706 lines
78 KiB
XML
706 lines
78 KiB
XML
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<mods:title id="794915031D29FC0B7192A98217AB14A1">Additions to Philippine Slender Skinks of the Brachymeles bonitae Complex (Reptilia: Squamata: Scincidae) III: a new species from Tablas Island</mods:title>
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<mods:namePart id="D4FD2CA36C85968B8AC76879B9BB9F9E">Sanguila, Marites B.</mods:namePart>
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<mods:namePart id="37128D9A6C5B1EBA52D1EF11191C9FB1">Siler, Cameron D.</mods:namePart>
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<mods:typeOfResource id="05D89F898BCE6B3436680CCE74E4E4BF">text</mods:typeOfResource>
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<mods:title id="24CEA260465BC9EF47043F0510E01AE9">Zootaxa</mods:title>
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<mods:part id="9A7ABFDAFDA74F312F3970F96E2030A3">
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<mods:date id="BF146590B0F850F28FB50EF717CD5E78">2016</mods:date>
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<mods:detail id="7429E2F5D6FC970D6519632ABCDAD86E" type="volume">
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<mods:number id="93545BB3D58B5638FD4BAB197CF41AA3">4132</mods:number>
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<mods:detail id="89C173DEDDB06CE206A0B660AC0D4EF2" type="issue">
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<mods:number id="6EAB232F23EC08D0BA0E7FE11975387D">1</mods:number>
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<mods:classification id="A8CA65BDC3AFF1DD48F5002FDF22F6AC">journal volume</mods:classification>
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<mods:identifier id="09CAF339C23C12907E49CE3CF6F5D17C" type="CLB-Dataset">38701</mods:identifier>
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<mods:identifier id="42449FBF327C7B7EF2210185F51EE920" type="DOI">10.11646/zootaxa.4132.1.3</mods:identifier>
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<mods:identifier id="B3DB8B9D706C0AA845F82F404FF91357" type="GBIF-Dataset">01ffc22a-a8dd-40f9-97f3-503df084c471</mods:identifier>
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<mods:identifier id="F266D1BBC82B14449F304A518EFB68F7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B856F678A1DFE6AEFC99A401938B0699" type="Zenodo-Dep">258936</mods:identifier>
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<treatment id="663D8783FF89FFA09A97CFF9C4FC531D" ID-DOI="http://doi.org/10.5281/zenodo.6090697" ID-GBIF-Taxon="121068730" ID-Zenodo-Dep="6090697" LSID="urn:lsid:plazi:treatment:663D8783FF89FFA09A97CFF9C4FC531D" httpUri="http://treatment.plazi.org/id/663D8783FF89FFA09A97CFF9C4FC531D" lastPageId="8" pageId="3">
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<subSubSection id="A68E651EFF89FFAB9A97CFF9C4035692" pageId="3" type="nomenclature">
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<paragraph id="EE2B3695FF89FFAB9A97CFF9C74456F5" blockId="3.[151,598,401,461]" box="[151,598,401,427]" pageId="3">
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<heading id="B56381F9FF89FFAB9A97CFF9C74456F5" bold="true" box="[151,598,401,427]" fontSize="11" level="1" pageId="3" reason="1">
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<emphasis id="DCE0EA87FF89FFAB9A97CFF9C74456F5" bold="true" box="[151,598,401,427]" pageId="3">
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<taxonomicName id="29944D16FF89FFAB9A97CFF9C4E256F5" box="[151,496,401,427]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
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<emphasis id="DCE0EA87FF89FFAB9A97CFF9C4E256F5" bold="true" box="[151,496,401,427]" italics="true" pageId="3">Brachymeles dalawangdaliri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C7D357FCFF89FFAB9BF8CFF9C74456F5" box="[504,598,401,427]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="EE2B3695FF89FFAB9A97CFDCC4035692" blockId="3.[151,598,401,461]" box="[151,273,436,461]" pageId="3">
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(
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<figureCitation id="76AF2A10FF89FFAB9A9FCFDCC5FC5692" box="[159,238,436,461]" captionStart="FIGURE 2" captionStartId="7.[151,250,1849,1871]" captionTargetBox="[390,1187,193,1819]" captionTargetId="figure@7.[389,1188,193,1820]" captionTargetPageId="7" captionText="FIGURE 2. Illustration of the head of Brachymeles dalawangdaliri sp. nov. (CAS 137149, holotype) in dorsal, lateral, and ventral views. Taxonomically diagnostic head scales are labeled as follows: C, chin shield; F, frontal; FN, frontonasal; FP, frontoparietal; IL, infralabial; IP, interparietal; L, loreal; M, mental; N, nasal; Nu, nuchal; P, parietal; PF, prefrontal; PM, postmental; PO, preocular; PSO, presubocular; R, rostral; SC, supraciliary; SL, supralabial; SN, supranasal; and SO, supraocular. Roman numerals indicate scales in the supraocular series, with Arabic numbers indicating scales in the supraciliary series. Illustrations by MLP and CDS." httpUri="https://zenodo.org/record/258938/files/figure.png" pageId="3">Figs. 2</figureCitation>
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,
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<figureCitation id="76AF2A10FF89FFAB9AFACFDCC41A5692" box="[250,264,436,460]" captionStart="FIGURE 3" captionStartId="8.[151,250,609,631]" captionTargetBox="[150,1436,193,587]" captionTargetId="figure@8.[150,1437,193,588]" captionTargetPageId="8" captionText="FIGURE 3. Photograph of lateral and ventral views of head of preserved holotype of Brachymeles dalawangdaliri sp. nov. (CAS 137149). Photographs taken by JLW and MLP." httpUri="https://zenodo.org/record/258939/files/figure.png" pageId="3">3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A68E651EFF89FFAB9A97CF94C11F556E" pageId="3" type="reference_group">
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<paragraph id="EE2B3695FF89FFAB9A97CF94C11F556E" blockId="3.[151,1436,508,560]" pageId="3">
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<taxonomicName id="29944D16FF89FFAB9A97CF94C778554C" authority="Dumeril & Bibron 1839" authorityName="Dumeril & Bibron" authorityYear="1839" box="[151,618,508,530]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="bonitae">
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<emphasis id="DCE0EA87FF89FFAB9A97CF94C47E554C" box="[151,364,508,530]" italics="true" pageId="3">Brachymeles bonitae</emphasis>
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<bibRefCitation id="8A054B64FF89FFAB9B73CF94C778554C" author="Dumeril" box="[371,618,508,530]" pageId="3" refString="Dumeril, A. M. C. & Bibron, G. (1839) Erpetologie General ou, Histoire Naturelle Complete des Reptiles. Librairie Encylopedique de Roret, Paris, 488 pp." type="book" year="1839">Duméril & Bibron 1839</bibRefCitation>
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</taxonomicName>
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;
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<bibRefCitation id="8A054B64FF89FFAB9875CF94C7E1554C" author="Taylor" box="[629,755,508,530]" pageId="3" refString="Taylor, E. H. (1917) Brachymeles, a genus of Philippine lizards. Philippine Journal of Science, 12, 267 - 279." type="journal article" year="1917">Taylor 1917</bibRefCitation>
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; Brown 1956:5;
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<bibRefCitation id="8A054B64FF89FFAB999CCF95C16F554C" author="Brown" box="[924,1149,508,530]" pageId="3" refString="Brown, W. C. & Rabor, D. S. (1967) Review of the genus Bracheymeles (Scincidae), with descriptions of new species and subspecies. Proceedings of the California Academy of Sciences, Fourth Series, 15, 525 - 548." type="journal article" year="1967">Brown & Rabor 1967</bibRefCitation>
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:526;
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<bibRefCitation id="8A054B64FF89FFAB9EB4CF95C08A554C" author="Brown" box="[1204,1432,508,530]" pageId="3" refString="Brown, W. C. & Alcala, A. C. (1970) The zoogeography of the herpetofauna of the Philippine Islands, a fringing archipelago. Proceedings of the California Academy of Sciences, Series 4, 18, 105 - 130." type="journal article" year="1970">Brown & Alcala 1970</bibRefCitation>
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;
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<bibRefCitation id="8A054B64FF89FFAB9AC7CC73C4B9556E" author="Brown" box="[199,427,538,560]" pageId="3" refString="Brown, W. C. & Alcala, A. C. (1980) Philippine Lizards of the Family Scincidae. Philippine University Press, Dumaguete City, 264 pp." type="book" year="1980">Brown & Alcala 1980</bibRefCitation>
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||
:20; Davis
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<emphasis id="DCE0EA87FF89FFAB9816CC73C75B556E" box="[534,585,538,560]" italics="true" pageId="3">et al.</emphasis>
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2014; Geheber
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<emphasis id="DCE0EA87FF89FFAB98EBCC73C60C556E" box="[747,798,538,560]" italics="true" pageId="3">et al.</emphasis>
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2016; Siler
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<emphasis id="DCE0EA87FF89FFAB999ACC73C6DF556E" box="[922,973,538,560]" italics="true" pageId="3">et al.</emphasis>
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2016.
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</paragraph>
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</subSubSection>
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<subSubSection id="A68E651EFF89FFAB9A97CC35C63654C8" pageId="3" type="materials_examined">
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<paragraph id="EE2B3695FF89FFAB9A97CC35C6C555E0" blockId="3.[151,1437,605,2006]" pageId="3">
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<emphasis id="DCE0EA87FF89FFAB9A97CC35C4195528" bold="true" box="[151,267,605,630]" pageId="3">
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<typeStatus id="312F8837FF89FFAB9A97CC35C4145528" box="[151,262,605,630]" pageId="3" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="8885AE50FF89FFAB9B14CC36C4585528" box="[276,330,606,630]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3">CAS</collectionCode>
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||
137149, adult male, collected among rotting leaves in early secondary growth forest on
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<date id="9A2A1055FF89FFAB9F44CC36C5C055C4" pageId="3" value="1972-05-24">24 May 1972</date>
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, in Barrio Dubduban, Municipality of San Augustin,
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<taxonomicName id="29944D16FF89FFAB9974CCE9C6CE55C4" box="[884,988,641,666]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="romblon">Romblon</taxonomicName>
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Province, Tablas Island,
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<collectingCountry id="96837605FF89FFAB9F1ECCE9C08E55C4" box="[1310,1436,641,666]" name="Philippines" pageId="3">Philippines</collectingCountry>
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(
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,
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<geoCoordinate id="8BA05052FF89FFAB9B3DCCCEC4C955E3" box="[317,475,678,702]" direction="east" orientation="longitude" pageId="3" precision="1" value="122.1">122.10000° E</geoCoordinate>
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; WGS 84;
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. elev.), by L.C. Alcala.
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</paragraph>
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|
||
<emphasis id="DCE0EA87FF89FFAB9AC7CCA1C45455BC" bold="true" box="[199,326,713,738]" pageId="3">
|
||
<typeStatus id="312F8837FF89FFAB9AC7CCA1C45355BC" box="[199,321,713,738]" pageId="3" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Five adult females (
|
||
<collectionCode id="8885AE50FF89FFAB9837CCA2C77D55BC" box="[567,623,714,738]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3">CAS</collectionCode>
|
||
137148, 137150, 137151, 137153, 137154) and one adult male (
|
||
<collectionCode id="8885AE50FF89FFAB9F64CCA2C08E55BC" box="[1380,1436,714,738]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3">CAS</collectionCode>
|
||
137152) were collected in Barrio Dubduban, Municipality of San Augustin,
|
||
<taxonomicName id="29944D16FF89FFAB9E15CC85C16F5458" box="[1045,1149,749,774]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="romblon">Romblon</taxonomicName>
|
||
Province, Tablas Island,
|
||
<collectingCountry id="96837605FF89FFAB9A97CD79C4075474" box="[151,277,785,810]" name="Philippines" pageId="3">Philippines</collectingCountry>
|
||
(
|
||
<geoCoordinate id="8BA05052FF89FFAB9B29CD79C4D05477" box="[297,450,785,810]" direction="north" orientation="latitude" pageId="3" precision="1" value="12.58333">12.58333° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8BA05052FF89FFAB9BD2CD7AC7665477" box="[466,628,786,810]" direction="east" orientation="longitude" pageId="3" precision="1" value="122.1">122.10000° E</geoCoordinate>
|
||
; WGS 84), by L.C. Alcala.
|
||
<collectionCode id="8885AE50FF89FFAB99CBCD7AC1135474" box="[971,1025,786,810]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3">CAS</collectionCode>
|
||
137148 was collected in a rotting coconut trunk on
|
||
<date id="9A2A1055FF89FFAB9B5ECD5EC4E45410" box="[350,502,822,846]" pageId="3" value="1972-05-23">23 May 1972</date>
|
||
(
|
||
<quantity id="296C9B70FF89FFAB9805CD5DC7975410" box="[517,645,821,846]" metricMagnitude="1" metricUnit="m" metricValue="6.096" metricValueMax="7.62" metricValueMin="4.572" pageId="3" unit="ft" value="200.0" valueMax="250.0" valueMin="150.0">150–250 ft</quantity>
|
||
. elev.),
|
||
<collectionCode id="8885AE50FF89FFAB98DDCD5EC6015410" box="[733,787,822,846]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3">CAS</collectionCode>
|
||
137150–137152 were collected in humus under a rotting log in original forest on
|
||
<date id="9A2A1055FF89FFAB9BB2CD32C72F542C" box="[434,573,858,882]" pageId="3" value="1972-06-01">1 June 1972</date>
|
||
(
|
||
<quantity id="296C9B70FF89FFAB984ECD31C78A542C" box="[590,664,857,882]" metricMagnitude="2" metricUnit="m" metricValue="1.3716" pageId="3" unit="ft" value="450.0">450 ft</quantity>
|
||
. elev.), and
|
||
<collectionCode id="8885AE50FF89FFAB9924CD32C648542C" box="[804,858,858,882]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3">CAS</collectionCode>
|
||
137153, 137154 were collected in humus under a rotting log in original forest on
|
||
<date id="9A2A1055FF89FFAB9BF8CD16C76D54C8" box="[504,639,894,918]" pageId="3" value="1972-06-02">2 June 1972</date>
|
||
(
|
||
<quantity id="296C9B70FF89FFAB988ECD15C7C754C8" box="[654,725,893,918]" metricMagnitude="2" metricUnit="m" metricValue="1.6764" pageId="3" unit="ft" value="550.0">550 ft</quantity>
|
||
. elev.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68E651EFF89FFAB9AC7CDC9C7195218" pageId="3" type="diagnosis">
|
||
<paragraph id="EE2B3695FF89FFAB9AC7CDC9C19F5314" blockId="3.[151,1437,605,2006]" pageId="3">
|
||
<emphasis id="DCE0EA87FF89FFAB9AC7CDC9C45354E4" bold="true" box="[199,321,929,954]" pageId="3">Diagnosis.</emphasis>
|
||
Following recent taxonomic revisions of
|
||
<taxonomicName id="29944D16FF89FFAB9914CDC9C6B754E7" box="[788,933,929,953]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="DCE0EA87FF89FFAB9914CDC9C6B754E7" box="[788,933,929,953]" italics="true" pageId="3">Brachymeles</emphasis>
|
||
</taxonomicName>
|
||
(Siler
|
||
<emphasis id="DCE0EA87FF89FFAB99F1CDCAC13B54E7" box="[1009,1065,929,953]" italics="true" pageId="3">et al.</emphasis>
|
||
2011; Davis
|
||
<emphasis id="DCE0EA87FF89FFAB9EBDCDCAC1E754E7" box="[1213,1269,929,953]" italics="true" pageId="3">et al.</emphasis>
|
||
2014) the new species is assigned to the
|
||
<taxonomicName id="29944D16FF89FFAB9BBACDAEC73F5483" box="[442,557,965,989]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="bonitae">
|
||
<emphasis id="DCE0EA87FF89FFAB9BBACDAEC73F5483" box="[442,557,965,989]" italics="true" pageId="3">B. bonitae</emphasis>
|
||
</taxonomicName>
|
||
Complex based on the following suite of morphological characters: (1) limbs present, (2) non-pentadactyl, (3) fore-limbs with 0–3 fingers. (4) hind limbs with 0–2 toes, (5) paravertebral scale rows ≥ 91, (6) presacral vertebrae 47–53, (7) supraoculars four, (8) enlarged, differentiated nuchals present, (9) longitudinal rows of dark spots around the body absent, and (10) auricular opening absent.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF89FFAB9AC7CA3DC7195218" blockId="3.[151,1437,605,2006]" pageId="3">
|
||
<taxonomicName id="29944D16FF89FFAB9AC7CA3DC71C5333" box="[199,526,1109,1133]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF89FFAB9AC7CA3DC71C5333" box="[199,526,1109,1133]" italics="true" pageId="3">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF89FFAB9816CA3DC7605330" box="[534,626,1109,1134]" pageId="3" rank="species">
|
||
<emphasis id="DCE0EA87FF89FFAB9816CA3DC7605330" bold="true" box="[534,626,1109,1134]" pageId="3">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from congeners by the following combination of characters: (1) body size small (SVL 66.0–
|
||
<quantity id="296C9B70FF89FFAB9879CA11C7F253CC" box="[633,736,1145,1170]" metricMagnitude="-2" metricUnit="m" metricValue="8.09" pageId="3" unit="mm" value="80.9">80.9 mm</quantity>
|
||
), (2) fore-limbs bidactyl, (3) hind limbs digitless, unidactyl, or bidactyl, (4) limb length short, (5) supralabials six or seven, (6) infralabials six or seven, (7) supraciliaries five, (8) supraoculars four, (9) midbody scale rows 24 or 25, (10) axilla–groin scale rows 80–83, (11) paravertebral scale rows 97–101, (12) prefrontal contact absent, (13) frontoparietal contact present or absent, (14) enlarged chin shields in two pairs, (15) nuchals enlarged, (16) auricular opening absent, (17) presacral vertebrae 49, and (18) uniform body color (Tables 1, 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68E651EFF89FFAC9AC7CB39C1F25059" lastPageId="4" pageId="3" type="description">
|
||
<paragraph id="EE2B3695FF89FFAC9AC7CB39C7D55789" blockId="3.[151,1437,605,2006]" lastBlockId="4.[151,1437,151,1979]" lastPageId="4" pageId="3">
|
||
<emphasis id="DCE0EA87FF89FFAB9AC7CB39C47E5234" bold="true" box="[199,364,1361,1386]" pageId="3">Comparisons.</emphasis>
|
||
<taxonomicName id="29944D16FF89FFAB9B73CB39C7A55237" box="[371,695,1361,1385]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF89FFAB9B73CB39C7A55237" box="[371,695,1361,1385]" italics="true" pageId="3">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF89FFAB98BECB39C6055234" box="[702,791,1361,1386]" pageId="3" rank="species">
|
||
<emphasis id="DCE0EA87FF89FFAB98BECB39C6055234" bold="true" box="[702,791,1361,1386]" pageId="3">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="29944D16FF89FFAB9E46CB3AC1DC5237" box="[1094,1230,1361,1385]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="ilocandia">
|
||
<emphasis id="DCE0EA87FF89FFAB9E46CB3AC1DC5237" box="[1094,1230,1361,1385]" italics="true" pageId="3">B. ilocandia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="29944D16FF89FFAB9EDCCB3AC0645237" box="[1244,1398,1361,1385]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="isangdaliri">
|
||
<emphasis id="DCE0EA87FF89FFAB9EDCCB3AC0645237" box="[1244,1398,1361,1385]" italics="true" pageId="3">B. isangdaliri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="DCE0EA87FF89FFAB9F84CB3AC45452D3" italics="true" pageId="3">
|
||
<taxonomicName id="29944D16FF89FFAB9F84CB3AC45052D3" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="mapalonggaon">B. mapalonggaon</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="29944D16FF89FFAB9B81CB1EC70952D3" box="[385,539,1397,1421]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tridactylus">
|
||
<emphasis id="DCE0EA87FF89FFAB9B81CB1EC70952D3" box="[385,539,1397,1421]" italics="true" pageId="3">B. tridactylus</emphasis>
|
||
</taxonomicName>
|
||
by having bidactyl fore-limbs and 0–2 digits on the hind limbs (
|
||
<emphasis id="DCE0EA87FF89FFAB9F0CCB1EC04652D3" box="[1292,1364,1398,1421]" italics="true" pageId="3">versus</emphasis>
|
||
being digitless [
|
||
<taxonomicName id="29944D16FF89FFAB9B05CBF2C49C52EF" box="[261,398,1433,1457]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="ilocandia">
|
||
<emphasis id="DCE0EA87FF89FFAB9B05CBF2C49C52EF" box="[261,398,1433,1457]" italics="true" pageId="3">B. ilocandia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="29944D16FF89FFAB9B9CCBF2C77652EF" box="[412,612,1433,1457]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="mapalanggaon">
|
||
<emphasis id="DCE0EA87FF89FFAB9B9CCBF2C77652EF" box="[412,612,1433,1457]" italics="true" pageId="3">B. mapalanggaon</emphasis>
|
||
</taxonomicName>
|
||
], unidactyl [
|
||
<taxonomicName id="29944D16FF89FFAB98F4CBF2C69C52EF" box="[756,910,1433,1457]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="isangdaliri">
|
||
<emphasis id="DCE0EA87FF89FFAB98F4CBF2C69C52EF" box="[756,910,1433,1457]" italics="true" pageId="3">B. isangdaliri</emphasis>
|
||
</taxonomicName>
|
||
], or tridactyl [
|
||
<taxonomicName id="29944D16FF89FFAB9E32CBF2C1DB52EF" box="[1074,1225,1433,1457]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tridactylus">
|
||
<emphasis id="DCE0EA87FF89FFAB9E32CBF2C1DB52EF" box="[1074,1225,1433,1457]" italics="true" pageId="3">B. tridactylus</emphasis>
|
||
</taxonomicName>
|
||
]); from
|
||
<taxonomicName id="29944D16FF89FFAB9F28CBF2C08E52EF" box="[1320,1436,1433,1457]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="bonitae">
|
||
<emphasis id="DCE0EA87FF89FFAB9F28CBF2C08E52EF" box="[1320,1436,1433,1457]" italics="true" pageId="3">B. bonitae</emphasis>
|
||
</taxonomicName>
|
||
by having fewer number of presacral vertebrae (49
|
||
<emphasis id="DCE0EA87FF89FFAB98EACBD6C620528B" box="[746,818,1470,1493]" italics="true" pageId="3">versus</emphasis>
|
||
53), fewer number of axilla–groin scale rows (80–83
|
||
<emphasis id="DCE0EA87FF89FFAB9A97CB8AC5CD52A7" box="[151,223,1506,1529]" italics="true" pageId="3">versus</emphasis>
|
||
83–90), fewer number of paravertebral scale rows (97–101
|
||
<emphasis id="DCE0EA87FF89FFAB99CECB8AC10752A7" box="[974,1045,1506,1529]" italics="true" pageId="3">versus</emphasis>
|
||
103–110), a greater number of infralabials (6, 7
|
||
<emphasis id="DCE0EA87FF89FFAB9B5EC861C4B4517E" box="[350,422,1545,1568]" italics="true" pageId="3">versus</emphasis>
|
||
5), and by the absence (
|
||
<emphasis id="DCE0EA87FF89FFAB98C6C861C61F517E" box="[710,781,1545,1568]" italics="true" pageId="3">versus</emphasis>
|
||
presence) of a 3rd chin shield and fused mental; from
|
||
<taxonomicName id="29944D16FF89FFAB9F84C861C413511A" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="ilocandia">
|
||
<emphasis id="DCE0EA87FF89FFAB9F84C861C413511A" italics="true" pageId="3">B. ilocandia</emphasis>
|
||
</taxonomicName>
|
||
by having a longer fore-limb (
|
||
<quantity id="296C9B70FF89FFAB9857C844C7CD511A" box="[599,735,1580,1604]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="3" unit="mm" value="1.8" valueMax="2.2" valueMin="1.4">1.4–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB98E5C845C63E511A" box="[741,812,1581,1604]" italics="true" pageId="3">versus</emphasis>
|
||
1.1–1.4) and hind limb length (
|
||
<quantity id="296C9B70FF89FFAB9E8EC844C004511A" box="[1166,1302,1580,1604]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB9F1CC845C076511A" box="[1308,1380,1581,1604]" italics="true" pageId="3">versus</emphasis>
|
||
1.6– 1.9), fewer number of presacral vertebrae (49
|
||
<emphasis id="DCE0EA87FF89FFAB98ACC83CC7E65135" box="[684,756,1620,1643]" italics="true" pageId="3">versus</emphasis>
|
||
50–53), and by the absence (
|
||
<emphasis id="DCE0EA87FF89FFAB9E4BC83CC1815135" box="[1099,1171,1620,1643]" italics="true" pageId="3">versus</emphasis>
|
||
presence) of a 3rd chin shield pair; from
|
||
<taxonomicName id="29944D16FF89FFAB9B5DC810C4EA51D1" box="[349,504,1655,1679]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="isangdaliri">
|
||
<emphasis id="DCE0EA87FF89FFAB9B5DC810C4EA51D1" box="[349,504,1655,1679]" italics="true" pageId="3">B. isangdaliri</emphasis>
|
||
</taxonomicName>
|
||
by having a longer snout–vent length (66.0–
|
||
<quantity id="296C9B70FF89FFAB99F9C81EC14D51D1" box="[1017,1119,1654,1679]" metricMagnitude="-2" metricUnit="m" metricValue="8.09" pageId="3" unit="mm" value="80.9">80.9 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB9E67C810C1BC51D1" box="[1127,1198,1656,1679]" italics="true" pageId="3">versus</emphasis>
|
||
59.5), a longer forelimb length (
|
||
<quantity id="296C9B70FF89FFAB9B2BC8F3C4A451ED" box="[299,438,1691,1715]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="3" unit="mm" value="1.8" valueMax="2.2" valueMin="1.4">1.4–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB9BBFC8F4C71451ED" box="[447,518,1692,1715]" italics="true" pageId="3">versus</emphasis>
|
||
1.3), a greater number of presacral vertebrae (49
|
||
<emphasis id="DCE0EA87FF89FFAB9E48C8F4C18251ED" box="[1096,1168,1692,1715]" italics="true" pageId="3">versus</emphasis>
|
||
47), greater number of midbody scale rows (24, 25
|
||
<emphasis id="DCE0EA87FF89FFAB9BD9C8A8C7335189" box="[473,545,1728,1751]" italics="true" pageId="3">versus</emphasis>
|
||
22), a greater number of axilla–groin scale rows (80–83
|
||
<emphasis id="DCE0EA87FF89FFAB9EA8C8A8C1E25189" box="[1192,1264,1728,1751]" italics="true" pageId="3">versus</emphasis>
|
||
73), and fewer number of supraciliaries (5
|
||
<emphasis id="DCE0EA87FF89FFAB9BCFC88CC70551A5" box="[463,535,1764,1787]" italics="true" pageId="3">versus</emphasis>
|
||
6); from
|
||
<taxonomicName id="29944D16FF89FFAB9883C88CC7F351A5" box="[643,737,1763,1787]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="ligtas">
|
||
<emphasis id="DCE0EA87FF89FFAB9883C88CC7F351A5" box="[643,737,1763,1787]" italics="true" pageId="3">B. ligtas</emphasis>
|
||
</taxonomicName>
|
||
by having a longer fore-limb length (
|
||
<quantity id="296C9B70FF89FFAB9E8BC88BC00651A5" box="[1163,1300,1763,1787]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="3" unit="mm" value="1.8" valueMax="2.2" valueMin="1.4">1.4–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB9F1CC88CC07651A5" box="[1308,1380,1764,1787]" italics="true" pageId="3">versus</emphasis>
|
||
1.2– 1.4), fewer number of presacral vertebrae (49
|
||
<emphasis id="DCE0EA87FF89FFAB98A4C960C7FE5041" box="[676,748,1800,1823]" italics="true" pageId="3">versus</emphasis>
|
||
50), a greater number of midbody scale rows (24, 25
|
||
<emphasis id="DCE0EA87FF89FFAB9F54C960C0895041" box="[1364,1435,1800,1823]" italics="true" pageId="3">versus</emphasis>
|
||
22), a greater number of axilla–groin scale rows (80–83
|
||
<emphasis id="DCE0EA87FF89FFAB9927C944C67C501D" box="[807,878,1836,1859]" italics="true" pageId="3">versus</emphasis>
|
||
74–76), a greater number of paravertebral scale rows (97–101
|
||
<emphasis id="DCE0EA87FF89FFAB9B38C93AC4925037" box="[312,384,1874,1897]" italics="true" pageId="3">versus</emphasis>
|
||
91–93), and by the absence (
|
||
<emphasis id="DCE0EA87FF89FFAB98C8C93AC6025037" box="[712,784,1874,1897]" italics="true" pageId="3">versus</emphasis>
|
||
presence) of a 3rd chin shield pair; from
|
||
<taxonomicName id="29944D16FF89FFAB9ED5C93AC08E5034" box="[1237,1436,1874,1898]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="mapalanggaon">
|
||
<emphasis id="DCE0EA87FF89FFAB9ED5C93AC08E5034" box="[1237,1436,1874,1898]" italics="true" pageId="3">B. mapalanggaon</emphasis>
|
||
</taxonomicName>
|
||
by having a longer fore-limb (
|
||
<quantity id="296C9B70FF89FFAB9BF9C91EC79450D3" box="[505,646,1910,1934]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="3" unit="mm" value="1.8" valueMax="2.2" valueMin="1.4">1.4–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB9890C91EC7C550D3" box="[656,727,1910,1933]" italics="true" pageId="3">versus</emphasis>
|
||
0.8–1.0) and hind limb length (
|
||
<quantity id="296C9B70FF89FFAB9E4FC91EC1C950D3" box="[1103,1243,1910,1934]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB9EE5C91EC03F50D3" box="[1253,1325,1910,1933]" italics="true" pageId="3">versus</emphasis>
|
||
1.2–1.6), fewer number of presacral vertebrae (49
|
||
<emphasis id="DCE0EA87FF89FFAB985DC9F2C7B650EF" box="[605,676,1946,1969]" italics="true" pageId="3">versus</emphasis>
|
||
51), and a greater number of midbody scale rows (24, 25
|
||
<emphasis id="DCE0EA87FF89FFAB9F2BC9F2C06050EF" box="[1323,1394,1946,1969]" italics="true" pageId="3">versus</emphasis>
|
||
22, 23); from
|
||
<taxonomicName id="29944D16FF89FFAB9B0DC9D6C4B4508B" box="[269,422,1981,2005]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tridactylus">
|
||
<emphasis id="DCE0EA87FF89FFAB9B0DC9D6C4B4508B" box="[269,422,1981,2005]" italics="true" pageId="3">B. tridactylus</emphasis>
|
||
</taxonomicName>
|
||
by having a shorter hind limb length (
|
||
<quantity id="296C9B70FF89FFAB9966C9D6C6E3508B" box="[870,1009,1982,2006]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
|
||
<emphasis id="DCE0EA87FF89FFAB99FAC9D6C153508B" box="[1018,1089,1982,2005]" italics="true" pageId="3">versus</emphasis>
|
||
2.6–3.6), a greater number of presacral vertebrae (49
|
||
<emphasis id="DCE0EA87FF8EFFAC9BA2CEF1C4F857EE" box="[418,490,153,176]" italics="true" pageId="4">versus</emphasis>
|
||
47), a greater number of axilla–groin scale rows (80–83
|
||
<emphasis id="DCE0EA87FF8EFFAC9E76CEF1C1AF57EE" box="[1142,1213,153,176]" italics="true" pageId="4">versus</emphasis>
|
||
72–79), and by the absence (
|
||
<emphasis id="DCE0EA87FF8EFFAC9B00CEA8C45A5789" box="[256,328,192,215]" italics="true" pageId="4">versus</emphasis>
|
||
presence) of a 3rd chin shield pair.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7CE8BC465561D" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<taxonomicName id="29944D16FF8EFFAC9AC7CE8BC71957A5" box="[199,523,227,251]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7CE8BC71957A5" box="[199,523,227,251]" italics="true" pageId="4">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF8EFFAC9812CE8AC77957A5" box="[530,619,226,251]" pageId="4" rank="species">
|
||
<emphasis id="DCE0EA87FF8EFFAC9812CE8AC77957A5" bold="true" box="[530,619,226,251]" pageId="4">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from all limbless species of
|
||
<taxonomicName id="29944D16FF8EFFAC9E95CE8BC03457A5" box="[1173,1318,227,251]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="DCE0EA87FF8EFFAC9E95CE8BC03457A5" box="[1173,1318,227,251]" italics="true" pageId="4">Brachymeles</emphasis>
|
||
</taxonomicName>
|
||
by having limbs, and from all pentadactyl species of
|
||
<taxonomicName id="29944D16FF8EFFAC987CCF6FC61F5641" box="[636,781,263,287]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="DCE0EA87FF8EFFAC987CCF6FC61F5641" box="[636,781,263,287]" italics="true" pageId="4">Brachymeles</emphasis>
|
||
</taxonomicName>
|
||
by having bidactyl fore-limbs and digitless, unidactyl, or bidactyl hind limbs.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7CF26C63E55D9" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7CF26C4F65639" bold="true" box="[199,484,334,359]" pageId="4">
|
||
Description of
|
||
<typeStatus id="312F8837FF8EFFAC9B75CF26C4CC5639" box="[373,478,334,359]" pageId="4" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Details of the head scalation are shown in
|
||
<figureCitation id="76AF2A10FF8EFFAC99C0CF26C1335639" box="[960,1057,334,359]" captionStart="FIGURE 2" captionStartId="7.[151,250,1849,1871]" captionTargetBox="[390,1187,193,1819]" captionTargetId="figure@7.[389,1188,193,1820]" captionTargetPageId="7" captionText="FIGURE 2. Illustration of the head of Brachymeles dalawangdaliri sp. nov. (CAS 137149, holotype) in dorsal, lateral, and ventral views. Taxonomically diagnostic head scales are labeled as follows: C, chin shield; F, frontal; FN, frontonasal; FP, frontoparietal; IL, infralabial; IP, interparietal; L, loreal; M, mental; N, nasal; Nu, nuchal; P, parietal; PF, prefrontal; PM, postmental; PO, preocular; PSO, presubocular; R, rostral; SC, supraciliary; SL, supralabial; SN, supranasal; and SO, supraocular. Roman numerals indicate scales in the supraocular series, with Arabic numbers indicating scales in the supraciliary series. Illustrations by MLP and CDS." httpUri="https://zenodo.org/record/258938/files/figure.png" pageId="4">Figure 2</figureCitation>
|
||
. Adult, male, body small, slender, SVL
|
||
<quantity id="296C9B70FF8EFFAC9AD5CF1BC42C56D4" box="[213,318,371,395]" metricMagnitude="-2" metricUnit="m" metricValue="7.49" pageId="4" unit="mm" value="74.9">74.9 mm</quantity>
|
||
; head weakly differentiated from neck, nearly as wide as body, HW 7.9% SVL, 107.3% HL; HL 37.1% SnFa; SnFa 19.7% SVL; snout narrow, sharply rounded in dorsal and lateral profile, SNL 60.2% HL; ear completely hidden by scales; eyes small, ED 17.9% HL, 40.5%
|
||
<collectingCountry id="96837605FF8EFFAC9972CFD3C6BC568C" box="[882,942,443,466]" name="India" pageId="4">END</collectingCountry>
|
||
, pupil subcircular; body slightly depressed, nearly uniform in thickness, MBW 100.0% MBH; scales smooth, glossy, imbricate; longitudinal scale rows at midbody 24; paravertebral scale rows 97; axilla–groin scale rows 80; limbs short, diminutive, bluntly rounded, with digits reduced to two small digit growths on fore-limbs, single digit growth on hind limb; finger and toe lamellae absent; FLL 3.6% AGD, 2.8% SVL; HLL 3.9% AGD, 3.0% SVL; tail not as wide as body, TW 72.6% MBW, autotomized posterior to cloaca, length unavailable.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7CCFAC6005321" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
Rostral projecting onto dorsal snout to level in line with posterior edge of nasal, roughly equal in width and height, in contact with frontonasal; frontonasal wider than long; nostril small, ovoid, in center of single trapezoidal nasal, longer axis directed posterodorsally and anteroventrally; supranasals present; postnasals absent; prefrontals moderately separated; frontal roughly hexagonal-shaped, its anterior margin in moderate contact with frontonasal, in contact with first two anterior supraoculars, 4× wider than anterior supraocular; supraoculars four; frontoparietals moderate in size, in moderate medial contact, frontoparietals in contact with supraoculars two and three; interparietal moderate in size, its length roughly equal to 2× midline length of frontoparietal, longer than wide, diamond-shaped, wider anteriorly, pineal eyespot not visible; parietals wider than frontoparietals, in moderate contact behind interparietal; head scales damaged in nuchal region; enlarged nuchals present; loreals two, anterior loreal about as long as and slightly higher than posterior loreal; preoculars two; presubocular one; supraciliaries five, the anteriormost contacting prefrontal and separating posterior loreal from first supraocular, posteriormost extending to midline edge of fourth supraocular; subocular scale row single, incomplete, in contact with supralabials; lower eyelid with one row of scales; supralabials seven, first twice the width of others, third, fourth and fifth subocular; infralabials seven (
|
||
<figureCitation id="76AF2A10FF8EFFAC9899CA0EC7FA5321" box="[665,744,1126,1151]" captionStart="FIGURE 2" captionStartId="7.[151,250,1849,1871]" captionTargetBox="[390,1187,193,1819]" captionTargetId="figure@7.[389,1188,193,1820]" captionTargetPageId="7" captionText="FIGURE 2. Illustration of the head of Brachymeles dalawangdaliri sp. nov. (CAS 137149, holotype) in dorsal, lateral, and ventral views. Taxonomically diagnostic head scales are labeled as follows: C, chin shield; F, frontal; FN, frontonasal; FP, frontoparietal; IL, infralabial; IP, interparietal; L, loreal; M, mental; N, nasal; Nu, nuchal; P, parietal; PF, prefrontal; PM, postmental; PO, preocular; PSO, presubocular; R, rostral; SC, supraciliary; SL, supralabial; SN, supranasal; and SO, supraocular. Roman numerals indicate scales in the supraocular series, with Arabic numbers indicating scales in the supraciliary series. Illustrations by MLP and CDS." httpUri="https://zenodo.org/record/258938/files/figure.png" pageId="4">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="76AF2A10FF8EFFAC98F4CA0FC6165321" box="[756,772,1127,1151]" captionStart="FIGURE 3" captionStartId="8.[151,250,609,631]" captionTargetBox="[150,1436,193,587]" captionTargetId="figure@8.[150,1437,193,588]" captionTargetPageId="8" captionText="FIGURE 3. Photograph of lateral and ventral views of head of preserved holotype of Brachymeles dalawangdaliri sp. nov. (CAS 137149). Photographs taken by JLW and MLP." httpUri="https://zenodo.org/record/258939/files/figure.png" pageId="4">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7CAE2C6295251" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
Mental wider than long, in contact with first infralabial on left side of head, fused with first infralabial on right side of head; postmental single, enlarged, its width equal to width of mental; followed by two pairs of enlarged chin shields, first pair moderately separated, second pair slightly smaller than first, broadly separated by three medial scales (
|
||
<figureCitation id="76AF2A10FF8EFFAC9AE9CA9EC42A5251" box="[233,312,1270,1295]" captionStart="FIGURE 2" captionStartId="7.[151,250,1849,1871]" captionTargetBox="[390,1187,193,1819]" captionTargetId="figure@7.[389,1188,193,1820]" captionTargetPageId="7" captionText="FIGURE 2. Illustration of the head of Brachymeles dalawangdaliri sp. nov. (CAS 137149, holotype) in dorsal, lateral, and ventral views. Taxonomically diagnostic head scales are labeled as follows: C, chin shield; F, frontal; FN, frontonasal; FP, frontoparietal; IL, infralabial; IP, interparietal; L, loreal; M, mental; N, nasal; Nu, nuchal; P, parietal; PF, prefrontal; PM, postmental; PO, preocular; PSO, presubocular; R, rostral; SC, supraciliary; SL, supralabial; SN, supranasal; and SO, supraocular. Roman numerals indicate scales in the supraocular series, with Arabic numbers indicating scales in the supraciliary series. Illustrations by MLP and CDS." httpUri="https://zenodo.org/record/258938/files/figure.png" pageId="4">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="76AF2A10FF8EFFAC9B44CA9FC4465251" box="[324,340,1271,1295]" captionStart="FIGURE 3" captionStartId="8.[151,250,609,631]" captionTargetBox="[150,1436,193,587]" captionTargetId="figure@8.[150,1437,193,588]" captionTargetPageId="8" captionText="FIGURE 3. Photograph of lateral and ventral views of head of preserved holotype of Brachymeles dalawangdaliri sp. nov. (CAS 137149). Photographs taken by JLW and MLP." httpUri="https://zenodo.org/record/258939/files/figure.png" pageId="4">3</figureCitation>
|
||
). Scales on limbs smaller than body scales.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7CB72C7F85155" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7CB72C429526D" bold="true" box="[199,315,1306,1331]" pageId="4">Variation.</emphasis>
|
||
Differences in digit number and scalation patterns were observed between the
|
||
<typeStatus id="312F8837FF8EFFAC9F04CB72C074526D" box="[1284,1382,1306,1331]" pageId="4" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="312F8837FF8EFFAC9A97CB28C5E55206" box="[151,247,1344,1368]" pageId="4" type="paratype">paratype</typeStatus>
|
||
series. The
|
||
<typeStatus id="312F8837FF8EFFAC9B7FCB56C4F35209" box="[383,481,1342,1367]" pageId="4" type="holotype">holotype</typeStatus>
|
||
(CAS 137149) and three
|
||
<typeStatus id="312F8837FF8EFFAC9904CB28C67D5206" box="[772,879,1344,1368]" pageId="4" type="paratype">paratypes</typeStatus>
|
||
(CAS 137148, 137153, 137154) possessed seven supralabials and seven infralabials, with three other
|
||
<typeStatus id="312F8837FF8EFFAC98E6CB0BC6435225" box="[742,849,1379,1403]" pageId="4" type="paratype">paratypes</typeStatus>
|
||
(CAS 137150–137152) having six supralabials and six infralabials. Unlike the
|
||
<typeStatus id="312F8837FF8EFFAC9BCDCBEEC73D52C1" box="[461,559,1414,1439]" pageId="4" type="holotype">holotype</typeStatus>
|
||
and majority of
|
||
<typeStatus id="312F8837FF8EFFAC98EFCBE0C64F52FE" box="[751,861,1416,1440]" pageId="4" type="paratype">paratypes</typeStatus>
|
||
, the frontoparietal scales were observed to contact medially in two specimens (CAS 137153, 137154). Observed digit number variation included body plans with two fingers and one toe (CAS 137149 [
|
||
<typeStatus id="312F8837FF8EFFAC9825CBA6C79D52B9" box="[549,655,1486,1511]" pageId="4" type="holotype">holotype</typeStatus>
|
||
], 137151, 137153, 137154), two fingers and two toes (CAS 137148, 137150), and two fingers and no toes (CAS 137152).
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7C87EC6E35129" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7C87EC7355171" bold="true" box="[199,551,1558,1583]" pageId="4">
|
||
Coloration of
|
||
<typeStatus id="312F8837FF8EFFAC9B6EC87EC4C75171" box="[366,469,1558,1583]" pageId="4" type="holotype">holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
Coloration in life is unrecorded; however, because
|
||
<taxonomicName id="29944D16FF8EFFAC9E6CC87FC1EF5171" box="[1132,1277,1559,1583]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="DCE0EA87FF8EFFAC9E6CC87FC1EF5171" box="[1132,1277,1559,1583]" italics="true" pageId="4">Brachymeles</emphasis>
|
||
</taxonomicName>
|
||
specimens do not change significantly during preservation (Siler & Brown 2010;
|
||
<bibRefCitation id="8A054B64FF8EFFAC9987C852C15A510D" author="Siler" box="[903,1096,1594,1619]" pageId="4" refString="Siler, C. D., Fuiten, A. M., Jones, R. M., Alcala, A. C. & Brown, R. M. (2011 a) Phylogeny-based species delimitation in Philippine slender skinks (Reptilia: Squamata: Scincidae) II: taxonomic revision of Brachymeles samarensis and description of five new species. Herpetological Monographs, 25, 76 - 112. http: // dx. doi. org / 10.1655 / herpmonographs-d- 10 - 00013.1" type="journal article" year="2011" yearSuffix="a">
|
||
Siler
|
||
<emphasis id="DCE0EA87FF8EFFAC99C2C853C6E8510D" box="[962,1018,1595,1619]" italics="true" pageId="4">et al.</emphasis>
|
||
2011a
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="8A054B64FF8EFFAC9E52C853C188510D" author="Siler" box="[1106,1178,1595,1619]" pageId="4" refString="Siler, C. D., Jones, R. M., Diesmos, A. C., Diesmos, M. L. & Brown, R. M. (2012 a) Phylogeny-based species delimitation in Philippine slender skinks (Reptilia: Squamata: Scincidae) III: taxonomic revision of the Brachymeles gracilis Complex, with descriptions of three new species. Herpetological Monographs, 26, 135 - 172. http: // dx. doi. org / 10.1655 / herpmonographs-d- 11 - 00006.1" type="journal article" year="2012" yearSuffix="a">2012a</bibRefCitation>
|
||
; Davis
|
||
<emphasis id="DCE0EA87FF8EFFAC9EEDC853C037510D" box="[1261,1317,1595,1619]" italics="true" pageId="4">et al.</emphasis>
|
||
2014), we suspect that the preserved coloration and patterns are much like those in life.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7C8EAC1F25059" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7C8EAC78B51C5" bold="true" box="[199,665,1666,1691]" pageId="4">
|
||
Coloration of
|
||
<typeStatus id="312F8837FF8EFFAC9B70C8EAC4C551C5" box="[368,471,1666,1691]" pageId="4" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
The background color on the dorsal, lateral and ventral sides of the body and tail is a solid Prout’s Brown (Color 47;
|
||
<bibRefCitation id="8A054B64FF8EFFAC98BFC8CEC64351E1" author="Kohler" box="[703,849,1702,1727]" pageId="4" refString="Kohler, G. (2012) Color Catalog for Field Biologists. Herpeton, Offenbach, 49 pp." type="book" year="2012">Köhler 2012</bibRefCitation>
|
||
). The ventral side of the head is the same solid color as the ventral side of the body (
|
||
<figureCitation id="76AF2A10FF8EFFAC9BF2C8A2C72451BD" box="[498,566,1738,1763]" captionStart="FIGURE 3" captionStartId="8.[151,250,609,631]" captionTargetBox="[150,1436,193,587]" captionTargetId="figure@8.[150,1437,193,588]" captionTargetPageId="8" captionText="FIGURE 3. Photograph of lateral and ventral views of head of preserved holotype of Brachymeles dalawangdaliri sp. nov. (CAS 137149). Photographs taken by JLW and MLP." httpUri="https://zenodo.org/record/258939/files/figure.png" pageId="4">Fig. 3</figureCitation>
|
||
; Prout’s Brown; Color 47;
|
||
<bibRefCitation id="8A054B64FF8EFFAC9963C8A2C6E651BD" author="Kohler" box="[867,1012,1738,1763]" pageId="4" refString="Kohler, G. (2012) Color Catalog for Field Biologists. Herpeton, Offenbach, 49 pp." type="book" year="2012">Köhler 2012</bibRefCitation>
|
||
). On the dorsal and lateral sides of the head, a single Fuscous splotch can be seen just above each orbit (
|
||
<figureCitation id="76AF2A10FF8EFFAC9972C886C6A45059" box="[882,950,1774,1799]" captionStart="FIGURE 3" captionStartId="8.[151,250,609,631]" captionTargetBox="[150,1436,193,587]" captionTargetId="figure@8.[150,1437,193,588]" captionTargetPageId="8" captionText="FIGURE 3. Photograph of lateral and ventral views of head of preserved holotype of Brachymeles dalawangdaliri sp. nov. (CAS 137149). Photographs taken by JLW and MLP." httpUri="https://zenodo.org/record/258939/files/figure.png" pageId="4">Fig. 3</figureCitation>
|
||
; Color 283;
|
||
<bibRefCitation id="8A054B64FF8EFFAC9E42C886C1C65059" author="Kohler" box="[1090,1236,1774,1799]" pageId="4" refString="Kohler, G. (2012) Color Catalog for Field Biologists. Herpeton, Offenbach, 49 pp." type="book" year="2012">Köhler 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68E651EFF8EFFAC9AC7C97AC19A502D" pageId="4" type="etymology">
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7C97AC19A502D" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7C97AC45C5075" bold="true" box="[199,334,1810,1835]" pageId="4">Etymology.</emphasis>
|
||
The name of the new species presented in this paper is derived from the Tagalog (Filipino) terms ‘dalawa,’ meaning two, and ‘daliri,’ meaning finger, referencing one of the defining characteristics of the new species, the two digits on the fore-limbs. Suggested common name: Tablas Slender Skink.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68E651EFF8EFFAD9AC7C916C0965447" lastPageId="5" pageId="4" type="distribution">
|
||
<paragraph id="EE2B3695FF8EFFAC9AC7C916C46550E5" blockId="4.[151,1437,151,1979]" pageId="4">
|
||
<emphasis id="DCE0EA87FF8EFFAC9AC7C916C47250C9" bold="true" box="[199,352,1918,1943]" pageId="4">Distribution.</emphasis>
|
||
<taxonomicName id="29944D16FF8EFFAC9B66C917C7B850C9" box="[358,682,1919,1943]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF8EFFAC9B66C917C7B850C9" box="[358,682,1919,1943]" italics="true" pageId="4">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF8EFFAC98B1C916C61850C9" box="[689,778,1918,1943]" pageId="4" rank="species">
|
||
<emphasis id="DCE0EA87FF8EFFAC98B1C916C61850C9" bold="true" box="[689,778,1918,1943]" pageId="4">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is known only from Tablas Island,
|
||
<taxonomicName id="29944D16FF8EFFAC9E94C916C1EE50C9" box="[1172,1276,1918,1943]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="romblon">Romblon</taxonomicName>
|
||
Island Group,
|
||
<collectingCountry id="96837605FF8EFFAC9A97C9CAC40750E5" box="[151,277,1954,1979]" name="Philippines" pageId="4">Philippines</collectingCountry>
|
||
(
|
||
<figureCitation id="76AF2A10FF8EFFAC9B24C9CAC47750E5" box="[292,357,1954,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1101,1123]" captionTargetBox="[151,1436,193,1079]" captionTargetId="figure@2.[151,1437,193,1080]" captionTargetPageId="2" captionText="FIGURE 1. (Left) Map of the Philippine islands, with labels provided for islands with representative samples used for this study. The five recognized major faunal regions (PAICs), major island groups, and additional deep-water islands are labeled for reference. Current islands in the Philippines are shown in medium grey to black; light gray areas enclosed in black 120 m bathymetric contours indicate the hypothesized maximum extent of land during the mid- to late Pleistocene. Location of the Romblon Island Group outlined by a square; Tablas Island highlighted in black. (Right) Map of Romblon Province in relation to the remainder of the Philippines. Known distribution of Brachymeles dalawangdaliri sp. nov. on Tablas Island is indicated with a star." httpUri="https://zenodo.org/record/258937/files/figure.png" pageId="4">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A77CCDBC0965447" blockId="5.[119,1861,691,865]" pageId="5">
|
||
<emphasis id="DCE0EA87FF8FFFAD9A77CCDBC5F65596" bold="true" box="[119,228,691,712]" pageId="5">TABLE ·.</emphasis>
|
||
Summary of meristic anđ mensural characters among species of the
|
||
<taxonomicName id="29944D16FF8FFFAD99A3CCDBC1675596" box="[931,1141,691,712]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="bonitae">
|
||
<emphasis id="DCE0EA87FF8FFFAD99A3CCDBC1675596" box="[931,1141,691,712]" italics="true" pageId="5">Brachymeles bonitae</emphasis>
|
||
</taxonomicName>
|
||
Complex. Sample size, bođy length anđ total length among males anđ females, anđ general geographical đistribution (PAIC = Pleistocene Aggregate Islanđ Complexes, sensu
|
||
<bibRefCitation id="8A054B64FF8FFFAD9E7ECCB4C09255AF" author="Brown" box="[1150,1408,732,753]" pageId="5" refString="Brown, R. M. & Diesmos, A. C. (2002) Application of lineage-based species concepts to oceanic island frog populations: the effects of differing taxonomic philosophies on the estimation of Philippine biodiversity. Silliman Journal, 42, 133 - 162." type="journal article" year="2002">Brown & Diesmos [2002]</bibRefCitation>
|
||
) are incluđeđ for reference (SVL, TotL, FLL, HLL given as range over mean ± stanđarđ đeviation; all bođy proportions given as percentage over mean ± stanđarđ đeviation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68E651EFF8FFFA09B23CD59C4FC531D" lastPageId="8" pageId="5" type="description">
|
||
<paragraph id="EE2B3695FF8FFFAD9B23CD59C211543F" blockId="5.[119,1861,691,865]" pageId="5">
|
||
<emphasis id="DCE0EA87FF8FFFAD9B23CD59C7B65418" box="[291,676,817,838]" italics="true" pageId="5">
|
||
<taxonomicName id="29944D16FF8FFFAD9B23CD59C47F5418" box="[291,365,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="bonitae">bonitae</taxonomicName>
|
||
<taxonomicName id="29944D16FF8FFFAD980CCD59C7B65418" box="[524,676,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">dalawangdaliri</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="C7D357FCFF8FFFAD98AACD59C7EA5418" box="[682,760,817,838]" pageId="5" rank="species">
|
||
<emphasis id="DCE0EA87FF8FFFAD98AACD59C7EA5418" bold="true" box="[682,760,817,838]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="DCE0EA87FF8FFFAD9937CD59C2115418" box="[823,1795,817,838]" italics="true" pageId="5">
|
||
<taxonomicName id="29944D16FF8FFFAD9937CD59C6865418" box="[823,916,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="ilocandia">ilocandia</taxonomicName>
|
||
<taxonomicName id="29944D16FF8FFFAD99DFCD59C1595418" box="[991,1099,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="isangdaliri">isangdaliri</taxonomicName>
|
||
<taxonomicName id="29944D16FF8FFFAD9EC8CD59C1ED5418" box="[1224,1279,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="ligtas">ligtas</taxonomicName>
|
||
<taxonomicName id="29944D16FF8FFFAD9FB1CD59C3575418" box="[1457,1605,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="mapalanggaon">mapalanggaon</taxonomicName>
|
||
<taxonomicName id="29944D16FF8FFFAD9C9ACD59C2115418" box="[1690,1795,817,838]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tridactylus">tridactylus</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<quantity id="296C9B70FF8FFFAD9B2BCD24C441543F" box="[299,339,844,865]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="5" unit="m" value="3.0">3 m</quantity>
|
||
, 1 f) (
|
||
<quantity id="296C9B70FF8FFFAD9813CD24C72E543F" box="[531,572,844,865]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="5" unit="m" value="2.0">2 m</quantity>
|
||
, 5 f) (9 f) (1 f) (
|
||
<quantity id="296C9B70FF8FFFAD9ED0CD24C1EA543F" box="[1232,1272,844,865]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="5" unit="m" value="3.0">3 m</quantity>
|
||
, 2 f) (
|
||
<quantity id="296C9B70FF8FFFAD9FB9CD24C0F3543F" box="[1465,1505,844,865]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="5" unit="m" value="3.0">3 m</quantity>
|
||
, 6 f) (
|
||
<quantity id="296C9B70FF8FFFAD9CA2CD24C3C4543F" box="[1698,1750,844,865]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="5" unit="m" value="12.0">12 m</quantity>
|
||
, 9 f)
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCDEBC3C354ED" blockId="5.[127,1827,899,1510]" pageId="5">Range Luzon & Polillo Tablas Islanđ Luzon & Luzon Islanđ Lubang Islanđ Masbate Islanđ West Visayan islanđs Camiguin Norte PAIC</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9937CDD1C6695490" blockId="5.[127,1827,899,1510]" box="[823,891,953,974]" pageId="5">islanđs</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCDB3C21C5355" blockId="5.[127,1827,899,1510]" pageId="5">SVL (f) 69.4 69.8=80.9 65.7=77.6 59.5 60.7=69.2 61.7=75.8 59.9=82.3 (73.6 ± 4.4) (73.7 ± 3.7) (65.0 ± 6.0) (67.2 ± 5.4) (71.4 ± 6.9)</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCA79C21C531F" blockId="5.[127,1827,899,1510]" pageId="5">
|
||
SVL (m) 69.7=78.4 66.0=
|
||
<geoCoordinate id="8BA05052FF8FFFAD9844CA79C65A5378" box="[580,840,1041,1062]" direction="north" orientation="latitude" pageId="5" precision="5555" value="74.9">74.9 N</geoCoordinate>
|
||
/A N/A 69.4=79.6 65.1=72.7 60.7=77.6 (72.8 ± 4.8) (70.4 ± 6.3) (74.5 ± 5.1) (68.4 ± 3.9) (69.0 ± 6.0)
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCA26C3CB5309" blockId="5.[127,1827,899,1510]" box="[127,1753,1097,1123]" pageId="5">TotL (f) N/A N/A 122.3=146.0 106.1 119.4 120.2 133.6 1</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9937CA01C6A55320" blockId="5.[127,1827,899,1510]" box="[823,951,1129,1150]" pageId="5">(134.1 ± 8.0)</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCAECC20853EA" blockId="5.[127,1827,899,1510]" pageId="5">
|
||
TotL (m)
|
||
<geoCoordinate id="8BA05052FF8FFFAD9B23CAECC70F53C7" box="[291,541,1156,1177]" direction="north" orientation="latitude" pageId="5" precision="5555" value="122.0">122.0 N</geoCoordinate>
|
||
/A N/A N/A 160.6 112.6=118.6 120.9=154.1 (115.6 ± 4.3) (136.0 ± 9.8)
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A9DCAA9C3E453AF" blockId="5.[127,1827,899,1510]" pageId="5">/SVL 73 N/A 81=90 78 97=102 67=84 85=112 (85 ± 3) (99 ± 4) (78 ± 9) (95 ± 10)</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCA96C2135270" blockId="5.[127,1827,899,1510]" pageId="5">FLL 1.0=1.7 1.4=2.2 1.1=1.4 1.3 1.2=1.4 0.8=1.0 1.5=2.5 (1.3 ± 0.3) (1.9 ± 0.3) (1.3 ± 0.1) (1.3 ± 0.1) (0.9 ± 0.1) (2.0 ± 0.3)</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCB54C3CE5235" blockId="5.[127,1827,899,1510]" pageId="5">FLL/SVL 1=2 2=3 1=2 2 2=2 1=2 2=4 (2 ± 0) (3 ± 0) (2 ± 0) (2 ± 0) (1 ± 0) (3 ± 0)</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAD9A7FCB10C21352F7" blockId="5.[127,1827,899,1510]" pageId="5">HLL 1.5=2.3 1.9=2.2 1.6=1.9 2.2 1.6=2.0 1.2=1.6 2.6=3.6 (1.9 ± 0.3) (2.1 ± 0.1) (1.8 ± 0.1) (1.8 ± 0.1) (1.4 ± 0.1) (3.1 ± 0.3)</paragraph>
|
||
<paragraph id="EE2B3695FF8FFFAE9A7FCBDEC62C545D" blockId="5.[127,1827,899,1510]" lastBlockId="6.[130,1829,668,841]" lastPageId="6" pageId="5">
|
||
HLL/SVL 2=3 2=3 2=3 4 2=3 2=2 3=6 (3 ± 0) (3 ± 0) (2 ± 0) (3 ± 0) (2 ± 0) (5 ± 1)
|
||
<emphasis id="DCE0EA87FF8CFFAE9A82CCF5C5FC55EC" bold="true" box="[130,238,669,690]" pageId="6">TABLE 2.</emphasis>
|
||
Summary of meristic anđ qualitative điagnostic characters (present, absent) among species of the
|
||
<taxonomicName id="29944D16FF8CFFAE9ECDCCF4C08855EF" box="[1229,1434,668,689]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="bonitae">
|
||
<emphasis id="DCE0EA87FF8CFFAE9ECDCCF4C08855EF" box="[1229,1434,668,689]" italics="true" pageId="6">Brachymeles bonitae</emphasis>
|
||
</taxonomicName>
|
||
Complex. The pairs of enlargeđ scales posterior to the postmental scale are abbreviateđ as chin shielđ pairs with reference to the 1st, 2nđ, anđ 3rđ pairs (when present). In cases of scale count variation within species, numbers of inđiviđuals showing specific counts are given in parentheses.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9BB8CD70C3F35417" blockId="6.[130,1829,668,841]" pageId="6">
|
||
<emphasis id="DCE0EA87FF8CFFAE9BB8CD70C6155473" box="[440,775,792,813]" italics="true" pageId="6">
|
||
<taxonomicName id="29944D16FF8CFFAE9BB8CD70C7125473" box="[440,512,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="bonitae">bonitae</taxonomicName>
|
||
<taxonomicName id="29944D16FF8CFFAE9873CD70C6155473" box="[627,775,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">dalawangdaliri</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="C7D357FCFF8CFFAE990DCD71C6485470" box="[781,858,793,814]" pageId="6" rank="species">
|
||
<emphasis id="DCE0EA87FF8CFFAE990DCD71C6485470" bold="true" box="[781,858,793,814]" pageId="6">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="DCE0EA87FF8CFFAE9976CD70C3F05473" box="[886,1762,792,813]" italics="true" pageId="6">
|
||
<taxonomicName id="29944D16FF8CFFAE9976CD70C6C25473" box="[886,976,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="ilocandia">ilocandia</taxonomicName>
|
||
<taxonomicName id="29944D16FF8CFFAE9E48CD70C1A35473" box="[1096,1201,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="isangdaliri">isangdaliri</taxonomicName>
|
||
<taxonomicName id="29944D16FF8CFFAE9F04CD70C02B5473" box="[1284,1337,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="ligtas">ligtas</taxonomicName>
|
||
<taxonomicName id="29944D16FF8CFFAE9FBFCD70C35D5473" box="[1471,1615,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="mapalanggaon">mapalanggaon</taxonomicName>
|
||
<taxonomicName id="29944D16FF8CFFAE9C7BCD70C3F05473" box="[1659,1762,792,813]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="tridactylus">tridactylus</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<quantity id="296C9B70FF8CFFAE9BC0CD5CC4F45417" box="[448,486,820,841]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" unit="m" value="3.0">3 m</quantity>
|
||
, 1 f) (
|
||
<quantity id="296C9B70FF8CFFAE9880CD5CC7BA5417" box="[640,680,820,841]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" unit="m" value="2.0">2 m</quantity>
|
||
, 5 f) (9 f) (1 f) (
|
||
<quantity id="296C9B70FF8CFFAE9F0CCD5CC0205417" box="[1292,1330,820,841]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" unit="m" value="3.0">3 m</quantity>
|
||
, 2 f) (
|
||
<quantity id="296C9B70FF8CFFAE9FC7CD5CC0FF5417" box="[1479,1517,820,841]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" unit="m" value="3.0">3 m</quantity>
|
||
, 6 f) (
|
||
<quantity id="296C9B70FF8CFFAE9C82CD5CC3A45417" box="[1666,1718,820,841]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="6" unit="m" value="12.0">12 m</quantity>
|
||
, 9 f)
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCD1DC4CC54C7" blockId="6.[125,1829,880,1554]" pageId="6">Number of đigits (fingers/toes) 0/0 1 2/0=2 0/0 1/1 2/0 0/0 3/3 2/1 1</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCDC6C381549D" blockId="6.[125,1829,880,1554]" box="[125,1683,942,963]" pageId="6">PSV 53 49 50=53 47 50 51 47</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCDA4C3A554BF" blockId="6.[125,1829,880,1554]" box="[125,1719,972,993]" pageId="6">MBSR 21=24 24, 25 22=24 22 22 22, 23 22=24</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCD83C3AA535E" blockId="6.[125,1829,880,1554]" box="[125,1720,1003,1024]" pageId="6">AGSR 83=90 80=83 80=82 73 74=76 80=84 72=79</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCA61C3955390" blockId="6.[125,1829,880,1554]" pageId="6">
|
||
PVSR 103=110 97=101 97=100 97 91=93 99=102 90=
|
||
<date id="9A2A1055FF8CFFAE9CA0CA61C76D5363" pageId="6" value="1998-06-06">98 6 6</date>
|
||
(3) 6 6 6 6 6 (13) 7 (4) 7 (8) 5 6 (3) 5 (3) 6 6 5 (8) 6 (13) 7 (4) 6 (6) 6 (1) 7 (8) 5 5 5 6 5 5 5 4 4 4 4 4 4 4
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCABFC6A95256" blockId="6.[125,1829,880,1554]" pageId="6">Prefrontal contact Absent Absent Absent or Point Absent Absent Absent Absent contact</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCB79C310521F" blockId="6.[125,1829,880,1554]" pageId="6">Frontoparietal contact Absent Present or Absent Absent Absent Present Point contact or Absent Absent</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A99CB22C2375201" blockId="6.[125,1829,880,1554]" box="[153,1829,1354,1375]" pageId="6">chin shielđ pair contact Absent Absent Absent Absent Present or Absent Absent Present or Absent</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A9CCB01C3D35220" blockId="6.[125,1829,880,1554]" box="[156,1729,1385,1406]" pageId="6">chin shielđ pair Present Absent Present Absent Present Present or Absent Present</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCBEFC3AD52C2" blockId="6.[125,1829,880,1554]" box="[125,1727,1410,1436]" pageId="6">Mental/1st IFL fusion Present Absent Present or Absent Absent Absent Present or Absent Absent</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCBCEC3D352E5" blockId="6.[125,1829,880,1554]" box="[125,1729,1446,1467]" pageId="6">Enlargeđ nuchals Present Present Present Present Present Present Present</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A7DCBACC3AD5287" blockId="6.[125,1829,880,1554]" box="[125,1727,1476,1497]" pageId="6">Longituđinal rows of đark Absent Absent Absent Absent Absent Absent Absent</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A95CBB7C5D452AA" blockId="6.[125,1829,880,1554]" box="[149,198,1503,1524]" pageId="6">spots</paragraph>
|
||
<paragraph id="EE2B3695FF8CFFAE9A8BCB95C4B8514C" blockId="6.[125,1829,880,1554]" box="[139,426,1533,1554]" pageId="6">Observeđ for two inđiviđuals.</paragraph>
|
||
<caption id="BAEB661DFF8DFFAF9A97C951C78550B6" httpUri="https://zenodo.org/record/258938/files/figure.png" pageId="7" targetBox="[390,1187,193,1819]" targetPageId="7">
|
||
<paragraph id="EE2B3695FF8DFFAF9A97C951C78550B6" blockId="7.[151,1436,1849,2024]" pageId="7">
|
||
<emphasis id="DCE0EA87FF8DFFAF9A97C951C4075010" bold="true" box="[151,277,1849,1871]" pageId="7">FIGURE 2.</emphasis>
|
||
Illustration of the head of
|
||
<taxonomicName id="29944D16FF8DFFAF982FC951C6445011" box="[559,854,1849,1871]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF8DFFAF982FC951C6445011" box="[559,854,1849,1871]" italics="true" pageId="7">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF8DFFAF995FC951C6A35010" box="[863,945,1849,1870]" pageId="7" rank="species">
|
||
<emphasis id="DCE0EA87FF8DFFAF995FC951C6A35010" bold="true" box="[863,945,1849,1870]" pageId="7">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(CAS 137149, holotype) in dorsal, lateral, and ventral views. Taxonomically diagnostic head scales are labeled as follows: C, chin shield; F, frontal; FN, frontonasal; FP, frontoparietal; IL, infralabial; IP, interparietal; L, loreal; M, mental; N, nasal; Nu, nuchal; P, parietal; PF, prefrontal; PM, postmental; PO, preocular; PSO, presubocular; R, rostral; SC, supraciliary; SL, supralabial; SN, supranasal; and SO, supraocular. Roman numerals indicate scales in the supraocular series, with Arabic numbers indicating scales in the supraciliary series. Illustrations by MLP and CDS.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BAEB661DFF82FFA09A97CC09C7A155C8" httpUri="https://zenodo.org/record/258939/files/figure.png" pageId="8" targetBox="[150,1436,193,587]" targetPageId="8">
|
||
<paragraph id="EE2B3695FF82FFA09A97CC09C7A155C8" blockId="8.[151,1435,609,662]" pageId="8">
|
||
<emphasis id="DCE0EA87FF82FFA09A97CC09C4065528" bold="true" box="[151,276,609,631]" pageId="8">FIGURE 3.</emphasis>
|
||
Photograph of lateral and ventral views of head of preserved holotype of
|
||
<taxonomicName id="29944D16FF82FFA09E1ACC09C0525529" box="[1050,1344,609,631]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF82FFA09E1ACC09C0525529" box="[1050,1344,609,631]" italics="true" pageId="8">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF82FFA09F49CC09C0895528" box="[1353,1435,609,630]" pageId="8" rank="species">
|
||
<emphasis id="DCE0EA87FF82FFA09F49CC09C0895528" bold="true" box="[1353,1435,609,630]" pageId="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(CAS 137149). Photographs taken by JLW and MLP.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EE2B3695FF82FFA09AC7CCAAC4F054D1" blockId="8.[151,1436,706,1091]" pageId="8">
|
||
<emphasis id="DCE0EA87FF82FFA09AC7CCAAC4955585" bold="true" box="[199,391,706,731]" pageId="8">Natural history.</emphasis>
|
||
Presumably,
|
||
<taxonomicName id="29944D16FF82FFA09821CCABC67A5585" box="[545,872,707,731]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF82FFA09821CCABC67A5585" box="[545,872,707,731]" italics="true" pageId="8">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF82FFA09970CCAAC6D95585" box="[880,971,706,731]" pageId="8" rank="species">
|
||
<emphasis id="DCE0EA87FF82FFA09970CCAAC6D95585" bold="true" box="[880,971,706,731]" pageId="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
once occurred in primary growth forest; however, little to no primary forest remains on Tablas Island. This species has never been observed in high densities, a pattern consistent with many members of the
|
||
<taxonomicName id="29944D16FF82FFA09924CD63C68A547D" box="[804,920,779,803]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="bonitae">
|
||
<emphasis id="DCE0EA87FF82FFA09924CD63C68A547D" box="[804,920,779,803]" italics="true" pageId="8">B. bonitae</emphasis>
|
||
</taxonomicName>
|
||
Complex. Furthermore, no recent collections of this species have been made, with the last documented observation of
|
||
<taxonomicName id="29944D16FF82FFA099E3CD58C1A35419" box="[995,1201,815,839]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF82FFA099E3CD58C1A35419" box="[995,1201,815,839]" italics="true" pageId="8">B. dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF82FFA09EBACD46C0045419" box="[1210,1302,814,839]" pageId="8" rank="species">
|
||
<emphasis id="DCE0EA87FF82FFA09EBACD46C0045419" bold="true" box="[1210,1302,814,839]" pageId="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the wild occurring more than four decades ago.
|
||
<taxonomicName id="29944D16FF82FFA0985BCD3BC6B35435" box="[603,929,851,875]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF82FFA0985BCD3BC6B35435" box="[603,929,851,875]" italics="true" pageId="8">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF82FFA099ABCD3AC1145435" box="[939,1030,850,875]" pageId="8" rank="species">
|
||
<emphasis id="DCE0EA87FF82FFA099ABCD3AC1145435" bold="true" box="[939,1030,850,875]" pageId="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
may be found in sympatry with
|
||
<taxonomicName id="29944D16FF82FFA09F84CD3BC4CC54D1" authority="Siler & Brown 2010" authorityName="Siler & Brown" authorityYear="2010" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="talinis">
|
||
<emphasis id="DCE0EA87FF82FFA09F84CD3BC5CF54D1" italics="true" pageId="8">B. talinis</emphasis>
|
||
(Siler & Brown 2010)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EE2B3695FF82FFA09AC7CDF2C4FC531D" blockId="8.[151,1436,706,1091]" pageId="8">
|
||
Little is known about the ecology and distribution of
|
||
<taxonomicName id="29944D16FF82FFA09929CDF3C17D54ED" box="[809,1135,923,947]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF82FFA09929CDF3C17D54ED" box="[809,1135,923,947]" italics="true" pageId="8">Brachymeles dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF82FFA09E77CDF2C1C054ED" box="[1143,1234,922,947]" pageId="8" rank="species">
|
||
<emphasis id="DCE0EA87FF82FFA09E77CDF2C1C054ED" bold="true" box="[1143,1234,922,947]" pageId="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
on Tablas Island. However, this species is endemic to an island with complete or near complete deforestation and no recent observations of wild populations. Therefore, in evaluating
|
||
<taxonomicName id="29944D16FF82FFA09927CD8BC6E154A5" box="[807,1011,995,1019]" class="Reptilia" family="Scincidae" genus="Brachymeles" kingdom="Animalia" order="Squamata" pageId="8" phylum="Chordata" rank="species" species="dalawangdaliri" status="sp. nov.">
|
||
<emphasis id="DCE0EA87FF82FFA09927CD8BC6E154A5" box="[807,1011,995,1019]" italics="true" pageId="8">B. dalawangdaliri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7D357FCFF82FFA099FACD8AC14654A5" box="[1018,1108,994,1019]" pageId="8" rank="species">
|
||
<emphasis id="DCE0EA87FF82FFA099FACD8AC14654A5" bold="true" box="[1018,1108,994,1019]" pageId="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
against the IUCN criteria for classification, we find that it qualifies for the status of Vulnerable (VU) based on the following criteria: VU B2ab(iii,iv); D2 (IUCN 2015).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |