treatments-xml/data/8A/74/87/8A7487DBFFC7FFDBFF53050AFD7E6A16.xml
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<mods:title id="03E52C8B4AEACB1DB4FBA6A0E0D36F22">On two new deep-water snapping shrimps from the Indo-West Pacific (Decapoda Alpheidae: Alpheus)</mods:title>
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<emphasis id="30A9EADFFFC7FFD2FF53050AFDF16A32" bold="true" box="[151,501,295,322]" pageId="1" pageNumber="394">
<taxonomicName id="C5DD4D4EFFC7FFD2FF53050AFDA86A31" ID-CoL="874LB" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[151,428,295,322]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFC7FFD2FF53050AFDA86A31" bold="true" box="[151,428,295,322]" italics="true" pageId="1" pageNumber="394">Alpheus alaincrosnieri</emphasis>
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<paragraph id="026236CDFFC7FFD2FF530566FD046A16" blockId="1.[151,501,295,357]" box="[151,256,331,357]" pageId="1" pageNumber="394">
<figureCitation id="9AE62A48FFC7FFD2FF530566FD046A16" box="[151,256,331,357]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,250,1868,1892]" captionStartId-1="5.[151,250,1794,1818]" captionStartId-2="6.[151,250,1779,1803]" captionStartId-3="7.[151,250,1735,1759]" captionTargetBox-0="[224,1385,213,1826]" captionTargetBox-1="[161,1433,190,1763]" captionTargetBox-2="[158,1423,194,1745]" captionTargetBox-3="[156,1428,193,1702]" captionTargetId-0="figure-29@4.[184,1404,181,1844]" captionTargetId-1="figure-19@5.[151,1436,181,1770]" captionTargetId-2="figure-29@6.[151,1436,181,1755]" captionTargetId-3="figure-19@7.[151,1436,181,1712]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." captionText-1="FIGURE 2. Alpheus alaincrosnieri n. sp.: a—left (major) cheliped, distal part of merus, carpus and chela, lateral; b—same, mesial; c—same, chela, lateral; d—same, coxa to carpus, lateral; e—same, chela fingers open, mesial (setae omitted); f—same, lateral (setae omitted). Drawn from: holotype male, cl 7.6 mm, MNHN-IU-2011-2230, off Sepik River, Papua New Guinea [a]; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [bf]." captionText-2="FIGURE 3. Alpheus alaincrosnieri n. sp.: a—right (minor) male cheliped, distal part of merus, carpus and chela, lateral; b—left (minor) male cheliped, chela, lateral; c—same, coxa to carpus, lateral; d—same, ischium and merus, ventral; e—left (minor) female cheliped, distal part of merus, carpus and chela, mesial, pollex broken. Drawn from: holotype male, cl 7.6 mm, MNHN-IU-2011-2230 [a]; paratype male, cl 6.5 mm, MNHN-IU-2019-3168 [bf]; paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [e], all off Sepik River, Papua New Guinea." captionText-3="FIGURE 4. Alpheus alaincrosnieri n. sp.: a—second pereiopod, lateral; b—same, distal part of merus, carpus and chela, lateral; c—third pereiopod, lateral; d, e—same, dactylus, ventrolateral and mesial views, respectively (propodal setae omitted in d, all setae omitted in e); f—fourth pereiopod, lateral; g—same, dactylus, lateral (all setae omitted); h—fifth pereiopod, lateral. Drawn from: paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [a]; holotype male, cl 7.6 mm, MNHN-IU-2011- 2230 [b, h], both off Sepik River, Papua New Guinea; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [cf]." figureDoi-0="http://doi.org/10.5281/zenodo.4406731" figureDoi-1="http://doi.org/10.5281/zenodo.4406733" figureDoi-2="http://doi.org/10.5281/zenodo.4406735" figureDoi-3="http://doi.org/10.5281/zenodo.4406737" httpUri-0="https://zenodo.org/record/4406731/files/figure.png" httpUri-1="https://zenodo.org/record/4406733/files/figure.png" httpUri-2="https://zenodo.org/record/4406735/files/figure.png" httpUri-3="https://zenodo.org/record/4406737/files/figure.png" pageId="1" pageNumber="394">Figs. 14</figureCitation>
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<taxonomicName id="C5DD4D4EFFC7FFD2FF5305BDFD836AD4" ID-CoL="85LDD" authorityName="De Man" authorityYear="1909" box="[151,391,400,424]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="30A9EADFFFC7FFD2FF5305BDFD836AD4" box="[151,391,400,424]" italics="true" pageId="1" pageNumber="394">Alpheus acutocarinatus</emphasis>
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<bibRefCitation id="664C4B3CFFC7FFD2FE6505BDFE3B6AD4" author="Chace, F. A. Jr." box="[417,575,399,424]" pageId="1" pageNumber="394" pagination="1 - 99" refId="ref9190" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988: 15</bibRefCitation>
</treatmentCitation>
(part., specimen from
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) [not
<taxonomicName id="C5DD4D4EFFC7FFD2FC7E05BDF9016ADB" ID-CoL="85LDD" authority="De Man 1909" authorityName="De Man" authorityYear="1909" box="[954,1285,400,424]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="30A9EADFFFC7FFD2FC7E05BDF86A6AD4" box="[954,1134,400,423]" italics="true" pageId="1" pageNumber="394">A. acutocarinatus</emphasis>
<bibRefCitation id="664C4B3CFFC7FFD2FBB005BDF9016ADB" author="De Man, J. G." box="[1140,1285,400,424]" pageId="1" pageNumber="394" pagination="99 - 125" refId="ref9637" refString="De Man, J. G. (1909) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandse Dierkundige Vereeniging, 2 (9), 99 - 125." type="journal article" year="1909">De Man 1909</bibRefCitation>
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].
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(?)
<taxonomicName id="C5DD4D4EFFC7FFD2FF7E0583FE086AB5" authority="Miya 1974: 106" authorityName="Miya" authorityPageNumber="106" authorityYear="1974" box="[186,524,430,454]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="30A9EADFFFC7FFD2FF7E0583FD656AB5" box="[186,353,430,454]" italics="true" pageId="1" pageNumber="394">Alpheus miyakei</emphasis>
<bibRefCitation id="664C4B3CFFC7FFD2FEA30583FE086AB5" author="Miya, Y." box="[359,524,430,454]" pageId="1" pageNumber="394" pagination="103 - 195" refId="ref10255" refString="Miya, Y. (1974) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part II. Publications from the Amakusa Marine Biology Laboratory, 3, 103 - 195, pls. 15 - 31." type="journal article" year="1974">Miya 1974: 106</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="664C4B3CFFC7FFD2FDD20582FEB66AB5" author="Bruce, A. J." box="[534,690,430,454]" pageId="1" pageNumber="394" pagination="15 - 28" refId="ref9099" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28." type="journal article" year="1994">Bruce 1994: 23</bibRefCitation>
(key);
<bibRefCitation id="664C4B3CFFC7FFD2FD3F0582F83D6AB5" author="De Grave, S. &amp; Fransen, C. H. J. M." box="[763,1081,430,454]" pageId="1" pageNumber="394" pagination="195 - 589" refId="ref9512" refString="De Grave, S. &amp; Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85, 195 - 589." type="journal article" year="2011">De Grave &amp; Fransen 2011: 476</bibRefCitation>
(listed as
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).
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, off
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BIOPAPUA,
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“Alis”,
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3701,
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, depth
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, leg.
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&amp;
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,
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.
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:
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, cl
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,
<materialsCitation id="B2B53C90FFC7FFD2FDB80610FFD369B0" ID-GBIF-Occurrence="3026668304" collectingDate="2010-10-15" collectionCode="R, V" collectorName="S. Samadi &amp; L. Corbari" country="Papua New Guinea" elevation="220" latitude="-4.65" location="Campaign" longLatPrecision="1306" longitude="153.05" municipality="River" pageId="1" pageNumber="394" specimenCount="2" specimenCount-female="2" stateProvince="New Ireland" typeStatus="paratype">
<specimenCount id="14DBFD44FFC7FFD2FDB80610FEE86925" box="[636,748,572,599]" pageId="1" pageNumber="394" type="female">2 females</specimenCount>
, cl 6.8,
<quantity id="C5259B28FFC7FFD2FC810610FF986924" box="[837,924,573,599]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="394" unit="mm" value="7.2">7.2 mm</quantity>
(missing both P1 and major P1, respectively, both with bopyrid isopod parasites), MNHN-IU-2019-3168, same collection data as for holotype; male, cl 7.0 mm, MNHN-2011-3024,
<collectingCountry id="7ACA765DFFC7FFD2FE7206A8FE9569ED" box="[438,657,644,671]" name="Papua New Guinea" pageId="1" pageNumber="394">Papua New Guinea</collectingCountry>
, off
<collectingRegion id="C019F82FFFC7FFD2FD0606A8FF5469ED" box="[706,848,644,671]" country="Papua New Guinea" name="New Ireland" pageId="1" pageNumber="394">New Ireland</collectingRegion>
,
<location id="07026016FFC7FFD2FC9F06A9FFCA69ED" LSID="urn:lsid:plazi:treatment:8A7487DBFFC7FFDBFF53050AFD7E6A16:07026016FFC7FFD2FC9F06A9FFCA69ED" box="[859,974,644,670]" country="Papua New Guinea" latitude="-4.65" longLatPrecision="1306" longitude="153.05" name="Campaign" pageId="1" pageNumber="394" stateProvince="New Ireland">Campaign</location>
BIOPAPUA,
<collectionCode id="64CCAE08FFC7FFD2FBA806A8F87B69EC" box="[1132,1151,645,671]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="394">R</collectionCode>
/
<collectionCode id="64CCAE08FFC7FFD2FB4306A8F89E69EC" box="[1159,1178,645,671]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="394" type="Museum">V</collectionCode>
“Alis”,
<collectorName id="AF28531BFFC7FFD2FB3306A8F95269EC" box="[1271,1366,645,671]" pageId="1" pageNumber="394">Sta. DW</collectorName>
3764,
<geoCoordinate id="67E9500AFFC7FFD2FF530684FCF769B0" box="[151,243,681,707]" degrees="04" direction="south" minutes="39" orientation="latitude" pageId="1" pageNumber="394" precision="925" value="-4.65">04°39S</geoCoordinate>
<geoCoordinate id="67E9500AFFC7FFD2FF3E0685FD6C69B0" box="[250,360,680,707]" degrees="153" direction="east" minutes="03" orientation="longitude" pageId="1" pageNumber="394" precision="925" value="153.05">153°03E</geoCoordinate>
, depth
<quantity id="C5259B28FFC7FFD2FE7D0684FE0669B0" box="[441,514,681,707]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="394" unit="m" value="220.0">
<elevation id="89F0D1FEFFC7FFD2FE7D0684FE0669B0" box="[441,514,681,707]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="394" unit="m" value="220.0">220 m</elevation>
</quantity>
, leg.
<collectorName id="AF28531BFFC7FFD2FDFA0684FEB469B1" box="[574,688,680,707]" pageId="1" pageNumber="394">S. Samadi</collectorName>
&amp;
<collectorName id="AF28531BFFC7FFD2FD170684FF4F69B1" box="[723,843,680,707]" pageId="1" pageNumber="394">L. Corbari</collectorName>
,
<date id="7663100DFFC7FFD2FC920685FFD369B0" box="[854,983,680,707]" pageId="1" pageNumber="394" value="2010-10-15">
<collectingDate id="6627E9E5FFC7FFD2FC920685FFD369B0" box="[854,983,680,707]" pageId="1" pageNumber="394" value="2010-10-15">15.10.2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="026236CDFFC7FFD2FF0306E1FFFC6804" blockId="1.[151,1437,500,2039]" pageId="1" pageNumber="394">
<emphasis id="30A9EADFFFC7FFD2FF0306E1FDB76995" bold="true" box="[199,435,716,742]" pageId="1" pageNumber="394">Additional material</emphasis>
.
<materialsCitation id="B2B53C90FFC7FFD2FE0406E0FF21685C" ID-GBIF-Occurrence="3026668301" collectingDate="2005-10-09" collectionCode="R, V" collectorName="Sta. CP &amp; S. Samadi" country="Vanuatu" county="Campaign" elevation="342" latitude="-15.116667" location="Espiritu Santo Island" municipality="Big Bay" pageId="1" pageNumber="394" specimenCount="1" specimenCount-male="1">
<specimenCount id="14DBFD44FFC7FFD2FE0406E0FE0A6995" box="[448,526,716,743]" pageId="1" pageNumber="394" type="male">1 male</specimenCount>
, cl
<quantity id="C5259B28FFC7FFD2FDF006E0FE8E6994" box="[564,650,717,743]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="394" unit="mm" value="7.4">7.4 mm</quantity>
(missing most legs, including both P1), MNHN-IU-2017-1947,
<collectingCountry id="7ACA765DFFC7FFD2FA9D06E0FCB86878" name="Vanuatu" pageId="1" pageNumber="394">Vanuatu</collectingCountry>
,
<location id="07026016FFC7FFD2FF0C06DDFDB36879" LSID="urn:lsid:plazi:treatment:8A7487DBFFC7FFDBFF53050AFD7E6A16:07026016FFC7FFD2FF0C06DDFDB36879" box="[200,439,752,779]" country="Vanuatu" county="Campaign" latitude="-15.116667" municipality="Big Bay" name="Espiritu Santo Island" pageId="1" pageNumber="394">Espiritu Santo Island</location>
, inner
<collectingMunicipality id="E206ACB7FFC7FFD2FDC006DDFE666878" box="[516,610,752,779]" pageId="1" pageNumber="394">Big Bay</collectingMunicipality>
,
<collectingCounty id="EB034E41FFC7FFD2FDAA06DDFEE56879" box="[622,737,752,778]" pageId="1" pageNumber="394">Campaign</collectingCounty>
BOA 1,
<collectionCode id="64CCAE08FFC7FFD2FC8306DCFF5E6878" box="[839,858,753,779]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="394">R</collectionCode>
/
<collectionCode id="64CCAE08FFC7FFD2FCA606DCFF716878" box="[866,885,753,779]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="394" type="Museum">V</collectionCode>
“Alis”,
<collectorName id="AF28531BFFC7FFD2FC1706DCF8226878" box="[979,1062,753,779]" pageId="1" pageNumber="394">Sta. CP</collectorName>
2448,
<geoCoordinate id="67E9500AFFC7FFD2FBB106DDF8FD6878" box="[1141,1273,752,779]" degrees="15" direction="south" minutes="06" orientation="latitude" pageId="1" pageNumber="394" precision="15" seconds="60" value="-15.116667">15°0660”S</geoCoordinate>
<geoCoordinate id="67E9500AFFC7FFD2FAC506DDF9936878" box="[1281,1431,752,779]" pageId="1" pageNumber="394">166°5080”E</geoCoordinate>
, depth
<quantity id="C5259B28FFC7FFD2FF190738FD5A685C" box="[221,350,789,815]" metricMagnitude="2" metricUnit="m" metricValue="3.42" metricValueMax="3.87" metricValueMin="2.9699999999999998" pageId="1" pageNumber="394" unit="m" value="342.0" valueMax="387.0" valueMin="297.0">
<elevation id="89F0D1FEFFC7FFD2FF190738FD5A685C" box="[221,350,789,815]" metricMagnitude="2" metricUnit="m" metricValue="3.42" metricValueMax="3.87" metricValueMin="2.9699999999999998" pageId="1" pageNumber="394" unit="m" value="342.0" valueMax="387.0" valueMin="297.0">297387 m</elevation>
</quantity>
, leg.
<collectorName id="AF28531BFFC7FFD2FE5F0738FE09685D" box="[411,525,788,815]" pageId="1" pageNumber="394">S. Samadi</collectorName>
/ IRD
<emphasis id="30A9EADFFFC7FFD2FD980738FE91685D" box="[604,661,788,814]" italics="true" pageId="1" pageNumber="394">et al.</emphasis>
,
<date id="7663100DFFC7FFD2FD670739FF21685C" box="[675,805,788,815]" pageId="1" pageNumber="394" value="2005-10-09">
<collectingDate id="6627E9E5FFC7FFD2FD670739FF21685C" box="[675,805,788,815]" pageId="1" pageNumber="394" value="2005-10-09">10.09.2005</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="B2B53C90FFC7FFD2FCF40738FFF16804" ID-GBIF-Occurrence="3026668303" collectingDate="2009-07-14" collectorName="Sta. CP &amp; P. Bouchet" country="Madagascar" county="Campaign" elevation="338" latitude="-14.5" location="Channel" longLatPrecision="1288" longitude="47.433334" municipality="Bay" pageId="1" pageNumber="394" specimenCount="1">
1 ovigerous female, cl
<quantity id="C5259B28FFC7FFD2FBF60739F88E685C" box="[1074,1162,788,815]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="394" unit="mm" value="8.5">8.5 mm</quantity>
, MNHN-IU-2010-1278,
<collectingCountry id="7ACA765DFFC7FFD2FF530715FD246821" box="[151,288,824,850]" name="Madagascar" pageId="1" pageNumber="394">Madagascar</collectingCountry>
, northwest coast,
<collectingCountry id="7ACA765DFFC7FFD2FE2A0715FE866821" box="[494,642,824,850]" name="Mozambique" pageId="1" pageNumber="394">Mozambique</collectingCountry>
<location id="07026016FFC7FFD2FD4E0715FEE36821" LSID="urn:lsid:plazi:treatment:8A7487DBFFC7FFDBFF53050AFD7E6A16:07026016FFC7FFD2FD4E0715FEE36821" box="[650,743,824,850]" country="Madagascar" county="Campaign" latitude="-14.5" longLatPrecision="1288" longitude="47.433334" municipality="Bay" name="Channel" pageId="1" pageNumber="394">Channel</location>
off
<location id="07026016FFC7FFD2FCDD0715FF776821" LSID="urn:lsid:plazi:treatment:8A7487DBFFC7FFDBFF53050AFD7E6A16:07026016FFC7FFD2FCDD0715FF776821" box="[793,883,824,850]" country="Madagascar" county="Campaign" latitude="-14.5" longLatPrecision="1288" longitude="47.433334" municipality="Bay" name="Narinda" pageId="1" pageNumber="394">Narinda</location>
(“Nazendry”)
<collectingMunicipality id="E206ACB7FFC7FFD2FBDA0714F8486820" box="[1054,1100,825,851]" pageId="1" pageNumber="394">Bay</collectingMunicipality>
,
<collectingCounty id="EB034E41FFC7FFD2FB9D0715F8C86821" box="[1113,1228,824,850]" pageId="1" pageNumber="394">Campaign</collectingCounty>
MIRIKY,
<collectorName id="AF28531BFFC7FFD2FA830714F9986820" box="[1351,1436,825,851]" pageId="1" pageNumber="394">Sta. CP</collectorName>
3293,
<geoCoordinate id="67E9500AFFC7FFD2FF190770FD3E6804" box="[221,314,861,887]" degrees="14" direction="south" minutes="30" orientation="latitude" pageId="1" pageNumber="394" precision="925" value="-14.5">14°30S</geoCoordinate>
<geoCoordinate id="67E9500AFFC7FFD2FE840771FDA46805" box="[320,416,860,886]" degrees="47" direction="east" minutes="26" orientation="longitude" pageId="1" pageNumber="394" precision="925" value="47.433334">47°26E</geoCoordinate>
, depth
<quantity id="C5259B28FFC7FFD2FE350771FE766804" box="[497,626,860,887]" metricMagnitude="2" metricUnit="m" metricValue="3.38" metricValueMax="4.08" metricValueMin="2.68" pageId="1" pageNumber="394" unit="m" value="338.0" valueMax="408.0" valueMin="268.0">
<elevation id="89F0D1FEFFC7FFD2FE350771FE766804" box="[497,626,860,887]" metricMagnitude="2" metricUnit="m" metricValue="3.38" metricValueMax="4.08" metricValueMin="2.68" pageId="1" pageNumber="394" unit="m" value="338.0" valueMax="408.0" valueMin="268.0">268408 m</elevation>
</quantity>
, leg.
<collectorName id="AF28531BFFC7FFD2FD6A0770FF226805" box="[686,806,860,887]" pageId="1" pageNumber="394">P. Bouchet</collectorName>
<emphasis id="30A9EADFFFC7FFD2FCE90770FF5A6805" box="[813,862,860,886]" italics="true" pageId="1" pageNumber="394">et al</emphasis>
.,
<date id="7663100DFFC7FFD2FCB70770FFF16804" box="[883,1013,861,887]" pageId="1" pageNumber="394" value="2009-07-14">
<collectingDate id="6627E9E5FFC7FFD2FCB70770FFF16804" box="[883,1013,861,887]" pageId="1" pageNumber="394" value="2009-07-14">14.07.2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFC7FFD0FF0307ADFE4A6A6E" lastPageId="3" lastPageNumber="396" pageId="1" pageNumber="394" type="description">
<paragraph id="026236CDFFC7FFD2FF0307ADF9686FC8" blockId="1.[151,1437,500,2039]" pageId="1" pageNumber="394">
<emphasis id="30A9EADFFFC7FFD2FF0307ADFD5568E9" bold="true" box="[199,337,896,922]" pageId="1" pageNumber="394">Description</emphasis>
. Medium-sized (present material: cl
<quantity id="C5259B28FFC7FFD2FD2B07ADFF7368E8" box="[751,887,896,923]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="1" pageNumber="394" unit="mm" value="7.5" valueMax="8.5" valueMin="6.5">6.58.5 mm</quantity>
) species of
<taxonomicName id="C5DD4D4EFFC7FFD2FC3807ADF8DA68E9" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[1020,1246,896,922]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFC7FFD2FC3807ADF8DA68E9" box="[1020,1246,896,922]" italics="true" pageId="1" pageNumber="394">Alpheus brevirostris</emphasis>
</taxonomicName>
group. Carapace glabrous, neither setose nor pubescent. Rostrum well developed, slender, about twice as long as wide at base, subacute distally, reaching mid-length of first article of antennular peduncle, somewhat ascendant in lateral view; rostral carina strong, sharp, continuing on carapace well past orbital hoods, ending in conspicuous post-rostral tubercle in form of anteriorly elevated, gradually flattening tooth (
<figureCitation id="9AE62A48FFC7FFD2FCE7003DFF976F58" box="[803,915,1040,1067]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1a, b</figureCitation>
). Orbital hoods swollen, somewhat projecting anteriorly in lateral view; anterior margin of each orbital hood armed with small sharp tooth; frontal margin between rostrum and orbital hood shallowly concave; adrostral furrows rather shallow, distinct mainly in anterior portion of orbital hoods, between eyes (
<figureCitation id="9AE62A48FFC7FFD2FE250051FE546FE4" box="[481,592,1148,1175]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1a, b</figureCitation>
). Pterygostomial angle protruding anteriorly as stout sharp tooth (
<figureCitation id="9AE62A48FFC7FFD2FAFE0051F9886FE4" box="[1338,1420,1148,1175]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1b</figureCitation>
); cardiac notch well developed, deep. Eyes well developed, with large, normally pigmented corneas (
<figureCitation id="9AE62A48FFC7FFD2FB35008DF95B6FC8" box="[1265,1375,1184,1211]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1a, b</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC7FFD2FF0300E9F9896E1C" blockId="1.[151,1437,500,2039]" pageId="1" pageNumber="394">
Telson narrow, subrectangular, tapering distally, about 2.7 times as long as maximal width, with lateral margins distinctly constricted at about telson mid-length; dorsal surface with two pairs of stout cuspidate setae both inserted at some distance from lateral margin, first pair at about 0.4 telson length, second pair at about 0.7 of telson length; posterior margin broadly rounded, with row of shorter slender setae above long plumose setae; posterolateral angles each with one pair of spiniform setae, mesial ones stouter and more than twice as long as lateral ones (
<figureCitation id="9AE62A48FFC7FFD2FAD60179F9846E1C" box="[1298,1408,1364,1391]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1c, d</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC7FFD2FF030155FE266DC0" blockId="1.[151,1437,500,2039]" pageId="1" pageNumber="394">
Antennular peduncle elongate, slender; stylocerite with basal part broad, swollen laterally, distal part slender, ending in sharp point, latter falling short of distal margin of first article; ventromesial carina with large tooth (not illustrated but visible in
<figureCitation id="9AE62A48FFC7FFD2FE6E01EDFDFF6EA8" box="[426,507,1472,1499]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1b</figureCitation>
); second article elongate, almost five times as long as wide; lateral antennular flagellum with secondary ramus fused to main ramus over most of its length, with numerous groups of aesthetascs distally, starting from about 14th subdivision (
<figureCitation id="9AE62A48FFC7FFD2FD6B0225FF446D50" box="[687,832,1544,1571]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1a, b, e</figureCitation>
). Antenna with basicerite very stout, its distoventral margin armed with very large, anteriorly projecting, sharp tooth; scaphocerite well developed, with straight lateral margin, moderately broad blade and strong distolateral tooth, latter reaching far beyond distal margin of blade, but not reaching end of antennular peduncle; carpocerite reaching slightly beyond scaphocerite, also not reaching end of antennular peduncle (
<figureCitation id="9AE62A48FFC7FFD2FE6302B5FE116DC0" box="[423,533,1688,1715]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1a, b</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC7FFD2FF030291F87B6C84" blockId="1.[151,1437,500,2039]" pageId="1" pageNumber="394">
Mouthparts generally typical for genus in external observation. Mandible with incisor process bearing nine teeth. First maxilliped with well-developed caridean lobe and endopod subdivided into two articles (
<figureCitation id="9AE62A48FFC7FFD2FB2D02CDF9326D88" box="[1257,1334,1760,1787]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1f</figureCitation>
). Second maxilliped with third article of endopod armed with small sharp tooth on mesial (ventral) margin (
<figureCitation id="9AE62A48FFC7FFD2FB180329F94F6C6C" box="[1244,1355,1796,1823]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">
Fig.
<quantity id="C5259B28FFC7FFD2FAD40328F92A6C6C" box="[1296,1326,1797,1823]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="1" pageNumber="394" unit="g" value="1.0">1g</quantity>
, h
</figureCitation>
). Third maxilliped slender, elongate, very setose; coxa with bluntly projecting lateral plate; antepenultimate article flattened ventrolaterally, with distinct ridge running parallel to dorsal margin on lateral surface, mesial margin slightly rugose; penultimate article relatively long, about 3.5 times as long as wide proximally, distoventral surface noticeably bulging distally, strongly convex, furnished with very long stiff setae; ultimate article unarmed distally, with numerous rows of shorter serrulate setae and much longer stiff setae, some of more apically situated setae extremely elongate, much longer than ultimate article; arthrobranch moderately developed (
<figureCitation id="9AE62A48FFC7FFD2FBE003F1F86A6C84" box="[1060,1134,2012,2039]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="1" pageNumber="394">Fig. 1i</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC4FFD1FF0304BAFF8269BD" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="395">
Major cheliped not sexually dimorphic, slightly weaker in smaller females.
<collectingRegion id="C019F82FFFC4FFD1FBF904BAF8736BC2" box="[1085,1143,151,177]" country="Maldives" name="Maale" pageId="2" pageNumber="395">Male</collectingRegion>
major cheliped more robust, but only slightly longer than minor cheliped; ischium short, stout, ventrolateral surface with small granules, ventromesial margin with at least two elevations each carrying slender spiniform seta; merus elongate, trigonal in cross-section, about four times as long as wide at base, patchily covered by small granules, distodorsal margin blunt, without distal or subdistal tooth, ventromesial margin rugose, armed with two or three slender spiniform setae in proximal third, with stout sharp distal tooth; carpus subcylindrical, short, distally widening, cup-shaped in larger individuals, with more or less protruding blunt distodorsal process; chela extremely elongate, slender; palm strongly compressed, subrectangular in cross-section, length / height ratio ranging from about
<quantity id="C5259B28FFC4FFD1FB0005BEF9016ADE" box="[1220,1285,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.0668" pageId="2" pageNumber="395" unit="in" value="4.2">4.2 in</quantity>
smaller individuals to over 5.0 in larger individuals, ventral margin broadly concave distally, forming shallow sinus, without deep constriction, dorsal margin without transverse groove or notch subdistally, large sections of palmar surface covered by small granules; fingers unequal in length, with dactylus noticeably shorter than pollex, about 0.250.3 times (dactylus) to 0.30.35 times (pollex) length of palm, not twisted or significantly deviating from chela axis, largely gaping proximally when closed; dactylus distally rounded, plunger highly reduced, continuous with dactylar cutting edge anteriorly, abruptly delimited posteriorly; adhesive disks small; distodorsal surface of merus, ventral and dorsal margins of palm, and ventral margin of pollex fringed with long setae, those of pollex projecting forward, often forming conspicuous tuft of several parallel setae (
<figureCitation id="9AE62A48FFC4FFD1FCCB069EFF5569BD" box="[783,849,691,718]" captionStart="FIGURE 2" captionStartId="5.[151,250,1794,1818]" captionTargetBox="[161,1433,190,1763]" captionTargetId="figure-19@5.[151,1436,181,1770]" captionTargetPageId="5" captionText="FIGURE 2. Alpheus alaincrosnieri n. sp.: a—left (major) cheliped, distal part of merus, carpus and chela, lateral; b—same, mesial; c—same, chela, lateral; d—same, coxa to carpus, lateral; e—same, chela fingers open, mesial (setae omitted); f—same, lateral (setae omitted). Drawn from: holotype male, cl 7.6 mm, MNHN-IU-2011-2230, off Sepik River, Papua New Guinea [a]; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [bf]." figureDoi="http://doi.org/10.5281/zenodo.4406733" httpUri="https://zenodo.org/record/4406733/files/figure.png" pageId="2" pageNumber="395">Fig. 2</figureCitation>
af).
</paragraph>
<paragraph id="026236CDFFC4FFD1FF0306FAF90E6EEE" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="395">
Minor cheliped sexually dimorphic, balaeniceps in males, simple in females (
<figureCitation id="9AE62A48FFC4FFD1FBE006FAF8636982" box="[1060,1127,727,753]" captionStart="FIGURE 3" captionStartId="6.[151,250,1779,1803]" captionTargetBox="[158,1423,194,1745]" captionTargetId="figure-29@6.[151,1436,181,1755]" captionTargetPageId="6" captionText="FIGURE 3. Alpheus alaincrosnieri n. sp.: a—right (minor) male cheliped, distal part of merus, carpus and chela, lateral; b—left (minor) male cheliped, chela, lateral; c—same, coxa to carpus, lateral; d—same, ischium and merus, ventral; e—left (minor) female cheliped, distal part of merus, carpus and chela, mesial, pollex broken. Drawn from: holotype male, cl 7.6 mm, MNHN-IU-2011-2230 [a]; paratype male, cl 6.5 mm, MNHN-IU-2019-3168 [bf]; paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [e], all off Sepik River, Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.4406735" httpUri="https://zenodo.org/record/4406735/files/figure.png" pageId="2" pageNumber="395">Fig. 3</figureCitation>
).
