plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/8A/74/87/8A7487DBFFCEFFDDFF53059BFE886F0E.xml
maintenance 65511e311a added 8A
2024-06-21 12:43:10 +02:00

702 lines
116 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="20C8313A6EDABE533C2AC46B5BCE232C" ID-CLB-Dataset="8609" ID-DOI="10.11646/zootaxa.4845.3.5" ID-GBIF-Dataset="98f82204-695b-437b-95b3-c063bbb6d03b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4406729" ID-ZooBank="F00936AE-D1E5-4F0A-A147-28A9CBBA6C85" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609425130760" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2020" docId="8A7487DBFFCEFFDDFF53059BFE886F0E" docLanguage="en" docName="zootaxa.4845.3.5.pdf" docOrigin="Zootaxa 4845 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus vanuatu Anker 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="407" masterDocId="764DFFA3FFC6FFD3FFC4042DFC046B73" masterDocTitle="On two new deep-water snapping shrimps from the Indo-West Pacific (Decapoda Alpheidae: Alpheus)" masterLastPageNumber="409" masterPageNumber="393" pageNumber="401" updateTime="1698850504838" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="14ED4C03F7E2DBA0D5129758F169A49F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="964A166C547E47CD96AE2E8010F891FD">
<mods:title id="E148F50DC67CBBDF869AFDBCB042A58F">On two new deep-water snapping shrimps from the Indo-West Pacific (Decapoda Alpheidae: Alpheus)</mods:title>
</mods:titleInfo>
<mods:name id="90273679FEC78869786ED550547952CA" type="personal">
<mods:role id="F675D3EFA1564804643B7BBF33FDCEA3">
<mods:roleTerm id="E9D9E1376E9460910E5FC82D5FBEE7B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="88E0AFC34B819C8C3FE9EE719A2516C4">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="3620D98EC369C81D417398C351B2EEA9">text</mods:typeOfResource>
<mods:relatedItem id="43004CEF302A97DA450D6887A56D6E30" type="host">
<mods:titleInfo id="E1CBFB018510C347F644F3D9D970D697">
<mods:title id="DA9E13DF1B13A4D1594CAFBA2A1F02B7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5D64529E07BD02B312D0762E5D6C351B">
<mods:date id="78C8880EF8486CCBE536128E48D65648">2020</mods:date>
<mods:detail id="8369CBD6BB505BA6028851676BE2268F" type="pubDate">
<mods:number id="9FBB0A6035AFFB8A602854451940D312">2020-09-03</mods:number>
</mods:detail>
<mods:detail id="7F20E2523F4CADA9D80F609F8E0D1CAB" type="volume">
<mods:number id="E9BCA93F917B521A2BC67BB8E06C7A84">4845</mods:number>
</mods:detail>
<mods:detail id="F7D0E5AAC9C96D5C9886BFAEDBE96637" type="issue">
<mods:number id="F8A38A8A455304856F22F7F877740BB0">3</mods:number>
</mods:detail>
<mods:extent id="BA4EC6DC41F09B5B0DACBE168DCDEB4C" unit="page">
<mods:start id="22BF73EB37CB6FD9A77B9E4F6C9CA687">393</mods:start>
<mods:end id="FF3606552A29EA2B5DF36EF49E73F6AD">409</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BB909299B674740C322E0275ED892452">journal article</mods:classification>
<mods:identifier id="1A330DD181BFFAF708AB032DB8B26DB9" type="CLB-Dataset">8609</mods:identifier>
<mods:identifier id="C69FAB522D86025865C12760066E911C" type="DOI">10.11646/zootaxa.4845.3.5</mods:identifier>
<mods:identifier id="D0495BBC30CBB15A59E36D92F34C57BA" type="GBIF-Dataset">98f82204-695b-437b-95b3-c063bbb6d03b</mods:identifier>
<mods:identifier id="A59B152DEEF3663893E84F1D07B81FAC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3BB2574934DD749C448885CE4D3D9131" type="Zenodo-Dep">4406729</mods:identifier>
<mods:identifier id="56943420FE2414F6F973FBCA08E8D065" type="ZooBank">F00936AE-D1E5-4F0A-A147-28A9CBBA6C85</mods:identifier>
</mods:mods>
<treatment id="8A7487DBFFCEFFDDFF53059BFE886F0E" ID-DOI="http://doi.org/10.5281/zenodo.4407397" ID-GBIF-Taxon="177136758" ID-Zenodo-Dep="4407397" LSID="urn:lsid:plazi:treatment:8A7487DBFFCEFFDDFF53059BFE886F0E" httpUri="http://treatment.plazi.org/id/8A7487DBFFCEFFDDFF53059BFE886F0E" lastPageId="14" lastPageNumber="407" pageId="8" pageNumber="401">
<subSubSection id="4AC76546FFCEFFDBFF53059BFDAB6AA2" box="[151,431,438,465]" pageId="8" pageNumber="401" type="nomenclature">
<paragraph id="026236CDFFCEFFDBFF53059BFDAB6AA2" blockId="8.[151,431,438,501]" box="[151,431,438,465]" pageId="8" pageNumber="401">
<heading id="592A81A1FFCEFFDBFF53059BFDAB6AA2" bold="true" box="[151,431,438,465]" fontSize="11" level="1" pageId="8" pageNumber="401" reason="1">
<emphasis id="30A9EADFFFCEFFDBFF53059BFDAB6AA2" bold="true" box="[151,431,438,465]" pageId="8" pageNumber="401">
<taxonomicName id="C5DD4D4EFFCEFFDBFF53059BFD616AA2" ID-CoL="874GZ" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[151,357,438,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="401" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCEFFDBFF53059BFD616AA2" bold="true" box="[151,357,438,465]" italics="true" pageId="8" pageNumber="401">Alpheus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B9A57A4FFCEFFDBFEA9059AFDAB6AA2" box="[365,431,439,465]" pageId="8" pageNumber="401" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFCEFFDBFF5305F6FD046A86" box="[151,256,474,501]" pageId="8" pageNumber="401" type="description">
<paragraph id="026236CDFFCEFFDBFF5305F6FD046A86" blockId="8.[151,431,438,501]" box="[151,256,474,501]" pageId="8" pageNumber="401">
<figureCitation id="9AE62A48FFCEFFDBFF5305F6FD046A86" box="[151,256,474,501]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="10.[151,250,1864,1888]" captionStartId-1="11.[151,250,1403,1427]" captionStartId-2="12.[151,250,1916,1940]" captionStartId-3="13.[151,250,1836,1860]" captionTargetBox-0="[251,1334,190,1823]" captionTargetBox-1="[190,1392,199,1329]" captionTargetBox-2="[163,1423,354,1882]" captionTargetBox-3="[159,1423,534,1803]" captionTargetId-0="figure-29@10.[243,1344,181,1840]" captionTargetId-1="figure-19@11.[178,1410,181,1379]" captionTargetId-2="figure-120@12.[151,1436,345,1893]" captionTargetId-3="figure-208@13.[151,1436,525,1812]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." captionText-1="FIGURE 6. Alpheus vanuatu n. sp.: a—right (major) cheliped, lateral; b—same, carpus and chela, mesial; c—same, lateral; d—same, distal part of palm and fingers open, lateral; e—same, distal part of merus, mesial; f—dactylus, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu." captionText-2="FIGURE 7. Alpheus vanuatu n. sp.: a—left (minor) male cheliped, lateral; b—same, chela, lateral; c—right (minor) female cheliped, lateral; d—same, chela, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a, b]; paratype female, cl 3.6 mm, MNHN-IU-2017-1955 [c, d], both from Espiritu Santo, Vanuatu." captionText-3="FIGURE 8. Alpheus vanuatu n. sp.: a—second pereiopod, lateral; b—third pereiopod, lateral; c—same distal part of propodus and dactylus, lateral (slender setae omitted); d—fifth pereiopod, lateral; e—same, propodus and dactylus, mesial. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu." figureDoi-0="http://doi.org/10.5281/zenodo.4406739" figureDoi-1="http://doi.org/10.5281/zenodo.4406741" figureDoi-2="http://doi.org/10.5281/zenodo.4406743" figureDoi-3="http://doi.org/10.5281/zenodo.4406745" httpUri-0="https://zenodo.org/record/4406739/files/figure.png" httpUri-1="https://zenodo.org/record/4406741/files/figure.png" httpUri-2="https://zenodo.org/record/4406743/files/figure.png" httpUri-3="https://zenodo.org/record/4406745/files/figure.png" pageId="8" pageNumber="401">Figs. 5–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFCEFFDBFF53060EFDE66866" pageId="8" pageNumber="401" type="materials_examined">
<paragraph id="026236CDFFCEFFDBFF53060EFDE66866" blockId="8.[151,1437,546,2013]" pageId="8" pageNumber="401">
<emphasis id="30A9EADFFFCEFFDBFF53060EFD3A694F" bold="true" box="[151,318,546,573]" pageId="8" pageNumber="401">Type material</emphasis>
.
<materialsCitation id="B2B53C90FFCEFFDBFE89060EFD1D69F6" ID-GBIF-Occurrence="3026668308" collectingDate="2005-10-09" collectionCode="R, V" collectorName="Sta. CP &amp; S. Samadi" country="Vanuatu" county="Campaign" elevation="258" latitude="-15.133333" location="Espiritu Santo Island" longLatPrecision="21" longitude="166.88417" municipality="Big Bay" pageId="8" pageNumber="401" specimenCount="1" typeStatus="holotype">
<typeStatus id="DD66886FFFCEFFDBFE89060EFDBE694E" box="[333,442,547,573]" pageId="8" pageNumber="401" type="holotype">Holotype</typeStatus>
: male, cl 6.0 mm, MNHN-IU-2017-1953,
<collectingCountry id="7ACA765DFFCEFFDBFC51060EFFF0694E" box="[917,1012,547,573]" name="Vanuatu" pageId="8" pageNumber="401">Vanuatu</collectingCountry>
,
<location id="07026016FFCEFFDBFC3B060EF8EA694E" LSID="urn:lsid:plazi:treatment:8A7487DBFFCEFFDDFF53059BFE886F0E:07026016FFCEFFDBFC3B060EF8EA694E" box="[1023,1262,547,573]" country="Vanuatu" county="Campaign" latitude="-15.133333" longLatPrecision="21" longitude="166.88417" municipality="Big Bay" name="Espiritu Santo Island" pageId="8" pageNumber="401">Espiritu Santo Island</location>
, inner
<collectingMunicipality id="E206ACB7FFCEFFDBFAFE060EF99C694E" box="[1338,1432,547,573]" pageId="8" pageNumber="401">Big Bay</collectingMunicipality>
,
<collectingCounty id="EB034E41FFCEFFDBFF53066AFD0E6912" box="[151,266,583,609]" pageId="8" pageNumber="401">Campaign</collectingCounty>
BOA 1,
<collectionCode id="64CCAE08FFCEFFDBFEA8066AFD7B6912" box="[364,383,583,609]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="401">R</collectionCode>
/
<collectionCode id="64CCAE08FFCEFFDBFE43066AFD9E6912" box="[391,410,583,609]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="401" type="Museum">V</collectionCode>
“Alis”,
<collectorName id="AF28531BFFCEFFDBFE31066AFE436912" box="[501,583,583,609]" pageId="8" pageNumber="401">Sta. CP</collectorName>
2445,
<geoCoordinate id="67E9500AFFCEFFDBFD57066BFF0E6912" box="[659,778,582,609]" degrees="15" direction="south" minutes="08" orientation="latitude" pageId="8" pageNumber="401" precision="15" seconds="0" value="-15.133333">15°08’0”S</geoCoordinate>
<geoCoordinate id="67E9500AFFCEFFDBFCD4066BFF9C6912" box="[784,920,582,609]" degrees="166" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="401" precision="15" seconds="3" value="166.88417">166°53’3”E</geoCoordinate>
, depth
<quantity id="C5259B28FFCEFFDBFC23066BF86C6912" box="[999,1128,582,609]" metricMagnitude="2" metricUnit="m" metricValue="2.58" metricValueMax="2.85" metricValueMin="2.31" pageId="8" pageNumber="401" unit="m" value="258.0" valueMax="285.0" valueMin="231.0">
<elevation id="89F0D1FEFFCEFFDBFC23066BF86C6912" box="[999,1128,582,609]" metricMagnitude="2" metricUnit="m" metricValue="2.58" metricValueMax="2.85" metricValueMin="2.31" pageId="8" pageNumber="401" unit="m" value="258.0" valueMax="285.0" valueMin="231.0">231–285 m</elevation>
</quantity>
, leg.
