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<document id="8DEDAD4C0599F5E1B084B84723E3A16B" ID-CLB-Dataset="284154" ID-DOI="10.5852/ejt.2023.909.2351" ID-GBIF-Dataset="d0cc9c9f-6e84-4a88-a82f-96565be36a3d" ID-ISSN="2118-9773" ID-Zenodo-Dep="10462860" ID-ZooBank="A48BD2B3-DC40-45BD-9968-F04890A1C5C5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701248827785" checkinUser="plazi" docAuthor="Huber, Bernhard A., Meng, Guanliang, Dupérré, Nadine, Astrin, Jonas &amp; Herrera, Mauricio" docDate="2023" docId="8A6F2C53FFCBA643FDACFE806133FDFD" docLanguage="en" docName="EJT.2023.909.1-63.pdf" docOrigin="European Journal of Taxonomy 909" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2351/10271" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.33:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="32" docTitle="Priscula bonita Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera 2023, sp. nov." docType="treatment" docUuid="E8DC917E-0778-4C1B-AA97-F59DB99B3A6A" docUuidSource="ZooBank" docVersion="4" lastPageNumber="48" masterDocId="7656542BFFE1A66CFFC8FF88626EFF96" masterDocTitle="Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="43" updateTime="1704471815894" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="58ADF45918B0FBAEBAE5A3260F6537C2">Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="8C105612E63EB434B654C1290AA2B4B4">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="3A1C539854EAE88AC1E78CC72519A26D">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany.</mods:affiliation>
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<mods:namePart id="10D75E379655E9E9FFD755A2D5D7B7B9">Meng, Guanliang</mods:namePart>
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<mods:affiliation id="BEB483B3C31C82C4A09E8C5A97A54118">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany.</mods:affiliation>
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<mods:namePart id="626A448B5C0F3181068DE39D20780FD4">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="E0563A37946452D9FEB8019284C4AF9A">Museum of Nature Hamburg, LIB, Hamburg, Germany.</mods:affiliation>
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<mods:namePart id="8BE9587B282D71BA21CAC57CCBB62768">Astrin, Jonas</mods:namePart>
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<mods:namePart id="EF837B59A12E5B20B35089756B41CB82">Herrera, Mauricio</mods:namePart>
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<mods:affiliation id="7B8CCAF7FAC77557597AE62C30EEDEDA">Instituto Nacional de Biodiversidad, Sección Invertebrados, Quito, Ecuador.</mods:affiliation>
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<taxonomicName id="C5C6E6C6FFCBA646FDACFE806103FEB5" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[612,877,264,291]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="bonita" status="sp. nov.">
<emphasis id="30B24157FFCBA646FDACFE806170FEB5" bold="true" box="[612,798,264,291]" italics="true" pageId="42" pageNumber="43">Priscula bonita</emphasis>
Huber
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<figureCitation id="9AFD81C0FFCBA646FD97FEC7608FFEFF" box="[607,737,335,361]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[183,1404,633,1854]" captionTargetId="figure-130@12.[183,1404,633,1854]" captionTargetPageId="12" captionText="Fig. 6. Priscula spp., binghamae and andinensis groups, live specimens. A–B. P. pastaza Huber sp. nov.; male and female with egg-sac from Cavernas del Anzu Forest Reserve. C–D. P. bonita Huber sp. nov.; male from La Bonita-Santa Barbara and female with egg-sac from Gruta de los Tayos. E–F. P. lumbaqui Huber sp. nov.; male and female from near Lumbaqui." figureDoi="http://doi.org/10.5281/zenodo.10462882" httpUri="https://zenodo.org/record/10462882/files/figure.png" pageId="42" pageNumber="43">Figs 6C–D</figureCitation>
, 
<figureCitation id="9AFD81C0FFCBA646FD25FEC76157FEFF" box="[749,825,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="43.[189,232,1899,1925]" captionStartId-1="44.[189,231,1860,1886]" captionStartId-2="45.[189,233,1928,1954]" captionStartId-3="46.[189,232,1892,1918]" captionTargetBox-0="[190,1398,1056,1853]" captionTargetBox-1="[225,1346,348,1775]" captionTargetBox-2="[193,1396,1296,1887]" captionTargetBox-3="[183,1404,561,1852]" captionTargetId-0="figure-263@43.[183,1404,1056,1853]" captionTargetId-1="figure-45@44.[186,1401,348,1784]" captionTargetId-2="figure-370@45.[187,1401,1296,1887]" captionTargetId-3="figure-67@46.[183,1404,561,1852]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionTargetPageId-2="45" captionTargetPageId-3="46" captionText-0="Fig. 33. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm." captionText-1="Fig. 34. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Abbreviation: ds=distal sclerite. Scale lines: 0.5 mm." captionText-2="Fig. 35. Priscula bonita Huber sp. nov.; male and female paratypes from near La Bonita, ZFMK Ar 24104. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." captionText-3="Fig. 36. Priscula bonita Huber sp. nov.; female paratype from near La Bonita, ZFMK Ar 24104 (A–D) and female from Cavernas del Anzu Forest Reserve, ZFMK Ar 24109 (E–F). A–B, E–F. Abdomens, ventral and lateral views; arrows point at sclerite in front of spinnerets. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10462940" figureDoi-1="http://doi.org/10.5281/zenodo.10462942" figureDoi-2="http://doi.org/10.5281/zenodo.10462944" figureDoi-3="http://doi.org/10.5281/zenodo.10462946" httpUri-0="https://zenodo.org/record/10462940/files/figure.png" httpUri-1="https://zenodo.org/record/10462942/files/figure.png" httpUri-2="https://zenodo.org/record/10462944/files/figure.png" httpUri-3="https://zenodo.org/record/10462946/files/figure.png" pageId="42" pageNumber="43">33–36</figureCitation>
, 
<figureCitation id="9AFD81C0FFCBA646FC8DFEC761F7FEFF" box="[837,921,335,361]" captionStart="Fig" captionStartId="52.[189,232,1775,1801]" captionTargetBox="[181,1402,265,1699]" captionTargetId="figure-113@52.[181,1402,265,1699]" captionTargetPageId="52" captionText="Fig. 42. Priscula spp., SEM photographs of male pedipalps. A–C. Tip of left procursus in dorsal view (A) and hair-like processes on its prolateral (B) and retrolateral (C) sides; P. esmeraldas Huber sp. nov. D–E. Tip of left procursus in retrolateral view (D) and hair-like processes in more detail (E); P. pastaza Huber sp. nov. F–G. Left procursus and bulb of P. bonita Huber sp. nov., showing position of tarsal organ (arrow in F) and position of sperm duct opening (arrow in G). H. Sperm duct opening (arrow) on bulbal process in P. pastaza. Scale lines: 100 µm (A, F–G); 10 µm (B–C, E); 20 µm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.10462958" httpUri="https://zenodo.org/record/10462958/files/figure.png" pageId="42" pageNumber="43">42F–G</figureCitation>
, 
<figureCitation id="9AFD81C0FFCBA646FC6DFEC761BBFEFF" box="[933,981,335,361]" captionStart="Fig" captionStartId="54.[189,231,1774,1800]" captionTargetBox="[183,1404,265,1696]" captionTargetId="figure-96@54.[183,1404,265,1696]" captionTargetPageId="54" captionText="Fig. 44. Priscula spp., SEM photographs of female legs. A–B. Oval plates of unknown function, in Priscula lumbaqui Huber sp. nov. (A, metatarsus 4) and P. pastaza Huber sp. nov. (B, metatarsus 4). C–F. Tarsal claws of leg 1 (C, P. pastaza), leg 2 (D, P. pastaza), leg 3 (E, P. bonita Huber sp. nov.), and leg 4 (F, P. pastaza). G. Microstructure of claw tines (P. chapintza Huber sp. nov.). H. Basis of mechanoreceptive hair (P. chapintza; metatarsus 4), showing distal bulge (on right side). Scale lines: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10462964" httpUri="https://zenodo.org/record/10462964/files/figure.png" pageId="42" pageNumber="43">44E</figureCitation>
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<treatmentCitationGroup id="22D6BA6BFFCBA646FF75FE196673FE3D" box="[189,1053,401,427]" pageId="42" pageNumber="43">
<taxonomicName id="C5C6E6C6FFCBA646FF75FE19632BFE3D" ID-CoL="6VVR" box="[189,325,401,427]" class="Arachnida" family="Pholcidae" genus="Priscula" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="30B24157FFCBA646FF75FE19634EFE3D" box="[189,288,401,427]" italics="true" pageId="42" pageNumber="43">Priscula</emphasis>
<taxonomicNameLabel id="2B81FC2CFFCBA646FEE0FE196308FE3D" box="[296,358,401,427]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</taxonomicName>
‘Ecu93’ – 
<treatmentCitation id="8367BB54FFCBA646FE2FFE196161FE3D" author="Dederichs T. M. &amp; Huber B. A. &amp; Michalik P." box="[487,783,401,427]" page="46" pageId="42" pageNumber="43" year="2022">
<bibRefCitation id="6657E0B4FFCBA646FE2FFE196161FE3D" author="Dederichs T. M. &amp; Huber B. A. &amp; Michalik P." box="[487,783,401,427]" pageId="42" pageNumber="43" pagination="52" refId="ref24079" refString="Dederichs T. M., Huber B. A. &amp; Michalik P. 2022. Evolutionary morphology of sperm in pholcid spiders (Pholcidae, Synspermiata). BMC Zoology 7: 52. https: // doi. org / 10.1186 / s 40850 - 022 - 00148 - 3" type="journal article" year="2022">
Dederichs 
<emphasis id="30B24157FFCBA646FDAEFE1A60CEFE3D" box="[614,672,401,427]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2022: 46
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(sperm ultrastructure).