<collectingRegion id="C019F82FFFC4FFD1FBB806FAF8B26982" box="[1148,1206,727,753]" country="Maldives" name="Maale" pageId="2" pageNumber="395">Male</collectingRegion>
minor cheliped with ischium short, stout, its ventromesial margin rugose, with at least two slender spiniform setae; merus elongate, trigonal in cross-section, about 4.5 times as long as wide at base, patchily covered by small granules, some organised in row on ventral surface, especially in larger individuals, distodorsal margin blunt, without distal or subdistal tooth, ventromesial margin rugose, armed with four or so slender spiniform setae in proximal third, with stout sharp distal tooth; carpus subcylindrical, short, distally widening, cup-shaped in larger individuals, with protruding blunt distodorsal process, dorsal and ventral surfaces more or less rugose or covered with small granules; palm compressed, subrectangular in cross-section, without grooves or notches, length / height ratio ranging from about
<quantity id="C5259B28FFC4FFD1FAC707FFF947689E" box="[1283,1347,978,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" pageId="2" pageNumber="395" unit="in" value="5.5">5.5 in</quantity>
smaller individuals to over
<quantity id="C5259B28FFC4FFD1FEAA07DBFDAA6F62" box="[366,430,1014,1041]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" pageId="2" pageNumber="395" unit="in" value="6.5">6.5 in</quantity>
larger individuals, surface slightly rugose in smaller individuals, covered with small granules in addition to rugosities in larger individuals; fingers about 0.60.7 length of palm, not twisted, slightly gaping when closed, subequal in length, strongly crossing distally; dactylus more or less flattened dorsoventrally and with margins slightly broadened, strongly bent distally, with two balaeniceps ridges, one on mesial side and one on lateral side, each furnished with densely inserted, thickened, plumose setae; both balaeniceps ridges extending from about half-length of dactylus to point of its curvature; pollex more gradually curving distally, without balaeniceps setae on either side; proximal surface of both dactylus and pollex covered with small granules; adhesive disks reduced; dorsal margin of palm, and ventral margin of pollex fringed with long setae, those of pollex projecting forward, often as conspicuous tuft of several parallel setae (
<figureCitation id="9AE62A48FFC4FFD1FD03013AFF0C6E41" box="[711,776,1303,1330]" captionStart="FIGURE 3" captionStartId="6.[151,250,1779,1803]" captionTargetBox="[158,1423,194,1745]" captionTargetId="figure-29@6.[151,1436,181,1755]" captionTargetPageId="6" captionText="FIGURE 3. Alpheus alaincrosnieri n. sp.: a—right (minor) male cheliped, distal part of merus, carpus and chela, lateral; b—left (minor) male cheliped, chela, lateral; c—same, coxa to carpus, lateral; d—same, ischium and merus, ventral; e—left (minor) female cheliped, distal part of merus, carpus and chela, mesial, pollex broken. Drawn from: holotype male, cl 7.6 mm, MNHN-IU-2011-2230 [a]; paratype male, cl 6.5 mm, MNHN-IU-2019-3168 [bf]; paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [e], all off Sepik River, Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.4406735" httpUri="https://zenodo.org/record/4406735/files/figure.png" pageId="2" pageNumber="395">Fig. 3</figureCitation>
ad). Female minor cheliped more slender than male minor cheliped; carpus longer, vase-shaped, with rugosities or granules and elongate setae; chela with fingers about 0.8 length of palm, slightly gaping; dactylus not expanded, simple, without balaeniceps setae; ventral and dorsal margins of palm, ventral margin of pollex and dorsal margin of dactylus fringed with long setae (
<figureCitation id="9AE62A48FFC4FFD1FB6F01AEF8F96EEE" box="[1195,1277,1411,1437]" captionStart="FIGURE 3" captionStartId="6.[151,250,1779,1803]" captionTargetBox="[158,1423,194,1745]" captionTargetId="figure-29@6.[151,1436,181,1755]" captionTargetPageId="6" captionText="FIGURE 3. Alpheus alaincrosnieri n. sp.: a—right (minor) male cheliped, distal part of merus, carpus and chela, lateral; b—left (minor) male cheliped, chela, lateral; c—same, coxa to carpus, lateral; d—same, ischium and merus, ventral; e—left (minor) female cheliped, distal part of merus, carpus and chela, mesial, pollex broken. Drawn from: holotype male, cl 7.6 mm, MNHN-IU-2011-2230 [a]; paratype male, cl 6.5 mm, MNHN-IU-2019-3168 [bf]; paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [e], all off Sepik River, Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.4406735" httpUri="https://zenodo.org/record/4406735/files/figure.png" pageId="2" pageNumber="395">Fig. 3e</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC4FFD1FF03018AF8226CE5" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="395">
Second pereiopod very slender, especially in larger individuals; ischium and merus subequal in length; carpus with five subdivisions, first longest, ratio of carpal subdivisions approximately equal to 3.0/2.2/1/0.9/1.2; chela longer than distal-most carpal subdivision (
<figureCitation id="9AE62A48FFC4FFD1FD4501C2FEF56D79" box="[641,753,1519,1546]" captionStart="FIGURE 4" captionStartId="7.[151,250,1735,1759]" captionTargetBox="[156,1428,193,1702]" captionTargetId="figure-19@7.[151,1436,181,1712]" captionTargetPageId="7" captionText="FIGURE 4. Alpheus alaincrosnieri n. sp.: a—second pereiopod, lateral; b—same, distal part of merus, carpus and chela, lateral; c—third pereiopod, lateral; d, e—same, dactylus, ventrolateral and mesial views, respectively (propodal setae omitted in d, all setae omitted in e); f—fourth pereiopod, lateral; g—same, dactylus, lateral (all setae omitted); h—fifth pereiopod, lateral. Drawn from: paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [a]; holotype male, cl 7.6 mm, MNHN-IU-2011- 2230 [b, h], both off Sepik River, Papua New Guinea; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [cf]." figureDoi="http://doi.org/10.5281/zenodo.4406737" httpUri="https://zenodo.org/record/4406737/files/figure.png" pageId="2" pageNumber="395">Fig. 4a, b</figureCitation>
). Third pereiopod generally slender; ischium with stout cuspidate seta on ventrolateral surface; merus slender, about 10.5 times as long as maximal width, unarmed distoventrally; carpus about 0.6 length of merus, much slenderer than merus, unarmed; propodus noticeably longer than carpus, with numerous long stiff setae, ventral margin without spiniform setae; dactylus about half-length of propodus, gradually curving distally, spatulate, broadened, flattened ventrally, with distinct longitudinal keel (
<figureCitation id="9AE62A48FFC4FFD1FAAB0252FCA26DCE" captionStart="FIGURE 4" captionStartId="7.[151,250,1735,1759]" captionTargetBox="[156,1428,193,1702]" captionTargetId="figure-19@7.[151,1436,181,1712]" captionTargetPageId="7" captionText="FIGURE 4. Alpheus alaincrosnieri n. sp.: a—second pereiopod, lateral; b—same, distal part of merus, carpus and chela, lateral; c—third pereiopod, lateral; d, e—same, dactylus, ventrolateral and mesial views, respectively (propodal setae omitted in d, all setae omitted in e); f—fourth pereiopod, lateral; g—same, dactylus, lateral (all setae omitted); h—fifth pereiopod, lateral. Drawn from: paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [a]; holotype male, cl 7.6 mm, MNHN-IU-2011- 2230 [b, h], both off Sepik River, Papua New Guinea; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [cf]." figureDoi="http://doi.org/10.5281/zenodo.4406737" httpUri="https://zenodo.org/record/4406737/files/figure.png" pageId="2" pageNumber="395">Fig. 4</figureCitation>
ce). Fourth pereiopod generally similar to third, but slenderer; ischium longer than in third pereiopod, with stout cuspidate seta on ventrolateral surface; merus about 11.4 times as long as maximal width; propodus and dactylus as in third pereiopod (
<figureCitation id="9AE62A48FFC4FFD1FE5702C6FE046C76" box="[403,512,1771,1797]" captionStart="FIGURE 4" captionStartId="7.[151,250,1735,1759]" captionTargetBox="[156,1428,193,1702]" captionTargetId="figure-19@7.[151,1436,181,1712]" captionTargetPageId="7" captionText="FIGURE 4. Alpheus alaincrosnieri n. sp.: a—second pereiopod, lateral; b—same, distal part of merus, carpus and chela, lateral; c—third pereiopod, lateral; d, e—same, dactylus, ventrolateral and mesial views, respectively (propodal setae omitted in d, all setae omitted in e); f—fourth pereiopod, lateral; g—same, dactylus, lateral (all setae omitted); h—fifth pereiopod, lateral. Drawn from: paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [a]; holotype male, cl 7.6 mm, MNHN-IU-2011- 2230 [b, h], both off Sepik River, Papua New Guinea; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [cf]." figureDoi="http://doi.org/10.5281/zenodo.4406737" httpUri="https://zenodo.org/record/4406737/files/figure.png" pageId="2" pageNumber="395">Fig. 4f, g</figureCitation>
). Fifth pereiopod much slenderer than third pereiopod and slightly slenderer than fourth pereiopod; ischium longer than in third and fourth pereiopods, unarmed; merus about 10.3 times as long as wide; carpus slenderer than merus, about 0.8 length of merus; propodus longer than carpus, with numerous long stiff setae, ventral margin without spiniform setae, distal ventrolateral surface with rows of serrulate setae forming cleaning brush; dactylus similar to that of third and fourth pereiopods (
<figureCitation id="9AE62A48FFC4FFD1FC740356F81C6CE5" box="[944,1048,1915,1942]" captionStart="FIGURE 4" captionStartId="7.[151,250,1735,1759]" captionTargetBox="[156,1428,193,1702]" captionTargetId="figure-19@7.[151,1436,181,1712]" captionTargetPageId="7" captionText="FIGURE 4. Alpheus alaincrosnieri n. sp.: a—second pereiopod, lateral; b—same, distal part of merus, carpus and chela, lateral; c—third pereiopod, lateral; d, e—same, dactylus, ventrolateral and mesial views, respectively (propodal setae omitted in d, all setae omitted in e); f—fourth pereiopod, lateral; g—same, dactylus, lateral (all setae omitted); h—fifth pereiopod, lateral. Drawn from: paratype ovigerous female, cl 7.2 mm, MNHN-IU-2019-3168 [a]; holotype male, cl 7.6 mm, MNHN-IU-2011- 2230 [b, h], both off Sepik River, Papua New Guinea; paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [cf]." figureDoi="http://doi.org/10.5281/zenodo.4406737" httpUri="https://zenodo.org/record/4406737/files/figure.png" pageId="2" pageNumber="395">Fig. 4h, i</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC4FFD0FF0303B2F95F6B89" blockId="2.[151,1437,151,2014]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="396" pageId="2" pageNumber="395">
<collectingRegion id="C019F82FFFC4FFD1FF0303B2FD056CCA" box="[199,257,1951,1977]" country="Maldives" name="Maale" pageId="2" pageNumber="395">Male</collectingRegion>
second pleopod with appendix masculina slightly longer than appendix interna, with long stiff setae on apex and along margins (
<figureCitation id="9AE62A48FFC4FFD1FE6F03EEFDF16CAD" box="[427,501,1987,2014]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="2" pageNumber="395">Fig. 1j</figureCitation>
). Uropod with both mesial and lateral lobes of protopod ending bluntly; exopod fairly broad, somewhat truncate distally, with short triangular distolateral tooth; diaeresis straight for most part, except for blunt lobe adjacent to slender distolateral spiniform seta, latter reaching level of distal margin of exopod; endopod much narrower than exopod; neither exopod nor endopod with spiniform setae on their distal margins (
<figureCitation id="9AE62A48FFC5FFD0FB3904F2F9496B89" box="[1277,1357,223,250]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="3" pageNumber="396">Fig. 1k</figureCitation>
).
</paragraph>
<paragraph id="026236CDFFC5FFD0FF03052EFE4A6A6E" blockId="3.[151,1437,151,2013]" box="[199,590,259,285]" pageId="3" pageNumber="396">Colour pattern presently unknown.</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFC5FFD0FF03050AF9706AF9" pageId="3" pageNumber="396" type="etymology">
<paragraph id="026236CDFFC5FFD0FF03050AF9706AF9" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="396">
<emphasis id="30A9EADFFFC5FFD0FF03050AFD4C6A32" bold="true" box="[199,328,295,321]" pageId="3" pageNumber="396">Etymology</emphasis>
. This spectacular deep-water species is named after Dr. Alain Crosnier (formerly at ORSTOM /
<collectingRegion id="C019F82FFFC5FFD0FF530566FCF56A16" box="[151,241,331,357]" country="Ireland" name="Monaghan" pageId="3" pageNumber="396">MNHN</collectingRegion>
), in recognition of his invaluable contribution to taxonomy of various groups of
<taxonomicName id="C5DD4D4EFFC5FFD0FB4D0566F8F86A16" box="[1161,1276,331,357]" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="order">Decapoda</taxonomicName>
, including the presently most complete monograph of the
<taxonomicName id="C5DD4D4EFFC5FFD0FDB80542FEEB6AFA" authorityName="Rafinesque" authorityYear="1815" box="[636,751,367,393]" class="Malacostraca" family="Alpheidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
of the tropical eastern Atlantic (
<bibRefCitation id="664C4B3CFFC5FFD0FB9C0542F9636AF9" author="Crosnier, A. &amp; Forest, J." box="[1112,1383,367,394]" pageId="3" pageNumber="396" pagination="199 - 314" refId="ref9379" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFC5FFD0FF0305BFFF8B6AA2" pageId="3" pageNumber="396" type="distribution">
<paragraph id="026236CDFFC5FFD0FF0305BFFF8B6AA2" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="396">
<emphasis id="30A9EADFFFC5FFD0FF0305BFFD5D6ADF" bold="true" box="[199,345,402,428]" pageId="3" pageNumber="396">Distribution</emphasis>
. Indo-West Pacific:
<collectingCountry id="7ACA765DFFC5FFD0FD8605BEFF196ADE" box="[578,797,403,429]" name="Papua New Guinea" pageId="3" pageNumber="396">Papua New Guinea</collectingCountry>
(off
<collectingRegion id="C019F82FFFC5FFD0FC9205BEFF926ADE" box="[854,918,403,429]" country="Papua New Guinea" name="West Sepik" pageId="3" pageNumber="396">Sepik</collectingRegion>
River,
<collectingRegion id="C019F82FFFC5FFD0FC2E05BEF87F6ADD" box="[1002,1147,403,430]" country="Papua New Guinea" name="New Ireland" pageId="3" pageNumber="396">New Ireland</collectingRegion>
),
<collectingCountry id="7ACA765DFFC5FFD0FB4A05BEF8EE6ADE" box="[1166,1258,403,429]" name="Vanuatu" pageId="3" pageNumber="396">Vanuatu</collectingCountry>
(Espiritu Santo Island) and
<collectingCountry id="7ACA765DFFC5FFD0FEDF059AFDA66AA2" box="[283,418,439,465]" name="Madagascar" pageId="3" pageNumber="396">Madagascar</collectingCountry>
(off Narinda Bay), at depths of
<quantity id="C5259B28FFC5FFD0FCCD059AFF8F6AA2" box="[777,907,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.03" metricValueMax="4.08" metricValueMin="1.98" pageId="3" pageNumber="396" unit="m" value="303.0" valueMax="408.0" valueMin="198.0">
198
<specimenCount id="14DBFD44FFC5FFD0FC84059AFF8F6AA2" box="[832,907,439,465]" pageId="3" pageNumber="396" type="generic">408 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4AC76546FFC5FFDBFF0305F6FD7E6A16" lastPageId="8" lastPageNumber="401" pageId="3" pageNumber="396" type="discussion">
<paragraph id="026236CDFFC5FFD0FF0305F6FFE8682E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="396">
<emphasis id="30A9EADFFFC5FFD0FF0305F6FD306A86" bold="true" box="[199,308,475,501]" pageId="3" pageNumber="396">Remarks</emphasis>
.
<taxonomicName id="C5DD4D4EFFC5FFD0FE8605F6FE456A86" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[322,577,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFC5FFD0FE8605F6FE456A86" box="[322,577,475,501]" italics="true" pageId="3" pageNumber="396">Alpheus alaincrosnieri</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC5FFD0FD8C05F6FE836A86" bold="true" box="[584,647,475,501]" pageId="3" pageNumber="396">
<taxonomicNameLabel id="2B9A57A4FFC5FFD0FD8C05F6FE836A86" box="[584,647,475,501]" pageId="3" pageNumber="396" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
clearly belongs to the
<taxonomicName id="C5DD4D4EFFC5FFD0FC4105F6F8226A86" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[901,1062,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFC5FFD0FC4105F6F8226A86" box="[901,1062,475,501]" italics="true" pageId="3" pageNumber="396">A. brevirostris</emphasis>
</taxonomicName>
group based on the overall shape of the chelipeds. However, this new species is unique within the
<taxonomicName id="C5DD4D4EFFC5FFD0FCAB05D2F80B696A" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[879,1039,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFC5FFD0FCAB05D2F80B696A" box="[879,1039,511,537]" italics="true" pageId="3" pageNumber="396">A. brevirostris</emphasis>
</taxonomicName>
group by the presenting two small, but well-developed, acute orbital teeth, projecting from the anterior margin of each orbital hood (
<figureCitation id="9AE62A48FFC5FFD0FB1C060EF942694D" box="[1240,1350,547,574]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="3" pageNumber="396">Fig. 1a, b</figureCitation>
). In addition, the pterygostomial angle of
<taxonomicName id="C5DD4D4EFFC5FFD0FDE5066AFEDB6912" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[545,735,583,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFC5FFD0FDE5066AFEDB6912" box="[545,735,583,609]" italics="true" pageId="3" pageNumber="396">A. alaincrosnieri</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC5FFD0FD22066AFF206912" bold="true" box="[742,804,583,609]" pageId="3" pageNumber="396">
<taxonomicNameLabel id="2B9A57A4FFC5FFD0FD22066AFF206912" box="[742,804,583,609]" pageId="3" pageNumber="396" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is strongly projecting anteriorly in form of a stout sharp tooth (
<figureCitation id="9AE62A48FFC5FFD0FF260646FD3169F5" box="[226,309,619,646]" captionStart="FIGURE 1" captionStartId="4.[151,250,1868,1892]" captionTargetBox="[224,1385,213,1826]" captionTargetId="figure-29@4.[184,1404,181,1844]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus alaincrosnieri n. sp.: a—frontal region, dorsal; b—same, lateral; c—telson, dorsal; d—same, posterior margin, dorsal (setae omitted); e—lateral flagellum of antennule, lateral; f—first maxilliped, lateral; g, h—second maxilliped, lateral; i—third maxilliped, lateral; j—appendix masculina and appendix interna of male second pleopod, anterior (lateral); k—uropod, dorsal. Drawn from: paratype male, cl 7.0 mm, MNHN-2011-3024, off New Ireland, Papua New Guinea [ag, ik]; non-type ovigerous female, cl 8.5 mm, MNHN-IU-2010-1278, from Mozambique Channel off Madagascar [h]." figureDoi="http://doi.org/10.5281/zenodo.4406731" httpUri="https://zenodo.org/record/4406731/files/figure.png" pageId="3" pageNumber="396">Fig. 1b</figureCitation>
), a feature not commonly seen in the genus
<taxonomicName id="C5DD4D4EFFC5FFD0FCF20646FF9469F6" authorityName="Fabricius" authorityYear="1798" box="[822,912,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="30A9EADFFFC5FFD0FCF20646FF9469F6" box="[822,912,619,645]" italics="true" pageId="3" pageNumber="396">Alpheus</emphasis>
</taxonomicName>
. Within the
<taxonomicName id="C5DD4D4EFFC5FFD0FBE50646F8C769F6" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[1057,1219,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFC5FFD0FBE50646F8C769F6" box="[1057,1219,619,645]" italics="true" pageId="3" pageNumber="396">A. brevirostris</emphasis>
</taxonomicName>
group, only
<taxonomicName id="C5DD4D4EFFC5FFD0FA930646FE3369DA" authority="Hayashi &amp; Nagata, 2000" authorityName="Hayashi &amp; Nagata" authorityYear="2000" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="kagoshimanus">
<emphasis id="30A9EADFFFC5FFD0FA930646FD1969DA" italics="true" pageId="3" pageNumber="396">A. kagoshimanus</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FEE006A2FE3369DA" author="Hayashi, K. - I. &amp; Nagata, M." box="[292,567,655,681]" pageId="3" pageNumber="396" pagination="1109 - 1120" refId="ref9907" refString="Hayashi, K. - I. &amp; Nagata, M. (2000) A new species of Alpheus (Decapoda, Caridea, Alpheidae) from Kagoshima Bay, Japan. Crustaceana, 73, 1109 - 1120. https: // doi. org / 10.1163 / 156854000505119" type="journal article" year="2000">Hayashi &amp; Nagata, 2000</bibRefCitation>
</taxonomicName>
has a pterygostomial tooth, which, however, is much smaller than in the present new species.All other species of the
<taxonomicName id="C5DD4D4EFFC5FFD0FD92069EFEF269BE" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[598,758,691,717]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFC5FFD0FD92069EFEF269BE" box="[598,758,691,717]" italics="true" pageId="3" pageNumber="396">A. brevirostris</emphasis>
</taxonomicName>
group characterised by the significantly elongated chelipeds, viz.