<collectorName id="AF28531BFFCEFFDBFB66066AF9176912" box="[1186,1299,583,609]" pageId="8" pageNumber="401">S. Samadi</collectorName>
/ IRD
<emphasis id="30A9EADFFFCEFFDBFA9A066AF9926912" box="[1374,1430,583,609]" italics="true" pageId="8" pageNumber="401">et al.</emphasis>
,
<date id="7663100DFFCEFFDBFF530647FD1D69F6" box="[151,281,618,645]" pageId="8" pageNumber="401" value="2005-10-09">
<collectingDate id="6627E9E5FFCEFFDBFF530647FD1D69F6" box="[151,281,618,645]" pageId="8" pageNumber="401" value="2005-10-09">10.09.2005</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="B2B53C90FFCEFFDBFEE00646F93969F6" ID-GBIF-Occurrence="3026668306" box="[292,1341,619,645]" collectingDate="2005-10-09" collectionCode="R, V" collectorName="Sta. CP &amp; S. Samadi" country="Vanuatu" county="Campaign" elevation="258" latitude="-15.133333" location="Espiritu Santo Island" longLatPrecision="21" longitude="166.88417" municipality="Big Bay" pageId="8" pageNumber="401" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="DD66886FFFCEFFDBFEE00646FD9169F6" box="[292,405,619,645]" pageId="8" pageNumber="401" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="14DBFD44FFCEFFDBFE650646FE0269F6" box="[417,518,619,645]" pageId="8" pageNumber="401" type="female">1 female</specimenCount>
, cl
<quantity id="C5259B28FFCEFFDBFDEA0646FE8369F6" box="[558,647,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="401" unit="mm" value="4.8">4.8 mm</quantity>
, MNHN-IU-2019-3169, same collection data as for holotype
</materialsCitation>
;
<materialsCitation id="B2B53C90FFCEFFDBFA8D0646F8ED69BE" ID-GBIF-Occurrence="3026668307" collectingDate="2005-10-09" collectionCode="R, V" collectorName="Sta. CP &amp; S. Samadi" country="Vanuatu" county="Campaign" elevation="291" location="Espiritu Santo Island" municipality="Big Bay" pageId="8" pageNumber="401" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="14DBFD44FFCEFFDBFA8D0646F99C69F6" box="[1353,1432,619,645]" pageId="8" pageNumber="401" type="male">1 male</specimenCount>
, cl
<quantity id="C5259B28FFCEFFDBFF7006A2FD0969DA" box="[180,269,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="401" unit="mm" value="4.2">4.2 mm</quantity>
, MNHN-IU-2017-1952,
<collectingCountry id="7ACA765DFFCEFFDBFDEC06A2FE8369DA" box="[552,647,655,681]" name="Vanuatu" pageId="8" pageNumber="401">Vanuatu</collectingCountry>
,
<location id="07026016FFCEFFDBFD5706A2FF8769DA" LSID="urn:lsid:plazi:treatment:8A7487DBFFCEFFDDFF53059BFE886F0E:07026016FFCEFFDBFD5706A2FF8769DA" box="[659,899,655,681]" country="Vanuatu" county="Campaign" municipality="Big Bay" name="Espiritu Santo Island" pageId="8" pageNumber="401">Espiritu Santo Island</location>
, inner
<collectingMunicipality id="E206ACB7FFCEFFDBFC1506A2F83469DA" box="[977,1072,655,681]" pageId="8" pageNumber="401">Big Bay</collectingMunicipality>
,
<collectingCounty id="EB034E41FFCEFFDBFBF906A2F8B469DA" box="[1085,1200,655,681]" pageId="8" pageNumber="401">Campaign</collectingCounty>
BOA 1,
<collectionCode id="64CCAE08FFCEFFDBFAD306A2F92E69DA" box="[1303,1322,655,681]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="401">R</collectionCode>
/
<collectionCode id="64CCAE08FFCEFFDBFAF606A2F94169DA" box="[1330,1349,655,681]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="401" type="Museum">V</collectionCode>
“Alis”,
<collectorName id="AF28531BFFCEFFDBFF53069EFCEE69BE" box="[151,234,691,717]" pageId="8" pageNumber="401">Sta. CP</collectorName>
2444, 15°07’80”S 166°53’70”E, depth
<quantity id="C5259B28FFCEFFDBFD6C069FFF2D69BE" box="[680,809,690,717]" metricMagnitude="2" metricUnit="m" metricValue="2.905" metricValueMax="3.31" metricValueMin="2.5" pageId="8" pageNumber="401" unit="m" value="290.5" valueMax="331.0" valueMin="250.0">
<elevation id="89F0D1FEFFCEFFDBFD6C069FFF2D69BE" box="[680,809,690,717]" metricMagnitude="2" metricUnit="m" metricValue="2.905" metricValueMax="3.31" metricValueMin="2.5" pageId="8" pageNumber="401" unit="m" value="290.5" valueMax="331.0" valueMin="250.0">250–331 m</elevation>
</quantity>
, leg.
<collectorName id="AF28531BFFCEFFDBFCA0069EFFD269BE" box="[868,982,691,717]" pageId="8" pageNumber="401">S. Samadi</collectorName>
/ IRD
<emphasis id="30A9EADFFFCEFFDBFBE6069EF85E69BE" box="[1058,1114,691,717]" italics="true" pageId="8" pageNumber="401">et al.</emphasis>
,
<date id="7663100DFFCEFFDBFBA3069FF8ED69BE" box="[1127,1257,690,717]" pageId="8" pageNumber="401" value="2005-10-09">
<collectingDate id="6627E9E5FFCEFFDBFBA3069FF8ED69BE" box="[1127,1257,690,717]" pageId="8" pageNumber="401" value="2005-10-09">10.09.2005</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="14DBFD44FFCEFFDBFB30069EF94869BE" box="[1268,1356,691,717]" pageId="8" pageNumber="401" type="male">4 males</specimenCount>
, cl 3.5, 3.5, 4.0,
<quantity id="C5259B28FFCEFFDBFF3C06FBFD546982" box="[248,336,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="401" unit="mm" value="4.5">4.5 mm</quantity>
,
<materialsCitation id="B2B53C90FFCEFFDBFE9F06FAFD776866" ID-GBIF-Occurrence="3026668305" collectingDate="2005-10-09" collectionCode="R, V" collectorName="Sta. CP &amp; S. Samadi" country="Vanuatu" county="Campaign" elevation="291" location="Espiritu Santo Island" municipality="Big Bay" pageId="8" pageNumber="401" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="14DBFD44FFCEFFDBFE9F06FAFDCD6982" box="[347,457,727,753]" pageId="8" pageNumber="401" type="female">2 females</specimenCount>
, cl 3.6,
<quantity id="C5259B28FFCEFFDBFDDB06FAFE726982" box="[543,630,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="401" unit="mm" value="3.8">3.8 mm</quantity>
, 1 ovigerous female, cl 4.0 mm, MNHN-IU-2017-1955, same collection data as for previous
</materialsCitation>
<typeStatus id="DD66886FFFCEFFDBFEBE06D6FDDA6866" box="[378,478,763,789]" pageId="8" pageNumber="401" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFCEFFDAFF030733FDE56D7A" lastPageId="9" lastPageNumber="402" pageId="8" pageNumber="401" type="description">
<paragraph id="026236CDFFCEFFDBFF030733F8586F29" blockId="8.[151,1437,546,2013]" pageId="8" pageNumber="401">
<emphasis id="30A9EADFFFCEFFDBFF030733FD55684B" bold="true" box="[199,337,798,824]" pageId="8" pageNumber="401">Description</emphasis>
. Small to medium-sized (present material: cl 3.5–6.0 mm) species of
<taxonomicName id="C5DD4D4EFFCEFFDBFB9A0732F8BC684A" authorityName="Fabricius" authorityYear="1798" box="[1118,1208,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="401" phylum="Arthropoda" rank="genus">
<emphasis id="30A9EADFFFCEFFDBFB9A0732F8BC684A" box="[1118,1208,799,825]" italics="true" pageId="8" pageNumber="401">Alpheus</emphasis>
</taxonomicName>
. Carapace glabrous, neither setose nor pubescent. Rostrum well developed, moderately slender, about twice as long as wide at base, subacute distally, reaching mid-length of first article of antennular peduncle, slightly descendant in lateral view; rostral carina strong, rounded, continuing and noticeably widening and flattening between orbital hoods, abruptly delimited from adrostral furrows, not extending past orbital hoods, disappearing posterior to level of eyes (
<figureCitation id="9AE62A48FFCEFFDBFB080782F93D68B9" box="[1228,1337,942,970]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">Fig. 5a, b</figureCitation>
). Orbital hoods swollen, slightly projecting anteriorly in lateral view, with rounded anterior margin; frontal margin between rostrum and orbital hood very shallowly concave; adrostral furrows shallow, distinct mainly in anterior portion of orbital hoods, between eyes (
<figureCitation id="9AE62A48FFCEFFDBFE190036FE4F6F45" box="[477,587,1050,1078]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">Fig. 5a, b</figureCitation>
). Pterygostomial angle broadly rounded (
<figureCitation id="9AE62A48FFCEFFDBFBDE0036F86A6F46" box="[1050,1134,1050,1077]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">Fig. 5b</figureCitation>
); cardiac notch well developed, deep. Eyes well developed, with large, normally pigmented corneas (
<figureCitation id="9AE62A48FFCEFFDBFC250012F84A6F29" box="[993,1102,1086,1114]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">Fig. 5a, b</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFCEFFDBFF03004EFDA56E42" blockId="8.[151,1437,546,2013]" pageId="8" pageNumber="401">
Telson broad, subrectangular, tapering distally, about 2.0–2.2 times as long as maximal width, with lateral margins slightly concave posterior to telson mid-length; dorsal surface typically with two pairs of stout cuspidate setae both inserted far from lateral margin, first pair at about telson mid-length, second pair at about 0.7–0.8 of telson length, rarely with several cuspidate missing (as in one
<typeStatus id="DD66886FFFCEFFDBFCC000E2FF6E6F9A" box="[772,874,1231,1257]" pageId="8" pageNumber="401" type="paratype">paratype</typeStatus>
); posterior margin broadly rounded; posterolateral angles each with one pair of slender spiniform setae, mesial ones much stouter and almost three times as long as lateral ones (
<figureCitation id="9AE62A48FFCEFFDBFEE3013AFD6C6E42" box="[295,360,1302,1329]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">Fig. 5</figureCitation>
c–e).
</paragraph>
<paragraph id="026236CDFFCEFFDBFF030116FE006DE9" blockId="8.[151,1437,546,2013]" pageId="8" pageNumber="401">
Antennular peduncle moderately stout; stylocerite with basal part moderately broad, slightly swollen laterally, distal part slender, ending in sharp point, latter slightly exceeding distal margin of first article; ventromesial carina with large, anteriorly directed tooth; second article about twice times as long as wide; lateral antennular flagellum with secondary ramus fused to main ramus over most of its length, with numerous groups of aesthetascs distally starting from about eighth subdivision (
<figureCitation id="9AE62A48FFCEFFDBFD9D01E6FEE66E95" box="[601,738,1482,1510]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">Fig. 5a, b, f</figureCitation>
). Antenna with basicerite stout, its distoventral margin armed with anteriorly projecting, slender, sharp tooth; scaphocerite well developed, with straight to faintly concave lateral margin, broad blade and very strong distolateral tooth, latter reaching far beyond distal margin of blade, also typically reaching or slightly overreaching end of antennular peduncle (distolateral tooth of left scaphocerite curved and shorter, slightly abnormal, in illustrated specimen); carpocerite reaching well beyond scaphocerite and end of antennular peduncle (
<figureCitation id="9AE62A48FFCEFFDBFE4D0252FDF36DE9" box="[393,503,1663,1690]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [a–g, i–k]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="8" pageNumber="401">Fig. 1a, b</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFCEFFDBFF03028EFE0F6C02" blockId="8.[151,1437,546,2013]" pageId="8" pageNumber="401">
Mouthparts typical for genus in external observation. Mandible with incisor process bearing about 10 teeth. Third maxilliped relatively stout, moderately setose; coxa with subacutely projecting lateral plate; antepenultimate article flattened ventrolaterally, with distinct ridge running parallel to dorsal margin on lateral surface, mesial margin slightly rugose; penultimate article relatively short, distally widening, at most twice as long as maximal width; ultimate article unarmed distally, with numerous rows of shorter serrulate setae and much longer stiff setae, especially on and near apex (
<figureCitation id="9AE62A48FFCEFFDBFE6E037AFDF96C02" box="[426,509,1878,1905]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="8" pageNumber="401">
Fig.
<quantity id="C5259B28FFCEFFDBFE1B037BFDF96C02" box="[479,509,1878,1905]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="8" pageNumber="401" unit="g" value="5.0">5g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFCEFFDAFF030356FEF36AF9" blockId="8.[151,1437,546,2013]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="402" pageId="8" pageNumber="401">
Major cheliped feebly sexually dimorphic, weaker in females.
<collectingRegion id="C019F82FFFCEFFDBFC5B0356FFDD6CE6" box="[927,985,1915,1941]" country="Maldives" name="Maale" pageId="8" pageNumber="401">Male</collectingRegion>
major cheliped much more robust and longer than minor cheliped; ischium short, stout, smooth, unarmed; merus moderately stout, subtrigonal in crosssection, less than three times as long as wide at base, smooth, distodorsal margin bluntly protruding, without strong tooth, ventromesial margin smooth proximally, faintly rugose distally, with very stout sharp subdistal tooth; carpus very short, cup-shaped, smooth; chela slightly elongate; palm not noticeably compressed, subcylindrical in crosssection, length / height ratio more or less equal to 2.0, without grooves, notches, smooth (no granules), ventral margin faintly concave distally, forming shallow sinus, dorsal margin somewhat convex near mid-length; fingers subequal in length, about half-length of palm, slightly twisted laterally, gaping proximally when closed in lateral view; dactylus distally rounded, plunger somewhat reduced, continuous with dactylar cutting edge anteriorly, abruptly delimited posteriorly; adhesive disks large (
<figureCitation id="9AE62A48FFCFFFDAFD440542FEC76AF9" box="[640,707,367,394]" captionStart="FIGURE 6" captionStartId="11.[151,250,1403,1427]" captionTargetBox="[190,1392,199,1329]" captionTargetId="figure-19@11.[178,1410,181,1379]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus vanuatu n. sp.: a—right (major) cheliped, lateral; b—same, carpus and chela, mesial; c—same, lateral; d—same, distal part of palm and fingers open, lateral; e—same, distal part of merus, mesial; f—dactylus, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406741" httpUri="https://zenodo.org/record/4406741/files/figure.png" pageId="9" pageNumber="402">Fig. 6</figureCitation>
a–f).
</paragraph>
<paragraph id="026236CDFFCFFFDAFF0305BEFE446849" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="402">
Minor cheliped sexually dimorphic, balaeniceps in males, simple in females (
<figureCitation id="9AE62A48FFCFFFDAFBE005BEF8636ADE" box="[1060,1127,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1916,1940]" captionTargetBox="[163,1423,354,1882]" captionTargetId="figure-120@12.[151,1436,345,1893]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus vanuatu n. sp.: a—left (minor) male cheliped, lateral; b—same, chela, lateral; c—right (minor) female cheliped, lateral; d—same, chela, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a, b]; paratype female, cl 3.6 mm, MNHN-IU-2017-1955 [c, d], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406743" httpUri="https://zenodo.org/record/4406743/files/figure.png" pageId="9" pageNumber="402">Fig. 7</figureCitation>
).
<collectingRegion id="C019F82FFFCFFFDAFBB805BEF8B26ADE" box="[1148,1206,403,429]" country="Maldives" name="Maale" pageId="9" pageNumber="402">Male</collectingRegion>
minor cheliped with ischium short, smooth; merus moderately slender, subtrigonal in cross-section, about four times as long as wide at base, smooth, distodorsal margin blunt, ventromesial margin smooth over most of its length, with stout sharp subdistal tooth; carpus short, cup-shaped, smooth; palm slender, subcylindrical in cross-section, smooth, without grooves or notches, length / height ratio about 2.5; fingers about 0.9 length of palm, slightly twisted, gaping when closed, subequal in length, strongly crossing distally; dactylus strongly flattened ventrodorsally, with lateral and mesial margins expanded, strongly curved distally, with two balaeniceps ridges, one on mesial side and one on lateral side, each furnished with densely inserted, thickened, plumose setae; both balaeniceps ridges extending from about 0.4 to 0.7 of dactylar length; mesial and lateral sides of pollex each with row of balaeniceps setae on proximal half; adhesive disks modestly developed (
<figureCitation id="9AE62A48FFCFFFDAFDF706FAFEA56981" box="[563,673,727,754]" captionStart="FIGURE 7" captionStartId="12.[151,250,1916,1940]" captionTargetBox="[163,1423,354,1882]" captionTargetId="figure-120@12.[151,1436,345,1893]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus vanuatu n. sp.: a—left (minor) male cheliped, lateral; b—same, chela, lateral; c—right (minor) female cheliped, lateral; d—same, chela, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a, b]; paratype female, cl 3.6 mm, MNHN-IU-2017-1955 [c, d], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406743" httpUri="https://zenodo.org/record/4406743/files/figure.png" pageId="9" pageNumber="402">Fig. 7a, b</figureCitation>
). Female minor cheliped smaller and more slender than male minor cheliped; chela with fingers slightly longer than palm, not gaping when closed; dactylus not expanded, simple, without balaeniceps setae (
<figureCitation id="9AE62A48FFCFFFDAFE010732FE366849" box="[453,562,799,826]" captionStart="FIGURE 7" captionStartId="12.[151,250,1916,1940]" captionTargetBox="[163,1423,354,1882]" captionTargetId="figure-120@12.[151,1436,345,1893]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus vanuatu n. sp.: a—left (minor) male cheliped, lateral; b—same, chela, lateral; c—right (minor) female cheliped, lateral; d—same, chela, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a, b]; paratype female, cl 3.6 mm, MNHN-IU-2017-1955 [c, d], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406743" httpUri="https://zenodo.org/record/4406743/files/figure.png" pageId="9" pageNumber="402">Fig. 7c, d</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFCFFFDAFF03076EFF106E7E" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="402">
Second pereiopod slender; ischium and merus subequal in length; carpus with five subdivisions, first longest, ratio of carpal subdivisions approximately equal to 3.5/2/1/1/1.6; chela longer than distal-most carpal subdivision (
<figureCitation id="9AE62A48FFCFFFDAFF6407A6FCF068D6" box="[160,244,907,933]" captionStart="FIGURE 8" captionStartId="13.[151,250,1836,1860]" captionTargetBox="[159,1423,534,1803]" captionTargetId="figure-208@13.[151,1436,525,1812]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus vanuatu n. sp.: a—second pereiopod, lateral; b—third pereiopod, lateral; c—same distal part of propodus and dactylus, lateral (slender setae omitted); d—fifth pereiopod, lateral; e—same, propodus and dactylus, mesial. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406745" httpUri="https://zenodo.org/record/4406745/files/figure.png" pageId="9" pageNumber="402">Fig. 8a</figureCitation>
). Third pereiopod moderately slender; ischium with stout cuspidate seta on ventrolateral surface; merus about 6.5 times as long as maximal width, unarmed distoventrally; carpus about half-length of merus, slightly slenderer than merus, unarmed; propodus much longer than carpus, with numerous (some elongate) stiff setae, ventral margin with five or six smaller or larger spiniform setae, in addition to one pair of longer slender spiniform setae near propodo-dactylar articulation; dactylus slightly more than half-length of propodus, faintly curving distally, subconical, noticeably flattened ventrally (
<figureCitation id="9AE62A48FFCFFFDAFDB30012FEE26F2A" box="[631,742,1087,1113]" captionStart="FIGURE 8" captionStartId="13.[151,250,1836,1860]" captionTargetBox="[159,1423,534,1803]" captionTargetId="figure-208@13.[151,1436,525,1812]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus vanuatu n. sp.: a—second pereiopod, lateral; b—third pereiopod, lateral; c—same distal part of propodus and dactylus, lateral (slender setae omitted); d—fifth pereiopod, lateral; e—same, propodus and dactylus, mesial. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406745" httpUri="https://zenodo.org/record/4406745/files/figure.png" pageId="9" pageNumber="402">Fig. 8b, c</figureCitation>
). Fourth pereiopod generally similar to third, somewhat slenderer. Fifth pereiopod much slenderer than third pereiopod; ischium longer than in third pereiopod, unarmed; merus slender, almost seven times as long as wide; carpus slenderer than merus, about 0.8 length of merus; propodus much longer than carpus, distal ventrolateral surface with rows of serrulate setae forming cleaning brush, ventromesial margin with about six slender spiniform setae, including longest one near propodo-dactylar articulation; dactylus similar to that of third and fourth pereiopods (
<figureCitation id="9AE62A48FFCFFFDAFD5D00DEFF026E7E" box="[665,774,1267,1293]" captionStart="FIGURE 8" captionStartId="13.[151,250,1836,1860]" captionTargetBox="[159,1423,534,1803]" captionTargetId="figure-208@13.[151,1436,525,1812]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus vanuatu n. sp.: a—second pereiopod, lateral; b—third pereiopod, lateral; c—same distal part of propodus and dactylus, lateral (slender setae omitted); d—fifth pereiopod, lateral; e—same, propodus and dactylus, mesial. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406745" httpUri="https://zenodo.org/record/4406745/files/figure.png" pageId="9" pageNumber="402">Fig. 8d, e</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFCFFFDAFF03013AFE3E6E96" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="402">
<collectingRegion id="C019F82FFFCFFFDAFF03013AFD056E42" box="[199,257,1303,1329]" country="Maldives" name="Maale" pageId="9" pageNumber="402">Male</collectingRegion>
second pleopod with appendix masculina shorter than appendix interna, with long stiff setae on apex and along margins (
<figureCitation id="9AE62A48FFCFFFDAFE870116FD926E26" box="[323,406,1338,1365]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="9" pageNumber="402">Fig. 5h</figureCitation>
). Uropod with both mesial and lateral lobes of protopod ending in sharp tooth; exopod broad, broadly rounded distally, with stout triangular distolateral tooth; diaeresis sinuous for most part, with two rounded lobes in its lateral section, one of them adjacent to slender distolateral spiniform seta, latter not reaching level of distal margin of exopod; endopod much narrower than exopod, ovate; neither exopod nor endopod with spiniform setae on their distal margins (
<figureCitation id="9AE62A48FFCFFFDAFE2401E6FE296E96" box="[480,557,1482,1509]" captionStart="FIGURE 5" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[251,1334,190,1823]" captionTargetId="figure-29@10.[243,1344,181,1840]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus vanuatu n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="9" pageNumber="402">Fig. 5i</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFCFFFDAFF0301C2FDE56D7A" blockId="9.[151,1437,151,2013]" box="[199,481,1519,1545]" pageId="9" pageNumber="402">Colour pattern unknown.</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFCFFFDAFF03023EFE086D22" pageId="9" pageNumber="402" type="etymology">
<paragraph id="026236CDFFCFFFDAFF03023EFE086D22" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="402">
<emphasis id="30A9EADFFFCFFFDAFF03023EFD4C6D5E" bold="true" box="[199,328,1555,1581]" pageId="9" pageNumber="402">Etymology</emphasis>
. This interesting new species is named after the country of the
<typeStatus id="DD66886FFFCFFFDAFBCA023EF83A6D5E" box="[1038,1086,1555,1581]" pageId="9" pageNumber="402">type</typeStatus>
locality, the
<collectingCountry id="7ACA765DFFCFFFDAFB0B023EFCB96D22" name="Vanuatu" pageId="9" pageNumber="402">Republic of Vanuatu</collectingCountry>
; used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFCFFFDAFF030277FCB06DEA" pageId="9" pageNumber="402" type="distribution">
<paragraph id="026236CDFFCFFFDAFF030277FCB06DEA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="402">
<emphasis id="30A9EADFFFCFFFDAFF030277FD5D6D07" bold="true" box="[199,345,1626,1652]" pageId="9" pageNumber="402">Distribution</emphasis>
. West Pacific: presently known only from
<collectingCountry id="7ACA765DFFCFFFDAFCF90276FF9D6D06" box="[829,921,1627,1653]" name="Vanuatu" pageId="9" pageNumber="402">Vanuatu</collectingCountry>
(Espiritu Santo Island), at depths of
<quantity id="C5259B28FFCFFFDAFAFE0276FCB46DEA" metricMagnitude="2" metricUnit="m" metricValue="2.81" metricValueMax="3.31" metricValueMin="2.31" pageId="9" pageNumber="402" unit="m" value="281.0" valueMax="331.0" valueMin="231.0">
231–
<specimenCount id="14DBFD44FFCFFFDAFAB70276FCB46DEA" pageId="9" pageNumber="402" type="generic">331 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFCFFFDDFF03028EFE886F0E" lastPageId="14" lastPageNumber="407" pageId="9" pageNumber="402" type="discussion">
<paragraph id="026236CDFFCFFFDAFF03028EF98F6CAE" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="402">
<emphasis id="30A9EADFFFCFFFDAFF03028EFD306DCE" bold="true" box="[199,308,1699,1725]" pageId="9" pageNumber="402">Remarks</emphasis>
.
<taxonomicName id="C5DD4D4EFFCFFFDAFE81028EFE006DCF" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[325,516,1699,1725]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCFFFDAFE81028EFE006DCF" box="[325,516,1699,1725]" italics="true" pageId="9" pageNumber="402">Alpheus vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFCFFFDAFDCB028EFE556DCE" bold="true" box="[527,593,1699,1725]" pageId="9" pageNumber="402">
<taxonomicNameLabel id="2B9A57A4FFCFFFDAFDCB028EFE556DCE" box="[527,593,1699,1725]" pageId="9" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
appears to be morphologically closest to the pantropical
<taxonomicName id="C5DD4D4EFFCFFFDAFB2B028EFD246D92" authority="Miers, 1881" authorityName="sensu Miers" authorityYear="1881" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCFFFDAFB2B028EF9986DCF" box="[1263,1436,1699,1725]" italics="true" pageId="9" pageNumber="402">A. paracrinitus</emphasis>
<bibRefCitation id="664C4B3CFFCFFFDAFF5302EAFD246D92" author="Miers, E. J." box="[151,288,1735,1761]" pageId="9" pageNumber="402" pagination="204 - 220" refId="ref10190" refString="Miers, E. J. (1881) On a collection of Crustacea made by Baron Hermann-Maltzan at Goree island, Senegambia. The Annals and Magazine of Natural History, Series 5, 8, 204 - 220 + 259 - 281 + 364 - 377, pls. 13 - 16. https: // doi. org / 10.1080 / 00222938109487471" type="journal article" year="1881">Miers, 1881</bibRefCitation>
</taxonomicName>
species complex, which contains species found mostly in waters shallower than
<quantity id="C5259B28FFCFFFDAFB7202EBF8F46D92" box="[1206,1264,1734,1761]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="402" unit="m" value="50.0">50 m</quantity>
(A. Anker, unpublished data). The assignment of
<taxonomicName id="C5DD4D4EFFCFFFDAFDE102C6FECA6C77" authorityName="sensu Miers" authorityYear="1881" box="[549,718,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCFFFDAFDE102C6FECA6C77" box="[549,718,1771,1797]" italics="true" pageId="9" pageNumber="402">A. paracrinitus</emphasis>
</taxonomicName>
and related taxa to the
<taxonomicName id="C5DD4D4EFFCFFFDAFC1602C6F8DC6C76" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[978,1240,1770,1797]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="30A9EADFFFCFFFDAFC1602C6F8546C76" box="[978,1104,1771,1797]" italics="true" pageId="9" pageNumber="402">A. diadema</emphasis>
<bibRefCitation id="664C4B3CFFCFFFDAFB9C02C6F8DC6C76" author="Dana, J. D." box="[1112,1240,1770,1797]" pageId="9" pageNumber="402" pagination="10 - 29" refId="ref9459" refString="Dana, J. D. (1852) Conspectus Crustaceorum &amp; c. Conspectus of the Crustacea of the Exploring Expedition under Capt. C. Wilkes, U. S. N. Macroura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1852, 10 - 29." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
group dates back to
<bibRefCitation id="664C4B3CFFCFFFDAFF700322FD626C5A" author="Coutiere, H." box="[180,358,1806,1833]" pageId="9" pageNumber="402" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière (1905)</bibRefCitation>
and was followed by most subsequent workers (e.g.
<bibRefCitation id="664C4B3CFFCFFFDAFC7D0322F8326C5A" author="Miya, Y." box="[953,1078,1807,1833]" pageId="9" pageNumber="402" pagination="103 - 195" refId="ref10255" refString="Miya, Y. (1974) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part II. Publications from the Amakusa Marine Biology Laboratory, 3, 103 - 195, pls. 15 - 31." type="journal article" year="1974">Miya 1974</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCFFFDAFB860322F94E6C5A" author="Banner, D. M. &amp; Banner, A. H." box="[1090,1354,1807,1833]" pageId="9" pageNumber="402" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCFFFDAFA920322FCD66C3E" author="Chace, F. A. Jr." pageId="9" pageNumber="402" pagination="1 - 99" refId="ref9190" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCFFFDAFF1B031EFDBE6C3E" author="Kim, W. &amp; Abele, L. G." box="[223,442,1843,1869]" pageId="9" pageNumber="402" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
). However, the
<taxonomicName id="C5DD4D4EFFCFFFDAFDA8031EFEEF6C3E" authorityName="Dana" authorityYear="1852" box="[620,747,1843,1869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="30A9EADFFFCFFFDAFDA8031EFEEF6C3E" box="[620,747,1843,1869]" italics="true" pageId="9" pageNumber="402">A. diadema</emphasis>
</taxonomicName>
group
<emphasis id="30A9EADFFFCFFFDAFCF8031EFF7E6C3F" box="[828,890,1843,1868]" italics="true" pageId="9" pageNumber="402">sensu</emphasis>
<bibRefCitation id="664C4B3CFFCFFFDAFC46031EF8AA6C3E" author="Banner, D. M. &amp; Banner, A. H." box="[898,1198,1843,1869]" pageId="9" pageNumber="402" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner and Banner (1982)</bibRefCitation>
is rather vaguely defined and contains morphologically diverse taxa, including several species that are obviously not closely related to the remaining species traditionally assigned to this group. Therefore, the present author does not recognise the
<taxonomicName id="C5DD4D4EFFCFFFDAFA400356FD256CCB" authorityName="sensu Miers" authorityYear="1881" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCFFFDAFA400356FD256CCB" italics="true" pageId="9" pageNumber="402">A. paracrinitus</emphasis>
</taxonomicName>
complex as part of the
<taxonomicName id="C5DD4D4EFFCFFFDAFDE603B2FEA46CCA" authorityName="Dana" authorityYear="1852" box="[546,672,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="30A9EADFFFCFFFDAFDE603B2FEA46CCA" box="[546,672,1951,1977]" italics="true" pageId="9" pageNumber="402">A. diadema</emphasis>
</taxonomicName>
group. In fact, it seems most reasonable to treat this morphologically well-defined species complex as a separate species group of
<taxonomicName id="C5DD4D4EFFCFFFDAFCFA03EEFF9C6CAE" authorityName="Fabricius" authorityYear="1798" box="[830,920,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="30A9EADFFFCFFFDAFCFA03EEFF9C6CAE" box="[830,920,1987,2013]" italics="true" pageId="9" pageNumber="402">Alpheus</emphasis>
</taxonomicName>
, hereafter the
<taxonomicName id="C5DD4D4EFFCFFFDAFBFE03EEF8E76CAF" authorityName="sensu Miers" authorityYear="1881" box="[1082,1251,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCFFFDAFBFE03EEF8E76CAF" box="[1082,1251,1987,2013]" italics="true" pageId="9" pageNumber="402">A. paracrinitus</emphasis>
</taxonomicName>
species group.
</paragraph>
<caption id="56A26645FFCCFFD9FF530365FE5F6C83" ID-DOI="http://doi.org/10.5281/zenodo.4406739" ID-Zenodo-Dep="4406739" httpUri="https://zenodo.org/record/4406739/files/figure.png" pageId="10" pageNumber="403" startId="10.[151,250,1864,1888]" targetBox="[251,1334,190,1823]" targetPageId="10">
<paragraph id="026236CDFFCCFFD9FF530365FE5F6C83" blockId="10.[151,1437,1864,2033]" pageId="10" pageNumber="403">
<emphasis id="30A9EADFFFCCFFD9FF530365FE106C12" bold="true" box="[151,532,1864,1889]" pageId="10" pageNumber="403">
FIGURE 5.
<taxonomicName id="C5DD4D4EFFCCFFD9FED90365FDD56C13" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[285,465,1864,1888]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="403" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCCFFD9FED90365FDD56C13" bold="true" box="[285,465,1864,1888]" italics="true" pageId="10" pageNumber="403">Alpheus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B9A57A4FFCCFFD9FE1D0364FE106C12" box="[473,532,1865,1889]" pageId="10" pageNumber="403" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: a—frontal region, dorsal; b—same, lateral; c—telson with abnormal number of cuspidate setae, dorsal; d—same, posterior margin, dorsal (setae omitted); e—telson with normal number of cuspidate setae, dorsal; g—third maxilliped, lateral; h—appendix masculina and appendix interna of male second pleopod, anterior (lateral); i—uropod, dorsal. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a–d, f–i]; paratype female, cl 3.6 mm, MNHN-IU-2017- 1955 [e], both from Espiritu Santo, Vanuatu.
</paragraph>
</caption>
<caption id="56A26645FFCDFFD8FF530156FFC36EA9" ID-DOI="http://doi.org/10.5281/zenodo.4406741" ID-Zenodo-Dep="4406741" httpUri="https://zenodo.org/record/4406741/files/figure.png" pageId="11" pageNumber="404" startId="11.[151,250,1403,1427]" targetBox="[190,1392,199,1329]" targetPageId="11">
<paragraph id="026236CDFFCDFFD8FF530156FFC36EA9" blockId="11.[151,1436,1402,1499]" pageId="11" pageNumber="404">
<emphasis id="30A9EADFFFCDFFD8FF530156FE146EE0" bold="true" box="[151,528,1402,1427]" pageId="11" pageNumber="404">
FIGURE 6.
<taxonomicName id="C5DD4D4EFFCDFFD8FEDF0157FDCB6EE1" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[283,463,1402,1426]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCDFFD8FEDF0157FDCB6EE1" bold="true" box="[283,463,1402,1426]" italics="true" pageId="11" pageNumber="404">Alpheus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B9A57A4FFCDFFD8FE120156FE146EE0" box="[470,528,1403,1427]" pageId="11" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: a—right (major) cheliped, lateral; b—same, carpus and chela, mesial; c—same, lateral; d—same, distal part of palm and fingers open, lateral; e—same, distal part of merus, mesial; f—dactylus, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu.
</paragraph>
</caption>
<paragraph id="026236CDFFCDFFDFFF030224FE886A32" blockId="11.[151,1437,1545,2040]" lastBlockId="12.[151,1437,151,321]" lastPageId="12" lastPageNumber="405" pageId="11" pageNumber="404">
The
<taxonomicName id="C5DD4D4EFFCDFFD8FF3F0224FDA16D50" authorityName="sensu Miers" authorityYear="1881" box="[251,421,1545,1571]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCDFFD8FF3F0224FDA16D50" box="[251,421,1545,1571]" italics="true" pageId="11" pageNumber="404">A. paracrinitus</emphasis>
</taxonomicName>
group can be defined by the following combination of morphological characters: (1) rostrum present, often small; (2) rostral carina, if present, not flattened and abruptly delimited from adrostral furrows, rostro-orbital region, including orbital hoods, without teeth; (3) third maxilliped with antepenultimate article often projecting distodorsally; (4) cheliped merus more or less slender, ventromesial margin with strong distal or subdistal tooth; (5) major chela subcylindrical, smooth, without grooves or notches; (6) minor cheliped simple or balaeniceps in males, simple in females; (7) third pereiopod merus lacking distoventral tooth and spiniform setae on ventral margin; and (8) third pereiopod dactylus typically simple, not strongly spatulate or biunguiculate. The following species can be presently assigned to the
<taxonomicName id="C5DD4D4EFFCDFFD8FD9B0328FF0D6C6C" authorityName="sensu Miers" authorityYear="1881" box="[607,777,1797,1823]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCDFFD8FD9B0328FF0D6C6C" box="[607,777,1797,1823]" italics="true" pageId="11" pageNumber="404">A. paracrinitus</emphasis>
</taxonomicName>
group:
<taxonomicName id="C5DD4D4EFFCDFFD8FCA70328F8996C6C" authority="Miers, 1881" authorityName="sensu Miers" authorityYear="1881" box="[867,1181,1797,1823]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCDFFD8FCA70328F8096C6C" box="[867,1037,1797,1823]" italics="true" pageId="11" pageNumber="404">A. paracrinitus</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FBD10328F8996C6C" author="Miers, E. J." box="[1045,1181,1797,1823]" pageId="11" pageNumber="404" pagination="204 - 220" refId="ref10190" refString="Miers, E. J. (1881) On a collection of Crustacea made by Baron Hermann-Maltzan at Goree island, Senegambia. The Annals and Magazine of Natural History, Series 5, 8, 204 - 220 + 259 - 281 + 364 - 377, pls. 13 - 16. https: // doi. org / 10.1080 / 00222938109487471" type="journal article" year="1881">Miers, 1881</bibRefCitation>
</taxonomicName>
(currently regarded as pantropical species, but probably restricted to the East and West Atlantic, A. Anker, unpublished data);
<taxonomicName id="C5DD4D4EFFCDFFD8FADD0304FD776C14" authority="Kim &amp; Abele, 1988" authorityName="Kim &amp; Abele" authorityYear="1988" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="30A9EADFFFCDFFD8FADD0304F9986C30" box="[1305,1436,1833,1859]" italics="true" pageId="11" pageNumber="404">A. rostratus</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FF530360FD776C14" author="Kim, W. &amp; Abele, L. G." box="[151,371,1869,1895]" pageId="11" pageNumber="404" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
</taxonomicName>
(East Pacific);
<taxonomicName id="C5DD4D4EFFCDFFD8FDE40360FF856C1B" authority="Coutiere, 1905" authorityName="Coutiere" authorityYear="1905" box="[544,897,1869,1896]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="alpheopsides">
<emphasis id="30A9EADFFFCDFFD8FDE40360FED56C14" box="[544,721,1869,1895]" italics="true" pageId="11" pageNumber="404">A. alpheopsides</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FD1C0360FF856C1B" author="Coutiere, H." box="[728,897,1869,1896]" pageId="11" pageNumber="404" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière, 1905</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="C5DD4D4EFFCDFFD8FC490360F9116C1B" authority="Coutiere, 1905" authorityName="Coutiere" authorityYear="1905" box="[909,1301,1869,1896]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paralpheopsides">
<emphasis id="30A9EADFFFCDFFD8FC490360F8606C14" box="[909,1124,1869,1895]" italics="true" pageId="11" pageNumber="404">A. paralpheopsides</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FBAF0360F9116C1B" author="Coutiere, H." box="[1131,1301,1869,1896]" pageId="11" pageNumber="404" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière, 1905</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="C5DD4D4EFFCDFFD8FAE40360FD3E6CF8" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="30A9EADFFFCDFFD8FAE40360F9986C14" box="[1312,1436,1869,1895]" italics="true" pageId="11" pageNumber="404">A. tenuipes</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FF53035FFD3E6CF8" author="De Man, J. G." box="[151,314,1905,1932]" pageId="11" pageNumber="404" pagination="287 - 319" refId="ref9676" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="C5DD4D4EFFCDFFD8FE81035CFEA96CF8" authority="Banner &amp; Banner, 1982" authorityName="Banner &amp; Banner" authorityYear="1982" box="[325,685,1905,1932]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFCDFFD8FE81035CFD9E6CF8" box="[325,410,1905,1931]" italics="true" pageId="11" pageNumber="404">A. labis</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FE65035FFEA96CF8" author="Banner, D. M. &amp; Banner, A. H." box="[417,685,1905,1932]" pageId="11" pageNumber="404" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner, 1982</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="C5DD4D4EFFCDFFD8FD7D035CFF9C6CFF" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[697,920,1905,1932]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFCDFFD8FD7D035CFF096CF8" box="[697,781,1905,1931]" italics="true" pageId="11" pageNumber="404">A. mitis</emphasis>
<bibRefCitation id="664C4B3CFFCDFFD8FCD0035FFF9C6CFF" author="Dana, J. D." box="[788,920,1905,1932]" pageId="11" pageNumber="404" pagination="10 - 29" refId="ref9459" refString="Dana, J. D. (1852) Conspectus Crustaceorum &amp; c. Conspectus of the Crustacea of the Exploring Expedition under Capt. C. Wilkes, U. S. N. Macroura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1852, 10 - 29." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="C5DD4D4EFFCDFFD8FC16035CF84E6CF8" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[978,1098,1905,1931]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCDFFD8FC16035CF84E6CF8" box="[978,1098,1905,1931]" italics="true" pageId="11" pageNumber="404">A. vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFCDFFD8FB95035CF8946CF8" bold="true" box="[1105,1168,1905,1931]" pageId="11" pageNumber="404">
<taxonomicNameLabel id="2B9A57A4FFCDFFD8FB95035CF8946CF8" box="[1105,1168,1905,1931]" pageId="11" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(all from the Indo-West Pacific). Two junior synonyms of
<taxonomicName id="C5DD4D4EFFCDFFD8FDD703B8FEB86CDC" authorityName="sensu Miers" authorityYear="1881" box="[531,700,1941,1967]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCDFFD8FDD703B8FEB86CDC" box="[531,700,1941,1967]" italics="true" pageId="11" pageNumber="404">A. paracrinitus</emphasis>
</taxonomicName>
will need to be resurrected, in addition to descriptions of possibly two cryptic or pseudocryptic species in the
<taxonomicName id="C5DD4D4EFFCDFFD8FD540394FF3F6CA0" authorityName="sensu Miers" authorityYear="1881" box="[656,827,1977,2003]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCDFFD8FD540394FF3F6CA0" box="[656,827,1977,2003]" italics="true" pageId="11" pageNumber="404">A. paracrinitus</emphasis>
</taxonomicName>
complex
<emphasis id="30A9EADFFFCDFFD8FC6B0397F83B6CA0" box="[943,1087,1978,2003]" italics="true" pageId="11" pageNumber="404">sensu stricto</emphasis>
. The taxonomic identity of
<taxonomicName id="C5DD4D4EFFCDFFD8FA400394FCC96C84" authorityName="Dana" authorityYear="1852" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFCDFFD8FA400394FCC96C84" italics="true" pageId="11" pageNumber="404">A. mitis</emphasis>
</taxonomicName>
remains problematic (
<bibRefCitation id="664C4B3CFFCDFFD8FE0D03F0FED76C84" author="Banner, A. H. &amp; Banner, D. M." box="[457,723,2013,2040]" pageId="11" pageNumber="404" pagination="1 - 164" refId="ref8845" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
), although the species seems to be distinguishable from both
<taxonomicName id="C5DD4D4EFFCDFFDFFA4003F0FE066BC2" authority="sensu Miers (1881)" authorityName="sensu Miers" authorityYear="1881" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" lastPageId="12" lastPageNumber="405" order="Decapoda" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCDFFDFFA4003F0FD636BC3" italics="true" lastPageId="12" lastPageNumber="405" pageId="11" pageNumber="404">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFCAFFDFFEAB04BAFE066BC2" author="Miers, E. J." box="[367,514,151,177]" pageId="12" pageNumber="405" pagination="204 - 220" refId="ref10190" refString="Miers, E. J. (1881) On a collection of Crustacea made by Baron Hermann-Maltzan at Goree island, Senegambia. The Annals and Magazine of Natural History, Series 5, 8, 204 - 220 + 259 - 281 + 364 - 377, pls. 13 - 16. https: // doi. org / 10.1080 / 00222938109487471" type="journal article" year="1881">Miers (1881)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFCAFFDFFDFE04BAF84A6BC2" authority="sensu Banner &amp; Banner (1982)" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[570,1102,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCAFFDFFDFE04BAFF2E6BC3" box="[570,810,151,177]" italics="true" pageId="12" pageNumber="405">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFCAFFDFFCF604BAF84A6BC2" author="Banner, D. M. &amp; Banner, A. H." box="[818,1102,151,177]" pageId="12" pageNumber="405" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
</taxonomicName>
. The recent record of
<taxonomicName id="C5DD4D4EFFCAFFDFFA8204BAF9986BC3" authorityName="Dana" authorityYear="1852" box="[1350,1436,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFCAFFDFFA8204BAF9986BC3" box="[1350,1436,151,177]" italics="true" pageId="12" pageNumber="405">A. mitis</emphasis>
</taxonomicName>
from
<collectingCountry id="7ACA765DFFCAFFDFFF110496FD066BA6" box="[213,258,187,213]" name="Iran" pageId="12" pageNumber="405">Iran</collectingCountry>
(
<bibRefCitation id="664C4B3CFFCAFFDFFED50496FDED6BA6" author="Ashrafi, H. &amp; Dehghani, A. &amp; Sari, A. &amp; Naderloo, R." box="[273,489,187,213]" pageId="12" pageNumber="405" pagination="521 - 534" refId="ref8601" refString="Ashrafi, H., Dehghani, A., Sari, A. &amp; Naderloo, R. (2020) An updated checklist of caridean shrimps of the Persian Gulf and Gulf of Oman. Zootaxa, 4747 (3), 521 - 534." type="journal article" year="2020">
Ashrafi
<emphasis id="30A9EADFFFCAFFDFFEAB0496FDAC6BA6" box="[367,424,187,213]" italics="true" pageId="12" pageNumber="405">et al.</emphasis>
2020
</bibRefCitation>
) has to be treated with caution. Thus, the
<taxonomicName id="C5DD4D4EFFCAFFDFFC060496F8686BA7" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[962,1132,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCAFFDFFC060496F8686BA7" box="[962,1132,187,213]" italics="true" pageId="12" pageNumber="405">A. paracrinitus</emphasis>
</taxonomicName>
group, when fully revised, may include at least 12 species (A. Anker, under study). Apart from
<taxonomicName id="C5DD4D4EFFCAFFDFFC4804F2F8006B8B" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[908,1028,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCAFFDFFC4804F2F8006B8B" box="[908,1028,223,249]" italics="true" pageId="12" pageNumber="405">A. vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFCAFFDFFBCF04F2F84D6B8A" bold="true" box="[1035,1097,223,249]" pageId="12" pageNumber="405">
<taxonomicNameLabel id="2B9A57A4FFCAFFDFFBCF04F2F84D6B8A" box="[1035,1097,223,249]" pageId="12" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, all the species of the
<taxonomicName id="C5DD4D4EFFCAFFDFFAFB04F2FCE96A6F" authorityName="sensu Banner &amp; Banner" authorityYear="1982" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCAFFDFFAFB04F2FCE96A6F" italics="true" pageId="12" pageNumber="405">A. paracrinitus</emphasis>
</taxonomicName>
group are distributed in shallow waters, although the habitat and depth of the single known specimen of
<taxonomicName id="C5DD4D4EFFCAFFDFFA40052EFCC96A32" authorityName="Banner &amp; Banner" authorityYear="1982" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFCAFFDFFA40052EFCC96A32" italics="true" pageId="12" pageNumber="405">A. labis</emphasis>
</taxonomicName>
are unknown (
<bibRefCitation id="664C4B3CFFCAFFDFFEB1050AFE7B6A32" author="Banner, D. M. &amp; Banner, A. H." box="[373,639,295,321]" pageId="12" pageNumber="405" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
).
</paragraph>
<caption id="56A26645FFCAFFDFFF530351FF9F6CAF" ID-DOI="http://doi.org/10.5281/zenodo.4406743" ID-Zenodo-Dep="4406743" httpUri="https://zenodo.org/record/4406743/files/figure.png" pageId="12" pageNumber="405" startId="12.[151,250,1916,1940]" targetBox="[163,1423,354,1882]" targetPageId="12">
<paragraph id="026236CDFFCAFFDFFF530351FF9F6CAF" blockId="12.[151,1437,1916,2013]" pageId="12" pageNumber="405">
<emphasis id="30A9EADFFFCAFFDFFF530351FE156CE6" bold="true" box="[151,529,1916,1941]" pageId="12" pageNumber="405">
FIGURE 7.
<taxonomicName id="C5DD4D4EFFCAFFDFFED80351FDCB6CE7" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[284,463,1916,1940]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCAFFDFFED80351FDCB6CE7" bold="true" box="[284,463,1916,1940]" italics="true" pageId="12" pageNumber="405">Alpheus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B9A57A4FFCAFFDFFE130350FE156CE6" box="[471,529,1917,1941]" pageId="12" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: a—left (minor) male cheliped, lateral; b—same, chela, lateral; c—right (minor) female cheliped, lateral; d—same, chela, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a, b]; paratype female, cl 3.6 mm, MNHN-IU-2017-1955 [c, d], both from Espiritu Santo, Vanuatu.
</paragraph>
</caption>
<paragraph id="026236CDFFCBFFDEFF0304BAFDC16A86" blockId="13.[151,1437,151,501]" pageId="13" pageNumber="406">
<taxonomicName id="C5DD4D4EFFCBFFDEFF0304BAFD876BC3" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[199,387,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCBFFDEFF0304BAFD876BC3" box="[199,387,151,177]" italics="true" pageId="13" pageNumber="406">Alpheus vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFCBFFDEFE4E04BAFDCD6BC2" bold="true" box="[394,457,151,177]" pageId="13" pageNumber="406">
<taxonomicNameLabel id="2B9A57A4FFCBFFDEFE4E04BAFDCD6BC2" box="[394,457,151,177]" pageId="13" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be easily separated from all of the species listed above by the strongly balaeniceps male major chela, with the dactylus strongly flattened and expanded laterally and mesially (
<figureCitation id="9AE62A48FFCBFFDEFB510496F9046BA5" box="[1173,1280,187,214]" captionStart="FIGURE 7" captionStartId="12.[151,250,1916,1940]" captionTargetBox="[163,1423,354,1882]" captionTargetId="figure-120@12.[151,1436,345,1893]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus vanuatu n. sp.: a—left (minor) male cheliped, lateral; b—same, chela, lateral; c—right (minor) female cheliped, lateral; d—same, chela, lateral. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955 [a, b]; paratype female, cl 3.6 mm, MNHN-IU-2017-1955 [c, d], both from Espiritu Santo, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4406743" httpUri="https://zenodo.org/record/4406743/files/figure.png" pageId="13" pageNumber="406">Fig. 7a, b</figureCitation>
) (cf.
<bibRefCitation id="664C4B3CFFCBFFDEFAF80496FCD66B8A" author="Coutiere, H." pageId="13" pageNumber="406" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière 1905</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCBFFDEFF1904F2FD7C6B8A" author="De Man, J. G." box="[221,376,223,249]" pageId="13" pageNumber="406" pagination="1 - 23" refId="ref9715" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. In: Siboga Expeditie. 39 a 1. Brill, Leiden, pp. 133 - 465; (1915) Supplement-Explanation of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCBFFDEFE4004F2FE8E6B8A" author="Banner, D. M. &amp; Banner, A. H." box="[388,650,223,249]" pageId="13" pageNumber="406" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCBFFDEFD5104F2FF696B8A" author="Kim, W. &amp; Abele, L. G." box="[661,877,223,249]" pageId="13" pageNumber="406" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
). The only species of the
<taxonomicName id="C5DD4D4EFFCBFFDEFB4C04F2F9356B8B" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[1160,1329,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCBFFDEFB4C04F2F9356B8B" box="[1160,1329,223,249]" italics="true" pageId="13" pageNumber="406">A. paracrinitus</emphasis>
</taxonomicName>
group, in which the minor cheliped is feebly balaeniceps, is
<taxonomicName id="C5DD4D4EFFCBFFDEFD10052EF8E56A6E" authority="sensu Banner &amp; Banner (1982" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[724,1249,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCBFFDEFD10052EFFC06A6F" box="[724,964,259,285]" italics="true" pageId="13" pageNumber="406">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFCBFFDEFC08052EF8E56A6E" author="Banner, D. M. &amp; Banner, A. H." box="[972,1249,259,285]" pageId="13" pageNumber="406" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982</bibRefCitation>
</taxonomicName>
: fig. 36d). However, this taxon (currently under study) differs from
<taxonomicName id="C5DD4D4EFFCBFFDEFD21050AFF5B6A33" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[741,863,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCBFFDEFD21050AFF5B6A33" box="[741,863,295,321]" italics="true" pageId="13" pageNumber="406">A. vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFCBFFDEFCA3050AFFA26A32" bold="true" box="[871,934,295,321]" pageId="13" pageNumber="406">
<taxonomicNameLabel id="2B9A57A4FFCBFFDEFCA3050AFFA26A32" box="[871,934,295,321]" pageId="13" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in several other morphological features (see below). The new species also differs from
<taxonomicName id="C5DD4D4EFFCBFFDEFDB20566F8046A16" authority="sensu Miers (1881)" authorityName="sensu Miers" authorityYear="1881" box="[630,1024,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFCBFFDEFDB20566FF616A17" box="[630,869,331,357]" italics="true" pageId="13" pageNumber="406">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFCBFFDEFCA90566F8046A16" author="Miers, E. J." box="[877,1024,331,357]" pageId="13" pageNumber="406" pagination="204 - 220" refId="ref10190" refString="Miers, E. J. (1881) On a collection of Crustacea made by Baron Hermann-Maltzan at Goree island, Senegambia. The Annals and Magazine of Natural History, Series 5, 8, 204 - 220 + 259 - 281 + 364 - 377, pls. 13 - 16. https: // doi. org / 10.1080 / 00222938109487471" type="journal article" year="1881">Miers (1881)</bibRefCitation>
</taxonomicName>
and
<emphasis id="30A9EADFFFCBFFDEFBF30566F8716A17" box="[1079,1141,331,356]" italics="true" pageId="13" pageNumber="406">sensu</emphasis>
<bibRefCitation id="664C4B3CFFCBFFDEFBB90566F9936A16" author="Banner, D. M. &amp; Banner, A. H." box="[1149,1431,331,357]" pageId="13" pageNumber="406" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
, as well as
<taxonomicName id="C5DD4D4EFFCBFFDEFEC30542FDB36AFA" authorityName="Coutiere" authorityYear="1905" box="[263,439,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="alpheopsides">
<emphasis id="30A9EADFFFCBFFDEFEC30542FDB36AFA" box="[263,439,367,393]" italics="true" pageId="13" pageNumber="406">A. alpheopsides</emphasis>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFCBFFDEFE000542FE9E6AFA" authorityName="Coutiere" authorityYear="1905" box="[452,666,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="paralpheopsides">
<emphasis id="30A9EADFFFCBFFDEFE000542FE9E6AFA" box="[452,666,367,393]" italics="true" pageId="13" pageNumber="406">A. paralpheopsides</emphasis>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFCBFFDEFD630542FF2D6AFB" authorityName="Kim &amp; Abele" authorityYear="1988" box="[679,809,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="30A9EADFFFCBFFDEFD630542FF2D6AFB" box="[679,809,367,393]" italics="true" pageId="13" pageNumber="406">A. rostratus</emphasis>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFCBFFDEFCF10542FF8D6AFA" authorityName="Banner &amp; Banner" authorityYear="1982" box="[821,905,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFCBFFDEFCF10542FF8D6AFA" box="[821,905,367,393]" italics="true" pageId="13" pageNumber="406">A. labis</emphasis>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFCBFFDEFC520542FFEE6AFB" authorityName="Dana" authorityYear="1852" box="[918,1002,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFCBFFDEFC520542FFEE6AFB" box="[918,1002,367,393]" italics="true" pageId="13" pageNumber="406">A. mitis</emphasis>
</taxonomicName>
by the presence of a strong, posteriorly extending rostral carina, which is not distinct in the other species (cf.
<bibRefCitation id="664C4B3CFFCBFFDEFC6305BEF84E6ADE" author="Coutiere, H." box="[935,1098,402,429]" pageId="13" pageNumber="406" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière 1905</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCBFFDEFB9205BEF95A6ADE" author="Banner, D. M. &amp; Banner, A. H." box="[1110,1374,403,429]" pageId="13" pageNumber="406" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFCBFFDEFAAE05BEFD3D6AA2" author="Kim, W. &amp; Abele, L. G." pageId="13" pageNumber="406" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
). In
<taxonomicName id="C5DD4D4EFFCBFFDEFEA8059AFDEC6AA3" authorityName="De Man" authorityYear="1910" box="[364,488,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="30A9EADFFFCBFFDEFEA8059AFDEC6AA3" box="[364,488,439,465]" italics="true" pageId="13" pageNumber="406">A. tenuipes</emphasis>
</taxonomicName>
, a rostral carina is present, but is less developed and not extending beyond the level of eyes (cf.
<bibRefCitation id="664C4B3CFFCBFFDEFEDF05F6FDBC6A86" author="De Man, J. G." box="[283,440,475,501]" pageId="13" pageNumber="406" pagination="1 - 23" refId="ref9715" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. In: Siboga Expeditie. 39 a 1. Brill, Leiden, pp. 133 - 465; (1915) Supplement-Explanation of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
).
</paragraph>
<caption id="56A26645FFCBFFDEFF530301FFC36CF8" ID-DOI="http://doi.org/10.5281/zenodo.4406745" ID-Zenodo-Dep="4406745" httpUri="https://zenodo.org/record/4406745/files/figure.png" pageId="13" pageNumber="406" startId="13.[151,250,1836,1860]" targetBox="[159,1423,534,1803]" targetPageId="13">
<paragraph id="026236CDFFCBFFDEFF530301FFC36CF8" blockId="13.[151,1436,1835,1932]" pageId="13" pageNumber="406">
<emphasis id="30A9EADFFFCBFFDEFF530301FE0C6C37" bold="true" box="[151,520,1835,1860]" pageId="13" pageNumber="406">
FIGURE 8.
<taxonomicName id="C5DD4D4EFFCBFFDEFEDC0306FDCE6C30" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[280,458,1835,1859]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFCBFFDEFEDC0306FDCE6C30" bold="true" box="[280,458,1835,1859]" italics="true" pageId="13" pageNumber="406">Alpheus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B9A57A4FFCBFFDEFE0B0301FE0C6C37" box="[463,520,1836,1860]" pageId="13" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: a—second pereiopod, lateral; b—third pereiopod, lateral; c—same distal part of propodus and dactylus, lateral (slender setae omitted); d—fifth pereiopod, lateral; e—same, propodus and dactylus, mesial. Drawn from paratype male, cl 4.5 mm, MNHN-IU-2017-1955, from Espiritu Santo, Vanuatu.
</paragraph>
</caption>
<paragraph id="026236CDFFC8FFDDFF0304BAF8A56F62" blockId="14.[151,1437,151,1149]" pageId="14" pageNumber="407">
In addition to the strong balaeniceps condition of the male minor chela and well-developed rostral carina,
<taxonomicName id="C5DD4D4EFFC8FFDDFA4004BAFCF56BA7" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFC8FFDDFA4004BAFCF56BA7" italics="true" pageId="14" pageNumber="407">A. vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC8FFDDFF330496FD306BA6" bold="true" box="[247,308,187,213]" pageId="14" pageNumber="407">
<taxonomicNameLabel id="2B9A57A4FFC8FFDDFF330496FD306BA6" box="[247,308,187,213]" pageId="14" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs specifically from each of the above-mentioned species, e.g. from
<taxonomicName id="C5DD4D4EFFC8FFDDFB9D0496FD5C6B8A" authority="sensu Banner &amp; Banner (1982)" authorityName="sensu Banner &amp; Banner" authorityYear="1982" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFC8FFDDFB9D0496F9416BA7" box="[1113,1349,187,213]" italics="true" pageId="14" pageNumber="407">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFC8FFDDFA8F0496FD5C6B8A" author="Banner, D. M. &amp; Banner, A. H." pageId="14" pageNumber="407" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
</taxonomicName>
by the major cheliped merus with the tooth on the ventromesial margin in subdistal position (
<emphasis id="30A9EADFFFC8FFDDFABA04F2F9986B8B" box="[1406,1436,223,248]" italics="true" pageId="14" pageNumber="407">vs.</emphasis>
more proximal, closer to merus mid-length in
<taxonomicName id="C5DD4D4EFFC8FFDDFD56052EFF3E6A6F" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[658,826,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFC8FFDDFD56052EFF3E6A6F" box="[658,826,259,285]" italics="true" pageId="14" pageNumber="407">A. paracrinitus</emphasis>
</taxonomicName>
of
<bibRefCitation id="664C4B3CFFC8FFDDFC99052EF8656A6E" author="Banner, D. M. &amp; Banner, A. H." box="[861,1121,259,285]" pageId="14" pageNumber="407" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC8FFDDFB74052EFD286A32" authority="sensu Miers (1881)" authorityName="sensu Miers" authorityYear="1881" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFC8FFDDFB74052EF9986A6F" box="[1200,1436,259,285]" italics="true" pageId="14" pageNumber="407">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFC8FFDDFF53050AFD286A32" author="Miers, E. J." box="[151,300,295,321]" pageId="14" pageNumber="407" pagination="204 - 220" refId="ref10190" refString="Miers, E. J. (1881) On a collection of Crustacea made by Baron Hermann-Maltzan at Goree island, Senegambia. The Annals and Magazine of Natural History, Series 5, 8, 204 - 220 + 259 - 281 + 364 - 377, pls. 13 - 16. https: // doi. org / 10.1080 / 00222938109487471" type="journal article" year="1881">Miers (1881)</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="664C4B3CFFC8FFDDFEF2050AFE4A6A31" author="Crosnier, A. &amp; Forest, J." box="[310,590,295,322]" pageId="14" pageNumber="407" pagination="199 - 314" refId="ref9379" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest (1966)</bibRefCitation>
and
<bibRefCitation id="664C4B3CFFC8FFDDFD46050AFF636A32" author="Kim, W. &amp; Abele, L. G." box="[642,871,295,321]" pageId="14" pageNumber="407" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele (1988)</bibRefCitation>
by the longer stylocerite, exceeding the distal margin of the first article of the antennular peduncle (
<emphasis id="30A9EADFFFC8FFDDFD070566FEDE6A17" box="[707,730,331,356]" italics="true" pageId="14" pageNumber="407">vs</emphasis>
. not reaching it in
<taxonomicName id="C5DD4D4EFFC8FFDDFC690566F8516A17" authorityName="sensu Miers" authorityYear="1881" box="[941,1109,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFC8FFDDFC690566F8516A17" box="[941,1109,331,357]" italics="true" pageId="14" pageNumber="407">A. paracrinitus</emphasis>
</taxonomicName>
) and the third maxilliped not protruding distodorsally (cf.