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</paragraph>
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<subSubSection id="4ADCCECEFFCBA646FF75FE5A614CFD78" pageId="42" pageNumber="43" type="diagnosis">
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<heading id="59312A29FFCBA646FF75FE5A6358FE7A" bold="true" box="[189,310,466,492]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="30B24157FFCBA646FF75FE5A6358FE7A" bold="true" box="[189,310,466,492]" pageId="42" pageNumber="43">Diagnosis</emphasis>
</heading>
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Distinguished from known congeners by details of procursus (
<figureCitation id="9AFD81C0FFCBA646FC79FD89662AFD8D" box="[945,1092,513,539]" captionStart="Fig" captionStartId="44.[189,231,1860,1886]" captionTargetBox="[225,1346,348,1775]" captionTargetId="figure-45@44.[186,1401,348,1784]" captionTargetPageId="44" captionText="Fig. 34. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Abbreviation: ds=distal sclerite. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462942" httpUri="https://zenodo.org/record/10462942/files/figure.png" pageId="42" pageNumber="43">Fig. 34A–C</figureCitation>
; distinctive distal sclerite with two pointed processes; slender proximal part – similar to 
<taxonomicName id="C5C6E6C6FFCBA646FC09FDAD672EFDA8" authority="(Keyserling, 1877)" baseAuthorityName="Keyserling" baseAuthorityYear="1877" box="[961,1344,548,574]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="30B24157FFCBA646FC09FDAD663AFDA8" box="[961,1108,548,574]" italics="true" pageId="42" pageNumber="43">P. annulipes</emphasis>
(Keyserling, 1877)
</taxonomicName>
and 
<taxonomicName id="C5C6E6C6FFCBA646FF75FDC06302FDF7" authorityName="Simon" authorityYear="1893" box="[189,364,583,609]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="venezuelana">
<emphasis id="30B24157FFCBA646FF75FDC06302FDF7" box="[189,364,583,609]" italics="true" pageId="42" pageNumber="43">P. venezuelana</emphasis>
</taxonomicName>
), genital bulb (
<figureCitation id="9AFD81C0FFCBA646FDEBFDCF60C1FDF7" box="[547,687,583,609]" captionStart="Fig" captionStartId="44.[189,231,1860,1886]" captionTargetBox="[225,1346,348,1775]" captionTargetId="figure-45@44.[186,1401,348,1784]" captionTargetPageId="44" captionText="Fig. 34. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Abbreviation: ds=distal sclerite. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462942" httpUri="https://zenodo.org/record/10462942/files/figure.png" pageId="42" pageNumber="43">Fig. 34D–F</figureCitation>
; main bulbal process longer than in most known congeners, with pointed tip), and female internal genitalia (
<figureCitation id="9AFD81C0FFCBA646FCC7FDE26111FD13" box="[783,895,618,645]" captionStart="Fig" captionStartId="45.[189,233,1928,1954]" captionTargetBox="[193,1396,1296,1887]" captionTargetId="figure-370@45.[187,1401,1296,1887]" captionTargetPageId="45" captionText="Fig. 35. Priscula bonita Huber sp. nov.; male and female paratypes from near La Bonita, ZFMK Ar 24104. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462944" httpUri="https://zenodo.org/record/10462944/files/figure.png" pageId="42" pageNumber="43">Fig. 35C</figureCitation>
; pore plates roughly triangular, narrower posteriorly than anteriorly – similar to 
<taxonomicName id="C5C6E6C6FFCBA646FD5CFD06617FFD31" authorityName="Huber" authorityYear="2020" box="[660,785,653,679]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="30B24157FFCBA646FD5CFD06617FFD31" box="[660,785,653,679]" italics="true" pageId="42" pageNumber="43">P. bolivari</emphasis>
</taxonomicName>
). From most known congeners in 
<collectingCountry id="7AD1DDD5FFCBA646FB7CFD056778FD31" box="[1204,1302,653,679]" name="Ecuador" pageId="42" pageNumber="43">Ecuador</collectingCountry>
(except 
<taxonomicName id="C5C6E6C6FFCBA646FF75FD3A6327FD5D" authority="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera, 2023" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[189,329,689,715]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="lumbaqui" status="sp. nov.">
<emphasis id="30B24157FFCBA646FF75FD3A6327FD5D" box="[189,329,689,715]" italics="true" pageId="42" pageNumber="43">P. lumbaqui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFCBA646FE87FD3963C5FD5D" box="[335,427,689,715]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
) also by very large male palpal femur (
<figureCitation id="9AFD81C0FFCBA646FCBEFD39618FFD5D" box="[886,993,689,715]" captionStart="Fig" captionStartId="43.[189,232,1899,1925]" captionTargetBox="[190,1398,1056,1853]" captionTargetId="figure-263@43.[183,1404,1056,1853]" captionTargetPageId="43" captionText="Fig. 33. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462940" httpUri="https://zenodo.org/record/10462940/files/figure.png" pageId="42" pageNumber="43">Fig. 33C</figureCitation>
; 
<figureCitation id="9AFD81C0FFCBA646FC25FD39667CFD5D" box="[1005,1042,689,715]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[186,1404,890,1781]" captionTargetId="figure-231@6.[183,1404,890,1781]" captionTargetPageId="6" captionText="Fig. 1; continued in Fig. 2. Relationships of Priscula derived from an unpartitioned analysis of molecular data using IQ-Tree and the ClipKIT kpic-smart-gap trimming strategy. Numbers on the branches are SH-aLRT supports (%). The tree shows only the ingroup. For the complete tree (incl. outgroups), and for clade support using different partitioning and alignment strategies, see Supplementary Figures S1–S3. Photos on the right, from top: Priscula llaviucu Huber sp. nov.; P. espejoi Huber sp. nov.; P. binghamae (Chamberlin, 1916); P. pastaza Huber sp. nov.; P. “Tri18-11”." figureDoi="http://doi.org/10.5281/zenodo.10462862" httpUri="https://zenodo.org/record/10462862/files/figure.png" pageId="42" pageNumber="43">1.7</figureCitation>
times longer than palpal tibia; 
<taxonomicName id="C5C6E6C6FFCBA646FF75FD5D6327FD78" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[189,329,724,750]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="lumbaqui">
<emphasis id="30B24157FFCBA646FF75FD5D6327FD78" box="[189,329,724,750]" italics="true" pageId="42" pageNumber="43">P. lumbaqui</emphasis>
</taxonomicName>
: 2.0; other species in 
<collectingCountry id="7AD1DDD5FFCBA646FD82FD5C60C0FD78" box="[586,686,724,750]" name="Ecuador" pageId="42" pageNumber="43">Ecuador</collectingCountry>
. 1.1–1.3).
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFCBA646FF75FC9E6370F8D6" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="02799D45FFCBA646FF75FC9E6302FCB9" blockId="42.[189,364,789,816]" box="[189,364,789,816]" pageId="42" pageNumber="43">
<heading id="59312A29FFCBA646FF75FC9E6302FCB9" bold="true" box="[189,364,789,816]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="30B24157FFCBA646FF75FC9E6302FCB9" bold="true" box="[189,364,789,816]" pageId="42" pageNumber="43">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCBA646FF22FCCC6180FC32" blockId="42.[189,1399,836,933]" pageId="42" pageNumber="43">
<materialsCitation id="B2AE9718FFCBA646FF22FCCC6180FC32" collectedFrom="humid forest" collectingDate="2021-10-01" collectionCode="MECN" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1870" latitude="0.474" location="La Bonita" longLatPrecision="78" longitude="-77.558" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-41-T" specimenCount="1" specimenCount-male="1" stateProvince="Sucumbios" typeStatus="holotype">
<emphasis id="30B24157FFCBA646FF22FCCC6332FCC8" bold="true" box="[234,348,836,862]" pageId="42" pageNumber="43">
<typeStatus id="DD7D23E7FFCBA646FF22FCCC6332FCC8" box="[234,348,836,862]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7AD1DDD5FFCBA646FF75FCEF6325FC17" box="[189,331,871,897]" name="Ecuador" pageId="42" pageNumber="43">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFCBA646FEB9FCEE6394FC17" bold="true" box="[369,506,870,897]" pageId="42" pageNumber="43">
<collectingRegion id="C00253A7FFCBA646FEB9FCEE6394FC17" box="[369,506,870,897]" country="Ecuador" name="Sucumbios" pageId="42" pageNumber="43">Sucumbíos</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFCBA646FDD4FCE0605BFC14" box="[540,565,872,898]" count="1" pageId="42" pageNumber="43" type="male">♂</specimenCount>
; near 
<location id="0719CB9EFFCBA646FD4CFCE0616EFC17" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FD4CFCE0616EFC17" box="[644,768,871,898]" country="Ecuador" latitude="0.474" longLatPrecision="78" longitude="-77.558" name="La Bonita" pageId="42" pageNumber="43" stateProvince="Sucumbios">La Bonita</location>
; 
<geoCoordinate id="67F2FB82FFCBA646FCD8FCEF61ECFC14" box="[784,898,871,898]" degrees="0.474" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="55" value="0.474">0.474° N</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FC5AFCEF6676FC14" box="[914,1048,871,898]" degrees="77.558" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="55" value="-77.558">77.558° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FBE0FCEF66A4FC17" box="[1064,1226,871,898]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="42" pageNumber="43" unit="m" value="1870.0">
<quantity id="C53E30A0FFCBA646FBE0FCEF66E8FC14" box="[1064,1158,871,898]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="42" pageNumber="43" unit="m" value="1870.0">1870 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFCBA646FB14FCEF671CFC17" box="[1244,1394,871,897]" pageId="42" pageNumber="43" value="2021-10-01">
<collectingDate id="663C426DFFCBA646FB14FCEF671CFC17" box="[1244,1394,871,897]" pageId="42" pageNumber="43" value="2021-10-01">1 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FF75FC03632BFC32" box="[189,325,906,933]" pageId="42" pageNumber="43">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFCBA646FEB6FC03606FFC33" box="[382,513,907,933]" pageId="42" pageNumber="43">M. Herrera</collectorName>
leg.; 
<collectedFrom id="B1DADEB8FFCBA646FD8AFC0260B2FC32" box="[578,732,906,932]" pageId="42" pageNumber="43">humid forest</collectedFrom>
; 
<specimenCode id="5260353EFFCBA646FD2FFC026184FC32" box="[743,1002,906,932]" pageId="42" pageNumber="43">MECN–ARAC–41–T</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="02799D45FFCBA646FF22FC4463E2FBBA" blockId="42.[189,1399,972,1069]" pageId="42" pageNumber="43">
<materialsCitation id="B2AE9718FFCBA646FF22FC4463E9FBBA" collectedFrom="humid forest" collectingDate="2021-10-01" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1870" latitude="0.474" location="Sucumbios" longLatPrecision="78" longitude="-77.558" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-42-T" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sucumbios" typeStatus="paratype">
<emphasis id="30B24157FFCBA646FF22FC446309FC70" bold="true" box="[234,359,972,998]" pageId="42" pageNumber="43">
<typeStatus id="DD7D23E7FFCBA646FF22FC446309FC70" box="[234,359,972,998]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7AD1DDD5FFCBA646FF75FC676325FB9F" box="[189,331,1007,1033]" name="Ecuador" pageId="42" pageNumber="43">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFCBA646FEA4FC66639BFB9F" bold="true" box="[364,501,1006,1033]" pageId="42" pageNumber="43">
<collectingRegion id="C00253A7FFCBA646FEA4FC66639BFB9F" box="[364,501,1006,1033]" country="Ecuador" name="Sucumbios" pageId="42" pageNumber="43">Sucumbíos</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFCBA646FDD9FC67602FFB9C" box="[529,577,1007,1034]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FD98FC676010FB9C" box="[592,638,1007,1034]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; same collection data as for holotype; 
<specimenCode id="5260353EFFCBA646FB98FC67673CFB9F" box="[1104,1362,1007,1033]" pageId="42" pageNumber="43">MECN–ARAC–42–T</specimenCode>
, in 
<collectionCode id="64D70580FFCBA646FF75FB9B6360FBBB" box="[189,270,1043,1069]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ar 24104
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFCBA646FF75FBDB6397FBFB" blockId="42.[189,505,1107,1133]" box="[189,505,1107,1133]" pageId="42" pageNumber="43">
<heading id="59312A29FFCBA646FF75FBDB6397FBFB" bold="true" box="[189,505,1107,1133]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="30B24157FFCBA646FF75FBDB6397FBFB" bold="true" box="[189,505,1107,1133]" pageId="42" pageNumber="43">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCBA646FF75FB0A671EFA4F" blockId="42.[189,1399,1153,1497]" pageId="42" pageNumber="43">
<materialsCitation id="B2AE9718FFCBA646FF75FB0A6379FB29" collectedFrom="humid forest" collectingDate="2021-10-01" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1870" latitude="0.474" location="Sucumbios" longLatPrecision="78" longitude="-77.558" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" stateProvince="Sucumbios">
<collectingCountry id="7AD1DDD5FFCBA646FF75FB0A6325FB0A" box="[189,331,1154,1180]" name="Ecuador" pageId="42" pageNumber="43">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFCBA646FEA2FB09639DFB0A" bold="true" box="[362,499,1153,1180]" pageId="42" pageNumber="43">
<collectingRegion id="C00253A7FFCBA646FEA2FB09639DFB0A" box="[362,499,1153,1180]" country="Ecuador" name="Sucumbios" pageId="42" pageNumber="43">Sucumbíos</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFCBA646FDC5FB0A6054FB0B" box="[525,570,1154,1181]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FD82FB0A60E2FB0A" box="[586,652,1154,1180]" count="1" pageId="42" pageNumber="43" type="generic">1 juv.</specimenCount>
(in pure ethanol); same collection data as for holotype; 
<collectionCode id="64D70580FFCBA646FAEDFB0B6718FB0B" box="[1317,1398,1155,1181]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ecu214
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FEE5FB2D666CFB75" collectedFrom="humid forest" collectingDate="2021-10-01" collectionCode="MECN" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2720" latitude="0.647" location="between La Bonita and Santa Barbara" longLatPrecision="7" longitude="-77.491" municipality="La Bonita" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-43-T" specimenCount="1" specimenCount-male="1" stateProvince="Sucumbios">
<specimenCount id="14C056CCFFCBA646FEE5FB2D6334FB56" box="[301,346,1189,1216]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
; 
<location id="0719CB9EFFCBA646FEADFB2D6170FB29" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FEADFB2D6170FB29" box="[357,798,1189,1216]" country="Ecuador" latitude="0.647" longLatPrecision="7" longitude="-77.491" municipality="La Bonita" name="between La Bonita and Santa Barbara" pageId="42" pageNumber="43" stateProvince="Sucumbios">between La Bonita and Santa Barbara</location>
; 
<geoCoordinate id="67F2FB82FFCBA646FCE0FB2D61CDFB56" box="[808,931,1189,1216]" degrees="0.6470" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="0.647">0.6470° N</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FC65FB2D6652FB56" box="[941,1084,1189,1216]" degrees="77.4910" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="-77.491">77.4910° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FB8EFB2D66B2FB29" box="[1094,1244,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.7199999999999998" pageId="42" pageNumber="43" unit="m" value="2720.0">
<quantity id="C53E30A0FFCBA646FB8EFB2D66F0FB56" box="[1094,1182,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.7199999999999998" pageId="42" pageNumber="43" unit="m" value="2720.0">2720 m</quantity>
a.s.l.
</elevation>
, 
<date id="7678BB85FFCBA646FB2FFB2D671CFB29" box="[1255,1394,1189,1215]" pageId="42" pageNumber="43" value="2021-10-01">
<collectingDate id="663C426DFFCBA646FB2FFB2D671CFB29" box="[1255,1394,1189,1215]" pageId="42" pageNumber="43" value="2021-10-01">1 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FF75FB416326FB74" box="[189,328,1224,1251]" pageId="42" pageNumber="43">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFCBA646FE40FB416060FB75" box="[392,526,1225,1251]" pageId="42" pageNumber="43">M. Herrera</collectorName>
leg.; 
<collectedFrom id="B1DADEB8FFCBA646FD9EFB40609CFB74" box="[598,754,1224,1250]" pageId="42" pageNumber="43">humid forest</collectedFrom>
; 
<specimenCode id="5260353EFFCBA646FCC9FB41666CFB75" box="[769,1026,1225,1251]" pageId="42" pageNumber="43">MECN–ARAC–43–T</specimenCode>
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FBE9FB416614FA91" collectedFrom="humid forest" collectingDate="2021-10-01" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2720" latitude="0.647" location="Santa Barbara" longLatPrecision="7" longitude="-77.491" municipality="La Bonita" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-44-T" specimenCount="1" specimenCount-male="1" stateProvince="Sucumbios">
<specimenCount id="14C056CCFFCBA646FBE9FB41663EFB75" box="[1057,1104,1225,1251]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
(1 palp used for SEM); same collection data as for preceding; 
<specimenCode id="5260353EFFCBA646FD4CFB6461E8FA90" box="[644,902,1260,1286]" pageId="42" pageNumber="43">MECN–ARAC–44–T</specimenCode>
, in 
<collectionCode id="64D70580FFCBA646FC7AFB64666DFA90" box="[946,1027,1260,1286]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ar 24105
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FB5CFB6461DDFABF" collectedFrom="humid forest" collectingDate="2021-10-01" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2720" latitude="0.647" location="Santa Barbara" longLatPrecision="7" longitude="-77.491" municipality="La Bonita" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" stateProvince="Sucumbios">
<specimenCount id="14C056CCFFCBA646FB5CFB6466AFFA90" box="[1172,1217,1260,1286]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FB18FB64677CFA90" box="[1232,1298,1260,1286]" count="1" pageId="42" pageNumber="43" type="generic">1 juv.</specimenCount>
(in pure ethanol); same collection data as for preceding; 
<collectionCode id="64D70580FFCBA646FD34FA876123FABF" box="[764,845,1295,1321]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ecu216
</materialsCitation>
. 
<materialsCitation id="B2AE9718FFCBA646FC08FA87660CFAF9" collectingDate="2021-09-29" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1590" latitude="-0.2186" location="Gruta de Los Tayos" longLatPrecision="7" longitude="-77.7402" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-45-T" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Napo">
<emphasis id="30B24157FFCBA646FC08FA87666DFABF" bold="true" box="[960,1027,1295,1321]" pageId="42" pageNumber="43">
<collectingRegion id="C00253A7FFCBA646FC08FA87666DFABF" box="[960,1027,1295,1321]" country="Ecuador" name="Napo" pageId="42" pageNumber="43">Napo</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFCBA646FBE8FA876621FABF" box="[1056,1103,1295,1321]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FB96FA8766CCFABF" box="[1118,1186,1295,1321]" count="4" pageId="42" pageNumber="43" type="female">4 ♀♀</specimenCount>
(
<specimenCount id="14C056CCFFCBA646FB7BFA876756FABF" box="[1203,1336,1295,1321]" count="1" pageId="42" pageNumber="43" type="female">one female</specimenCount>
used for SEM); 
<location id="0719CB9EFFCBA646FEF3FABA604AFADB" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FEF3FABA604AFADB" box="[315,548,1330,1357]" country="Ecuador" latitude="-0.2186" longLatPrecision="7" longitude="-77.7402" name="Gruta de Los Tayos" pageId="42" pageNumber="43" stateProvince="Napo">Gruta de Los Tayos</location>
, in ‘cave’ (canyon); 
<geoCoordinate id="67F2FB82FFCBA646FCDAFABA61E9FADA" box="[786,903,1330,1356]" degrees="0.2186" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-0.2186">0.2186° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FC5CFABA664AFADB" box="[916,1060,1330,1357]" degrees="77.7402" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="-77.7402">77.7402° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FBF8FABA66A6FADA" box="[1072,1224,1330,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="42" pageNumber="43" unit="m" value="1590.0">
<quantity id="C53E30A0FFCBA646FBF8FABA66E6FADB" box="[1072,1160,1330,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="42" pageNumber="43" unit="m" value="1590.0">1590 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFCBA646FB1DFABA671CFADA" box="[1237,1394,1330,1356]" pageId="42" pageNumber="43" value="2021-09-29">
<collectingDate id="663C426DFFCBA646FB1DFABA671CFADA" box="[1237,1394,1330,1356]" pageId="42" pageNumber="43" value="2021-09-29">29 Sep. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FF75FADE632BFAF9" box="[189,325,1365,1392]" pageId="42" pageNumber="43">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFCBA646FEB6FADE606FFAE6" box="[382,513,1366,1392]" pageId="42" pageNumber="43">M. Herrera</collectorName>
leg.; 
<specimenCode id="5260353EFFCBA646FD8BFADD612BFAE6" box="[579,837,1365,1392]" pageId="42" pageNumber="43">MECN–ARAC–45–T</specimenCode>
, in 
<collectionCode id="64D70580FFCBA646FCA7FADE61AEFAE6" box="[879,960,1366,1392]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ar 24106–07
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FBB3FADD61FAFA05" collectingDate="2021-09-29" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1590" latitude="-0.2186" location="Gruta de Los Tayos" longLatPrecision="7" longitude="-77.7402" pageId="42" pageNumber="43" specimenCount="5" specimenCount-female="2" stateProvince="Napo">
<specimenCount id="14C056CCFFCBA646FBB3FADD66D0FAE6" box="[1147,1214,1365,1392]" count="2" pageId="42" pageNumber="43" type="female">2 ♀♀</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FB04FADD677DFAF9" box="[1228,1299,1365,1391]" pageId="42" pageNumber="43">3 juvs</specimenCount>
(in pure ethanol); same collection data as for preceding; 
<collectionCode id="64D70580FFCBA646FD2BFAF1615AFA05" box="[739,820,1401,1427]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ecu205
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FC61FAF163EDFA4F" collectedFrom="humid forest" collectingDate="2021-09-28" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2000" latitude="-0.6409" location="Antisana N. P." longLatPrecision="7" longitude="-77.8086" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-46-T" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="14C056CCFFCBA646FC61FAF161BBFA05" box="[937,981,1401,1427]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
; 
<location id="0719CB9EFFCBA646FC28FAF066ECFA05" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FC28FAF066ECFA05" box="[992,1154,1400,1427]" country="Ecuador" latitude="-0.6409" longLatPrecision="7" longitude="-77.8086" name="Antisana N. P." pageId="42" pageNumber="43" stateProvince="Napo">Antisana N.P.</location>
; 
<geoCoordinate id="67F2FB82FFCBA646FB44FAF0676EFA05" box="[1164,1280,1400,1427]" degrees="0.6409" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-0.6409">0.6409° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FAC3FAF062B5FA20" degrees="77.8086" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="-77.8086">77.8086° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FF2EFA146313FA20" box="[230,381,1436,1462]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="2000.0">
<quantity id="C53E30A0FFCBA646FF2EFA146350FA20" box="[230,318,1436,1462]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFCBA646FE42FA14604BFA20" box="[394,549,1436,1462]" pageId="42" pageNumber="43" value="2021-09-28">
<collectingDate id="663C426DFFCBA646FE42FA14604BFA20" box="[394,549,1436,1462]" pageId="42" pageNumber="43" value="2021-09-28">28 Sep. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FDF8FA1460D9FA20" box="[560,695,1436,1462]" pageId="42" pageNumber="43">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFCBA646FD26FA14611EFA20" box="[750,880,1436,1462]" pageId="42" pageNumber="43">M. Herrera</collectorName>
leg.; 
<collectedFrom id="B1DADEB8FFCBA646FC78FA146626FA20" box="[944,1096,1436,1462]" pageId="42" pageNumber="43">humid forest</collectedFrom>
; 
<specimenCode id="5260353EFFCBA646FB9BFA14673BFA20" box="[1107,1365,1436,1462]" pageId="42" pageNumber="43">MECN–ARAC–46–T</specimenCode>
, in 
<collectionCode id="64D70580FFCBA646FF75FA376360FA4F" box="[189,270,1471,1497]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ar 24108
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FE55FA376702FA4F" box="[413,1388,1471,1497]" collectedFrom="humid forest" collectingDate="2021-09-28" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="2000" latitude="-0.6409" location="Antisana N. P." longLatPrecision="7" longitude="-77.8086" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Napo">
<specimenCount id="14C056CCFFCBA646FE55FA3763A7FA4F" box="[413,457,1471,1497]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FE1FFA376077FA4F" box="[471,537,1471,1497]" count="1" pageId="42" pageNumber="43" type="juvenile">1 juv.</specimenCount>
(in pure ethanol); same collection data as for preceding; 
<collectionCode id="64D70580FFCBA646FB7EFA376769FA4F" box="[1206,1287,1471,1497]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ecu201
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFCBA646FF75F9886024F98C" blockId="42.[189,586,1536,1562]" box="[189,586,1536,1562]" pageId="42" pageNumber="43">
<heading id="59312A29FFCBA646FF75F9886024F98C" bold="true" box="[189,586,1536,1562]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="30B24157FFCBA646FF75F9886024F98C" bold="true" box="[189,586,1536,1562]" pageId="42" pageNumber="43">Assigned tentatively (see below)</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCBA646FF75F9A76370F8D6" blockId="42.[189,1399,1583,1856]" pageId="42" pageNumber="43">
<materialsCitation id="B2AE9718FFCBA646FF75F9A76141F9FB" collectingDate="2009-11-29" collectionCode="ZFMK" collectorName="P. Michalik" country="Ecuador" elevation="660" latitude="-0.955" location="Rio Hollin" longLatPrecision="78" longitude="-77.748" municipality="SE Archidona" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<collectingCountry id="7AD1DDD5FFCBA646FF75F9A76325F9DF" box="[189,331,1583,1609]" name="Ecuador" pageId="42" pageNumber="43">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFCBA646FEA4F9A763C1F9DF" bold="true" box="[364,431,1583,1609]" pageId="42" pageNumber="43">
<collectingRegion id="C00253A7FFCBA646FEA4F9A763C1F9DF" box="[364,431,1583,1609]" country="Ecuador" name="Napo" pageId="42" pageNumber="43">Napo</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFCBA646FE03F9A76397F9DF" box="[459,505,1583,1609]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
(in pure ethanol); 
<collectingMunicipality id="E21D073FFFCBA646FD13F9A761EAF9DF" box="[731,900,1583,1609]" pageId="42" pageNumber="43">SE Archidona</collectingMunicipality>
, 
<location id="0719CB9EFFCBA646FC59F9A7667AF9DF" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FC59F9A7667AF9DF" box="[913,1044,1583,1609]" country="Ecuador" latitude="-0.955" longLatPrecision="78" longitude="-77.748" municipality="SE Archidona" name="Rio Hollin" pageId="42" pageNumber="43" stateProvince="Napo">Rio Hollin</location>
; 
<geoCoordinate id="67F2FB82FFCBA646FBE9F9A766E4F9DF" box="[1057,1162,1583,1610]" degrees="0.955" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="55" value="-0.955">0.955° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FB51F9A76772F9DF" box="[1177,1308,1583,1609]" degrees="77.748" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="55" value="-77.748">77.748° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FAE2F9A76298F9FA" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="42" pageNumber="43" unit="m" value="660.0">
<quantity id="C53E30A0FFCBA646FAE2F9A76719F9DF" box="[1322,1399,1583,1609]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="42" pageNumber="43" unit="m" value="660.0">660 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFCBA646FECBF9DA63CAF9FA" box="[259,420,1618,1645]" pageId="42" pageNumber="43" value="2009-11-29">
<collectingDate id="663C426DFFCBA646FECBF9DA63CAF9FA" box="[259,420,1618,1645]" pageId="42" pageNumber="43" value="2009-11-29">29 Nov. 2009</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FE78F9DB6058F9FA" box="[432,566,1618,1645]" pageId="42" pageNumber="43">P. Michalik</collectorName>
leg.; 
<collectionCode id="64D70580FFCBA646FDBFF9DB60A6F9FB" box="[631,712,1619,1645]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Mich35
</materialsCitation>
. 
<materialsCitation id="B2AE9718FFCBA646FCF1F9DA6691F925" collectingDate="2021-09-26" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1140" latitude="-1.4054" location="Cueva Copa del Mundo" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" pageId="42" pageNumber="43" specimenCode="MECN-ARAC-47-T" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Pastaza">
<emphasis id="30B24157FFCBA646FCF1F9DA61F7F9FA" bold="true" box="[825,921,1618,1644]" pageId="42" pageNumber="43">
<collectingRegion id="C00253A7FFCBA646FCF1F9DA61F7F9FA" box="[825,921,1618,1644]" country="Ecuador" name="Pastaza" pageId="42" pageNumber="43">Pastaza</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFCBA646FC7AF9DA61B1F9FB" box="[946,991,1618,1645]" count="1" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
, 
<specimenCount id="14C056CCFFCBA646FC24F9DA6676F9FB" box="[1004,1048,1618,1645]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
abdomen; 
<collectingMunicipality id="E21D073FFFCBA646FB51F9DA631CF906" pageId="42" pageNumber="43">Cavernas del Anzu Forest Reserve</collectingMunicipality>
, at cave entrance of 
<location id="0719CB9EFFCBA646FDBAF9FD61F6F919" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FDBAF9FD61F6F919" box="[626,920,1653,1679]" country="Ecuador" latitude="-1.4054" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" name="Cueva Copa del Mundo" pageId="42" pageNumber="43" stateProvince="Pastaza">Cueva Copa del Mundo</location>
; 
<geoCoordinate id="67F2FB82FFCBA646FC6EF9FD6671F919" box="[934,1055,1653,1680]" degrees="1.4054" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-1.4054">1.4054° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FBF8F9FD66ADF906" box="[1072,1219,1653,1680]" degrees="78.0433" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="-78.0433">78.0433° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FB1AF9FD671EF919" box="[1234,1392,1653,1680]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="42" pageNumber="43" unit="m" value="1140.0">
<quantity id="C53E30A0FFCBA646FB1AF9FD6743F906" box="[1234,1325,1653,1680]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="42" pageNumber="43" unit="m" value="1140.0">1140 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFCBA646FF75F9106331F925" box="[189,351,1688,1715]" pageId="42" pageNumber="43" value="2021-09-26">
<collectingDate id="663C426DFFCBA646FF75F9106331F925" box="[189,351,1688,1715]" pageId="42" pageNumber="43" value="2021-09-26">26 Sep. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FEA6F9116396F924" box="[366,504,1688,1715]" pageId="42" pageNumber="43">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFCBA646FDFFF91160D2F925" box="[567,700,1689,1715]" pageId="42" pageNumber="43">M. Herrera</collectorName>
leg.; 
<specimenCode id="5260353EFFCBA646FCCAF911666BF925" box="[770,1029,1689,1715]" pageId="42" pageNumber="43">MECN–ARAC–47–T</specimenCode>
, in 
<collectionCode id="64D70580FFCBA646FBFCF91166EBF925" box="[1076,1157,1689,1715]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ar 24109
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FAD5F9116378F96F" collectingDate="2021-09-26" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1140" latitude="-1.4054" location="Cueva Copa del Mundo" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Pastaza">
<specimenCount id="14C056CCFFCBA646FAD5F9116722F925" box="[1309,1356,1689,1715]" count="1" pageId="42" pageNumber="43" type="female">1 ♀</specimenCount>
(in pure ethanol; abdomen transferred to 
<collectionCode id="64D70580FFCBA646FDB2F93460A5F940" box="[634,715,1724,1750]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ar 24109); same collection data as for preceding; 
<collectionCode id="64D70580FFCBA646FAEDF9346718F940" box="[1317,1398,1724,1750]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ecu184
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFCBA646FEFDF9576375F8D6" collectingDate="2021-09-26" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1205" latitude="-1.416" location="Cavernas del Anzu Forest Reserve" longLatPrecision="78" longitude="-78.049" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="3" stateProvince="Pastaza">
<specimenCount id="14C056CCFFCBA646FEFDF9576314F96F" box="[309,378,1759,1785]" count="3" pageId="42" pageNumber="43" type="female">3 ♀♀</specimenCount>
(in pure ethanol); 
<location id="0719CB9EFFCBA646FD97F957666FF96F" LSID="urn:lsid:plazi:treatment:8A6F2C53FFCBA643FDACFE806133FDFD:0719CB9EFFCBA646FD97F957666FF96F" box="[607,1025,1759,1785]" country="Ecuador" latitude="-1.416" longLatPrecision="78" longitude="-78.049" name="Cavernas del Anzu Forest Reserve" pageId="42" pageNumber="43" stateProvince="Pastaza">Cavernas del Anzu Forest Reserve</location>
; between 
<geoCoordinate id="67F2FB82FFCBA646FBB4F9576688F96F" box="[1148,1254,1759,1785]" degrees="1.406" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="55" value="-1.406">1.406° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FB3DF9576718F96F" box="[1269,1398,1759,1785]" degrees="78.043" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="55" value="-78.043">78.043° W</geoCoordinate>
and 
<geoCoordinate id="67F2FB82FFCBA646FF38F88A6337F88A" box="[240,345,1794,1820]" degrees="1.416" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="55" value="-1.416">1.416° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFCBA646FEAFF88A6384F88B" box="[359,490,1794,1821]" degrees="78.049" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="55" value="-78.049">78.049° W</geoCoordinate>
; 
<elevation id="89EB7A76FFCBA646FE3FF88A60B5F88A" box="[503,731,1794,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.27" metricValueMin="1.14" pageId="42" pageNumber="43" unit="m" value="1205.0" valueMax="1270.0" valueMin="1140.0">
<quantity id="C53E30A0FFCBA646FE3FF88A60F4F88B" box="[503,666,1794,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.27" metricValueMin="1.14" pageId="42" pageNumber="43" unit="m" value="1205.0" valueMax="1270.0" valueMin="1140.0">1140–1270 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFCBA646FD22F88A61E4F88A" box="[746,906,1794,1820]" pageId="42" pageNumber="43" value="2021-09-26">
<collectingDate id="663C426DFFCBA646FD22F88A61E4F88A" box="[746,906,1794,1820]" pageId="42" pageNumber="43" value="2021-09-26">26 Sep. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFCBA646FC50F88A664FF88A" box="[920,1057,1794,1820]" pageId="42" pageNumber="43">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFCBA646FB95F88B668FF88B" box="[1117,1249,1795,1821]" pageId="42" pageNumber="43">M. Herrera</collectorName>
leg., 
<collectionCode id="64D70580FFCBA646FAEDF88B6718F88B" box="[1317,1398,1795,1821]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="43" type="Museum">ZFMK</collectionCode>
Ecu185
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFCBA646FF75F8EF6195F839" box="[189,1019,1895,1968]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="02799D45FFCBA646FF75F8EF632AF817" blockId="42.[189,324,1895,1921]" box="[189,324,1895,1921]" pageId="42" pageNumber="43">
<heading id="59312A29FFCBA646FF75F8EF632AF817" bold="true" box="[189,324,1895,1921]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="30B24157FFCBA646FF75F8EF632AF817" bold="true" box="[189,324,1895,1921]" pageId="42" pageNumber="43">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCBA646FF75F81D6195F839" blockId="42.[189,1019,1941,1968]" box="[189,1019,1941,1968]" pageId="42" pageNumber="43">
The species name is derived from the 
<typeStatus id="DD7D23E7FFCBA646FDB3F81E60C0F826" box="[635,686,1942,1968]" pageId="42" pageNumber="43">type</typeStatus>
locality, noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFCAA642FF75FE806635FE1A" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44" type="description">
<paragraph id="02799D45FFCAA647FF75FE806320FEB4" blockId="43.[189,334,264,290]" box="[189,334,264,290]" pageId="43" pageNumber="44">
<heading id="59312A29FFCAA647FF75FE806320FEB4" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="30B24157FFCAA647FF75FE806320FEB4" bold="true" box="[189,334,264,290]" pageId="43" pageNumber="44">Description</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCAA647FF22FEBF63DEFEC7" blockId="43.[189,1399,311,479]" box="[234,432,311,337]" pageId="43" pageNumber="44">
<emphasis id="30B24157FFCAA647FF22FEBF63DEFEC7" bold="true" box="[234,432,311,337]" pageId="43" pageNumber="44">
Male (
<typeStatus id="DD7D23E7FFCAA647FEF1FEBF63C6FEC7" box="[313,424,311,337]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="02799D45FFCAA647FF75FED360E8FE49" blockId="43.[189,1399,311,479]" pageId="43" pageNumber="44">
<smallCapsWord id="049F0B99FFCAA647FF75FED36302FEE5" baselines="366,366" box="[189,364,347,373]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="43" pageNumber="44">MEASUREMENTS</smallCapsWord>
. Total body length 5.2, carapace width 2.1. Distance PME–PME 270 µm; diameter PME 170 µm; distance PME–ALE 180 µm; distance AME–AME 45 µm; diameter AME 45 µm. ALE and PLE larger than PME (diameters 250 µm). Leg 1: 46.1 (11.7 +0.9+11.5 + 18.7 +3.3), tibia 2: 8.1, tibia 3: 5.7, tibia 4: 7.3; tibia 1 L/d: 55.
</paragraph>
<paragraph id="02799D45FFCAA647FF75FD8D63D4FD59" blockId="43.[189,1399,517,719]" pageId="43" pageNumber="44">
<smallCapsWord id="049F0B99FFCAA647FF75FD8D6367FD88" baselines="536,537" box="[189,265,517,543]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="43" pageNumber="44">COLOR</smallCapsWord>
(in ethanol). Carapace pale ochre-yellow, with large brown median mark and three pairs of lateral marks, ocular area also brown, clypeus with large brown band narrowing towards chelicerae; sternum light brown with thin darker margins; legs pale ochre, with 3–4 variably distinct dark rings on each femur and tibia; abdomen dorsally and laterally densely covered with black marks, with some small white marks in-between, ventrally with large brown mark in front of gonopore and light brown sclerite in front of spinnerets.
</paragraph>
<paragraph id="02799D45FFCAA647FF75FD7F6380FCEC" blockId="43.[189,1399,759,890]" pageId="43" pageNumber="44">
<smallCapsWord id="049F0B99FFCAA647FF75FD7F6293FC99" baselines="778,778" box="[189,253,759,785]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="43" pageNumber="44">BODY</smallCapsWord>
. Habitus as in 
<figureCitation id="9AFD81C0FFCAA647FE74FD7F6072FC87" box="[444,540,759,785]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[183,1404,633,1854]" captionTargetId="figure-130@12.[183,1404,633,1854]" captionTargetPageId="12" captionText="Fig. 6. Priscula spp., binghamae and andinensis groups, live specimens. A–B. P. pastaza Huber sp. nov.; male and female with egg-sac from Cavernas del Anzu Forest Reserve. C–D. P. bonita Huber sp. nov.; male from La Bonita-Santa Barbara and female with egg-sac from Gruta de los Tayos. E–F. P. lumbaqui Huber sp. nov.; male and female from near Lumbaqui." figureDoi="http://doi.org/10.5281/zenodo.10462882" httpUri="https://zenodo.org/record/10462882/files/figure.png" pageId="43" pageNumber="44">Fig. 6C</figureCitation>
. Ocular area raised, without hump on posterior side, without comb of stronger hairs at median side of each ocular triad. Deep thoracic groove. Clypeus unmodified except sclerotized rim. Sternum wider than long (1.45/1.00), unmodified. Abdomen slightly higher than long, dorso-posteriorly pointed.
</paragraph>
<paragraph id="02799D45FFCAA647FF75FC2A6637FC49" blockId="43.[189,1398,930,991]" pageId="43" pageNumber="44">
<smallCapsWord id="049F0B99FFCAA647FF75FC2A632DFC2C" baselines="949,949" box="[189,323,930,956]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Chelicerae" pageId="43" pageNumber="44">CHELICERAE</smallCapsWord>
. As in 
<figureCitation id="9AFD81C0FFCAA647FE5EFC2A604DFC2B" box="[406,547,930,957]" captionStart="Fig" captionStartId="45.[189,233,1928,1954]" captionTargetBox="[193,1396,1296,1887]" captionTargetId="figure-370@45.[187,1401,1296,1887]" captionTargetPageId="45" captionText="Fig. 35. Priscula bonita Huber sp. nov.; male and female paratypes from near La Bonita, ZFMK Ar 24104. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462944" httpUri="https://zenodo.org/record/10462944/files/figure.png" pageId="43" pageNumber="44">Fig. 35A–B</figureCitation>
, with short entapophyses, with pair of very small frontal apophyses near fang joints, whitish lateral area slightly protruding; without stridulatory ridges.
</paragraph>
<caption id="56B9CDCDFFCAA647FF75F8E361AAF83F" ID-DOI="http://doi.org/10.5281/zenodo.10462940" ID-Zenodo-Dep="10462940" httpUri="https://zenodo.org/record/10462940/files/figure.png" pageId="43" pageNumber="44" startId="43.[189,232,1899,1925]" targetBox="[190,1398,1056,1853]" targetPageId="43">
<paragraph id="02799D45FFCAA647FF75F8E361AAF83F" blockId="43.[189,1398,1899,1961]" pageId="43" pageNumber="44">
<emphasis id="30B24157FFCAA647FF75F8E36374F813" bold="true" box="[189,282,1899,1925]" pageId="43" pageNumber="44">Fig. 33.</emphasis>
<taxonomicName id="C5C6E6C6FFCAA647FEE8F8E3604BF813" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[288,549,1899,1925]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="bonita" status="sp. nov.">
<emphasis id="30B24157FFCAA647FEE8F8E363BBF813" box="[288,469,1899,1925]" italics="true" pageId="43" pageNumber="44">Priscula bonita</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFCAA647FDE4F8E460E7F810" box="[556,649,1900,1926]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
; male paratype from near La Bonita, ZFMK Ar 24104; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm.
</paragraph>
</caption>
<caption id="56B9CDCDFFCDA640FF75F8CC6197F833" ID-DOI="http://doi.org/10.5281/zenodo.10462942" ID-Zenodo-Dep="10462942" httpUri="https://zenodo.org/record/10462942/files/figure.png" pageId="44" pageNumber="45" startId="44.[189,231,1860,1886]" targetBox="[225,1346,348,1775]" targetPageId="44">
<paragraph id="02799D45FFCDA640FF75F8CC6197F833" blockId="44.[189,1399,1860,1957]" pageId="44" pageNumber="45">
<emphasis id="30B24157FFCDA640FF75F8CC6379F8C8" bold="true" box="[189,279,1860,1886]" pageId="44" pageNumber="45">Fig. 34.</emphasis>
<taxonomicName id="C5C6E6C6FFCDA640FED6F8CC6071F8C8" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[286,543,1860,1886]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="bonita" status="sp. nov.">
<emphasis id="30B24157FFCDA640FED6F8CC63BFF8C8" box="[286,465,1860,1886]" italics="true" pageId="44" pageNumber="45">Priscula bonita</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFCDA640FDEEF8CC60ECF8C8" box="[550,642,1860,1886]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
; male paratype from near La Bonita, ZFMK Ar 24104. 
<emphasis id="30B24157FFCDA640FAC8F8CC6751F8C8" bold="true" box="[1280,1343,1860,1886]" pageId="44" pageNumber="45">A–C.</emphasis>
Left tarsus and procursus, prolateral, dorsal, and retrolateral views. 
<emphasis id="30B24157FFCDA640FC47F8EF61A6F817" bold="true" box="[911,968,1895,1921]" pageId="44" pageNumber="45">D–F.</emphasis>
Left genital bulb, dorsal, retrolateral, and ventral views. Abbreviation: ds=distal sclerite. Scale lines: 0.5 mm.
</paragraph>
</caption>
<paragraph id="02799D45FFCCA641FF75FE8166FCFD8F" blockId="45.[186,1399,265,538]" pageId="45" pageNumber="46">
<smallCapsWord id="049F0B99FFCCA641FF75FE816290FEB7" baselines="284,284" box="[189,254,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Palps" pageId="45" pageNumber="46">PALPS</smallCapsWord>
. As in 
<figureCitation id="9AFD81C0FFCCA641FE93FE816384FEB5" box="[347,490,265,291]" captionStart="Fig" captionStartId="43.[189,232,1899,1925]" captionTargetBox="[190,1398,1056,1853]" captionTargetId="figure-263@43.[183,1404,1056,1853]" captionTargetPageId="43" captionText="Fig. 33. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462940" httpUri="https://zenodo.org/record/10462940/files/figure.png" pageId="45" pageNumber="46">Fig. 33A–C</figureCitation>
; coxa unmodified, trochanter slightly protruding ventrally, femur very large, proximally with distinct retrolateral process and prolateral-ventral ridge, distal ventral rim not protruding; patella ventrally reduced to strongly sclerotized narrow rim; tibia small relative to femur; procursus (
<figureCitation id="9AFD81C0FFCCA641FF0DFEFA633FFE1B" box="[197,337,370,397]" captionStart="Fig" captionStartId="44.[189,231,1860,1886]" captionTargetBox="[225,1346,348,1775]" captionTargetId="figure-45@44.[186,1401,348,1784]" captionTargetPageId="44" captionText="Fig. 34. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Abbreviation: ds=distal sclerite. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462942" httpUri="https://zenodo.org/record/10462942/files/figure.png" pageId="45" pageNumber="46">Fig. 34A–C</figureCitation>
) slender, proximally with exposed tarsal organ (arrow in 
<figureCitation id="9AFD81C0FFCCA641FC24FEFA663CFE1B" box="[1004,1106,370,397]" captionStart="Fig" captionStartId="52.[189,232,1775,1801]" captionTargetBox="[181,1402,265,1699]" captionTargetId="figure-113@52.[181,1402,265,1699]" captionTargetPageId="52" captionText="Fig. 42. Priscula spp., SEM photographs of male pedipalps. A–C. Tip of left procursus in dorsal view (A) and hair-like processes on its prolateral (B) and retrolateral (C) sides; P. esmeraldas Huber sp. nov. D–E. Tip of left procursus in retrolateral view (D) and hair-like processes in more detail (E); P. pastaza Huber sp. nov. F–G. Left procursus and bulb of P. bonita Huber sp. nov., showing position of tarsal organ (arrow in F) and position of sperm duct opening (arrow in G). H. Sperm duct opening (arrow) on bulbal process in P. pastaza. Scale lines: 100 µm (A, F–G); 10 µm (B–C, E); 20 µm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.10462958" httpUri="https://zenodo.org/record/10462958/files/figure.png" pageId="45" pageNumber="46">Fig. 42F</figureCitation>
), distally with prolateral-dorsal band of whitish membranous and fringed elements, with strongly sclerotized distal sclerite apparently moveable against proximal part of procursus; genital bulb (
<figureCitation id="9AFD81C0FFCCA641FC4EFE31667BFE45" box="[902,1045,441,467]" captionStart="Fig" captionStartId="44.[189,231,1860,1886]" captionTargetBox="[225,1346,348,1775]" captionTargetId="figure-45@44.[186,1401,348,1784]" captionTargetPageId="44" captionText="Fig. 34. Priscula bonita Huber sp. nov.; male paratype from near La Bonita, ZFMK Ar 24104. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Abbreviation: ds=distal sclerite. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462942" httpUri="https://zenodo.org/record/10462942/files/figure.png" pageId="45" pageNumber="46">Fig. 34D–F</figureCitation>
) with small proximal sclerite connecting to tarsus, with large whitish area on retrolateral-ventral side, strong and slightly spiraling main bulbal process with subdistal sperm duct opening (arrow in 
<figureCitation id="9AFD81C0FFCCA641FCA1FE7761BEFD8F" box="[873,976,511,537]" captionStart="Fig" captionStartId="52.[189,232,1775,1801]" captionTargetBox="[181,1402,265,1699]" captionTargetId="figure-113@52.[181,1402,265,1699]" captionTargetPageId="52" captionText="Fig. 42. Priscula spp., SEM photographs of male pedipalps. A–C. Tip of left procursus in dorsal view (A) and hair-like processes on its prolateral (B) and retrolateral (C) sides; P. esmeraldas Huber sp. nov. D–E. Tip of left procursus in retrolateral view (D) and hair-like processes in more detail (E); P. pastaza Huber sp. nov. F–G. Left procursus and bulb of P. bonita Huber sp. nov., showing position of tarsal organ (arrow in F) and position of sperm duct opening (arrow in G). H. Sperm duct opening (arrow) on bulbal process in P. pastaza. Scale lines: 100 µm (A, F–G); 10 µm (B–C, E); 20 µm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.10462958" httpUri="https://zenodo.org/record/10462958/files/figure.png" pageId="45" pageNumber="46">Fig. 42G</figureCitation>
) and pointed tip.
</paragraph>
<paragraph id="02799D45FFCCA641FF75FDC963F5FD52" blockId="45.[189,1398,577,708]" pageId="45" pageNumber="46">
<smallCapsWord id="049F0B99FFCCA641FF75FDC96298FDCF" baselines="596,596" box="[189,246,577,603]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="45" pageNumber="46">LEGS</smallCapsWord>
. Without spines; with curved hairs on all tibiae and metatarsi, few weakly curved hairs also on femora; with few short vertical hairs; retrolateral trichobothrium of tibia 1 at 7%; prolateral trichobothrium present on all leg tibiae; tarsi without regular pseudosegmentation but rather with many indistinct platelets.
</paragraph>
<paragraph id="02799D45FFCCA641FF22FD6463D9FC93" blockId="45.[188,1399,747,1021]" box="[234,439,747,774]" pageId="45" pageNumber="46">
<emphasis id="30B24157FFCCA641FF22FD6463D9FC93" bold="true" box="[234,439,747,774]" pageId="45" pageNumber="46">Male (variation)</emphasis>
</paragraph>
<paragraph id="02799D45FFCCA641FF75FC876079FC6A" blockId="45.[188,1399,747,1021]" pageId="45" pageNumber="46">
Tibia 
<quantity id="C53E30A0FFCCA641FECEFC876356FCBF" box="[262,312,783,809]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="45" pageNumber="46" unit="in" value="1.0">1 in</quantity>
<specimenCount id="14C056CCFFCCA641FE8BFC9863B8FCBF" box="[323,470,783,810]" count="7" pageId="45" pageNumber="46" type="male">seven males</specimenCount>
(incl. 
<typeStatus id="DD7D23E7FFCCA641FDE4FC8760F7FCBF" box="[556,665,783,809]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
): 9.9–12.4 (mean 11.4). Lateral marks on carapace variably separated or connected with each other at carapace margin; sternum sometimes with light marks. Males from southern 
<collectingRegion id="C00253A7FFCCA641FEAFFCDE63C9FCE6" box="[359,423,854,880]" country="Ecuador" name="Napo" pageId="45" pageNumber="46">Napo</collectingRegion>
and 
<collectingRegion id="C00253A7FFCCA641FE15FCDE605BFCE6" box="[477,565,854,880]" country="Ecuador" name="Pastaza" pageId="45" pageNumber="46">Pastaza</collectingRegion>
are very similar but have slightly smaller palps and the distal sclerite on the procursus is slightly narrower in retrolateral view. This is also reflected in the relatively large CO1 distances between the sequenced specimen from 
<collectingRegion id="C00253A7FFCCA641FD30FC14613EFC20" box="[760,848,924,950]" country="Ecuador" name="Pastaza" pageId="45" pageNumber="46">Pastaza</collectingRegion>
and the sequenced specimens from 
<collectingRegion id="C00253A7FFCCA641FB3CFC146718FC20" box="[1268,1398,924,950]" country="Ecuador" name="Sucumbios" pageId="45" pageNumber="46">Sucumbíos</collectingRegion>
and central to northern 
<collectingRegion id="C00253A7FFCCA641FE04FC486062FC4C" box="[460,524,960,986]" country="Ecuador" name="Napo" pageId="45" pageNumber="46">Napo</collectingRegion>
(8.1–8.8%). Distances among the 
<collectingRegion id="C00253A7FFCCA641FC57FC37664FFC4F" box="[927,1057,959,985]" country="Ecuador" name="Sucumbios" pageId="45" pageNumber="46">Sucumbíos</collectingRegion>
and central to northern 
<collectingRegion id="C00253A7FFCCA641FAFFFC486719FC4C" box="[1335,1399,960,986]" country="Ecuador" name="Napo" pageId="45" pageNumber="46">Napo</collectingRegion>
specimens are 5.3% or lower.
</paragraph>
<paragraph id="02799D45FFCCA641FF22FBAC632BFBA8" blockId="45.[189,1399,1060,1227]" box="[234,325,1060,1086]" pageId="45" pageNumber="46">
<emphasis id="30B24157FFCCA641FF22FBAC632BFBA8" bold="true" box="[234,325,1060,1086]" pageId="45" pageNumber="46">Female</emphasis>
</paragraph>
<paragraph id="02799D45FFCCA642FF75FBCF6635FE1A" blockId="45.[189,1399,1060,1227]" lastBlockId="46.[189,1398,265,397]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
In general similar to male (
<figureCitation id="9AFD81C0FFCCA641FDDBFBCF601DFBF7" box="[531,627,1095,1121]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[183,1404,633,1854]" captionTargetId="figure-130@12.[183,1404,633,1854]" captionTargetPageId="12" captionText="Fig. 6. Priscula spp., binghamae and andinensis groups, live specimens. A–B. P. pastaza Huber sp. nov.; male and female with egg-sac from Cavernas del Anzu Forest Reserve. C–D. P. bonita Huber sp. nov.; male from La Bonita-Santa Barbara and female with egg-sac from Gruta de los Tayos. E–F. P. lumbaqui Huber sp. nov.; male and female from near Lumbaqui." figureDoi="http://doi.org/10.5281/zenodo.10462882" httpUri="https://zenodo.org/record/10462882/files/figure.png" pageId="45" pageNumber="46">Fig. 6D</figureCitation>
) but clypeus rim not sclerotized but with median ochre mark. Ventral sclerite in front of spinnerets variably divided medially or undivided. Tibia 
<quantity id="C53E30A0FFCCA641FB4BFBE266C1FB12" box="[1155,1199,1130,1156]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="45" pageNumber="46" unit="in" value="1.0">1 in</quantity>
<specimenCount id="14C056CCFFCCA641FB7EFBE26757FB12" box="[1206,1337,1130,1156]" count="13" pageId="45" pageNumber="46" type="female">13 females</specimenCount>
: 6.1– 7.7 (mean 7.2). ALS with one strongly widened spigot, one pointed spigot, and one large and four small cylindrical spigots; with distinctively sculptured area medially in front of ALS (similar to 
<figureCitation id="9AFD81C0FFCCA641FB33FB386704FB5D" box="[1275,1386,1200,1227]" captionStart="Fig" captionStartId="51.[189,233,1773,1799]" captionTargetBox="[183,1404,265,1703]" captionTargetId="figure-86@51.[183,1404,265,1703]" captionTargetPageId="51" captionText="Fig. 41. Priscula spp., SEM photographs of female ocular areas and spinnerets. A–B. Ocular areas, frontal views; Priscula pastaza Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C–E. ALS, P. pastaza (C), P. esmeraldas Huber sp. nov. (D), and P. lumbaqui sp. nov. (E). F–H. Distinctive structures in front of spinnerets (in place of colulus); P. chapintza Huber sp. nov. (F), P. esmeraldas (G), and P. pastaza (H). Scale lines: 100 µm (A–B); 10 µm (C–G); 2 µm (H)." figureDoi="http://doi.org/10.5281/zenodo.10462956" httpUri="https://zenodo.org/record/10462956/files/figure.png" pageId="45" pageNumber="46">Fig. 41G</figureCitation>
). Epigynum (
<figureCitation id="9AFD81C0FFCFA642FE8DFE8163A1FEB5" box="[325,463,265,291]" captionStart="Fig" captionStartId="46.[189,232,1892,1918]" captionTargetBox="[183,1404,561,1852]" captionTargetId="figure-67@46.[183,1404,561,1852]" captionTargetPageId="46" captionText="Fig. 36. Priscula bonita Huber sp. nov.; female paratype from near La Bonita, ZFMK Ar 24104 (A–D) and female from Cavernas del Anzu Forest Reserve, ZFMK Ar 24109 (E–F). A–B, E–F. Abdomens, ventral and lateral views; arrows point at sclerite in front of spinnerets. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462946" httpUri="https://zenodo.org/record/10462946/files/figure.png" pageId="46" pageNumber="47">Fig. 36A–B</figureCitation>
) main anterior plate trapezoidal, slightly protruding, posteriorly medially slightly indented; posterior epigynal plate reduced to pair of small lateral sclerites. Females from 
<collectingRegion id="C00253A7FFCFA642FB2AFEA56754FED1" box="[1250,1338,301,327]" country="Ecuador" name="Pastaza" pageId="46" pageNumber="47">Pastaza</collectingRegion>
with similar anterior plate but less reduced posterior plate (
<figureCitation id="9AFD81C0FFCFA642FCE6FEC761D9FEFF" box="[814,951,335,361]" captionStart="Fig" captionStartId="46.[189,232,1892,1918]" captionTargetBox="[183,1404,561,1852]" captionTargetId="figure-67@46.[183,1404,561,1852]" captionTargetPageId="46" captionText="Fig. 36. Priscula bonita Huber sp. nov.; female paratype from near La Bonita, ZFMK Ar 24104 (A–D) and female from Cavernas del Anzu Forest Reserve, ZFMK Ar 24109 (E–F). A–B, E–F. Abdomens, ventral and lateral views; arrows point at sclerite in front of spinnerets. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462946" httpUri="https://zenodo.org/record/10462946/files/figure.png" pageId="46" pageNumber="47">Fig. 36E–F</figureCitation>
). Internal genitalia (
<figureCitation id="9AFD81C0FFCFA642FB57FEC76763FEFC" box="[1183,1293,335,362]" captionStart="Fig" captionStartId="45.[189,233,1928,1954]" captionTargetBox="[193,1396,1296,1887]" captionTargetId="figure-370@45.[187,1401,1296,1887]" captionTargetPageId="45" captionText="Fig. 35. Priscula bonita Huber sp. nov.; male and female paratypes from near La Bonita, ZFMK Ar 24104. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462944" httpUri="https://zenodo.org/record/10462944/files/figure.png" pageId="46" pageNumber="47">Figs 35C</figureCitation>
, 
<figureCitation id="9AFD81C0FFCFA642FAD0FEC76700FEFF" box="[1304,1390,335,361]" captionStart="Fig" captionStartId="46.[189,232,1892,1918]" captionTargetBox="[183,1404,561,1852]" captionTargetId="figure-67@46.[183,1404,561,1852]" captionTargetPageId="46" captionText="Fig. 36. Priscula bonita Huber sp. nov.; female paratype from near La Bonita, ZFMK Ar 24104 (A–D) and female from Cavernas del Anzu Forest Reserve, ZFMK Ar 24109 (E–F). A–B, E–F. Abdomens, ventral and lateral views; arrows point at sclerite in front of spinnerets. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462946" httpUri="https://zenodo.org/record/10462946/files/figure.png" pageId="46" pageNumber="47">36C–D</figureCitation>
) with pair of large pore plates in almost parallel position, narrowing posteriorly.
</paragraph>
</subSubSection>
<caption id="56B9CDCDFFCCA641FF75F80063DCF87F" ID-DOI="http://doi.org/10.5281/zenodo.10462944" ID-Zenodo-Dep="10462944" httpUri="https://zenodo.org/record/10462944/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,233,1928,1954]" targetBox="[193,1396,1296,1887]" targetPageId="45">
<paragraph id="02799D45FFCCA641FF75F80063DCF87F" blockId="45.[189,1399,1928,2025]" pageId="45" pageNumber="46">
<emphasis id="30B24157FFCCA641FF75F8006372F834" bold="true" box="[189,284,1928,1954]" pageId="45" pageNumber="46">Fig. 35.</emphasis>
<taxonomicName id="C5C6E6C6FFCCA641FEEDF800605AF834" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[293,564,1928,1954]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bonita" status="sp. nov.">
<emphasis id="30B24157FFCCA641FEEDF80063B1F834" box="[293,479,1928,1954]" italics="true" pageId="45" pageNumber="46">Priscula bonita</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFCCA641FDF5F80060F3F834" box="[573,669,1928,1954]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
; male and female paratypes from near La Bonita, ZFMK Ar 24104. 
<emphasis id="30B24157FFCCA641FED2F8236335F853" bold="true" box="[282,347,1963,1989]" pageId="45" pageNumber="46">A–B.</emphasis>
Male chelicerae, frontal and lateral views. 
<emphasis id="30B24157FFCCA641FCB3F82361F6F853" bold="true" box="[891,920,1963,1989]" pageId="45" pageNumber="46">C.</emphasis>
Cleared female genitalia, dorsal view. Scale lines: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="4ADCCECEFFCFA642FF75FE3C66CEFE6B" box="[189,1184,436,509]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="02799D45FFCFA642FF75FE3C6338FE58" blockId="46.[189,342,436,462]" box="[189,342,436,462]" pageId="46" pageNumber="47">
<heading id="59312A29FFCFA642FF75FE3C6338FE58" bold="true" box="[189,342,436,462]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="30B24157FFCFA642FF75FE3C6338FE58" bold="true" box="[189,342,436,462]" pageId="46" pageNumber="47">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCFA642FF75FE6A66CEFE6B" blockId="46.[189,1184,482,509]" box="[189,1184,482,509]" pageId="46" pageNumber="47">
Apparently widely distributed along the eastern Andean slopes of 
<collectingCountry id="7AD1DDD5FFCFA642FC0CFE6A6648FE6A" box="[964,1062,482,508]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
(
<figureCitation id="9AFD81C0FFCFA642FBFDFE6A66FCFE6B" box="[1077,1170,482,509]" captionStart="Fig" captionStartId="9.[189,232,1821,1847]" captionTargetBox="[183,1405,1129,1775]" captionTargetId="figure-307@9.[184,1405,1129,1774]" captionTargetPageId="9" captionText="Fig. 4. Known distribution of Priscula in Ecuador. A. Representatives of the gularis group. Circles: P. gularis (TL=type locality) and specimens treated herein as “P. gularis?” and “P. cf. gularis”. Squares: other Ecuadorian representatives of the gularis group. B. Representatives of the andinensis group (red) and binghamae group (blue) in Ecuador. The background colors indicate biogeographic regions of Ecuador (as far as relevant for Priscula, i.e., Andean regions from western foothill forests to eastern foothill forests), taken from Torres-Carvajal et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10462874" httpUri="https://zenodo.org/record/10462874/files/figure.png" pageId="46" pageNumber="47">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="56B9CDCDFFCFA642FF75F8EC60EEF87F" ID-DOI="http://doi.org/10.5281/zenodo.10462946" ID-Zenodo-Dep="10462946" httpUri="https://zenodo.org/record/10462946/files/figure.png" pageId="46" pageNumber="47" startId="46.[189,232,1892,1918]" targetBox="[183,1404,561,1852]" targetPageId="46">
<paragraph id="02799D45FFCFA642FF75F8EC60EEF87F" blockId="46.[189,1398,1892,2025]" pageId="46" pageNumber="47">
<emphasis id="30B24157FFCFA642FF75F8EC6375F8E9" bold="true" box="[189,283,1892,1919]" pageId="46" pageNumber="47">Fig. 36.</emphasis>
<taxonomicName id="C5C6E6C6FFCFA642FEEAF8ED6046F8E9" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[290,552,1893,1919]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="bonita" status="sp. nov.">
<emphasis id="30B24157FFCFA642FEEAF8ED63B6F8E9" box="[290,472,1893,1919]" italics="true" pageId="46" pageNumber="47">Priscula bonita</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFCFA642FDF8F8ED60E0F8E9" box="[560,654,1893,1919]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
; female paratype from near La Bonita, ZFMK Ar 24104 (A–D) and female from Cavernas del Anzu Forest Reserve, ZFMK Ar 24109 (E–F). 
<emphasis id="30B24157FFCFA642FBAFF8006686F834" bold="true" box="[1127,1256,1928,1954]" pageId="46" pageNumber="47">A–B, E–F.</emphasis>
Abdomens, ventral and lateral views; arrows point at sclerite in front of spinnerets. 
<emphasis id="30B24157FFCFA642FBCEF8236628F853" bold="true" box="[1030,1094,1963,1989]" pageId="46" pageNumber="47">C–D.</emphasis>
Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="4ADCCECEFFCEA643FF75FE816133FDFD" pageId="47" pageNumber="48" type="biology_ecology">
<paragraph id="02799D45FFCEA643FF75FE816310FEB4" blockId="47.[189,382,264,291]" box="[189,382,264,291]" pageId="47" pageNumber="48">
<heading id="59312A29FFCEA643FF75FE816310FEB4" bold="true" box="[189,382,264,291]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="30B24157FFCEA643FF75FE816310FEB4" bold="true" box="[189,382,264,291]" pageId="47" pageNumber="48">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFCEA643FF75FEBF6133FDFD" blockId="47.[188,1400,311,619]" pageId="47" pageNumber="48">
At the 
<typeStatus id="DD7D23E7FFCEA643FEC7FEB0632CFEC4" box="[271,322,312,338]" pageId="47" pageNumber="48">type</typeStatus>
locality and between La Bonita and Santa Barbara, the spiders were found hidden in deep sheltered spaces at ground level. In Antisana National Park, 
<specimenCount id="14C056CCFFCEA643FC58FED3665CFEE2" box="[912,1074,346,373]" count="1" pageId="47" pageNumber="48" type="generic">one specimen</specimenCount>
was found in thick mosses covering a tree trunk, the second one in a crevice of a rock wall. In the canyon of the Gruta de Los Tayos, the spiders were fairly abundant, usually under small ledges on the rock wall, up to several meters above the ground; no specimens were found in the neighboring forest. In Cavernas del Anzu Forest Reserve, some specimens were found in the entrance areas of caves (whose deeper parts were occupied by 
<taxonomicName id="C5C6E6C6FFCEA643FF2CFD836333FDB2" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[228,349,523,548]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pastaza">
<emphasis id="30B24157FFCEA643FF2CFD836333FDB2" box="[228,349,523,548]" italics="true" pageId="47" pageNumber="48">P. pastaza</emphasis>
</taxonomicName>
). Others were found in the neighboring forest; among the latter, 
<specimenCount id="14C056CCFFCEA643FBA5FD83669FFDB2" box="[1133,1265,522,549]" count="1" pageId="47" pageNumber="48" type="female">one female</specimenCount>
was found in the mosses covering a tree trunk. 
<specimenCount id="14C056CCFFCEA643FDA5FDA660BAFDDE" box="[621,724,558,584]" count="2" pageId="47" pageNumber="48" type="egg">Two egg</specimenCount>
sacs had diameters of 3.3 and 
<quantity id="C53E30A0FFCEA643FB8DFDA566CAFDDE" box="[1093,1188,557,584]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="47" pageNumber="48" unit="mm" value="4.5">4.5 mm</quantity>
, respectively, and contained ~25/ 
<specimenCount id="14C056CCFFCEA643FEA5FDD863A6FDFD" box="[365,456,592,619]" count="65" pageId="47" pageNumber="48" type="egg">65 eggs</specimenCount>
with an egg diameter of 1.00 mm.
</paragraph>
</subSubSection>
</treatment>
</document>