<taxonomicName id="C5DD4D4EFFC5FFD0FF0D06FAFEAD6982" authority="Alcock &amp; Anderson, 1894" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[201,681,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="30A9EADFFFC5FFD0FF0D06FAFD776982" box="[201,371,727,753]" italics="true" pageId="3" pageNumber="396">A. macroskeles</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FEBE06FAFEAD6982" author="Alcock, A. &amp; Anderson, A. R. S." box="[378,681,727,753]" pageId="3" pageNumber="396" pagination="141 - 185" refId="ref8451" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer &quot; Investigator &quot;, Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article" year="1894">Alcock &amp; Anderson, 1894</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FD7106FAFFF66982" authority="Coutiere, 1898" authorityName="Coutiere" authorityYear="1898" box="[693,1010,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="30A9EADFFFC5FFD0FD7106FAFF3A6982" box="[693,830,727,753]" italics="true" pageId="3" pageNumber="396">A. talismani</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FC8306FAFFF66982" author="Coutiere, H." box="[839,1010,727,753]" pageId="3" pageNumber="396" pagination="31 - 33" refId="ref9248" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33. https: // doi. org / 10.5962 / bhl. part. 16849" type="journal article" year="1898">Coutière, 1898</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FC3A06FAF9736982" authority="De Man, 1909" authorityName="De Man" authorityYear="1909" box="[1022,1399,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="30A9EADFFFC5FFD0FC3A06FAF8C36983" box="[1022,1223,727,753]" italics="true" pageId="3" pageNumber="396">A. acutocarinatus</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FB1406FAF9736982" author="De Man, J. G." box="[1232,1399,727,753]" pageId="3" pageNumber="396" pagination="99 - 125" refId="ref9637" refString="De Man, J. G. (1909) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandse Dierkundige Vereeniging, 2 (9), 99 - 125." type="journal article" year="1909">De Man, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FA4006FAFDAE6866" authority="Stebbing, 1915" authorityName="Stebbing" authorityYear="1915" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="30A9EADFFFC5FFD0FA4006FAFCFD6866" italics="true" pageId="3" pageNumber="396">A. notabilis</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FF3B06D6FDAE6866" author="Stebbing, T. R. R." box="[255,426,762,789]" pageId="3" pageNumber="396" pagination="57 - 103" refId="ref10481" refString="Stebbing, T. R. R. (1915) South African Crustacea. Part VIII of S. A. Crustacea, for the marine investigations in South Africa. Annals of the South African Museum, 15, 57 - 103, pls. 13 - 25. https: // doi. org / 10.5962 / bhl. part. 22195" type="journal article" year="1915">Stebbing, 1915</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FE7006D6FEDB6866" authority="Boone, 1935" authorityName="Boone" authorityYear="1935" box="[436,735,762,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="30A9EADFFFC5FFD0FE7006D6FE4C6866" box="[436,584,763,789]" italics="true" pageId="3" pageNumber="396">A. explorator</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FD8A06D6FEDB6866" author="Boone, L." box="[590,735,762,789]" pageId="3" pageNumber="396" pagination="1 - 264" refId="ref9038" refString="Boone, L. (1935) The Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea of the &quot; Alva &quot; world cruise, 1931, William K. Vanderbilt, commanding. Bulletin of the Vanderbilt Marine Museum, 6, 1 - 264, pls. 1 - 96." type="journal article" year="1935">Boone, 1935</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FD2D06D6F8AB6866" authority="Lewinsohn &amp; Holthuis, 1978" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1978" box="[745,1199,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="30A9EADFFFC5FFD0FD2D06D6FF676867" box="[745,867,763,789]" italics="true" pageId="3" pageNumber="396">A. migrans</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FCAD06D6F8AB6866" author="Lewinsohn, C. &amp; Holthuis, L. B." box="[873,1199,763,789]" pageId="3" pageNumber="396" pagination="75 - 82" refId="ref10148" refString="Lewinsohn, C. &amp; Holthuis, L. B. (1978) On a new species of Alpheus (Crustacea Decapoda, Natantia) from the eastern Mediterranean. Zoologische Mededelingen, Leiden, 53, 75 - 82." type="journal article" year="1978">Lewinsohn &amp; Holthuis, 1978</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FB7D06D6FCD6684A" authority="Kensley, 1969" authorityName="Kensley" authorityYear="1969" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="30A9EADFFFC5FFD0FB7D06D6F9326866" box="[1209,1334,763,789]" italics="true" pageId="3" pageNumber="396">A. nonalter</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FAFF06D6FCD6684A" author="Kensley, B." pageId="3" pageNumber="396" pagination="149 - 181" refId="ref9958" refString="Kensley, B. (1969) Decapod Crustacea from the south-west Indian Ocean. Annals of the South African Museum, 52, 149 - 181." type="journal article" year="1969">Kensley, 1969</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="C5DD4D4EFFC5FFD0FEC90732FEC6684A" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[269,706,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="longipalma">
<emphasis id="30A9EADFFFC5FFD0FEC90732FDA8684A" box="[269,428,799,825]" italics="true" pageId="3" pageNumber="396">A. longipalma</emphasis>
<bibRefCitation id="664C4B3CFFC5FFD0FE770732FEC6684A" author="Komai, T. &amp; Ohtomi, J." box="[435,706,799,825]" pageId="3" pageNumber="396" pagination="1" refId="ref10087" refString="Komai, T. &amp; Ohtomi, J. (2018) A new deep-sea species of the snapping shrimp genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from Kagoshima Bay, Japan. Zootaxa, 4434 (1), 099 - 110. https: // doi. org / 10.11646 / zootaxa. 4434.1.6" type="journal article" year="2018">Komai &amp; Ohtomi, 2018</bibRefCitation>
</taxonomicName>
, can be easily separated from
<taxonomicName id="C5DD4D4EFFC5FFD0FBD00732F8D5684A" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[1044,1233,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFC5FFD0FBD00732F8D5684A" box="[1044,1233,799,825]" italics="true" pageId="3" pageNumber="396">A. alaincrosnieri</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC5FFD0FB1C0732F912684A" bold="true" box="[1240,1302,799,825]" pageId="3" pageNumber="396">
<taxonomicNameLabel id="2B9A57A4FFC5FFD0FB1C0732F912684A" box="[1240,1302,799,825]" pageId="3" pageNumber="396" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
both by the rounded orbital hoods and rounded to slightly angular pterygostomial angle.
</paragraph>
<paragraph id="026236CDFFC5FFD0FF03074AF8D26DEA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="396">
<taxonomicName id="C5DD4D4EFFC5FFD0FF03074AFDC268F2" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[199,454,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFC5FFD0FF03074AFDC268F2" box="[199,454,871,897]" italics="true" pageId="3" pageNumber="396">Alpheus alaincrosnieri</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC5FFD0FE09074AFE0868F2" bold="true" box="[461,524,871,897]" pageId="3" pageNumber="396">
<taxonomicNameLabel id="2B9A57A4FFC5FFD0FE09074AFE0868F2" box="[461,524,871,897]" pageId="3" pageNumber="396" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be separated from each of these species by at least one additional, taxonomically significant character, for instance, from
<taxonomicName id="C5DD4D4EFFC5FFD0FD4807A6FF4E68D6" authorityName="Hayashi &amp; Nagata" authorityYear="2000" box="[652,842,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="kagoshimanus">
<emphasis id="30A9EADFFFC5FFD0FD4807A6FF4E68D6" box="[652,842,907,933]" italics="true" pageId="3" pageNumber="396">A. kagoshimanus</emphasis>
</taxonomicName>
by the much stronger distolateral tooth of the scaphocerite, distinctly overreaching the distal margin of the blade (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FC9F0782F86F68BA" author="Hayashi, K. - I. &amp; Nagata, M." box="[859,1131,943,969]" pageId="3" pageNumber="396" pagination="1109 - 1120" refId="ref9907" refString="Hayashi, K. - I. &amp; Nagata, M. (2000) A new species of Alpheus (Decapoda, Caridea, Alpheidae) from Kagoshima Bay, Japan. Crustaceana, 73, 1109 - 1120. https: // doi. org / 10.1163 / 156854000505119" type="journal article" year="2000">Hayashi &amp; Nagata 2000</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FB7D0782F97368BA" authorityName="Hayashi &amp; Nagata" authorityYear="2000" box="[1209,1399,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="kagoshimanus">
<emphasis id="30A9EADFFFC5FFD0FB7D0782F97368BA" box="[1209,1399,943,969]" italics="true" pageId="3" pageNumber="396">A. kagoshimanus</emphasis>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FA400782FCFD689E" authorityName="Stebbing" authorityYear="1915" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="30A9EADFFFC5FFD0FA400782FCFD689E" italics="true" pageId="3" pageNumber="396">A. notabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFC5FFD0FEE807FEFDA2689F" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1978" box="[300,422,979,1005]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="30A9EADFFFC5FFD0FEE807FEFDA2689F" box="[300,422,979,1005]" italics="true" pageId="3" pageNumber="396">A. migrans</emphasis>
</taxonomicName>
(and possibly from most if not all other species) by the presence of a sharp tooth on the third article of the endopod of the second maxilliped (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FD1307DAF81C6F62" author="Lewinsohn, C. &amp; Holthuis, L. B." box="[727,1048,1015,1041]" pageId="3" pageNumber="396" pagination="75 - 82" refId="ref10148" refString="Lewinsohn, C. &amp; Holthuis, L. B. (1978) On a new species of Alpheus (Crustacea Decapoda, Natantia) from the eastern Mediterranean. Zoologische Mededelingen, Leiden, 53, 75 - 82." type="journal article" year="1978">Lewinsohn &amp; Holthuis 1978</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FBE707DAF9316F62" author="Hayashi, K. - I. &amp; Nagata, M." box="[1059,1333,1015,1041]" pageId="3" pageNumber="396" pagination="1109 - 1120" refId="ref9907" refString="Hayashi, K. - I. &amp; Nagata, M. (2000) A new species of Alpheus (Decapoda, Caridea, Alpheidae) from Kagoshima Bay, Japan. Crustaceana, 73, 1109 - 1120. https: // doi. org / 10.1163 / 156854000505119" type="journal article" year="2000">Hayashi &amp; Nagata 2000</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FA4007DAFD246F46" authorityName="Alcock &amp; Anderson" authorityYear="1894" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="30A9EADFFFC5FFD0FA4007DAFD246F46" italics="true" pageId="3" pageNumber="396">A. macroskeles</emphasis>
</taxonomicName>
,
<taxonomicName id="C5DD4D4EFFC5FFD0FEEA0036FDB06F46" authorityName="Coutiere" authorityYear="1898" box="[302,436,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="30A9EADFFFC5FFD0FEEA0036FDB06F46" box="[302,436,1051,1077]" italics="true" pageId="3" pageNumber="396">A. talismani</emphasis>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFC5FFD0FE2C0036FE836F46" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[488,647,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="longipalma">
<emphasis id="30A9EADFFFC5FFD0FE2C0036FE836F46" box="[488,647,1051,1077]" italics="true" pageId="3" pageNumber="396">A. longipalma</emphasis>
</taxonomicName>
by the cheliped merus without subdistal dorsal tooth (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FAC40036FD206F2A" author="Alcock, A. &amp; Anderson, A. R. S." pageId="3" pageNumber="396" pagination="141 - 185" refId="ref8451" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer &quot; Investigator &quot;, Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article" year="1894">Alcock &amp; Anderson 1894</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FEF40012FDD76F2A" author="Coutiere, H." box="[304,467,1087,1113]" pageId="3" pageNumber="396" pagination="31 - 33" refId="ref9248" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33. https: // doi. org / 10.5962 / bhl. part. 16849" type="journal article" year="1898">Coutière 1898</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FE1A0012FE756F2A" author="Alcock, A." box="[478,625,1087,1113]" pageId="3" pageNumber="396" refId="ref8380" refString="Alcock, A. (1901) A descriptive catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a revised account of the deep-sea species collected by the Royal Indian marine survey ship Investigator. Indian Museum, Calcutta, iv + 286 pp., 3 pls. https: // doi. org / 10.5962 / bhl. title. 30840" type="book" year="1901">Alcock 1901</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FDB90012FF8F6F2A" author="Crosnier, A. &amp; Forest, J." box="[637,907,1087,1113]" pageId="3" pageNumber="396" pagination="199 - 314" refId="ref9379" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FC530012F8A66F2A" author="Komai, T. &amp; Ohtomi, J." box="[919,1186,1087,1113]" pageId="3" pageNumber="396" pagination="1" refId="ref10087" refString="Komai, T. &amp; Ohtomi, J. (2018) A new deep-sea species of the snapping shrimp genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from Kagoshima Bay, Japan. Zootaxa, 4434 (1), 099 - 110. https: // doi. org / 10.11646 / zootaxa. 4434.1.6" type="journal article" year="2018">Komai &amp; Ohtomi 2018</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FB370012F9986F2A" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[1267,1436,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="30A9EADFFFC5FFD0FB370012F9986F2A" box="[1267,1436,1087,1113]" italics="true" pageId="3" pageNumber="396">A. macroskeles</emphasis>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFC5FFD0FF02004EFD486F0E" authorityName="Coutiere" authorityYear="1898" box="[198,332,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="30A9EADFFFC5FFD0FF02004EFD486F0E" box="[198,332,1123,1149]" italics="true" pageId="3" pageNumber="396">A. talismani</emphasis>
</taxonomicName>
by the minor chela strongly balaeniceps in males (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FC65004EF8C56F0E" author="Alcock, A. &amp; Anderson, A. R. S." box="[929,1217,1123,1149]" pageId="3" pageNumber="396" pagination="141 - 185" refId="ref8451" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer &quot; Investigator &quot;, Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article" year="1894">Alcock &amp; Anderson 1894</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FB08004EF9696F0E" author="Coutiere, H." box="[1228,1389,1123,1149]" pageId="3" pageNumber="396" pagination="31 - 33" refId="ref9248" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33. https: // doi. org / 10.5962 / bhl. part. 16849" type="journal article" year="1898">Coutière 1898</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FAB3004EFD096FD2" author="Alcock, A." pageId="3" pageNumber="396" refId="ref8380" refString="Alcock, A. (1901) A descriptive catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a revised account of the deep-sea species collected by the Royal Indian marine survey ship Investigator. Indian Museum, Calcutta, iv + 286 pp., 3 pls. https: // doi. org / 10.5962 / bhl. title. 30840" type="book" year="1901">Alcock 1901</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FEDD00AAFE206FD1" author="Crosnier, A. &amp; Forest, J." box="[281,548,1159,1186]" pageId="3" pageNumber="396" pagination="199 - 314" refId="ref9379" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FDB000AAFF186FD2" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[628,796,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="30A9EADFFFC5FFD0FDB000AAFF186FD2" box="[628,796,1159,1185]" italics="true" pageId="3" pageNumber="396">A. macroskeles</emphasis>
</taxonomicName>
by the better developed cornea of eyes (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FB3B00AAFD206FB6" author="Alcock, A. &amp; Anderson, A. R. S." pageId="3" pageNumber="396" pagination="141 - 185" refId="ref8451" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer &quot; Investigator &quot;, Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article" year="1894">Alcock &amp; Anderson 1894</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FEEA0086FDC46FB6" author="Alcock, A." box="[302,448,1195,1221]" pageId="3" pageNumber="396" refId="ref8380" refString="Alcock, A. (1901) A descriptive catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a revised account of the deep-sea species collected by the Royal Indian marine survey ship Investigator. Indian Museum, Calcutta, iv + 286 pp., 3 pls. https: // doi. org / 10.5962 / bhl. title. 30840" type="book" year="1901">Alcock 1901</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FE080086FED06FB6" author="Komai, T. &amp; Ohtomi, J." box="[460,724,1195,1221]" pageId="3" pageNumber="396" pagination="1" refId="ref10087" refString="Komai, T. &amp; Ohtomi, J. (2018) A new deep-sea species of the snapping shrimp genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from Kagoshima Bay, Japan. Zootaxa, 4434 (1), 099 - 110. https: // doi. org / 10.11646 / zootaxa. 4434.1.6" type="journal article" year="2018">Komai &amp; Ohtomi 2018</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FCE00086FFAF6FB6" authorityName="Coutiere" authorityYear="1898" box="[804,939,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="30A9EADFFFC5FFD0FCE00086FFAF6FB6" box="[804,939,1195,1221]" italics="true" pageId="3" pageNumber="396">A. talismani</emphasis>
</taxonomicName>
by the presence of an elevated rostral carina armed with a post-rostral tubercle (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FD8700E2FEE16F9A" author="Coutiere, H." box="[579,741,1231,1257]" pageId="3" pageNumber="396" pagination="31 - 33" refId="ref9248" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33. https: // doi. org / 10.5962 / bhl. part. 16849" type="journal article" year="1898">Coutière 1898</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FD3500E2FFF96F99" author="Crosnier, A. &amp; Forest, J." box="[753,1021,1231,1258]" pageId="3" pageNumber="396" pagination="199 - 314" refId="ref9379" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FB8A00E2F9116F9B" authorityName="De Man" authorityYear="1909" box="[1102,1301,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="30A9EADFFFC5FFD0FB8A00E2F9116F9B" box="[1102,1301,1231,1257]" italics="true" pageId="3" pageNumber="396">A. acutocarinatus</emphasis>
</taxonomicName>
by the third pereiopod ischium armed with a stout cuspidate seta (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FCD000DEFFB46E7E" author="De Man, J. G." box="[788,944,1267,1293]" pageId="3" pageNumber="396" pagination="1 - 23" refId="ref9715" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. In: Siboga Expeditie. 39 a 1. Brill, Leiden, pp. 133 - 465; (1915) Supplement-Explanation of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FBC500DEF8CC6E7F" authorityName="De Man" authorityYear="1909" box="[1025,1224,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="30A9EADFFFC5FFD0FBC500DEF8CC6E7F" box="[1025,1224,1267,1293]" italics="true" pageId="3" pageNumber="396">A. acutocarinatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFC5FFD0FB3B00DEF97E6E7F" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1978" box="[1279,1402,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="30A9EADFFFC5FFD0FB3B00DEF97E6E7F" box="[1279,1402,1267,1293]" italics="true" pageId="3" pageNumber="396">A. migrans</emphasis>
</taxonomicName>
by the absence of small tubercle on the mid-dorsal line of the posterior region of the carapace and the second pereiopod with the first carpal subdivision longer than the second (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FCFA0116FFD86E26" author="De Man, J. G." box="[830,988,1339,1365]" pageId="3" pageNumber="396" pagination="1 - 23" refId="ref9715" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. In: Siboga Expeditie. 39 a 1. Brill, Leiden, pp. 133 - 465; (1915) Supplement-Explanation of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FC2D0116F9346E26" author="Lewinsohn, C. &amp; Holthuis, L. B." box="[1001,1328,1339,1365]" pageId="3" pageNumber="396" pagination="75 - 82" refId="ref10148" refString="Lewinsohn, C. &amp; Holthuis, L. B. (1978) On a new species of Alpheus (Crustacea Decapoda, Natantia) from the eastern Mediterranean. Zoologische Mededelingen, Leiden, 53, 75 - 82." type="journal article" year="1978">Lewinsohn &amp; Holthuis 1978</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FA400116FCFD6E0A" authorityName="Stebbing" authorityYear="1915" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="30A9EADFFFC5FFD0FA400116FCFD6E0A" italics="true" pageId="3" pageNumber="396">A. notabilis</emphasis>
</taxonomicName>
by the broader and less constricted telson and the scaphocerite with a straight (not noticeably concave, as in
<taxonomicName id="C5DD4D4EFFC5FFD0FF7701AEFD306EEE" authorityName="Stebbing" authorityYear="1915" box="[179,308,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="30A9EADFFFC5FFD0FF7701AEFD306EEE" box="[179,308,1411,1437]" italics="true" pageId="3" pageNumber="396">A. notabilis</emphasis>
</taxonomicName>
) lateral margin (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FDD601AEFEBC6EEE" author="Stebbing, T. R. R." box="[530,696,1410,1437]" pageId="3" pageNumber="396" pagination="57 - 103" refId="ref10481" refString="Stebbing, T. R. R. (1915) South African Crustacea. Part VIII of S. A. Crustacea, for the marine investigations in South Africa. Annals of the South African Museum, 15, 57 - 103, pls. 13 - 25. https: // doi. org / 10.5962 / bhl. part. 22195" type="journal article" year="1915">Stebbing 1915</bibRefCitation>
); from both
<taxonomicName id="C5DD4D4EFFC5FFD0FC8501AEFFD26EEE" authorityName="Boone" authorityYear="1935" box="[833,982,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="30A9EADFFFC5FFD0FC8501AEFFD26EEE" box="[833,982,1411,1437]" italics="true" pageId="3" pageNumber="396">A. explorator</emphasis>
</taxonomicName>
and
<taxonomicName id="C5DD4D4EFFC5FFD0FBC801AEF88D6EEE" authorityName="Kensley" authorityYear="1969" box="[1036,1161,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="30A9EADFFFC5FFD0FBC801AEF88D6EEE" box="[1036,1161,1411,1437]" italics="true" pageId="3" pageNumber="396">A. nonalter</emphasis>
</taxonomicName>
by the major chela palm without a deep constriction near the base of the pollex (
<bibRefCitation id="664C4B3CFFC5FFD0FD33018AFF876EB2" author="Boone, L." box="[759,899,1446,1473]" pageId="3" pageNumber="396" pagination="1 - 264" refId="ref9038" refString="Boone, L. (1935) The Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea of the &quot; Alva &quot; world cruise, 1931, William K. Vanderbilt, commanding. Bulletin of the Vanderbilt Marine Museum, 6, 1 - 264, pls. 1 - 96." type="journal article" year="1935">Boone 1935</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FC4A018AF82E6EB2" author="Kensley, B." box="[910,1066,1447,1473]" pageId="3" pageNumber="396" pagination="149 - 181" refId="ref9958" refString="Kensley, B. (1969) Decapod Crustacea from the south-west Indian Ocean. Annals of the South African Museum, 52, 149 - 181." type="journal article" year="1969">Kensley 1969</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FBF1018AF8BF6EB2" author="Chace, F. A. Jr." box="[1077,1211,1447,1473]" pageId="3" pageNumber="396" pagination="1 - 99" refId="ref9190" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FB02018AF9576EB2" author="Komai, T." box="[1222,1363,1447,1473]" pageId="3" pageNumber="396" pagination="279 - 337" refId="ref10040" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, Central Japan. Memoirs of the National Museum of Nature and Science, 47, 279 - 337." type="journal article" year="2011">Komai 2011</bibRefCitation>
); from
<taxonomicName id="C5DD4D4EFFC5FFD0FF5301E6FD296E96" authorityName="Boone" authorityYear="1935" box="[151,301,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="30A9EADFFFC5FFD0FF5301E6FD296E96" box="[151,301,1483,1509]" italics="true" pageId="3" pageNumber="396">A. explorator</emphasis>
</taxonomicName>
by the position of the ventromesial tooth on the cheliped merus (distal in the new species
<emphasis id="30A9EADFFFC5FFD0FAEE01E6F9456E97" box="[1322,1345,1483,1508]" italics="true" pageId="3" pageNumber="396">vs</emphasis>
. clearly subdistal in
<taxonomicName id="C5DD4D4EFFC5FFD0FEDB01C2FDB16D7A" authorityName="Boone" authorityYear="1935" box="[287,437,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="30A9EADFFFC5FFD0FEDB01C2FDB16D7A" box="[287,437,1519,1545]" italics="true" pageId="3" pageNumber="396">A. explorator</emphasis>
</taxonomicName>
) (
<bibRefCitation id="664C4B3CFFC5FFD0FE0901C2FE5E6D7A" author="Boone, L." box="[461,602,1518,1545]" pageId="3" pageNumber="396" pagination="1 - 264" refId="ref9038" refString="Boone, L. (1935) The Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea of the &quot; Alva &quot; world cruise, 1931, William K. Vanderbilt, commanding. Bulletin of the Vanderbilt Marine Museum, 6, 1 - 264, pls. 1 - 96." type="journal article" year="1935">Boone 1935</bibRefCitation>
; A. Anker, in prep.); and from
<taxonomicName id="C5DD4D4EFFC5FFD0FC7001C2F8346D7B" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1978" box="[948,1072,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="30A9EADFFFC5FFD0FC7001C2F8346D7B" box="[948,1072,1519,1545]" italics="true" pageId="3" pageNumber="396">A. migrans</emphasis>
</taxonomicName>
by the much longer penultimate article of the third maxilliped and the lower, more reduced plunger of the major chela (cf.
<bibRefCitation id="664C4B3CFFC5FFD0FB5D023EFCD66D22" author="Lewinsohn, C. &amp; Holthuis, L. B." pageId="3" pageNumber="396" pagination="75 - 82" refId="ref10148" refString="Lewinsohn, C. &amp; Holthuis, L. B. (1978) On a new species of Alpheus (Crustacea Decapoda, Natantia) from the eastern Mediterranean. Zoologische Mededelingen, Leiden, 53, 75 - 82." type="journal article" year="1978">Lewinsohn &amp; Holthuis 1978</bibRefCitation>
;
<bibRefCitation id="664C4B3CFFC5FFD0FF1B021AFE346D22" author="Dworschak, P. C. &amp; Pervesler, P." box="[223,560,1591,1617]" pageId="3" pageNumber="396" pagination="351 - 357" refId="ref9772" refString="Dworschak, P. C. &amp; Pervesler, P. (2002) Alpheus migrans Lewinsohn &amp; Holthuis, 1978 (Decapoda, Caridea, Alpheidae): burrow morphology and first record from the Red Sea. Crustaceana, 75, 351 - 357. https: // doi. org / 10.1163 / 156854002760095426" type="journal article" year="2002">Dworschak &amp; Pervesler 2002</bibRefCitation>
). It must be noted that
<taxonomicName id="C5DD4D4EFFC5FFD0FCF7021AFFB06D22" authorityName="Stebbing" authorityYear="1915" box="[819,948,1591,1617]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="30A9EADFFFC5FFD0FCF7021AFFB06D22" box="[819,948,1591,1617]" italics="true" pageId="3" pageNumber="396">A. notabilis</emphasis>
</taxonomicName>
, described by
<bibRefCitation id="664C4B3CFFC5FFD0FB9D021AF90B6D22" author="Stebbing, T. R. R." box="[1113,1295,1590,1617]" pageId="3" pageNumber="396" pagination="57 - 103" refId="ref10481" refString="Stebbing, T. R. R. (1915) South African Crustacea. Part VIII of S. A. Crustacea, for the marine investigations in South Africa. Annals of the South African Museum, 15, 57 - 103, pls. 13 - 25. https: // doi. org / 10.5962 / bhl. part. 22195" type="journal article" year="1915">Stebbing (1915)</bibRefCitation>
on the basis of a heavily damaged specimen missing its major cheliped, with somewhat diagrammatic and possibly not very accurate illustrations, has not been reported since the original description (
<bibRefCitation id="664C4B3CFFC5FFD0FC7B0252F8CD6DEA" author="Banner, A. H. &amp; Banner, D. M." box="[959,1225,1663,1689]" pageId="3" pageNumber="396" pagination="1 - 164" refId="ref8845" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
).
</paragraph>
<paragraph id="026236CDFFC5FFD0FF03028EF8C56CE6" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="396">
<bibRefCitation id="664C4B3CFFC5FFD0FF03028EFD5A6DCE" author="Chace, F. A. Jr." box="[199,350,1699,1725]" pageId="3" pageNumber="396" pagination="1 - 99" refId="ref9190" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace (1988)</bibRefCitation>
listed a mutilated ovigerous female (cl
<quantity id="C5259B28FFC5FFD0FCD2028EFF696DCE" box="[790,877,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="3" pageNumber="396" unit="mm" value="8.3">8.3 mm</quantity>
) collected at a depth of
<quantity id="C5259B28FFC5FFD0FBB2028FF8B96DCE" box="[1142,1213,1698,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="396" unit="m" value="245.0">245 m</quantity>
near
<collectingRegion id="C019F82FFFC5FFD0FB3D028EF95F6DCE" box="[1273,1371,1699,1725]" country="Philippines" name="Masbate" pageId="3" pageNumber="396">Masbate</collectingRegion>
,
<collectingCountry id="7ACA765DFFC5FFD0FAA2028EFCEF6D92" name="Philippines" pageId="3" pageNumber="396">Philippines</collectingCountry>
, in the remarks under
<taxonomicName id="C5DD4D4EFFC5FFD0FE1B02EAFEA26D93" authorityName="De Man" authorityYear="1909" box="[479,678,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="30A9EADFFFC5FFD0FE1B02EAFEA26D93" box="[479,678,1735,1761]" italics="true" pageId="3" pageNumber="396">A. acutocarinatus</emphasis>
</taxonomicName>
. However, the authors notes show that he was well aware that it did not belong to that species. Based on
<bibRefCitation id="664C4B3CFFC5FFD0FDF502C6FED86C76" author="Chace, F. A. Jr." box="[561,732,1771,1797]" pageId="3" pageNumber="396" pagination="1 - 99" refId="ref9190" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chaces (1988)</bibRefCitation>
observations, the specimen from
<collectingRegion id="C019F82FFFC5FFD0FB9302C6F8B26C76" box="[1111,1206,1771,1797]" country="Philippines" name="Masbate" pageId="3" pageNumber="396">Masbate</collectingRegion>
had “the remnant of what must have been a larger mesial gastric tooth [= post-rostral tubercle], by a small spine on the frontal margin either side of the rostrum [= orbital teeth], and by a rather distinct branchiostegal spine [= pterygostomial tooth]”, which are the most diagnostic features of the carapace of
<taxonomicName id="C5DD4D4EFFC5FFD0FCDB037AFFD96C02" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[799,989,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFC5FFD0FCDB037AFFD96C02" box="[799,989,1879,1905]" italics="true" pageId="3" pageNumber="396">A. alaincrosnieri</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFC5FFD0FC20037AF8276C02" bold="true" box="[996,1059,1879,1905]" pageId="3" pageNumber="396">
<taxonomicNameLabel id="2B9A57A4FFC5FFD0FC20037AF8276C02" box="[996,1059,1879,1905]" pageId="3" pageNumber="396" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Therefore,
<bibRefCitation id="664C4B3CFFC5FFD0FB63037AF9566C02" author="Chace, F. A. Jr." box="[1191,1362,1879,1905]" pageId="3" pageNumber="396" pagination="1 - 99" refId="ref9190" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chaces (1988)</bibRefCitation>
specimen is here tentatively referred to the new species, extending its distribution to the
<collectingCountry id="7ACA765DFFC5FFD0FBF80356F8BA6CE6" box="[1084,1214,1915,1941]" name="Philippines" pageId="3" pageNumber="396">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="026236CDFFC5FFDBFF0303B2FD7E6A16" blockId="3.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,357]" lastPageId="8" lastPageNumber="401" pageId="3" pageNumber="396">
<bibRefCitation id="664C4B3CFFC5FFD0FF0303B2FD586CCA" author="Miya, Y." box="[199,348,1951,1977]" pageId="3" pageNumber="396" pagination="103 - 195" refId="ref10255" refString="Miya, Y. (1974) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part II. Publications from the Amakusa Marine Biology Laboratory, 3, 103 - 195, pls. 15 - 31." type="journal article" year="1974">Miya (1974)</bibRefCitation>
, while providing short diagnoses of the seven species groups of
<taxonomicName id="C5DD4D4EFFC5FFD0FBA803B2F8C26CCA" authorityName="Fabricius" authorityYear="1798" box="[1132,1222,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="30A9EADFFFC5FFD0FBA803B2F8C26CCA" box="[1132,1222,1951,1977]" italics="true" pageId="3" pageNumber="396">Alpheus</emphasis>
</taxonomicName>
, mentioned under the
<taxonomicName id="C5DD4D4EFFC5FFD0FF0703EEFD626CAE" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[195,358,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFC5FFD0FF0703EEFD626CAE" box="[195,358,1987,2013]" italics="true" pageId="3" pageNumber="396">A. brevirostris</emphasis>
</taxonomicName>
group “
<taxonomicName id="C5DD4D4EFFC5FFD0FE0303EEFE446CAE" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[455,576,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="miyakei" status="sp. nov.">
<emphasis id="30A9EADFFFC5FFD0FE0303EEFE446CAE" box="[455,576,1987,2013]" italics="true" pageId="3" pageNumber="396">A. miyakei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B9A57A4FFC5FFD0FD8E03EEFEA06CAE" box="[586,676,1987,2013]" pageId="3" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
which will be described in Part III”. However, Part III of Miyas monograph of the
<taxonomicName id="C5DD4D4EFFCEFFDBFEA504BAFDD06BC2" authorityName="Rafinesque" authorityYear="1815" box="[353,468,151,177]" class="Malacostraca" family="Alpheidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="401" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
of
<collectingCountry id="7ACA765DFFCEFFDBFE3304BAFE336BC2" box="[503,567,151,177]" name="Japan" pageId="8" pageNumber="401">Japan</collectingCountry>
was never published and Miyas name is therefore a
<emphasis id="30A9EADFFFCEFFDBFB4404BAF9266BC2" box="[1152,1314,151,177]" italics="true" pageId="8" pageNumber="401">Nomen nudum</emphasis>
(
<bibRefCitation id="664C4B3CFFCEFFDBFAF404BAFD566BA6" author="De Grave, S. &amp; Fransen, C. H. J. M." pageId="8" pageNumber="401" pagination="195 - 589" refId="ref9512" refString="De Grave, S. &amp; Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85, 195 - 589." type="journal article" year="2011">De Grave &amp; Fransen 2011</bibRefCitation>
). It remains unknown from which locality (or localities) Miyas material came from and if it was collected in shallow or in deep water. The only character mentioned by
<bibRefCitation id="664C4B3CFFCEFFDBFC6904F2F83C6B8A" author="Miya, Y." box="[941,1080,223,249]" pageId="8" pageNumber="401" pagination="103 - 195" refId="ref10255" refString="Miya, Y. (1974) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part II. Publications from the Amakusa Marine Biology Laboratory, 3, 103 - 195, pls. 15 - 31." type="journal article" year="1974">Miya (1974)</bibRefCitation>
for
<taxonomicName id="C5DD4D4EFFCEFFDBFBA104F2F8DE6B8A" authorityName="Anker" authorityYear="2020" box="[1125,1242,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="401" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="30A9EADFFFCEFFDBFBA104F2F8DE6B8A" box="[1125,1242,223,249]" italics="true" pageId="8" pageNumber="401">A. miyakei</emphasis>
</taxonomicName>
was the presence of orbital teeth, which was used by
<bibRefCitation id="664C4B3CFFCEFFDBFDEC052EFEB96A6E" author="Bruce, A. J." box="[552,701,259,285]" pageId="8" pageNumber="401" pagination="15 - 28" refId="ref9099" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28." type="journal article" year="1994">Bruce (1994)</bibRefCitation>
in the first couplet of his key to the
<taxonomicName id="C5DD4D4EFFCEFFDBFB93052EF8FC6A6E" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[1111,1272,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="401" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="30A9EADFFFCEFFDBFB93052EF8FC6A6E" box="[1111,1272,259,285]" italics="true" pageId="8" pageNumber="401">A. brevirostris</emphasis>
</taxonomicName>
group. Miyas invalid name is here tentatively listed in the synonymy of
<taxonomicName id="C5DD4D4EFFCEFFDBFCF7050AFFF66A32" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[819,1010,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="401" phylum="Arthropoda" rank="species" species="alaincrosnieri" status="sp. nov.">
<emphasis id="30A9EADFFFCEFFDBFCF7050AFFF66A32" box="[819,1010,295,321]" italics="true" pageId="8" pageNumber="401">A. alaincrosnieri</emphasis>
</taxonomicName>
<emphasis id="30A9EADFFFCEFFDBFC3F050AF83F6A32" bold="true" box="[1019,1083,295,321]" pageId="8" pageNumber="401">
<taxonomicNameLabel id="2B9A57A4FFCEFFDBFC3F050AF83F6A32" box="[1019,1083,295,321]" pageId="8" pageNumber="401" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, although its true identity may never be uncovered.
</paragraph>
</subSubSection>
</treatment>
</document>