<bibRefCitation id="664C4B3CFFC8FFDDFE170542FEDE6AFA" author="Crosnier, A. &amp; Forest, J." box="[467,730,367,393]" pageId="14" pageNumber="407" pagination="199 - 314" refId="ref9379" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC8FFDDFD270542FFB36AFA" author="Kim, W. &amp; Abele, L. G." box="[739,951,367,393]" pageId="14" pageNumber="407" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
); from the form described as
<taxonomicName id="C5DD4D4EFFC8FFDDFB310542FDCD6ADE" authority="Coutiere, 1905" authorityName="Coutiere" authorityYear="1905" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="subSpecies" species="paracrinitus" subSpecies="bengalensis">
<emphasis id="30A9EADFFFC8FFDDFB310542FD1F6ADE" italics="true" pageId="14" pageNumber="407">A. paracrinitus bengalensis</emphasis>
<bibRefCitation id="664C4B3CFFC8FFDDFEE605BEFDCD6ADE" author="Coutiere, H." box="[290,457,402,429]" pageId="14" pageNumber="407" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière, 1905</bibRefCitation>
</taxonomicName>
by the longer stylocerite, as well as the proportions of the major chela and telson; from
<taxonomicName id="C5DD4D4EFFC8FFDDFF53059AFD4C6AA2" authorityName="Coutiere" authorityYear="1905" box="[151,328,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="alpheopsides">
<emphasis id="30A9EADFFFC8FFDDFF53059AFD4C6AA2" box="[151,328,439,465]" italics="true" pageId="14" pageNumber="407">A. alpheopsides</emphasis>
</taxonomicName>
by the relatively longer second article of the antennular peduncle, the stronger distolateral tooth of the scaphocerite and the much narrower telson; from
<taxonomicName id="C5DD4D4EFFC8FFDDFD2B05F6FFCC6A86" authorityName="Coutiere" authorityYear="1905" box="[751,968,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paralpheopsides">
<emphasis id="30A9EADFFFC8FFDDFD2B05F6FFCC6A86" box="[751,968,475,501]" italics="true" pageId="14" pageNumber="407">A. paralpheopsides</emphasis>
</taxonomicName>
by the shape of the rostro-orbital margin, especially by the margin between the rostrum and orbital hood not as deeply incised as in
<taxonomicName id="C5DD4D4EFFC8FFDDFB4905D2F961696A" authorityName="Coutiere" authorityYear="1905" box="[1165,1381,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paralpheopsides">
<emphasis id="30A9EADFFFC8FFDDFB4905D2F961696A" box="[1165,1381,511,537]" italics="true" pageId="14" pageNumber="407">A. paralpheopsides</emphasis>
</taxonomicName>
, and by the second pereiopod carpus with the first subdivision much longer than the second (
<emphasis id="30A9EADFFFC8FFDDFBB7060EF88E694F" box="[1139,1162,547,572]" italics="true" pageId="14" pageNumber="407">vs</emphasis>
. the two being subequal in
<taxonomicName id="C5DD4D4EFFC8FFDDFF70066AFD886912" authorityName="Coutiere" authorityYear="1905" box="[180,396,583,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paralpheopsides">
<emphasis id="30A9EADFFFC8FFDDFF70066AFD886912" box="[180,396,583,609]" italics="true" pageId="14" pageNumber="407">A. paralpheopsides</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="664C4B3CFFC8FFDDFE0E066AFE6A6912" author="Coutiere, H." box="[458,622,582,609]" pageId="14" pageNumber="407" pagination="70 - 87" refId="ref9305" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière 1905</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC8FFDDFD7B066AFF476913" authorityName="Kim &amp; Abele" authorityYear="1988" box="[703,835,583,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="30A9EADFFFC8FFDDFD7B066AFF476913" box="[703,835,583,609]" italics="true" pageId="14" pageNumber="407">A. rostratus</emphasis>
</taxonomicName>
by the longer stylocerite, exceeding the distal margin of the first article of the antennular peduncle (
<emphasis id="30A9EADFFFC8FFDDFD570646FEAE69F7" box="[659,682,619,644]" italics="true" pageId="14" pageNumber="407">vs</emphasis>
. not reaching it in
<taxonomicName id="C5DD4D4EFFC8FFDDFCBD0646FFF869F7" authorityName="Kim &amp; Abele" authorityYear="1988" box="[889,1020,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="30A9EADFFFC8FFDDFCBD0646FFF869F7" box="[889,1020,619,645]" italics="true" pageId="14" pageNumber="407">A. rostratus</emphasis>
</taxonomicName>
), the distomesial margin of the major chela palm protruding forward, forming almost a right angle, the major cheliped merus with the tooth on the ventromesial margin in subdistal position (
<emphasis id="30A9EADFFFC8FFDDFD88069EFE6F69BF" box="[588,619,691,716]" italics="true" pageId="14" pageNumber="407">vs.</emphasis>
distal in
<taxonomicName id="C5DD4D4EFFC8FFDDFD15069EFF5069BF" authorityName="Kim &amp; Abele" authorityYear="1988" box="[721,852,691,717]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="30A9EADFFFC8FFDDFD15069EFF5069BF" box="[721,852,691,717]" italics="true" pageId="14" pageNumber="407">A. rostratus</emphasis>
</taxonomicName>
), and the third pereiopod dactylus noticeably longer and slenderer (cf.
<bibRefCitation id="664C4B3CFFC8FFDDFE9A06FAFE316982" author="Kim, W. &amp; Abele, L. G." box="[350,565,727,753]" pageId="14" pageNumber="407" pagination="1 - 119" refId="ref9986" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC8FFDDFD4006FAFEDD6982" authorityName="Banner &amp; Banner" authorityYear="1982" box="[644,729,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFC8FFDDFD4006FAFEDD6982" box="[644,729,727,753]" italics="true" pageId="14" pageNumber="407">A. labis</emphasis>
</taxonomicName>
by the distolateral tooth of the scaphocerite by far overreaching the distal margin of the blade (
<emphasis id="30A9EADFFFC8FFDDFE3406D6FE036867" box="[496,519,763,788]" italics="true" pageId="14" pageNumber="407">vs</emphasis>
. not reaching it in
<taxonomicName id="C5DD4D4EFFC8FFDDFD1806D6FF356866" authorityName="Banner &amp; Banner" authorityYear="1982" box="[732,817,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFC8FFDDFD1806D6FF356866" box="[732,817,763,789]" italics="true" pageId="14" pageNumber="407">A. labis</emphasis>
</taxonomicName>
), the non-expanded antepenultimate article of the third maxilliped (noticeably expanded in
<taxonomicName id="C5DD4D4EFFC8FFDDFDE30732FE7F684A" authorityName="Banner &amp; Banner" authorityYear="1982" box="[551,635,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFC8FFDDFDE30732FE7F684A" box="[551,635,799,825]" italics="true" pageId="14" pageNumber="407">A. labis</emphasis>
</taxonomicName>
), the major chela with a much higher palm length / dactylus length ratio (i.e. with the relatively longer dactylus in the new species), and the major cheliped merus without spiniform setae on the ventromesial margin; from
<taxonomicName id="C5DD4D4EFFC8FFDDFDD1074AFFC268F2" authority="sensu Banner &amp; Banner (1982" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[533,966,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFC8FFDDFDD1074AFEAB68F3" box="[533,687,871,897]" italics="true" pageId="14" pageNumber="407">A. mitis sensu</emphasis>
<bibRefCitation id="664C4B3CFFC8FFDDFD72074AFFC268F2" author="Banner, D. M. &amp; Banner, A. H." box="[694,966,871,897]" pageId="14" pageNumber="407" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982</bibRefCitation>
</taxonomicName>
, but see
<bibRefCitation id="664C4B3CFFC8FFDDFBE2074AF92A68F2" author="Banner, A. H. &amp; Banner, D. M." box="[1062,1326,871,897]" pageId="14" pageNumber="407" pagination="1 - 164" refId="ref8845" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
: p. 55) by the longer stylocerite, exceeding the distal margin of the first article of the antennular peduncle (
<emphasis id="30A9EADFFFC8FFDDFB7907A6F8D068D7" box="[1213,1236,907,932]" italics="true" pageId="14" pageNumber="407">vs</emphasis>
. not reaching it in
<taxonomicName id="C5DD4D4EFFC8FFDDFF530782FCE868BB" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[151,236,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFC8FFDDFF530782FCE868BB" box="[151,236,943,969]" italics="true" pageId="14" pageNumber="407">A. mitis</emphasis>
</taxonomicName>
) and the major cheliped merus less stout and with the tooth on the ventromesial margin clearly in subdistal position (
<emphasis id="30A9EADFFFC8FFDDFEC507FEFD24689F" box="[257,288,979,1004]" italics="true" pageId="14" pageNumber="407">vs.</emphasis>
more distal in
<taxonomicName id="C5DD4D4EFFC8FFDDFE0107FEFE1E689F" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[453,538,979,1005]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="30A9EADFFFC8FFDDFE0107FEFE1E689F" box="[453,538,979,1005]" italics="true" pageId="14" pageNumber="407">A. mitis</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="664C4B3CFFC8FFDDFD9307FEFF5A689E" author="Banner, D. M. &amp; Banner, A. H." box="[599,862,979,1005]" pageId="14" pageNumber="407" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
); and from
<taxonomicName id="C5DD4D4EFFC8FFDDFC1907FEF85D689F" authorityName="De Man" authorityYear="1910" box="[989,1113,979,1005]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="30A9EADFFFC8FFDDFC1907FEF85D689F" box="[989,1113,979,1005]" italics="true" pageId="14" pageNumber="407">A. tenuipes</emphasis>
</taxonomicName>
by the major cheliped merus much stouter and without spiniform setae along the ventromesial margin (cf.
<bibRefCitation id="664C4B3CFFC8FFDDFC3307DAF8906F62" author="De Man, J. G." box="[1015,1172,1015,1041]" pageId="14" pageNumber="407" pagination="1 - 23" refId="ref9715" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. In: Siboga Expeditie. 39 a 1. Brill, Leiden, pp. 133 - 465; (1915) Supplement-Explanation of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
).
</paragraph>
<paragraph id="026236CDFFC8FFDDFF030036FE886F0E" blockId="14.[151,1437,151,1149]" pageId="14" pageNumber="407">
Based on the comparisons above,
<taxonomicName id="C5DD4D4EFFC8FFDDFD820036FEC46F47" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[582,704,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="30A9EADFFFC8FFDDFD820036FEC46F47" box="[582,704,1051,1077]" italics="true" pageId="14" pageNumber="407">A. vanuatu</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC8FFDDFD0C0036FF036F46" bold="true" box="[712,775,1051,1077]" pageId="14" pageNumber="407">
<taxonomicNameLabel id="2B9A57A4FFC8FFDDFD0C0036FF036F46" box="[712,775,1051,1077]" pageId="14" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
does not seem to be very closely related to any of the species of the
<taxonomicName id="C5DD4D4EFFC8FFDDFED00012FDBA6F2B" authorityName="sensu Miers" authorityYear="1881" box="[276,446,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFC8FFDDFED00012FDBA6F2B" box="[276,446,1087,1113]" italics="true" pageId="14" pageNumber="407">A. paracrinitus</emphasis>
</taxonomicName>
group, the morphologically closest taxa being
<taxonomicName id="C5DD4D4EFFC8FFDDFC170012F8556F2B" authorityName="De Man" authorityYear="1910" box="[979,1105,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="30A9EADFFFC8FFDDFC170012F8556F2B" box="[979,1105,1087,1113]" italics="true" pageId="14" pageNumber="407">A. tenuipes</emphasis>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFC8FFDDFB4D0012F8DB6F2A" authorityName="Banner &amp; Banner" authorityYear="1982" box="[1161,1247,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="30A9EADFFFC8FFDDFB4D0012F8DB6F2A" box="[1161,1247,1087,1113]" italics="true" pageId="14" pageNumber="407">A. labis</emphasis>
</taxonomicName>
, and possibly
<taxonomicName id="C5DD4D4EFFC8FFDDFA400012FE8C6F0E" authority="sensu Banner &amp; Banner (1982)" authorityName="sensu Banner &amp; Banner" authorityYear="1982" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="30A9EADFFFC8FFDDFA400012FD636F0F" italics="true" pageId="14" pageNumber="407">A. paracrinitus sensu</emphasis>
<bibRefCitation id="664C4B3CFFC8FFDDFEAA004EFE8C6F0E" author="Banner, D. M. &amp; Banner, A. H." box="[366,648,1123,1149]" pageId="14" pageNumber="407" pagination="1 - 357" refId="ref8927" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink