0" pageNumber="205">Genitalia</emphasis>
|
||
: Spermathecal plate
|
||
<emphasis bold="true" pageId="0" pageNumber="205">sp</emphasis>
|
||
rather small, pouch
|
||
<emphasis bold="true" pageId="0" pageNumber="205">gcp</emphasis>
|
||
rather small (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)" figureDoi="10.3897/asp.79.e67569.figure19" httpUri="https://binary.pensoft.net/fig/563288" pageId="0" pageNumber="205">19B</figureCitation>
|
||
). In laterosternal shelf sclerite (Fig.
|
||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G)." figureDoi="10.3897/asp.79.e67569.figure22" httpUri="https://binary.pensoft.net/fig/563291" pageId="0" pageNumber="205">22C, D</figureCitation>
|
||
) central part (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">c</emphasis>
|
||
) moderately long and posteriorly transversally cut, arms (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">a</emphasis>
|
||
) very narrow (narrower than in
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. aptera" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="aptera">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. aptera</emphasis>
|
||
</taxonomicName>
|
||
), wing parts (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">w</emphasis>
|
||
) moderately wide, their base reaching far into anterior half of tubes (blue arrowhead); tubes (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">lst</emphasis>
|
||
) straight, of fairly uniform width throughout, anterior end widely rounded. Mesal gonangulum sclerites (
|
||
<emphasis bold="true" pageId="0" pageNumber="205">gg-m</emphasis>
|
||
) distinct, lateral ones absent (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)" figureDoi="10.3897/asp.79.e67569.figure19" httpUri="https://binary.pensoft.net/fig/563288" pageId="0" pageNumber="205">19B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="host">
|
||
<paragraph pageId="0" pageNumber="205">Host species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">Acromyrmex lundii</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
|
||
, 1838); host species of var.
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii var. minor" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="lundii" variety="minor">Acromyrmex lundii minor</taxonomicName>
|
||
according to
|
||
<bibRefCitation author="Bruch, C" journalOrPublisher="Insectes Sociaux" pageId="0" pageNumber="205" refId="B19" refString="Bruch, C, 1916. Contribucion de las Hormigas de la Provincia de San Luis. Revista del Museo de La Plata 2. Ser., Tomo 10, 23: 291-357." title="Contribucion de las Hormigas de la Provincia de San Luis. Revista del Museo de La Plata 2. Ser., Tomo 10, 23: 291 - 357." year="1916">Bruch (1916)</bibRefCitation>
|
||
<taxonomicName genus="Acr." lsidName="Acr. lobicornis" pageId="0" pageNumber="205" rank="species" species="lobicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acr. lobicornis</emphasis>
|
||
</taxonomicName>
|
||
(Emery, 1888), and
|
||
<taxonomicName authority="" authorityName="" class="Insecta" family="Formicidae" genus="Amoimyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amoimyrmex silvestrii" order="Mantophasmatodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="silvestrii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Amoimyrmex silvestrii</emphasis>
|
||
</taxonomicName>
|
||
(Emery, 1905).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="distribution">
|
||
<paragraph pageId="0" pageNumber="205">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
Argentina: Prov. Buenos Aires (Castelar, La Plata, Moreno), Prov. Entre Rios (Paranacito), Prov. Santa
|
||
<normalizedToken originalValue="Fé">Fe</normalizedToken>
|
||
(Rosario). The
|
||
<taxonomicName class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila bergi subsp. var. var. minor" order="Isoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="bergi" subSpecies="var." variety="minor">Attaphila bergi var. minor</taxonomicName>
|
||
is reported from the Provinces San Luis and Catamarca; the report of
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. bergi" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="bergi">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. bergi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<normalizedToken originalValue="Córdoba">Cordoba</normalizedToken>
|
||
(
|
||
<bibRefCitation author="Bruch, C" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="205" pagination="421 - 437" refId="B20" refString="Bruch, C, 1929. Neue myrmekophile Histeriden und Verzeichnis der aus Argentinien bekannten Ameisengaeste. Zoologischer Anzeiger 82: 421 - 437" title="Neue myrmekophile Histeriden und Verzeichnis der aus Argentinien bekannten Ameisengaeste." volume="82" year="1929">Bruch 1929</bibRefCitation>
|
||
) most likely is a mistake and refers to the occurrence of
|
||
<taxonomicName class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila bergi subsp. var. var. minor" order="Isoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="bergi" subSpecies="var." variety="minor">Attaphila bergi var. minor</taxonomicName>
|
||
in San Luis, not mentioned in this paper; Uruguay.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="205" type="remarks">
|
||
<paragraph pageId="0" pageNumber="205">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
The loss of the type specimens of
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. bergi" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="bergi">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. bergi</emphasis>
|
||
</taxonomicName>
|
||
and the insufficient original description of the species prevent an unequivocal identification. But for several reasons the determination of the available Argentine specimens as
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. bergi" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="bergi">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. bergi</emphasis>
|
||
</taxonomicName>
|
||
is most probably correct: They have the same host ant species,
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
; no other species of
|
||
<taxonomicName class="Insecta" family="Ectobiidae" genus="Attaphila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attaphila" order="Isoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Attaphila</emphasis>
|
||
</taxonomicName>
|
||
has so far been found in nests of this ant. The specimens studied are from at least 5 different localities in rather close distance, not very far from the localities of the type specimens (Argentina: Buenos Aires?, Uruguay), without showing remarkable differences among each other.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="205">
|
||
<bibRefCitation author="Bruch, C" journalOrPublisher="Insectes Sociaux" pageId="0" pageNumber="205" refId="B19" refString="Bruch, C, 1916. Contribucion de las Hormigas de la Provincia de San Luis. Revista del Museo de La Plata 2. Ser., Tomo 10, 23: 291-357." title="Contribucion de las Hormigas de la Provincia de San Luis. Revista del Museo de La Plata 2. Ser., Tomo 10, 23: 291 - 357." year="1916">Bruch (1916)</bibRefCitation>
|
||
described a
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex silvestrii subsp. var. var. minor" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="silvestrii" subSpecies="var." variety="minor">Acromyrmex silvestrii var. minor</taxonomicName>
|
||
occurring in nests of
|
||
<taxonomicName authority="" authorityName="" class="Insecta" family="Formicidae" genus="Amoimyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amoimyrmex silvestrii" order="Mantophasmatodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="silvestrii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Amoimyrmex silvestrii</emphasis>
|
||
</taxonomicName>
|
||
(Prov. San Luis) and
|
||
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1888" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lobicornis" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lobicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lobicornis</emphasis>
|
||
</taxonomicName>
|
||
(Prov. Catamarca; belonging to
|
||
<taxonomicName authority="" authorityName="" class="Insecta" family="Formicidae" genus="Amoimyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amoimyrmex" order="Mantophasmatodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Amoimyrmex</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex</emphasis>
|
||
</taxonomicName>
|
||
clade not yet phylogenetically tested), of smaller size and paler colour than the nominate form. This characterisation and the missing of males raise the suspicion that the type specimens were larvae, possibly of normal
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. bergi" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="bergi">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. bergi</emphasis>
|
||
</taxonomicName>
|
||
. On the other hand, the complete or partial association with
|
||
<taxonomicName authority="" authorityName="" class="Insecta" family="Formicidae" genus="Amoimyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amoimyrmex" order="Mantophasmatodea" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Amoimyrmex</emphasis>
|
||
</taxonomicName>
|
||
hosts, which form the sister taxon of
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atta" order="Embiidina" pageId="0" pageNumber="205" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Atta</emphasis>
|
||
</taxonomicName>
|
||
and are thus phylogenetically far remote from
|
||
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="species" species="lundii">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">Acromyrmex lundii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.1111/aen.12493" author="Cristiano, MP" editor="Cristiano, MP" journalOrPublisher="Austral Entomology" pageId="0" pageNumber="205" pagination="643 - 676" refId="B25" refString="Cristiano, MP, Cardoso, DC, Sandoval-Gomez, VE, 2020. Amoimyrmex gen. nov. and relevant taxonomic changes attributed to Cristiano, M. P.; Cardoso, D. C.; Sandoval-Gomez, V. E. Pp. 647-674. In: Cristiano, MP, Cardoso, DC, Sandoval-Gomez, VE, Simoes-Gomes, F, Eds., nov. (Hymenoptera: Formicidae): a new genus of leaf-cutting ants revealed by multilocus molecular phylogenetic and morphological analyses. Austral Entomology 59: 643 - 676, DOI: https://doi.org/10.1111/aen.12493" title="Amoimyrmex gen. nov. and relevant taxonomic changes attributed to Cristiano, M. P.; Cardoso, D. C.; Sandoval-Gomez, V. E. Pp. 647 - 674" url="https://doi.org/10.1111/aen.12493" volume="59" volumeTitle="nov. (Hymenoptera: Formicidae): a new genus of leaf-cutting ants revealed by multilocus molecular phylogenetic and morphological analyses." year="2020">Cristiano et al. 2020</bibRefCitation>
|
||
) raises doubt on
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex lundii subsp. var. var. minor" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="lundii" subSpecies="var." variety="minor">Acromyrmex lundii var. minor</taxonomicName>
|
||
being conspecific with
|
||
<taxonomicName family="Ectobiidae" genus="A." kingdom="Animalia" lsidName="A. bergi" order="Blattodea" pageId="0" pageNumber="205" rank="species" species="bergi">
|
||
<emphasis italics="true" pageId="0" pageNumber="205">A. bergi</emphasis>
|
||
</taxonomicName>
|
||
. The type specimens of
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex bergi subsp. var. var. minor" order="Orthoptera" pageId="0" pageNumber="205" phylum="Arthropoda" rank="variety" species="bergi" subSpecies="var." variety="minor">Acromyrmex bergi var. minor</taxonomicName>
|
||
, formerly deposited in the Museo de La Plata, are no longer present there and presumably lost.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |