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<document id="D098EC717B4F4E7B25CDA5842F17B6B9" ID-CLB-Dataset="302078" ID-DOI="10.11646/phytotaxa.234.3.3" ID-GBIF-Dataset="c1ee2a05-2056-4a94-a5bd-ee2a845c917a" ID-ISSN="1179-3163" ID-Zenodo-Dep="13632963" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725319495140" checkinUser="felipe" docAuthor="Frankovich, Thomas A., Sullivan, Michael J. &amp; Stacy, Nicole I." docDate="2015" docId="038BD034154C650EFF103AEA6B17F87C" docLanguage="en" docName="phytotaxa.234.3.3.pdf" docOrigin="Phytotaxa 234 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.234.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Tursiocola denysii Frankovich &amp; M. J. Sullivan 2015, sp. nov." docType="treatment" docVersion="3" lastPageNumber="231" masterDocId="FFB2A84C154D650AFF983F3C680EFFC4" masterDocTitle="Tursiocola denysii sp. nov. (Bacillariophyta) from the neck skin of Loggerhead sea turtles (Caretta caretta)" masterLastPageNumber="236" masterPageNumber="227" pageNumber="228" updateTime="1725325296098" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3F4F3C46CC0BE1298276346361EB507E">Tursiocola denysii sp. nov. (Bacillariophyta) from the neck skin of Loggerhead sea turtles (Caretta caretta)</mods:title>
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<treatment id="038BD034154C650EFF103AEA6B17F87C" ID-DOI="http://doi.org/10.5281/zenodo.13632967" ID-GBIF-Taxon="240066361" ID-Zenodo-Dep="13632967" LSID="urn:lsid:plazi:treatment:038BD034154C650EFF103AEA6B17F87C" httpUri="http://treatment.plazi.org/id/038BD034154C650EFF103AEA6B17F87C" lastPageId="4" lastPageNumber="231" pageId="1" pageNumber="228">
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<heading id="D0D5D64E154C650BFF103AEA6BACFA2B" bold="true" box="[136,930,1494,1519]" fontSize="11" level="1" pageId="1" pageNumber="228" reason="1">
<emphasis id="B956BD30154C650BFF103AEA6BACFA2B" bold="true" box="[136,930,1494,1519]" pageId="1" pageNumber="228">
<taxonomicName id="4C221AA1154C650BFF103AEA6ABAFA2B" authority="Frankovich &amp; M.J. Sullivan" authorityName="Frankovich &amp; M. J. Sullivan" authorityYear="2015" box="[136,692,1494,1519]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="1" pageNumber="228" phylum="Ochrophyta" rank="species" species="denysii" status="sp. nov.">
<emphasis id="B956BD30154C650BFF103AEA6955FA2B" bold="true" box="[136,347,1494,1519]" italics="true" pageId="1" pageNumber="228">Tursiocola denysii</emphasis>
Frankovich &amp; M.J. Sullivan
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,
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<emphasis id="B956BD30154C650BFD583AEA6AD7FA2B" bold="true" box="[704,729,1494,1519]" italics="true" pageId="1" pageNumber="228">sp</emphasis>
.
<emphasis id="B956BD30154C650BFD7F3AEA6B1FFA2B" bold="true" box="[743,785,1494,1519]" italics="true" pageId="1" pageNumber="228">nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13197DA7154C650BFCBE3AEA6B94FA2B" box="[806,922,1494,1519]" captionStart-0="FIGURES 114" captionStart-1="FIGURES 1521" captionStart-2="FIGURES 2224" captionStart-3="FIGURES 2530" captionStartId-0="1.[136,243,1941,1962]" captionStartId-1="2.[136,243,1487,1508]" captionStartId-2="3.[136,243,1900,1921]" captionStartId-3="4.[136,243,1305,1326]" captionTargetBox-0="[276,1312,1548,1911]" captionTargetBox-1="[169,1418,190,1461]" captionTargetBox-2="[382,1205,1252,1875]" captionTargetBox-3="[374,1213,191,1280]" captionTargetId-0="figure-496@1.[270,1317,1543,1916]" captionTargetId-1="figure-17@2.[168,1419,190,1462]" captionTargetId-2="figure-698@3.[381,1205,1251,1875]" captionTargetId-3="figure-17@4.[373,1213,190,1281]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." captionText-1="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." captionText-2="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." captionText-3="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13632965" figureDoi-1="http://doi.org/10.5281/zenodo.13632969" figureDoi-2="http://doi.org/10.5281/zenodo.13632971" figureDoi-3="http://doi.org/10.5281/zenodo.13632975" httpUri-0="https://zenodo.org/record/13632965/files/figure.png" httpUri-1="https://zenodo.org/record/13632969/files/figure.png" httpUri-2="https://zenodo.org/record/13632971/files/figure.png" httpUri-3="https://zenodo.org/record/13632975/files/figure.png" pageId="1" pageNumber="228">Figs 130</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<caption id="DF5D31AA154C650BFF1038A96A87F836" ID-DOI="http://doi.org/10.5281/zenodo.13632965" ID-Zenodo-Dep="13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="1" pageNumber="228" startId="1.[136,243,1941,1962]" targetBox="[276,1312,1548,1911]" targetPageId="1" targetType="figure">
<paragraph id="8B9D6122154C650BFF1038A96A87F836" blockId="1.[136,1453,1941,2034]" pageId="1" pageNumber="228">
<emphasis id="B956BD30154C650BFF1038A96926F86E" bold="true" box="[136,296,1941,1962]" pageId="1" pageNumber="228">FIGURES 114</emphasis>
.
<taxonomicName id="4C221AA1154C650BFEAB38A969EDF86E" authorityName="Frankovich &amp; M. J. Sullivan" authorityYear="2015" box="[307,483,1941,1962]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="1" pageNumber="228" phylum="Ochrophyta" rank="species" species="denysii">
<emphasis id="B956BD30154C650BFEAB38A969EDF86E" box="[307,483,1941,1962]" italics="true" pageId="1" pageNumber="228">Tursiocola denysii</emphasis>
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. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci.
</paragraph>
</caption>
<caption id="DF5D31AA154F6508FF103AF369D4F95C" ID-DOI="http://doi.org/10.5281/zenodo.13632969" ID-Zenodo-Dep="13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="2" pageNumber="229" startId="2.[136,243,1487,1508]" targetBox="[169,1418,190,1461]" targetPageId="2" targetType="figure">
<paragraph id="8B9D6122154F6508FF103AF369D4F95C" blockId="2.[136,1452,1487,1688]" pageId="2" pageNumber="229">
<emphasis id="B956BD30154F6508FF103AF36939FA20" bold="true" box="[136,311,1487,1508]" pageId="2" pageNumber="229">FIGURES 1521</emphasis>
.
<taxonomicName id="4C221AA1154F6508FEDE3AF369F7FA20" authorityName="Frankovich &amp; M. J. Sullivan" authorityYear="2015" box="[326,505,1487,1508]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="2" pageNumber="229" phylum="Ochrophyta" rank="species" species="denysii">
<emphasis id="B956BD30154F6508FEDE3AF369F7FA20" box="[326,505,1487,1508]" italics="true" pageId="2" pageNumber="229">Tursiocola denysii</emphasis>
</taxonomicName>
. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm.
</paragraph>
</caption>
<paragraph id="8B9D6122154F6509FF1039F769D4FB99" blockId="2.[136,1452,1739,2084]" lastBlockId="3.[136,1453,160,1225]" lastPageId="3" lastPageNumber="230" pageId="2" pageNumber="229">
LM morphology: frustules are rectangular in girdle view with bluntly rounded ends and porose girdle copulae (
<figureCitation id="13197DA7154F6508FB3239F76D0FF924" box="[1194,1281,1739,1760]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="2" pageNumber="229">Figs 14</figureCitation>
). Pseudosepta are evident in girdle view and are deflected towards the interior of the frustule at their termination (
<figureCitation id="13197DA7154F6508FBC139D36CC7F8C0" box="[1113,1225,1775,1796]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="2" pageNumber="229">Figs 12, 4</figureCitation>
). Valves are isopolar to slightly heteropolar and narrowly lanceolate, tapering gradually from the middle of the valves to the juncture of the pseudosepta and the valve margin and then tapering more rapidly towards sub-acute apices (
<figureCitation id="13197DA7154F6508FC0B380B6BF9F888" box="[915,1015,1847,1868]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="2" pageNumber="229">Figs 513</figureCitation>
). Valve morphometrics are as follows: length 1020 μm, width 2.02.9 μm, length to width ratio 4.17.4, n = 30. Heteropolarity of the valve, when present, is evidenced by small differences in the lengths of the valve halves and differences in the shape of the apices (
<emphasis id="B956BD30154F6508FC6738436C08F850" box="[1023,1030,1919,1940]" italics="true" pageId="2" pageNumber="229">i</emphasis>
.
<emphasis id="B956BD30154F6508FB9438436C19F850" box="[1036,1047,1919,1940]" italics="true" pageId="2" pageNumber="229">e</emphasis>
., one valve apex, usually the shorter valve half, may be more bluntly rounded than the opposing end). The valve face is slightly asymmetric around a narrow axial area with 1/2 of the valve face slightly wider than the other (
<figureCitation id="13197DA7154F6508FD0738FB6B5BF818" box="[671,853,1991,2012]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="2" pageNumber="229">Figs 57, 9, 1113</figureCitation>
). The raphe is straight and the central area is formed by a wide stauros that extends to the valve margins (
<figureCitation id="13197DA7154F6508FDC038D76B1DF7C4" box="[600,787,2027,2048]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="2" pageNumber="229">Figs 5, 7, 9, 1113</figureCitation>
). The transapical striae are radiate around the central area becoming undiscernible in LM at the apices,
<quantity id="4CDACCC7154F6508FD9237336A51F7E0" box="[522,607,2063,2084]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" metricValueMax="8.636" metricValueMin="6.604" pageId="2" pageNumber="229" unit="in" value="30.0" valueMax="34.0" valueMin="26.0">2634 in</quantity>
10 μm, n = 30 (
<figureCitation id="13197DA7154F6508FD6237336BBEF7E0" box="[762,944,2063,2084]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="2" pageNumber="229">Figs 5, 7, 9, 1113</figureCitation>
). By focusing through the valve, each pseudoseptum can be seen to extend over
<emphasis id="B956BD30154E6509FE5F3F9C69D0FF71" box="[455,478,160,181]" italics="true" pageId="3" pageNumber="230">ca</emphasis>
. 1/6 of the valve length from the apices and then continues as narrow strips along the valve margin. These strips then widen at the central area where they fuse with the stauros (
<figureCitation id="13197DA7154E6509FC083FF86BFCFF1D" box="[912,1010,196,217]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="3" pageNumber="230">Figs 8, 10</figureCitation>
). SEM morphology: Externally, the valve face has uniseriate transapical striae composed of transapically elongated areolae (
<figureCitation id="13197DA7154E6509FC3F3FD46C16FF39" box="[935,1048,232,253]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1621</figureCitation>
). The striae are radiate and more distantly spaced in the middle of the valve,
<quantity id="4CDACCC7154E6509FD9C3E306A56FEE5" box="[516,600,268,289]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" metricValueMax="8.636" metricValueMin="6.604" pageId="3" pageNumber="230" unit="in" value="30.0" valueMax="34.0" valueMin="26.0">2634 in</quantity>
10 μm. n = 5 (
<figureCitation id="13197DA7154E6509FD7C3E306B78FEE5" box="[740,886,268,289]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1618, 20</figureCitation>
), becoming less radiate to parallel and more closely spaced towards the apices,
<quantity id="4CDACCC7154E6509FEE33E0C69DEFE81" box="[379,464,304,325]" metricMagnitude="0" metricUnit="m" metricValue="1.016" metricValueMax="1.0922" metricValueMin="0.9398" pageId="3" pageNumber="230" unit="in" value="40.0" valueMax="43.0" valueMin="37.0">3743 in</quantity>
10 μm, n = 5 (
<figureCitation id="13197DA7154E6509FDF83E0C6B16FE81" box="[608,792,304,325]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1617, 19, 21</figureCitation>
). The striae extend partly onto the valve mantle terminating before a hyaline mantle margin (
<figureCitation id="13197DA7154E6509FE3D3E6869E3FEAD" box="[421,493,340,361]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Fig. 15</figureCitation>
). The areolae are arranged in slightly undulate longitudinal rows along the transapical axis, 3337 areolae in 10 μm, n = 5 (
<figureCitation id="13197DA7154E6509FE333E446A15FE49" box="[427,539,376,397]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1621</figureCitation>
). The valve mantle slopes steeply without a sharp transition between the valve face and mantle (
<figureCitation id="13197DA7154E6509FF5D3EA06904FE75" box="[197,266,412,433]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Fig. 15</figureCitation>
).The mantle margin is undulate (
<figureCitation id="13197DA7154E6509FDCC3EA06A92FE75" box="[596,668,412,433]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Fig. 15</figureCitation>
). A straight and narrow rib lies within the axial area (
<figureCitation id="13197DA7154E6509FB363EA06D2EFE75" box="[1198,1312,412,433]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1621</figureCitation>
). The raphe is slightly eccentric and straight (
<figureCitation id="13197DA7154E6509FE723EFC6A52FE11" box="[490,604,448,469]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1621</figureCitation>
). A large, non-punctate, rectangular to bow tie-shaped stauros that extends to the valve margins is located in the central area (
<figureCitation id="13197DA7154E6509FDA33ED86ADAFE3D" box="[571,724,484,505]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1618, 20</figureCitation>
). The external proximal raphe ends are straight, opening into asymmetric spathulate grooves, and terminate where the stauros extends towards the valve margin (
<figureCitation id="13197DA7154E6509FB883D346CA8FDD9" box="[1040,1190,520,541]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 1618, 20</figureCitation>
). The distal raphe ends are hooked but are obscured by overhanging siliceous flaps that bend towards the same side of the valve at both apices (
<figureCitation id="13197DA7154E6509FAB73D106D93FD85" box="[1327,1437,556,577]" captionStart="FIGURES 1521" captionStartId="2.[136,243,1487,1508]" captionTargetBox="[169,1418,190,1461]" captionTargetId="figure-17@2.[168,1419,190,1462]" captionTargetPageId="2" captionText="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632969" httpUri="https://zenodo.org/record/13632969/files/figure.png" pageId="3" pageNumber="230">Figs 19, 21</figureCitation>
). Internal views of the valve reveal the butterfly-like structure that connects the pseudosepta to the central area and stauros (
<figureCitation id="13197DA7154E6509FA1A3D6C68F3FD4D" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 2224</figureCitation>
). The pseudosepta extend from the apices as siliceous plates for
<emphasis id="B956BD30154E6509FCFA3D486B77FD4D" box="[866,889,628,649]" italics="true" pageId="3" pageNumber="230">ca</emphasis>
. 1/6 of the valve length, and then continue as narrow strips that run along the valve margins before expanding into the “wings” of the butterfly-like structure in the central area (
<figureCitation id="13197DA7154E6509FAA83DA46DAFFD69" box="[1328,1441,664,685]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 2224</figureCitation>
). The wings fuse with the large oval central area close to the raphe sternum (
<figureCitation id="13197DA7154E6509FC1A3D806BC7FD15" box="[898,969,700,721]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Fig. 23</figureCitation>
). The wings of the butterfly widen at their junction with the central area forming narrow concave “wings” on each side of the raphe sternum (
<figureCitation id="13197DA7154E6509FB973DDC6C70FD31" box="[1039,1150,736,757]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 2224</figureCitation>
). Internally, the raphe slits open along the middle of a strong siliceous rib that widens slightly in the central area (
<figureCitation id="13197DA7154E6509FC5A3C386C3CFCDD" box="[962,1074,772,793]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 2224</figureCitation>
). A single knob-like structure is present on the rib at the valve center (
<figureCitation id="13197DA7154E6509FE783C146A5CFCF9" box="[480,594,808,829]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 2224</figureCitation>
). The bands of the cingulum are differentiated into
<specimenCount id="9D24AAAB154E6509FBDD3C146CABFCF9" box="[1093,1189,808,829]" count="2" pageId="3" pageNumber="230" type="generic" typeStatus="types">two types</specimenCount>
: closed valvocopulae (
<figureCitation id="13197DA7154E6509FA1A3C1468F4FCA5" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Figs 2527</figureCitation>
) and multiple copulae (up to 4 observed) that are open at one end (
<figureCitation id="13197DA7154E6509FCE33C706C00FCA5" box="[891,1038,844,865]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Figs 25, 29, 30</figureCitation>
). The valvocopula is flanged inward on the pars interior where it attaches to the valve (
<figureCitation id="13197DA7154E6509FDC73C4C6AA9FC41" box="[607,679,880,901]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Fig. 26</figureCitation>
). The flange widens into 3 pairs of opposing tabs (
<figureCitation id="13197DA7154E6509FB173C4C6D0EFC41" box="[1167,1280,880,901]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Figs 2627</figureCitation>
). One pair of tabs is located at the valve middle underneath the butterfly structure, whereas the other two pairs are located near the poles underneath the valve where the pseudosepta start to widen at the valve apices (
<figureCitation id="13197DA7154E6509FCA23C846B8AFC09" box="[826,900,952,973]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 24</figureCitation>
,
<figureCitation id="13197DA7154E6509FC163C846BC0FC09" box="[910,974,952,973]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">2627</figureCitation>
). The valvocopula has a single row of ovoid pores around the perimeter, pore density =
<quantity id="4CDACCC7154E6509FDBB3CE06A76FC35" box="[547,632,988,1009]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.143" metricValueMin="0.9398" pageId="3" pageNumber="230" unit="in" value="41.0" valueMax="45.0" valueMin="37.0">3745 in</quantity>
10 μm (
<figureCitation id="13197DA7154E6509FD533CE06B35FC35" box="[715,827,988,1009]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Figs 2628</figureCitation>
), and a second row of similarly spaced pores located abvalvar to the complete row only at the poles (
<figureCitation id="13197DA7154E6509FD803B3C6A6FFBD1" box="[536,609,1024,1045]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Fig. 28</figureCitation>
). Views of the pars exterior side of the valvocopula show a thickening of the polar tabs and extension of the pores through the tabs (
<figureCitation id="13197DA7154E6509FDF13B186ABAFBFD" box="[617,692,1060,1081]" captionStart="FIGURES 2224" captionStartId="3.[136,243,1900,1921]" captionTargetBox="[382,1205,1252,1875]" captionTargetId="figure-698@3.[381,1205,1251,1875]" captionTargetPageId="3" captionText="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230">Figs 24</figureCitation>
,
<figureCitation id="13197DA7154E6509FD263B186AD7FBFD" box="[702,729,1060,1081]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">27</figureCitation>
). The copulae have only a single row of ovoid pores, pore density =
<quantity id="4CDACCC7154E6509FAF73B1868DEFB99" metricMagnitude="0" metricUnit="m" metricValue="1.1811" metricValueMax="1.3716" metricValueMin="0.9906" pageId="3" pageNumber="230" unit="in" value="46.5" valueMax="54.0" valueMin="39.0">3954 in</quantity>
10 μm, n = 2 (
<figureCitation id="13197DA7154E6509FEC73B7469C1FB99" box="[351,463,1096,1117]" captionStart="FIGURES 2530" captionStartId="4.[136,243,1305,1326]" captionTargetBox="[374,1213,191,1280]" captionTargetId="figure-17@4.[373,1213,190,1281]" captionTargetPageId="4" captionText="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">Figs 2930</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33832A9154E6509FF103B506CA8FB0D" pageId="3" pageNumber="230" type="materials_examined">
<paragraph id="8B9D6122154E6509FF103B506CA8FB0D" blockId="3.[136,1453,160,1225]" pageId="3" pageNumber="230">
<emphasis id="B956BD30154E6509FF103B5068D4FB45" bold="true" box="[136,218,1132,1153]" pageId="3" pageNumber="230">Type:—</emphasis>
<collectingCountry id="F33521B2154E6509FF423B50699EFB45" box="[218,400,1132,1153]" name="United States of America" pageId="3" pageNumber="230">UNITED STATES</collectingCountry>
.
<collectingRegion id="49E6AFC0154E6509FE003B5069EFFB45" box="[408,481,1132,1153]" country="United States of America" name="Florida" pageId="3" pageNumber="230">Florida</collectingRegion>
:
<collectingRegion id="49E6AFC0154E6509FE723B506A21FB45" box="[490,559,1132,1153]" country="United States of America" name="Florida" pageId="3" pageNumber="230">Florida</collectingRegion>
Bay, samples removed from the skin in the dorsal neck area of loggerhead sea turtles
<taxonomicName id="4C221AA1154E6509FAFB3B50690EFB61" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Testudines" family="Cheloniidae" genus="Caretta" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="230" phylum="Chordata" rank="species" species="caretta">
<emphasis id="B956BD30154E6509FAFB3B50690EFB61" italics="true" pageId="3" pageNumber="230">Caretta caretta</emphasis>
</taxonomicName>
,
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,
<geoCoordinate id="EE1607E5154E6509FE303BAC6A36FB61" box="[424,568,1168,1189]" degrees="80" direction="west" minutes="48" orientation="longitude" pageId="3" pageNumber="230" precision="15" seconds="28" value="-80.80778">80º 48 28” W</geoCoordinate>
,
<emphasis id="B956BD30154E6509FDDC3BAC6A5DFB61" box="[580,595,1168,1189]" italics="true" pageId="3" pageNumber="230">B</emphasis>
.
<emphasis id="B956BD30154E6509FDC03BAC6A69FB61" box="[600,615,1168,1189]" italics="true" pageId="3" pageNumber="230">A</emphasis>
.
<emphasis id="B956BD30154E6509FDED3BAC6AA7FB61" box="[629,681,1168,1189]" italics="true" pageId="3" pageNumber="230">Stacy</emphasis>
,
<emphasis id="B956BD30154E6509FD203BAC6B30FB61" box="[696,830,1168,1189]" italics="true" pageId="3" pageNumber="230">
<date id="FF9C47E2154E6509FD203BAC6B30FB61" box="[696,830,1168,1189]" pageId="3" pageNumber="230" value="2015-06-24">24 June 2015</date>
</emphasis>
(
<typeStatus id="5499DF80154E6509FCD53BAC6BADFB61" box="[845,931,1168,1189]" pageId="3" pageNumber="230" type="holotype">holotype</typeStatus>
<collectionCode id="ED33F9E7154E6509FC323BAC6BD0FB61" box="[938,990,1168,1189]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="230" type="Museum">CAS</collectionCode>
! 223049, illustrated as
<figureCitation id="13197DA7154E6509FB5D3BAC6D53FB61" box="[1221,1373,1168,1189]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="3" pageNumber="230">Figs 14, 6, 12</figureCitation>
,
<figureCitation id="13197DA7154E6509FAF13BAC6DA6FB61" box="[1385,1448,1168,1189]" captionStart-0="FIGURES 1521" captionStart-1="FIGURES 2224" captionStart-2="FIGURES 2530" captionStartId-0="2.[136,243,1487,1508]" captionStartId-1="3.[136,243,1900,1921]" captionStartId-2="4.[136,243,1305,1326]" captionTargetBox-0="[169,1418,190,1461]" captionTargetBox-1="[382,1205,1252,1875]" captionTargetBox-2="[374,1213,191,1280]" captionTargetId-0="figure-17@2.[168,1419,190,1462]" captionTargetId-1="figure-698@3.[381,1205,1251,1875]" captionTargetId-2="figure-17@4.[373,1213,190,1281]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1521. Tursiocola denysii. Type population, SEM. 15. Single valve with attached cingulum in girdle view. Note undulate mantle margin with three tabs. 1617. External view of whole valves showing axial and central areas. 18. Detail of external central area of valve depicted in Fig. 16 showing bow tie-shaped stauros and proximal raphe ends within spathulate grooves. 19. Detail of external valve apex of valve depicted in Fig. 16 showing hooked distal raphe end obscured by overhanging siliceous flap. 20. Detail of external central area of valve depicted in Fig. 17 showing areolae structure. 21. Detail of external valve apex of valve depicted in Fig. 17. Scale bars: Figs 1517 = 2 μm, Figs 1821 = 1 μm." captionText-1="FIGURES 2224. Tursiocola denysii. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm." captionText-2="FIGURES 2530. Tursiocola denysii. Type population, SEM.25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13632969" figureDoi-1="http://doi.org/10.5281/zenodo.13632971" figureDoi-2="http://doi.org/10.5281/zenodo.13632975" httpUri-0="https://zenodo.org/record/13632969/files/figure.png" httpUri-1="https://zenodo.org/record/13632971/files/figure.png" httpUri-2="https://zenodo.org/record/13632975/files/figure.png" pageId="3" pageNumber="230">1530</figureCitation>
,
<typeStatus id="5499DF80154E6509FF253B886917FB0D" box="[189,281,1204,1225]" pageId="3" pageNumber="230" type="paratype">paratypes</typeStatus>
<collectionCode id="ED33F9E7154E6509FE863B88696EFB0D" box="[286,352,1204,1225]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="3" pageNumber="230">ANSP</collectionCode>
!
<collectionCode id="ED33F9E7154E6509FEF13B886985FB0D" box="[361,395,1204,1225]" country="0" name="Goucher College" pageId="3" pageNumber="230">GC</collectionCode>
59142,
<collectionCode id="ED33F9E7154E6509FE4B3B8869F3FB0D" box="[467,509,1204,1225]" country="United Kingdom" name="Bristol Museum" pageId="3" pageNumber="230">BM</collectionCode>
! 101 808, illustrated as
<figureCitation id="13197DA7154E6509FD783B886B6BFB0D" box="[736,869,1204,1225]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="3" pageNumber="230">Figs 710, 14</figureCitation>
,
<collectionCode id="ED33F9E7154E6509FCF73B886BA7FB0D" box="[879,937,1204,1225]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14903" lsid="urn:lsid:biocol.org:col:14903" name="Alfred-Wegener-Institut für Polar- und Meeresforschung" pageId="3" pageNumber="230" type="Herbarium">BRM</collectionCode>
! ZU10/31,
<figureCitation id="13197DA7154E6509FB8D3B886C94FB0D" box="[1045,1178,1204,1225]" captionStart="FIGURES 114" captionStartId="1.[136,243,1941,1962]" captionTargetBox="[276,1312,1548,1911]" captionTargetId="figure-496@1.[270,1317,1543,1916]" captionTargetPageId="1" captionText="FIGURES 114. Tursiocola denysii. Type population, LM. 14. Frustules in girdle view showing size range and morphological variation. Arrows in Figs 1, 2, 8, 10 indicate pseudosepta. 513. Specimens in valve view showing size range. 14. Valvocopula. Scale bar: Figs 114 = 10 μm. = indicates same specimen at different foci." figureDoi="http://doi.org/10.5281/zenodo.13632965" httpUri="https://zenodo.org/record/13632965/files/figure.png" pageId="3" pageNumber="230">Figs 5, 11, 13</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5D31AA154E6509FF1038506981F829" ID-DOI="http://doi.org/10.5281/zenodo.13632971" ID-Zenodo-Dep="13632971" httpUri="https://zenodo.org/record/13632971/files/figure.png" pageId="3" pageNumber="230" startId="3.[136,243,1900,1921]" targetBox="[382,1205,1252,1875]" targetPageId="3" targetType="figure">
<paragraph id="8B9D6122154E6509FF1038506981F829" blockId="3.[136,1452,1900,2029]" pageId="3" pageNumber="230">
<emphasis id="B956BD30154E6509FF1038506937F845" bold="true" box="[136,313,1900,1921]" pageId="3" pageNumber="230">FIGURES 2224</emphasis>
.
<taxonomicName id="4C221AA1154E6509FED0385069F3F845" authorityName="Frankovich &amp; M. J. Sullivan" authorityYear="2015" box="[328,509,1900,1921]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="3" pageNumber="230" phylum="Ochrophyta" rank="species" species="denysii">
<emphasis id="B956BD30154E6509FED0385069F3F845" box="[328,509,1900,1921]" italics="true" pageId="3" pageNumber="230">Tursiocola denysii</emphasis>
</taxonomicName>
. Type population, SEM. 22. Internal view of whole valve showing pseudosepta and butterfly structure. 23. Detail of butterfly structure and internal central area with one knob on the raphe rib. 24. Internal view of whole valve with attached valvocopula. Note one central and 2 polar tabs underlying undulate valve margin on each side of the valve. Scale bars: Figs 22, 24 = 2 μm, Fig. 23 = 1 μm.
</paragraph>
</caption>
<caption id="DF5D31AA1549650EFF103A256945FA7A" ID-DOI="http://doi.org/10.5281/zenodo.13632975" ID-Zenodo-Dep="13632975" httpUri="https://zenodo.org/record/13632975/files/figure.png" pageId="4" pageNumber="231" startId="4.[136,243,1305,1326]" targetBox="[374,1213,191,1280]" targetPageId="4" targetType="figure">
<paragraph id="8B9D61221549650EFF103A256945FA7A" blockId="4.[136,1452,1305,1470]" pageId="4" pageNumber="231">
<emphasis id="B956BD301549650EFF103A25693AFAEA" bold="true" box="[136,308,1305,1326]" pageId="4" pageNumber="231">FIGURES 2530</emphasis>
.
<taxonomicName id="4C221AA11549650EFEA63A2569E0FAEA" authorityName="Frankovich &amp; M. J. Sullivan" authorityYear="2015" box="[318,494,1305,1326]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="denysii">
<emphasis id="B956BD301549650EFEA63A2569E0FAEA" box="[318,494,1305,1326]" italics="true" pageId="4" pageNumber="231">Tursiocola denysii</emphasis>
</taxonomicName>
. Type population, SEM. 25. Cingulum showing valvocopula (V) with four attached abvalvar copulae (numbered arrows) in side view. 26. Pars interior of valvocopula showing flanged margin and three pairs of opposing tabs. 27. Pars exterior of valvocopula showing thickening of the two pairs of polar tabs. 28. Detail of apex of valvocopula showing second row of abvalvar pores (arrow) located only at the poles. 2930. Copulae showing open ends and only a single row of pores. Scale bars: Figs 2527, 2930= 2 μm, Fig. 28 = 1 μm.
</paragraph>
</caption>
<subSubSection id="C33832A91549650EFF10392F6B17F87C" pageId="4" pageNumber="231" type="etymology">
<paragraph id="8B9D61221549650EFF10392F6B06F994" blockId="4.[136,1452,1555,1976]" pageId="4" pageNumber="231">
<emphasis id="B956BD301549650EFF10392F6920F9E8" bold="true" box="[136,302,1555,1580]" pageId="4" pageNumber="231">Etymology:—</emphasis>
the epithet honours Dr. Luc Denys (Research Institute for Nature and Forest,
<collectingRegion id="49E6AFC01549650EFB34392F6D00F9E8" box="[1196,1294,1555,1580]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="4" pageNumber="231">Brussels</collectingRegion>
,
<collectingCountry id="F33521B21549650EFA83392F6D8EF9E8" box="[1307,1408,1555,1580]" name="Belgium" pageId="4" pageNumber="231">Belgium</collectingCountry>
), in recognition of his important research on epizoic diatoms.
</paragraph>
<paragraph id="8B9D61221549650EFF2539676B17F87C" blockId="4.[136,1452,1555,1976]" pageId="4" pageNumber="231">
<emphasis id="B956BD301549650EFF2539676A1DF9B0" bold="true" box="[189,531,1627,1652]" pageId="4" pageNumber="231">Taxa relative abundances:—</emphasis>
46 taxa from 23 genera were observed from the neck skin of 4 loggerhead turtles. The relative abundances of the newly described
<taxonomicName id="4C221AA11549650EFD0239436B69F95C" authorityName="Frankovich &amp; M. J. Sullivan" authorityYear="2015" box="[666,871,1663,1688]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="denysii">
<emphasis id="B956BD301549650EFD0239436B69F95C" box="[666,871,1663,1688]" italics="true" pageId="4" pageNumber="231">Tursiocola denysii</emphasis>
</taxonomicName>
ranged from 5 to 43%. The most relatively abundant taxon was
<taxonomicName id="4C221AA11549650EFE98399F6CF0F978" authority="(Grunow 1877: 166) D. M. Williams &amp; F. E. Round (1986: 316)" authorityName="D. M. Williams &amp; F. E. Round" authorityPageNumber="316" authorityYear="1986" baseAuthorityName="Grunow" baseAuthorityPageNumber="166" baseAuthorityYear="1877" box="[256,1278,1699,1724]" class="Bacillariophyceae" family="Ulnariaceae" genus="Hyalosynedra" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Licmophorales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="laevigata">
<emphasis id="B956BD301549650EFE98399F6993F978" box="[256,413,1699,1724]" italics="true" pageId="4" pageNumber="231">Hyalosynedra</emphasis>
cf.
<emphasis id="B956BD301549650EFE51399F6A3FF978" box="[457,561,1699,1724]" italics="true" pageId="4" pageNumber="231">laevigata</emphasis>
(
<bibRefCitation id="EFB31CD31549650EFDD8399F6B19F978" author="Grunow, A." box="[576,791,1699,1724]" pageId="4" pageNumber="231" pagination="165 - 186" refId="ref7165" refString="Grunow, A. (1877) New diatoms from Honduras. The Monthly Microscopical Journal 18: 165 - 186. http: // dx. doi. org / 10.1111 / j. 1365 - 2818.1877. tb 00123. x" type="journal article" year="1877">Grunow 1877: 166</bibRefCitation>
) D.M. Williams &amp; F.E. Round (1986: 316)
</taxonomicName>
, comprising 2 62% of the valve counts. The next most relatively abundant taxon was an undescribed species of
<taxonomicName id="4C221AA11549650EFB5D39FB69CAF8C0" authority="Majewska, De Stefano &amp; Van de Vijver" authorityName="Majewska, De Stefano &amp; Van de Vijver" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Poulinea" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="genus">
<emphasis id="B956BD301549650EFB5D39FB6D24F924" box="[1221,1322,1735,1760]" italics="true" pageId="4" pageNumber="231">Poulinea</emphasis>
Majewska, De Stefano &amp; Van de Vijver
</taxonomicName>
(in press), comprising 1347% of the valve count. We were unable to identify
<emphasis id="B956BD301549650EFAA639D76D56F8C0" box="[1342,1368,1771,1796]" italics="true" pageId="4" pageNumber="231">ca</emphasis>
. 7% of the valves to genus or species. Only four other taxa (
<emphasis id="B956BD301549650EFD7D38336AE3F8EC" box="[741,749,1807,1832]" italics="true" pageId="4" pageNumber="231">i</emphasis>
.
<emphasis id="B956BD301549650EFD6C38336B0EF8EC" box="[756,768,1807,1832]" italics="true" pageId="4" pageNumber="231">e</emphasis>
.,
<taxonomicName id="4C221AA11549650EFC8F383368B3F888" authority="(Meister 1937: 268) Simonsen (1990: 134)" authorityName="Simonsen" authorityPageNumber="134" authorityYear="1990" baseAuthorityName="Meister" baseAuthorityPageNumber="268" baseAuthorityYear="1937" class="Bacillariophyceae" family="Naviculaceae" genus="Mastogloia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="cuneata">
<emphasis id="B956BD301549650EFC8F38336BF6F8EC" box="[791,1016,1807,1832]" italics="true" pageId="4" pageNumber="231">Mastogloia cuneata</emphasis>
(
<bibRefCitation id="EFB31CD31549650EFB9F38336CD5F8EC" author="Meister, F." box="[1031,1243,1807,1832]" pageId="4" pageNumber="231" pagination="258 - 276" refId="ref7670" refString="Meister, F. (1937) Seltene und neue Kieselalgen II. Berichte der Schweizerischen Botanischen Gesellschaft 48: 258 - 276." type="journal article" year="1937">Meister 1937: 268</bibRefCitation>
)
<bibRefCitation id="EFB31CD31549650EFB73383368B3F888" author="Simonsen, R." pageId="4" pageNumber="231" pagination="121 - 142" refId="ref7773" refString="Simonsen, R. (1990) On some diatoms of the genus Mastogloia. Beiheft zur Nova Hedwigia 100: 121 - 142." type="journal article" year="1990">Simonsen (1990: 134)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C221AA11549650EFF51380F6959F888" box="[201,343,1843,1868]" class="Bacillariophyceae" family="Naviculaceae" genus="Navicula" kingdom="Chromista" order="Naviculales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="undetermined">
<emphasis id="B956BD301549650EFF51380F6920F888" box="[201,302,1843,1868]" italics="true" pageId="4" pageNumber="231">Navicula</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C221AA11549650EFEFB380F6BEEF888" authority="Hendey (1964: 177)" authorityName="Hendey" authorityPageNumber="177" authorityYear="1964" box="[355,992,1843,1868]" class="Bacillariophyceae" family="Achnanthaceae" genus="Achnanthes" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Achnanthales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="pseudogroenlandica">
<emphasis id="B956BD301549650EFEFB380F69E8F888" box="[355,486,1843,1868]" italics="true" pageId="4" pageNumber="231">Achnanthes</emphasis>
cf.
<emphasis id="B956BD301549650EFD89380F6AFBF888" box="[529,757,1843,1868]" italics="true" pageId="4" pageNumber="231">pseudogroenlandica</emphasis>
<bibRefCitation id="EFB31CD31549650EFD64380F6BEEF888" author="Hendey, N. I." box="[764,992,1843,1868]" pageId="4" pageNumber="231" refId="ref7244" refString="Hendey, N. I. (1964) An introductory account of the smaller algae of British coastal waters. Part V. Bacillariophyceae (Diatoms). Ministry of Agriculture, Fisheries and Food, Fisheries Investigations, Seies IV, London, 317 pp." type="book" year="1964">Hendey (1964: 177)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C221AA11549650EFB83380F68A7F8B4" authority="Kutzing (1836: 3))" authorityName="Kutzing" authorityPageNumber="3" authorityYear="1836" class="Bacillariophyceae" family="Brachysiraceae" genus="Brachysira" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="aponina">
<emphasis id="B956BD301549650EFB83380F6CF4F888" box="[1051,1274,1843,1868]" italics="true" pageId="4" pageNumber="231">Brachysira aponina</emphasis>
<bibRefCitation id="EFB31CD31549650EFA99380F68A7F8B4" author="Kutzing, F. T." pageId="4" pageNumber="231" refId="ref7524" refString="Kutzing, F. T. (1836) Algarum aquae dulcis germanicarum, Decas XV 1. Halis Saxonum: in commissis C. A. Schwetschkii et fil, 25 pp." type="book" year="1836">Kützing (1836: 3))</bibRefCitation>
</taxonomicName>
exhibited mean relative abundances&gt;1%. The remaining taxa observed are common benthic forms. Only a single valve of
<taxonomicName id="4C221AA11549650EFF6938476B11F850" authority="Frankovich &amp; M. J. Sullivan" authorityName="Frankovich &amp; M. J. Sullivan" box="[241,799,1915,1940]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Tursiocola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="4" pageNumber="231" phylum="Ochrophyta" rank="species" species="ziemanii">
<emphasis id="B956BD301549650EFF69384769DCF850" box="[241,466,1915,1940]" italics="true" pageId="4" pageNumber="231">Tursiocola ziemanii</emphasis>
Frankovich &amp; M.J. Sullivan
</taxonomicName>
in
<bibRefCitation id="EFB31CD31549650EFCD238476C9CF850" author="Frankovich, T. A. &amp; Sullivan, M. J. &amp; Stacy, N. I." box="[842,1170,1915,1940]" pageId="4" pageNumber="231" pagination="33 - 48" refId="ref7101" refString="Frankovich, T. A., Sullivan, M. J. &amp; Stacy, N. I. (2015) Three new species of Tursiocola (Bacillariophyta) from the skin of the West Indian manatee. Phytotaxa 204 (1): 33 - 48. http: // dx. doi. org / 10.11646 / phytotaxa. 204.1.3" type="journal article" year="2015">
Frankovich
<emphasis id="B956BD301549650EFC4C38476C07F850" box="[980,1033,1915,1940]" italics="true" pageId="4" pageNumber="231">et al</emphasis>
. (2015: 34)
</bibRefCitation>
of the seven previously described
<taxonomicName id="4C221AA11549650EFF6238A369C4F87C" box="[250,458,1951,1976]" pageId="4" pageNumber="231">
<emphasis id="B956BD301549650EFF6238A3697CF87C" box="[250,370,1951,1976]" italics="true" pageId="4" pageNumber="231">Tursiocola</emphasis>
species
</taxonomicName>
was observed in the material.
</paragraph>
</subSubSection>
</treatment>
</document>