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<mods:title id="7815512EC47090C3A9D48747EEF24EFA">Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="8EEF0F591F61C34EAEF7CAFE335FB126">Meng, Guanliang</mods:namePart>
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<mods:namePart id="8A0B5A4542EDB159F573945AC3E3FF15">Astrin, Jonas</mods:namePart>
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<taxonomicName id="C5C6E6C6FFF7A67AFDA2FA0E61A7FA37" ID-CoL="4MHKG" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[618,969,1414,1441]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF7A67AFDA2FA0E6142FA37" bold="true" box="[618,812,1414,1441]" italics="true" pageId="22" pageNumber="23">Priscula gularis</emphasis>
Simon, 1893
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<treatmentCitation id="8367BB54FFF7A67AFF75FA40600BFA74" author="Simon E." box="[189,613,1480,1506]" page="319" pageId="22" pageNumber="23" year="1893">
<taxonomicName id="C5C6E6C6FFF7A67AFF75FA40600BFA74" ID-CoL="4MHKG" authority="Simon, 1893 a: 319" authorityName="Simon" authorityPageNumber="319" authorityYear="1893" box="[189,613,1480,1506]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF7A67AFF75FA406315FA74" box="[189,379,1480,1506]" italics="true" pageId="22" pageNumber="23">Priscula gularis</emphasis>
<bibRefCitation id="6657E0B4FFF7A67AFE4AFA40600BFA74" author="Simon E." box="[386,613,1480,1506]" pageId="22" pageNumber="23" pagination="299 - 330" refId="ref25255" refString="Simon E. 1893 a. Descriptions d'especes et de genres nouveaux de l'ordre des Araneae. Annales de la Societe Entomologique de France 62: 299 - 330." type="journal article" year="1893">Simon, 1893a: 319</bibRefCitation>
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.
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<taxonomicName id="C5C6E6C6FFF7A67AFF75FA636315F993" ID-CoL="4MHKG" authorityName="Simon" authorityYear="1893" box="[189,379,1515,1541]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gularis">Priscula gularis</taxonomicName>
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<treatmentCitation id="8367BB54FFF7A67AFE50FA63601FF993" author="Simon E." box="[408,625,1515,1541]" page="477" pageId="22" pageNumber="23" year="1893">
<bibRefCitation id="6657E0B4FFF7A67AFE50FA63601FF993" author="Simon E." box="[408,625,1515,1541]" pageId="22" pageNumber="23" pagination="256 - 488" refId="ref25285" refString="Simon E. 1893 b. Histoire Naturelle des Araignees. 2 nd edition, 1 (2): 256 - 488. Roret, Paris." type="journal article" year="1893">Simon 1893b: 477</bibRefCitation>
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<treatmentCitation id="8367BB54FFF7A67AFD55FA636109F990" author="Huber B. A." box="[669,871,1515,1542]" page="595" pageId="22" pageNumber="23" year="1997">
<bibRefCitation id="6657E0B4FFF7A67AFD55FA636109F990" author="Huber B. A." box="[669,871,1515,1542]" pageId="22" pageNumber="23" pagination="573 - 612" refId="ref24298" refString="Huber B. A. 1997. Redescriptions of Eugene Simon's neotropical pholcids (Araneae, Pholcidae). Zoosystema 19 (3): 573 - 612." type="journal article" year="1997">Huber 1997: 595</bibRefCitation>
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, figs 1719; 2000: 129, figs 495500.
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<taxonomicName id="C5C6E6C6FFF7A67AFF75F98663C6F9BE" authority="(Simon, 1893)" baseAuthorityName="Simon" baseAuthorityYear="1893" box="[189,424,1550,1576]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF7A67AFF75F98663C6F9BE" box="[189,424,1550,1576]" italics="true" pageId="22" pageNumber="23">Physocyclus gularis</emphasis>
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<bibRefCitation id="6657E0B4FFF7A67AFE0EF98660F7F9BE" author="Brignoli P. M." box="[454,665,1550,1576]" pageId="22" pageNumber="23" pagination="90 - 100" refId="ref23832" refString="Brignoli P. M. 1981. Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae). Bulletin of the American Museum of Natural History 170: 90 - 100." type="journal article" year="1981">Brignoli 1981: 96</bibRefCitation>
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, figs 810, 25.
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<emphasis id="30B24157FFF7A67AFF75F9D86341F9FC" bold="true" box="[189,303,1616,1642]" pageId="22" pageNumber="23">Remarks</emphasis>
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Our sample of sequenced material includes several specimens that appear very close to or even conspecific with the
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species
<taxonomicName id="C5C6E6C6FFF7A67AFE7DF92A604DF92D" box="[437,547,1697,1723]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF7A67AFE7DF92A604DF92D" box="[437,547,1697,1723]" italics="true" pageId="22" pageNumber="23">P. gularis</emphasis>
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. This species or species complex is badly in need of revision but this will require restudying the
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material (a loan request remained unanswered) and more focused collecting. Here, we only point out the problematic identity of the
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species, explaining the
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<emphasis id="30B24157FFF7A67AFB1CF9606749F894" box="[1236,1319,1768,1794]" italics="true" pageId="22" pageNumber="23">gularis</emphasis>
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?” and “
<taxonomicName id="C5C6E6C6FFF7A67AFF01F883632CF8B3" box="[201,322,1803,1829]" class="Arachnida" family="Pholcidae" genus="Physocyclus" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gularis">
cf.
<emphasis id="30B24157FFF7A67AFF27F883632CF8B3" box="[239,322,1803,1829]" italics="true" pageId="22" pageNumber="23">gularis</emphasis>
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” in
<figureCitation id="9AFD81C0FFF7A67AFEBDF88363D5F8B3" box="[373,443,1803,1829]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[186,1404,890,1781]" captionTargetId="figure-231@6.[183,1404,890,1781]" captionTargetPageId="6" captionText="Fig. 1; continued in Fig. 2. Relationships of Priscula derived from an unpartitioned analysis of molecular data using IQ-Tree and the ClipKIT kpic-smart-gap trimming strategy. Numbers on the branches are SH-aLRT supports (%). The tree shows only the ingroup. For the complete tree (incl. outgroups), and for clade support using different partitioning and alignment strategies, see Supplementary Figures S1S3. Photos on the right, from top: Priscula llaviucu Huber sp. nov.; P. espejoi Huber sp. nov.; P. binghamae (Chamberlin, 1916); P. pastaza Huber sp. nov.; P. “Tri18-11”." figureDoi="http://doi.org/10.5281/zenodo.10462862" httpUri="https://zenodo.org/record/10462862/files/figure.png" pageId="22" pageNumber="23">Fig. 1</figureCitation>
.
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No detailed illustrations are available of the type material of
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<emphasis id="30B24157FFF7A67AFC53F8C56664F8F0" box="[923,1034,1868,1894]" italics="true" pageId="22" pageNumber="23">P. gularis</emphasis>
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. The
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and the female
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originate from “Quito” (without further specification), but the drawings in
<bibRefCitation id="6657E0B4FFF7A67AFB0AF8F86717F81C" author="Brignoli P. M." box="[1218,1401,1904,1930]" pageId="22" pageNumber="23" pagination="90 - 100" refId="ref23832" refString="Brignoli P. M. 1981. Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae). Bulletin of the American Museum of Natural History 170: 90 - 100." type="journal article" year="1981">Brignoli (1981)</bibRefCitation>
and
<bibRefCitation id="6657E0B4FFF7A67AFF24F81B63E4F83B" author="Huber B. A." box="[236,394,1939,1965]" pageId="22" pageNumber="23" pagination="573 - 612" refId="ref24298" refString="Huber B. A. 1997. Redescriptions of Eugene Simon's neotropical pholcids (Araneae, Pholcidae). Zoosystema 19 (3): 573 - 612." type="journal article" year="1997">Huber (1997)</bibRefCitation>
are based on Eugène Simons specimens from “Nariguel” (or “Narigual”), a locality we were not able to identify.
<bibRefCitation id="6657E0B4FFF7A67AFE2FF83E60E1F846" author="Simon E." box="[487,655,1974,2000]" pageId="22" pageNumber="23" pagination="299 - 330" refId="ref25255" refString="Simon E. 1893 a. Descriptions d'especes et de genres nouveaux de l'ordre des Araneae. Annales de la Societe Entomologique de France 62: 299 - 330." type="journal article" year="1893">Simon (1893a</bibRefCitation>
, b) never mentioned these specimens, meaning that they cannot be considered types (contra
<bibRefCitation id="6657E0B4FFF6A67BFDDDFE8160AFFEB5" author="Brignoli P. M." box="[533,705,265,291]" pageId="23" pageNumber="24" pagination="90 - 100" refId="ref23832" refString="Brignoli P. M. 1981. Studies on the Pholcidae, I. Notes on the genera Artema and Physocyclus (Araneae). Bulletin of the American Museum of Natural History 170: 90 - 100." type="journal article" year="1981">Brignoli 1981</bibRefCitation>
). The drawings in
<bibRefCitation id="6657E0B4FFF6A67BFC61FE816622FEB5" author="Huber B. A." box="[937,1100,265,291]" pageId="23" pageNumber="24" pagination="1 - 348" refId="ref24327" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 &lt;0001: NWPSAP&gt; 2.0. CO; 2" type="journal article" year="2000">Huber (2000)</bibRefCitation>
are based on specimens from Baños (~
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S of Quito), which were collected in 1937 and which appear identical to a newly collected male from Desierto de Palmira (code of sequenced juvenile from this locality: EB083). We thus speculate that the male from Desierto de Palmira and the sequenced juvenile EB083 might be true
<taxonomicName id="C5C6E6C6FFF6A67BFF75FE1F6343FE26" box="[189,301,406,432]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF6A67BFF75FE1F6343FE26" box="[189,301,406,432]" italics="true" pageId="23" pageNumber="24">P. gularis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="02799D45FFF6A67BFF75FE5F63B7FCC7" blockId="23.[186,1399,471,849]" pageId="23" pageNumber="24">
The sequenced juvenile from Desierto de Palmira is genetically closest to a female specimen from between Guano and Ilapo (EB020; distance 5.5%) but we have no male from this locality. Morphologically, the Desierto de Palmira male is also very similar to a newly collected male from near Zhud (EB077), but in this case, the genetic distance is already at 7.2%. Surprisingly, a newly collected male from Baños (EB084) is morphologically quite clearly different from the old (1937) Baños males, but the genetic distances to the previous specimens are almost the same as among those specimens, i.e. ~78%. Similar distances were also found between these specimens and a further “
<taxonomicName id="C5C6E6C6FFF6A67BFC04FD226628FD52" box="[972,1094,682,708]" class="Arachnida" family="Pholcidae" genus="Physocyclus" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
cf.
<emphasis id="30B24157FFF6A67BFC3BFD226628FD52" box="[1011,1094,682,708]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
” specimen (EB127) from near El Ángel (~
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NE of Quito), a further locality from which we have no male. Genetic distances between all these
<taxonomicName id="C5C6E6C6FFF6A67BFE4FFD796389FC9D" box="[391,487,753,779]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF6A67BFE5CFD796389FC9D" box="[404,487,753,779]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
?” and “
<taxonomicName id="C5C6E6C6FFF6A67BFD89FD7960D8FC9D" box="[577,694,753,779]" class="Arachnida" family="Pholcidae" genus="Physocyclus" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
cf.
<emphasis id="30B24157FFF6A67BFDABFD7960D8FC9D" box="[611,694,753,779]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
” specimens and the morphologically very distinct
<taxonomicName id="C5C6E6C6FFF6A67BFB33FD7A6719FC9D" authority="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera, 2023" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[1275,1399,753,779]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="llaviucu" status="sp. nov.">
<emphasis id="30B24157FFF6A67BFB33FD7A6719FC9D" box="[1275,1399,753,779]" italics="true" pageId="23" pageNumber="24">P. llaviucu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFF6A67BFF75FC9D6379FCB9" box="[189,279,789,815]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
are only slightly higher (8.79.2%) suggesting that several species might be represented by our “
<taxonomicName id="C5C6E6C6FFF6A67BFF01FCBF632CFCC7" box="[201,322,823,849]" class="Arachnida" family="Pholcidae" genus="Physocyclus" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
cf.
<emphasis id="30B24157FFF6A67BFF27FCBF632CFCC7" box="[239,322,823,849]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
” specimens.
</paragraph>
<paragraph id="02799D45FFF6A67BFF75FCF1615DFBD5" blockId="23.[189,1399,888,1091]" pageId="23" pageNumber="24">
Two further closely related potential species (“
<emphasis id="30B24157FFF6A67BFD2BFCF16094FC04" box="[739,762,889,914]" italics="true" pageId="23" pageNumber="24">P.</emphasis>
Dup55” and “
<emphasis id="30B24157FFF6A67BFC6CFCF161D4FC04" box="[932,954,889,914]" italics="true" pageId="23" pageNumber="24">P.</emphasis>
Ecu5”) are included in our molecular dataset and shown in the distribution map (
<figureCitation id="9AFD81C0FFF6A67BFD08FC14614EFC20" box="[704,800,924,950]" captionStart="Fig" captionStartId="9.[189,232,1821,1847]" captionTargetBox="[183,1405,1129,1775]" captionTargetId="figure-307@9.[184,1405,1129,1774]" captionTargetPageId="9" captionText="Fig. 4. Known distribution of Priscula in Ecuador. A. Representatives of the gularis group. Circles: P. gularis (TL=type locality) and specimens treated herein as “P. gularis?” and “P. cf. gularis”. Squares: other Ecuadorian representatives of the gularis group. B. Representatives of the andinensis group (red) and binghamae group (blue) in Ecuador. The background colors indicate biogeographic regions of Ecuador (as far as relevant for Priscula, i.e., Andean regions from western foothill forests to eastern foothill forests), taken from Torres-Carvajal et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10462874" httpUri="https://zenodo.org/record/10462874/files/figure.png" pageId="23" pageNumber="24">Fig. 4A</figureCitation>
), but not formally described because no males are available. The CO1 K2Pdistance between the two sequenced specimens was 7.8%, and distances to other specimens of the
<taxonomicName id="C5C6E6C6FFF6A67BFE1AFC6A604BFC6A" box="[466,549,994,1020]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF6A67BFE1AFC6A604BFC6A" box="[466,549,994,1020]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
group ranged from 7.612.3%. The distribution map (
<figureCitation id="9AFD81C0FFF6A67BFB62FC6A6769FC6A" box="[1194,1287,994,1020]" captionStart="Fig" captionStartId="9.[189,232,1821,1847]" captionTargetBox="[183,1405,1129,1775]" captionTargetId="figure-307@9.[184,1405,1129,1774]" captionTargetPageId="9" captionText="Fig. 4. Known distribution of Priscula in Ecuador. A. Representatives of the gularis group. Circles: P. gularis (TL=type locality) and specimens treated herein as “P. gularis?” and “P. cf. gularis”. Squares: other Ecuadorian representatives of the gularis group. B. Representatives of the andinensis group (red) and binghamae group (blue) in Ecuador. The background colors indicate biogeographic regions of Ecuador (as far as relevant for Priscula, i.e., Andean regions from western foothill forests to eastern foothill forests), taken from Torres-Carvajal et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10462874" httpUri="https://zenodo.org/record/10462874/files/figure.png" pageId="23" pageNumber="24">Fig. 4A</figureCitation>
) shows a further potential species, “
<emphasis id="30B24157FFF6A67BFE3EFB8E6062FB89" box="[502,524,1030,1055]" italics="true" pageId="23" pageNumber="24">P.</emphasis>
Dup140” that we tentatively assign to the
<taxonomicName id="C5C6E6C6FFF6A67BFBC1FB8D6632FB89" box="[1033,1116,1029,1055]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF6A67BFBC1FB8D6632FB89" box="[1033,1116,1029,1055]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
group, but we were not able to extract DNA from the single available female.
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFF6A674FF75FBE26167FDCC" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="02799D45FFF6A67BFF75FBE26126FB12" blockId="23.[189,840,1130,1156]" box="[189,840,1130,1156]" pageId="23" pageNumber="24">
<heading id="59312A29FFF6A67BFF75FBE26126FB12" bold="true" box="[189,840,1130,1156]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="30B24157FFF6A67BFF75FBE26126FB12" bold="true" box="[189,840,1130,1156]" pageId="23" pageNumber="24">New material examined (identity unclear, see above)</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFF6A67BFF22FB106305FB24" blockId="23.[189,1399,1176,1414]" box="[234,363,1176,1203]" pageId="23" pageNumber="24">
<emphasis id="30B24157FFF6A67BFF22FB106305FB24" bold="true" box="[234,363,1176,1203]" pageId="23" pageNumber="24">
<taxonomicName id="C5C6E6C6FFF6A67BFF22FB106320FB25" box="[234,334,1176,1203]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
<emphasis id="30B24157FFF6A67BFF31FB106320FB25" bold="true" box="[249,334,1176,1203]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
?”
</emphasis>
</paragraph>
<paragraph id="02799D45FFF6A67BFF75FB34632FFA10" blockId="23.[189,1399,1176,1414]" pageId="23" pageNumber="24">
<materialsCitation id="B2AE9718FFF6A67BFF75FB346721FB6C" collectedFrom="under rocks" collectingDate="2017-02-27" collectionCode="ZMH" collectorName="E. Tapia &amp; A. Tapia &amp; I. Tapia" country="Ecuador" elevation="3250" latitude="-2.0322" location="Desierto de Palmira" longLatPrecision="7" longitude="-78.744" municipality="Guamote" pageId="23" pageNumber="24" specimenCode="A2590" specimenCount="1" specimenCount-male="1" stateProvince="Chimborazo">
<collectingCountry id="7AD1DDD5FFF6A67BFF75FB346325FB40" box="[189,331,1212,1238]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
<emphasis id="30B24157FFF6A67BFEA7FB346065FB40" bold="true" box="[367,523,1212,1238]" pageId="23" pageNumber="24">
<collectingRegion id="C00253A7FFF6A67BFEA7FB346065FB40" box="[367,523,1212,1238]" country="Ecuador" name="Chimborazo" pageId="23" pageNumber="24">Chimborazo</collectingRegion>
</emphasis>
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;
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,
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;
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,
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a.s.l.
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;
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, deserted region;
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;
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,
<collectorName id="AF33F893FFF6A67BFCB0FB5761B1FB6F" box="[888,991,1247,1273]" pageId="23" pageNumber="24">A. Tapia</collectorName>
, and
<collectorName id="AF33F893FFF6A67BFBD5FB686618FB6F" box="[1053,1142,1247,1274]" pageId="23" pageNumber="24">I. Tapia</collectorName>
leg.;
<collectionCode id="64D70580FFF6A67BFB70FB686697FB6C" box="[1208,1273,1248,1274]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="23" pageNumber="24" type="Museum">ZMH</collectionCode>
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, in pure ethanol;
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;
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a.s.l.
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;
<collectedFrom id="B1DADEB8FFF6A67BFB93FA8A637AFAA9" pageId="23" pageNumber="24">under dead wood on the ground</collectedFrom>
;
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;
<collectorName id="AF33F893FFF6A67BFE0CFAAE6024FAA9" box="[452,586,1317,1344]" pageId="23" pageNumber="24">B.A. Huber</collectorName>
and
<collectorName id="AF33F893FFF6A67BFD49FAAE616CFAD6" box="[641,770,1318,1344]" pageId="23" pageNumber="24">M. Herrera</collectorName>
leg;
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Ecu167
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;
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,
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;
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,
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;
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<quantity id="C53E30A0FFF6A67BFE15FAC06059FAF5" box="[477,567,1352,1379]" metricMagnitude="3" metricUnit="m" metricValue="2.68" pageId="23" pageNumber="24" unit="m" value="2680.0">2680 m</quantity>
a.s.l.
</elevation>
;
<date id="7678BB85FFF6A67BFD4DFAC16179FAF5" box="[645,791,1353,1379]" pageId="23" pageNumber="24" value="2020-03-07">
<collectingDate id="663C426DFFF6A67BFD4DFAC16179FAF5" box="[645,791,1353,1379]" pageId="23" pageNumber="24" value="2020-03-07">7 Mar. 2020</collectingDate>
</date>
;
<collectorName id="AF33F893FFF6A67BFCECFAC161C5FAF5" box="[804,939,1353,1379]" pageId="23" pageNumber="24">N. Dupérré</collectorName>
,
<collectorName id="AF33F893FFF6A67BFC7EFAC16654FAF5" box="[950,1082,1353,1379]" pageId="23" pageNumber="24">A.A. Tapia</collectorName>
, and
<collectorName id="AF33F893FFF6A67BFBB0FAC1669DFAF5" box="[1144,1267,1353,1379]" pageId="23" pageNumber="24">E.E. Tapia</collectorName>
leg.;
<specimenCode id="5260353EFFF6A67BFAFDFAC16353FA10" pageId="23" pageNumber="24">ZMH A148889</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFF6A67BFF22FA2563ECFA51" blockId="23.[189,1399,1453,1797]" box="[234,386,1453,1480]" pageId="23" pageNumber="24">
<emphasis id="30B24157FFF6A67BFF22FA2563ECFA51" bold="true" box="[234,386,1453,1480]" pageId="23" pageNumber="24">
<taxonomicName id="C5C6E6C6FFF6A67BFF31FA25631DFA5E" box="[249,371,1453,1480]" class="Arachnida" family="Pholcidae" genus="Physocyclus" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gularis">
cf.
<emphasis id="30B24157FFF6A67BFED6FA25631DFA5E" bold="true" box="[286,371,1453,1480]" italics="true" pageId="23" pageNumber="24">gularis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="02799D45FFF6A67BFF75FA596012F892" blockId="23.[189,1399,1453,1797]" pageId="23" pageNumber="24">
<materialsCitation id="B2AE9718FFF6A67BFF75FA596747F998" collectedFrom="degraded forest in ravine" collectingDate="2021-10-02" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="3240" latitude="0.6178" location="near El Angel" longLatPrecision="7" longitude="-77.9261" pageId="23" pageNumber="24" specimenCode="MECN-ARAC-31-T" specimenCount="1" specimenCount-female="1" stateProvince="Carchi">
<collectingCountry id="7AD1DDD5FFF6A67BFF75FA596325FA7D" box="[189,331,1489,1515]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
<emphasis id="30B24157FFF6A67BFEA2FA5863AEFA7C" bold="true" box="[362,448,1488,1514]" pageId="23" pageNumber="24">
<collectingRegion id="C00253A7FFF6A67BFEA2FA5863AEFA7C" box="[362,448,1488,1514]" country="Ecuador" name="Carchi" pageId="23" pageNumber="24">Carchi</collectingRegion>
</emphasis>
<specimenCount id="14C056CCFFF6A67BFE13FA596067FA7D" box="[475,521,1489,1515]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
;
<location id="0719CB9EFFF6A67BFDD1FA5960ACFA7B" LSID="urn:lsid:plazi:treatment:8A6F2C53FFF7A674FDA2FA0E6167FDCC:0719CB9EFFF6A67BFDD1FA5960ACFA7B" box="[537,706,1488,1517]" country="Ecuador" latitude="0.6178" longLatPrecision="7" longitude="-77.9261" name="near El Angel" pageId="23" pageNumber="24" stateProvince="Carchi">near El Ángel</location>
;
<geoCoordinate id="67F2FB82FFF6A67BFD07FA586122FA7D" box="[719,844,1488,1515]" degrees="0.6178" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="0.6178">0.6178° N</geoCoordinate>
,
<geoCoordinate id="67F2FB82FFF6A67BFC91FA586184FA7D" box="[857,1002,1488,1515]" degrees="77.9261" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="-77.9261">77.9261° W</geoCoordinate>
;
<elevation id="89EB7A76FFF6A67BFC3FFA5966FCFA7C" box="[1015,1170,1488,1515]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="23" pageNumber="24" unit="m" value="3240.0">
<quantity id="C53E30A0FFF6A67BFC3FFA59663FFA7D" box="[1015,1105,1489,1515]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="23" pageNumber="24" unit="m" value="3240.0">3240 m</quantity>
a.s.l.
</elevation>
;
<collectedFrom id="B1DADEB8FFF6A67BFB68FA586364F998" pageId="23" pageNumber="24">degraded forest in ravine</collectedFrom>
;
<date id="7678BB85FFF6A67BFEDEFA7C63CDF998" box="[278,419,1524,1550]" pageId="23" pageNumber="24" value="2021-10-02">
<collectingDate id="663C426DFFF6A67BFEDEFA7C63CDF998" box="[278,419,1524,1550]" pageId="23" pageNumber="24" value="2021-10-02">2 Oct. 2021</collectingDate>
</date>
;
<collectorName id="AF33F893FFF6A67BFE78FA7C6056F998" box="[432,568,1524,1550]" pageId="23" pageNumber="24">B.A. Huber</collectorName>
and
<collectorName id="AF33F893FFF6A67BFDB9FA7C609AF998" box="[625,756,1524,1550]" pageId="23" pageNumber="24">M. Herrera</collectorName>
leg.;
<specimenCode id="5260353EFFF6A67BFCFEFA7C6656F998" box="[822,1080,1524,1550]" pageId="23" pageNumber="24">MECNARAC31T</specimenCode>
, in
<collectionCode id="64D70580FFF6A67BFBAAFA7C66DDF998" box="[1122,1203,1524,1550]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ar 24091
</materialsCitation>
<materialsCitation id="B2AE9718FFF6A67BFA8BFA7C663EF9A7" collectedFrom="degraded forest in ravine" collectingDate="2021-10-02" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="3240" latitude="0.6178" location="Carchi" longLatPrecision="7" longitude="-77.9261" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" stateProvince="Carchi">
<specimenCount id="14C056CCFFF6A67BFA8BFA7C6701F998" box="[1347,1391,1524,1550]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
,
<specimenCount id="14C056CCFFF6A67BFF75F99F636CF9A7" box="[189,258,1559,1585]" count="1" pageId="23" pageNumber="24" type="generic">1 juv.</specimenCount>
, in pure ethanol; same collection data as for preceding;
<collectionCode id="64D70580FFF6A67BFC54F99F6183F9A7" box="[924,1005,1559,1585]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ecu222
</materialsCitation>
<materialsCitation id="B2AE9718FFF6A67BFBA7F99F6098F9E1" collectingDate="2021-09-22" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2960" latitude="-2.479" location="Zhud" longLatPrecision="7" longitude="-78.9978" pageId="23" pageNumber="24" specimenCode="MECN-ARAC-32-T" specimenCount="1" specimenCount-male="1" stateProvince="Canar">
<emphasis id="30B24157FFF6A67BFBA7F99F66D1F9A7" bold="true" box="[1135,1215,1559,1585]" pageId="23" pageNumber="24">
<collectingRegion id="C00253A7FFF6A67BFBA7F99F66D1F9A7" box="[1135,1215,1559,1585]" country="Ecuador" name="Canar" pageId="23" pageNumber="24">Cañar</collectingRegion>
</emphasis>
<specimenCount id="14C056CCFFF6A67BFB13F99F6765F9A7" box="[1243,1291,1559,1585]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; S
<location id="0719CB9EFFF6A67BFAF9F99F671CF9A7" LSID="urn:lsid:plazi:treatment:8A6F2C53FFF7A674FDA2FA0E6167FDCC:0719CB9EFFF6A67BFAF9F99F671CF9A7" box="[1329,1394,1559,1585]" country="Ecuador" latitude="-2.479" longLatPrecision="7" longitude="-78.9978" name="Zhud" pageId="23" pageNumber="24" stateProvince="Canar">Zhud</location>
, ravine above road;
<geoCoordinate id="67F2FB82FFF6A67BFE6DF9B26073F9C2" box="[421,541,1594,1620]" degrees="2.4790" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="-2.479">2.4790° S</geoCoordinate>
,
<geoCoordinate id="67F2FB82FFF6A67BFDE5F9B260D1F9C3" box="[557,703,1594,1621]" degrees="78.9978" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="-78.9978">78.9978° W</geoCoordinate>
;
<elevation id="89EB7A76FFF6A67BFD05F9B26104F9C2" box="[717,874,1594,1621]" metricMagnitude="3" metricUnit="m" metricValue="2.96" pageId="23" pageNumber="24" unit="m" value="2960.0">
<quantity id="C53E30A0FFF6A67BFD05F9B26146F9C3" box="[717,808,1594,1621]" metricMagnitude="3" metricUnit="m" metricValue="2.96" pageId="23" pageNumber="24" unit="m" value="2960.0">2960 m</quantity>
a.s.l.
</elevation>
;
<date id="7678BB85FFF6A67BFCB2F9B26672F9C2" box="[890,1052,1594,1620]" pageId="23" pageNumber="24" value="2021-09-22">
<collectingDate id="663C426DFFF6A67BFCB2F9B26672F9C2" box="[890,1052,1594,1620]" pageId="23" pageNumber="24" value="2021-09-22">22 Sep. 2021</collectingDate>
</date>
;
<collectorName id="AF33F893FFF6A67BFBE2F9B266DAF9C2" box="[1066,1204,1594,1620]" pageId="23" pageNumber="24">B.A. Huber</collectorName>
and
<collectorName id="AF33F893FFF6A67BFB3AF9B36719F9C3" box="[1266,1399,1595,1621]" pageId="23" pageNumber="24">M. Herrera</collectorName>
leg.;
<specimenCode id="5260353EFFF6A67BFF32F9D56392F9E1" box="[250,508,1629,1655]" pageId="23" pageNumber="24">MECNARAC32T</specimenCode>
, in
<collectionCode id="64D70580FFF6A67BFDE3F9D66012F9EE" box="[555,636,1630,1656]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ar 24092
</materialsCitation>
<materialsCitation id="B2AE9718FFF6A67BFCDCF9D56395F90C" collectingDate="2021-09-22" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2960" latitude="-2.479" location="Canar" longLatPrecision="7" longitude="-78.9978" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Canar">
<specimenCount id="14C056CCFFF6A67BFCDCF9D5610EF9E1" box="[788,864,1629,1655]" pageId="23" pageNumber="24">2 juvs</specimenCount>
, in pure ethanol; same collection data as for preceding;
<collectionCode id="64D70580FFF6A67BFE8DF90963F8F90D" box="[325,406,1665,1691]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ecu162
</materialsCitation>
<materialsCitation id="B2AE9718FFF6A67BFDEAF90961FEF977" collectedFrom="degraded forest along river" collectingDate="2021-09-24" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2100" latitude="-1.4134" location="Banos" longLatPrecision="7" longitude="-78.434" pageId="23" pageNumber="24" specimenCode="MECN-ARAC-33-T" specimenCount="1" specimenCount-male="1" stateProvince="Tungurahua">
<emphasis id="30B24157FFF6A67BFDEAF90960D3F90D" bold="true" box="[546,701,1665,1691]" pageId="23" pageNumber="24">
<collectingRegion id="C00253A7FFF6A67BFDEAF90960D3F90D" box="[546,701,1665,1691]" country="Ecuador" name="Tungurahua" pageId="23" pageNumber="24">Tungurahua</collectingRegion>
</emphasis>
<specimenCount id="14C056CCFFF6A67BFD28F909617DF90D" box="[736,787,1665,1691]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; S
<location id="0719CB9EFFF6A67BFC88F90961E3F90D" LSID="urn:lsid:plazi:treatment:8A6F2C53FFF7A674FDA2FA0E6167FDCC:0719CB9EFFF6A67BFC88F90961E3F90D" box="[832,909,1665,1691]" country="Ecuador" latitude="-1.4134" longLatPrecision="7" longitude="-78.434" name="Banos" pageId="23" pageNumber="24" stateProvince="Tungurahua">Baños</location>
;
<geoCoordinate id="67F2FB82FFF6A67BFC56F9096676F90D" box="[926,1048,1665,1691]" degrees="1.4134" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="-1.4134">1.4134° S</geoCoordinate>
,
<geoCoordinate id="67F2FB82FFF6A67BFBE1F90966D0F90D" box="[1065,1214,1665,1691]" degrees="78.4340" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="-78.434">78.4340° W</geoCoordinate>
;
<elevation id="89EB7A76FFF6A67BFB06F909671EF90C" box="[1230,1392,1664,1691]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="24" unit="m" value="2100.0">
<quantity id="C53E30A0FFF6A67BFB06F9096742F90D" box="[1230,1324,1665,1691]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="24" unit="m" value="2100.0">2100 m</quantity>
a.s.l.
</elevation>
;
<collectedFrom id="B1DADEB8FFF6A67BFF75F92C606DF928" box="[189,515,1700,1726]" pageId="23" pageNumber="24">degraded forest along river</collectedFrom>
;
<date id="7678BB85FFF6A67BFDD9F92C60DCF928" box="[529,690,1700,1726]" pageId="23" pageNumber="24" value="2021-09-24">
<collectingDate id="663C426DFFF6A67BFDD9F92C60DCF928" box="[529,690,1700,1726]" pageId="23" pageNumber="24" value="2021-09-24">24 Sep. 2021</collectingDate>
</date>
;
<collectorName id="AF33F893FFF6A67BFD08F92C6127F928" box="[704,841,1700,1726]" pageId="23" pageNumber="24">B.A. Huber</collectorName>
and
<collectorName id="AF33F893FFF6A67BFC4EF92C6665F928" box="[902,1035,1700,1726]" pageId="23" pageNumber="24">M. Herrera</collectorName>
leg.;
<specimenCode id="5260353EFFF6A67BFB98F92C673CF928" box="[1104,1362,1700,1726]" pageId="23" pageNumber="24">MECNARAC33T</specimenCode>
, in
<collectionCode id="64D70580FFF6A67BFF75F94F6360F977" box="[189,270,1735,1761]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ar 24093 and
<collectionCode id="64D70580FFF6A67BFE09F94F607CF977" box="[449,530,1735,1761]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ecu168 (2 legs in pure ethanol)
</materialsCitation>
<materialsCitation id="B2AE9718FFF6A67BFC63F94F6016F892" collectedFrom="degraded forest along river" collectingDate="2021-09-24" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2100" latitude="-1.4134" location="Tungurahua" longLatPrecision="7" longitude="-78.434" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Tungurahua">
<specimenCount id="14C056CCFFF6A67BFC63F94F619EF977" box="[939,1008,1735,1761]" count="1" pageId="23" pageNumber="24" type="generic">1 juv.</specimenCount>
, in pure ethanol; same collection data as for preceding;
<collectionCode id="64D70580FFF6A67BFE0AF963607DF893" box="[450,531,1771,1797]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ecu168
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFF6A67BFF22F8CA6329F8CA" blockId="23.[189,1399,1858,1990]" box="[234,327,1858,1884]" pageId="23" pageNumber="24">
<emphasis id="30B24157FFF6A67BFF22F8CA6329F8CA" bold="true" box="[234,327,1858,1884]" pageId="23" pageNumber="24">“Ecu5”</emphasis>
</paragraph>
<paragraph id="02799D45FFF6A67BFF75F8ED6657F850" blockId="23.[189,1399,1858,1990]" pageId="23" pageNumber="24">
<materialsCitation id="B2AE9718FFF6A67BFF75F8ED6725F835" collectedFrom="forest at roadside" collectingDate="2021-09-15" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1900" latitude="-0.2327" location="San Juan and Chiriboga" longLatPrecision="7" longitude="-78.7497" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha">
<collectingCountry id="7AD1DDD5FFF6A67BFF75F8ED6325F8E9" box="[189,331,1893,1919]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
<emphasis id="30B24157FFF6A67BFEA7F8ED6385F8E9" bold="true" box="[367,491,1893,1919]" pageId="23" pageNumber="24">
<collectingRegion id="C00253A7FFF6A67BFEA7F8ED6385F8E9" box="[367,491,1893,1919]" country="Ecuador" name="Pichincha" pageId="23" pageNumber="24">Pichincha</collectingRegion>
</emphasis>
<specimenCount id="14C056CCFFF6A67BFDC2F8ED6055F816" box="[522,571,1893,1920]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; between
<location id="0719CB9EFFF6A67BFD71F8ED61B3F8E9" LSID="urn:lsid:plazi:treatment:8A6F2C53FFF7A674FDA2FA0E6167FDCC:0719CB9EFFF6A67BFD71F8ED61B3F8E9" box="[697,989,1893,1920]" country="Ecuador" latitude="-0.2327" longLatPrecision="7" longitude="-78.7497" name="San Juan and Chiriboga" pageId="23" pageNumber="24" stateProvince="Pichincha">San Juan and Chiriboga</location>
, site 2;
<geoCoordinate id="67F2FB82FFF6A67BFB9DF8ED66A0F8E9" box="[1109,1230,1893,1919]" degrees="0.2327" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="-0.2327">0.2327° S</geoCoordinate>
,
<geoCoordinate id="67F2FB82FFF6A67BFB16F8ED671CF816" box="[1246,1394,1893,1920]" degrees="78.7497" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="-78.7497">78.7497° W</geoCoordinate>
;
<elevation id="89EB7A76FFF6A67BFF75F8016337F834" box="[189,345,1928,1955]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="m" value="1900.0">
<quantity id="C53E30A0FFF6A67BFF75F8016376F835" box="[189,280,1929,1955]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
;
<collectedFrom id="B1DADEB8FFF6A67BFEA0F8006057F834" box="[360,569,1928,1955]" pageId="23" pageNumber="24">forest at roadside</collectedFrom>
;
<date id="7678BB85FFF6A67BFD8EF8006088F835" box="[582,742,1928,1955]" pageId="23" pageNumber="24" value="2021-09-15">
<collectingDate id="663C426DFFF6A67BFD8EF8006088F835" box="[582,742,1928,1955]" pageId="23" pageNumber="24" value="2021-09-15">15 Sep. 2021</collectingDate>
</date>
;
<collectorName id="AF33F893FFF6A67BFD3CF8016113F834" box="[756,893,1928,1955]" pageId="23" pageNumber="24">B.A. Huber</collectorName>
and
<collectorName id="AF33F893FFF6A67BFC72F8016650F835" box="[954,1086,1929,1955]" pageId="23" pageNumber="24">M. Herrera</collectorName>
leg.;
<collectionCode id="64D70580FFF6A67BFB4BF80166BAF835" box="[1155,1236,1929,1955]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ar 24113
</materialsCitation>
<materialsCitation id="B2AE9718FFF6A67BFAA0F801665BF850" collectedFrom="forest at roadside" collectingDate="2021-09-15" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1900" latitude="-0.2327" location="San Juan" longLatPrecision="7" longitude="-78.7497" pageId="23" pageNumber="24" specimenCount="3" stateProvince="Pichincha">
<specimenCount id="14C056CCFFF6A67BFAA0F801629FF850" pageId="23" pageNumber="24">3 juvs</specimenCount>
, in pure ethanol; same collection data as for preceding;
<collectionCode id="64D70580FFF6A67BFCB7F82461BEF850" box="[895,976,1964,1990]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="24" type="Museum">ZFMK</collectionCode>
Ecu124
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFF9A674FF22FE806335FEB4" blockId="24.[189,1399,264,467]" box="[234,347,264,290]" pageId="24" pageNumber="25">
<emphasis id="30B24157FFF9A674FF22FE806335FEB4" bold="true" box="[234,347,264,290]" pageId="24" pageNumber="25">“Dup55”</emphasis>
</paragraph>
<paragraph id="02799D45FFF9A674FF75FEA4602AFE45" blockId="24.[189,1399,264,467]" pageId="24" pageNumber="25">
<materialsCitation id="B2AE9718FFF9A674FF75FEA4670CFEFF" collectingDate="2014-09-04" collectingDateMax="2014-09-07" collectingDateMin="2014-09-04" collectionCode="QCAZ" collectorName="N. Duperre &amp; E. E. Tapia &amp; C. A. Tapia" country="Ecuador" elevation="1995" latitude="-0.4199" location="OTONGA Natural Reserve" longLatPrecision="7" longitude="-79.0062" municipality="San Francisco de Las Pampas" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="2" stateProvince="Cotopaxi">
<collectingCountry id="7AD1DDD5FFF9A674FF75FEA46325FED0" box="[189,331,300,326]" name="Ecuador" pageId="24" pageNumber="25">ECUADOR</collectingCountry>
<emphasis id="30B24157FFF9A674FEA4FEA363B0FED3" bold="true" box="[364,478,299,325]" pageId="24" pageNumber="25">
<collectingRegion id="C00253A7FFF9A674FEA4FEA363B0FED3" box="[364,478,299,325]" country="Ecuador" name="Cotopaxi" pageId="24" pageNumber="25">Cotopaxi</collectingRegion>
</emphasis>
<specimenCount id="14C056CCFFF9A674FE33FEA4602EFED0" box="[507,576,300,326]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>
;
<collectingMunicipality id="E21D073FFFF9A674FD99FEA461D6FED0" box="[593,952,300,326]" pageId="24" pageNumber="25">San Francisco de Las Pampas</collectingMunicipality>
,
<location id="0719CB9EFFF9A674FC0EFEA46762FED0" LSID="urn:lsid:plazi:treatment:8A6F2C53FFF7A674FDA2FA0E6167FDCC:0719CB9EFFF9A674FC0EFEA46762FED0" box="[966,1292,300,326]" country="Ecuador" latitude="-0.4199" longLatPrecision="7" longitude="-79.0062" municipality="San Francisco de Las Pampas" name="OTONGA Natural Reserve" pageId="24" pageNumber="25" stateProvince="Cotopaxi">OTONGA Natural Reserve</location>
;
<geoCoordinate id="67F2FB82FFF9A674FAD2FEA462A1FEFF" degrees="0.4199" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="-0.4199">0.4199° S</geoCoordinate>
,
<geoCoordinate id="67F2FB82FFF9A674FF17FEC7631FFEFF" box="[223,369,335,361]" degrees="79.0062" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="-79.0062">79.0062° W</geoCoordinate>
;
<elevation id="89EB7A76FFF9A674FE48FEC76070FEFF" box="[384,542,335,362]" metricMagnitude="3" metricUnit="m" metricValue="1.9949999999999999" pageId="24" pageNumber="25" unit="m" value="1995.0">
<quantity id="C53E30A0FFF9A674FE48FEC763B2FEFF" box="[384,476,335,362]" metricMagnitude="3" metricUnit="m" metricValue="1.9949999999999999" pageId="24" pageNumber="25" unit="m" value="1995.0">1995 m</quantity>
a.s.l.
</elevation>
;
<date id="7678BB85FFF9A674FDE6FEC7608EFEFF" box="[558,736,335,361]" pageId="24" pageNumber="25" value="2014-09-04" valueMax="2014-09-07" valueMin="2014-09-04">
<collectingDate id="663C426DFFF9A674FDE6FEC7608EFEFF" box="[558,736,335,361]" pageId="24" pageNumber="25" value="2014-09-04" valueMax="2014-09-07" valueMin="2014-09-04">47 Sep. 2014</collectingDate>
</date>
;
<collectorName id="AF33F893FFF9A674FD27FEC76116FEFF" box="[751,888,335,361]" pageId="24" pageNumber="25">N. Dupérré</collectorName>
,
<collectorName id="AF33F893FFF9A674FC4FFEC76669FEFF" box="[903,1031,335,361]" pageId="24" pageNumber="25">E.E. Tapia</collectorName>
, and
<collectorName id="AF33F893FFF9A674FB81FEC766A5FEFF" box="[1097,1227,335,361]" pageId="24" pageNumber="25">C.A. Tapia</collectorName>
leg.;
<collectionCode id="64D70580FFF9A674FADAFEC7670CFEFF" box="[1298,1378,335,361]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="24" pageNumber="25" type="Museum">QCAZ</collectionCode>
</materialsCitation>
<materialsCitation id="B2AE9718FFF9A674FF75FEFA618BFE1A" box="[189,997,370,397]" collectingDate="2014-11-12" collectingDateMax="2014-09-07" collectingDateMin="2014-09-04" collectionCode="QCAZ" collectorName="N. Duperre &amp; E. E. Tapia &amp; C. A. Tapia" country="Ecuador" elevation="1995" latitude="-0.4199" location="Natural Reserve" longLatPrecision="7" longitude="-79.0062" municipality="San Francisco de Las Pampas" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Cotopaxi">
<specimenCount id="14C056CCFFF9A674FF75FEFA6285FE1B" box="[189,235,370,397]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="7678BB85FFF9A674FD2DFEFA61E6FE1A" box="[741,904,370,397]" pageId="24" pageNumber="25" value="2014-11-12">
<collectingDate id="663C426DFFF9A674FD2DFEFA61E6FE1A" box="[741,904,370,397]" pageId="24" pageNumber="25" value="2014-11-12">12 Nov. 2014</collectingDate>
</date>
;
<collectionCode id="64D70580FFF9A674FC5DFEFA618BFE1A" box="[917,997,370,396]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="24" pageNumber="25" type="Museum">QCAZ</collectionCode>
</materialsCitation>
<materialsCitation id="B2AE9718FFF9A674FC37FEFA6056FE39" collectingDate="2013-01" collectingDateMax="2014-09-07" collectingDateMin="2014-09-04" collectionCode="QCAZ" collectorName="N. Duperre &amp; E. E. Tapia &amp; C. A. Tapia" country="Ecuador" elevation="1995" latitude="-0.4199" location="Natural Reserve" longLatPrecision="7" longitude="-79.0062" municipality="San Francisco de Las Pampas" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Cotopaxi">
<specimenCount id="14C056CCFFF9A674FC37FEFA6642FE1B" box="[1023,1068,370,397]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="7678BB85FFF9A674FEAFFE1E63B5FE39" box="[359,475,405,432]" pageId="24" pageNumber="25" value="2013-01">
<collectingDate id="663C426DFFF9A674FEAFFE1E63B5FE39" box="[359,475,405,432]" pageId="24" pageNumber="25" value="2013-01">Jan. 2013</collectingDate>
</date>
;
<collectionCode id="64D70580FFF9A674FE20FE1D6056FE39" box="[488,568,405,431]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="24" pageNumber="25" type="Museum">QCAZ</collectionCode>
</materialsCitation>
<materialsCitation id="B2AE9718FFF9A674FD9BFE1D602EFE45" collectingDate="2014-09-05" collectingDateMax="2014-09-07" collectingDateMin="2014-09-05" collectionCode="ZFMK" collectorName="N. Duperre &amp; E. E. Tapia &amp; C. A. Tapia" country="Ecuador" elevation="1995" latitude="-0.4199" location="Natural Reserve" longLatPrecision="7" longitude="-79.0062" municipality="San Francisco de Las Pampas" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Cotopaxi">
<specimenCount id="14C056CCFFF9A674FD9BFE1D60EEFE26" box="[595,640,405,432]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
, sequenced specimen; same collection data as for preceding but
<date id="7678BB85FFF9A674FF75FE306307FE45" box="[189,361,440,467]" pageId="24" pageNumber="25" value="2014-09-05" valueMax="2014-09-07" valueMin="2014-09-05">
<collectingDate id="663C426DFFF9A674FF75FE306307FE45" box="[189,361,440,467]" pageId="24" pageNumber="25" value="2014-09-05" valueMax="2014-09-07" valueMin="2014-09-05">57 Sep. 2014</collectingDate>
</date>
;
<collectionCode id="64D70580FFF9A674FEBDFE3163A8FE45" box="[373,454,441,467]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="24" pageNumber="25" type="Museum">ZFMK</collectionCode>
Ar 24189
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFF9A674FF22FE726307FD82" blockId="24.[189,1398,506,603]" box="[234,361,506,532]" pageId="24" pageNumber="25">
<emphasis id="30B24157FFF9A674FF22FE726307FD82" bold="true" box="[234,361,506,532]" pageId="24" pageNumber="25">“Dup140”</emphasis>
</paragraph>
<paragraph id="02799D45FFF9A674FF75FD956167FDCC" blockId="24.[189,1398,506,603]" pageId="24" pageNumber="25">
<materialsCitation id="B2AE9718FFF9A674FF75FD95616AFDCC" collectingDate="2020-03-05" collectionCode="ZFMK" collectorName="N. Duperre &amp; A. A. Tapia &amp; E. E. Tapia" country="Ecuador" elevation="2700" latitude="-3.3289" location="Nabon" longLatPrecision="7" longitude="-79.0508" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Azuay">
<collectingCountry id="7AD1DDD5FFF9A674FF75FD956325FDA1" box="[189,331,541,567]" name="Ecuador" pageId="24" pageNumber="25">ECUADOR</collectingCountry>
<emphasis id="30B24157FFF9A674FEA3FD9563D5FDA1" bold="true" box="[363,443,541,567]" pageId="24" pageNumber="25">
<collectingRegion id="C00253A7FFF9A674FEA3FD9563D5FDA1" box="[363,443,541,567]" country="Ecuador" name="Azuay" pageId="24" pageNumber="25">Azuay</collectingRegion>
</emphasis>
<specimenCount id="14C056CCFFF9A674FE10FD956068FDAE" box="[472,518,541,568]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
;
<location id="0719CB9EFFF9A674FDDEFD956006FDA1" LSID="urn:lsid:plazi:treatment:8A6F2C53FFF7A674FDA2FA0E6167FDCC:0719CB9EFFF9A674FDDEFD956006FDA1" box="[534,616,541,567]" country="Ecuador" latitude="-3.3289" longLatPrecision="7" longitude="-79.0508" name="Nabon" pageId="24" pageNumber="25" stateProvince="Azuay">Nabon</location>
;
<geoCoordinate id="67F2FB82FFF9A674FDBEFD956083FDA1" box="[630,749,541,567]" degrees="3.3289" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="-3.3289">3.3289° S</geoCoordinate>
,
<geoCoordinate id="67F2FB82FFF9A674FD34FD9561E3FDAE" box="[764,909,541,568]" degrees="79.0508" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="-79.0508">79.0508° W</geoCoordinate>
;
<elevation id="89EB7A76FFF9A674FC53FD956659FDA1" box="[923,1079,541,568]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="24" pageNumber="25" unit="m" value="2700.0">
<quantity id="C53E30A0FFF9A674FC53FD956198FDAE" box="[923,1014,541,568]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="24" pageNumber="25" unit="m" value="2700.0">2700 m</quantity>
a.s.l.
</elevation>
;
<date id="7678BB85FFF9A674FB8EFD9566B2FDA1" box="[1094,1244,541,568]" pageId="24" pageNumber="25" value="2020-03-05">
<collectingDate id="663C426DFFF9A674FB8EFD9566B2FDA1" box="[1094,1244,541,568]" pageId="24" pageNumber="25" value="2020-03-05">5 Mar. 2020</collectingDate>
</date>
;
<collectorName id="AF33F893FFF9A674FB21FD96671CFDA1" box="[1257,1394,541,568]" pageId="24" pageNumber="25">N. Dupérré</collectorName>
,
<collectorName id="AF33F893FFF9A674FF75FDC9632FFDCC" box="[189,321,576,603]" pageId="24" pageNumber="25">A.A. Tapia</collectorName>
, and
<collectorName id="AF33F893FFF9A674FEB6FDC96396FDCC" box="[382,504,576,603]" pageId="24" pageNumber="25">E.E. Tapia</collectorName>
leg.;
<collectionCode id="64D70580FFF9A674FDF2FDC960E5FDCD" box="[570,651,577,603]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="24" pageNumber="25" type="Museum">ZFMK</collectionCode>
Ar 24190
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>