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<mods:title>A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Köhler, Gunther</mods:namePart>
<mods:nameIdentifier type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pérez, Raúl Gómez Trejo</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Petersen, Claus Bo P.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Méndez, Fausto R.</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cruz, De La</mods:namePart>
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<mods:date>2014</mods:date>
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<mods:number>2014-09-19</mods:number>
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<mods:number>3862</mods:number>
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<taxonomicName authority="Fitch 1978" authorityName="Fitch" authorityYear="1978" box="[151,508,476,502]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis bold="true" box="[151,508,476,502]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[151,370,476,502]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
<bibRefCitation author="Fitch, H. S." box="[377,508,476,502]" pageId="84" pageNumber="85" pagination="1 - 15" refId="ref89954" refString="Fitch, H. S. (1978) Two new anoles (Reptilia: Iguanidae) from Oaxaca with comments on other Mexican species. Contributions in Biology and Geology Milwaukee Public Museum, 20, 1 - 15." type="journal article" year="1978">Fitch 1978</bibRefCitation>
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<treatmentCitation author="Fitch, H. S." box="[151,484,546,568]" page="6" pageId="84" pageNumber="85" year="1978">
<taxonomicName authority="Fitch 1978: 6" authorityName="Fitch" authorityPageNumber="6" authorityYear="1978" box="[151,484,546,568]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[151,334,546,568]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
<bibRefCitation author="Fitch, H. S." box="[343,484,546,568]" pageId="84" pageNumber="85" pagination="1 - 15" refId="ref89954" refString="Fitch, H. S. (1978) Two new anoles (Reptilia: Iguanidae) from Oaxaca with comments on other Mexican species. Contributions in Biology and Geology Milwaukee Public Museum, 20, 1 - 15." type="journal article" year="1978">Fitch 1978: 6</bibRefCitation>
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; type locality: “ 
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SE Nochixtlán (
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NW Cuesta Blanca, Highway 190), 
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, 
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”. 
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: 
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. 
<bibRefCitation author="Lieb, C. S." box="[542,649,577,599]" pageId="84" pageNumber="85" refId="ref93186" refString="Lieb, C. S. (1981) Biochemical and Karyological Systematics of the Mexican Lizards of the Anolis gadovi and A. nebulosus Species Groups (Reptilia: Iguanidae). Unpublished Ph. D. Dissertation, University of California, Los Angeles, 308 pp." type="book" year="1981">Lieb 1981</bibRefCitation>
, 
<bibRefCitation author="Fitch, H. S. &amp; Hillis, D. M." box="[660,863,577,599]" pageId="84" pageNumber="85" pagination="315 - 323" refId="ref90154" refString="Fitch, H. S. &amp; Hillis, D. M. (1984) The Anolis dewlap: Interspecific variability and morphological associations with habitat. Copeia, 1984 (2), 315 - 323. http: // dx. doi. org / 10.2307 / 1445187" type="journal article" year="1984">Fitch &amp; Hillis 1984</bibRefCitation>
, 
<bibRefCitation author="Gorman, G. C. &amp; Lieb, C. S. &amp; Harwood, R. H." box="[874,1075,577,599]" pageId="84" pageNumber="85" pagination="145 - 152" refId="ref91071" refString="Gorman, G. C., Lieb, C. S. &amp; Harwood, R. H. (1984) The relationships of Anolis gadovi: Albumin immunological evidence. Caribbean Journal of Science, 20 (3 - 4), 145 - 152." type="journal article" year="1984">
Gorman 
<emphasis box="[964,1016,577,599]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
1984
</bibRefCitation>
, Flores-Villela 1993, 
<bibRefCitation author="Flores-Villela, O. &amp; Gerez, P." pageId="84" pageNumber="85" refId="ref90370" refString="Flores-Villela, O. &amp; Gerez, P. (1994) Biodiversidad y conservacion en Mexico: vertebrados, vegetacion y uso del suelo. Secunda Edicion. Conision Nacional para el Conocimiento y uso de la Biodiversidad y Universidad Nacional Autonoma de Mexico, Mexico, D. F., 439 pp." type="book" year="1994">Flores-Villela &amp; Gerez 1994</bibRefCitation>
, 
<bibRefCitation author="Lieb, C. S." box="[363,472,608,630]" pageId="84" pageNumber="85" pagination="146 - 157" refId="ref93236" refString="Lieb, C. S. (1995) Preliminary key to the anole lizards of Mexico. In: Flores-Villela, O. A., Mendoza Quijano, F. &amp; Gonzalez Porter, G. (Eds.), Recopilacion de Caves para la Determinacion de Anfibios y Reptiles de Mexico. Publicaciones Especiales del Museo de Zoologia, Universidad Nacional Autonoma de Mexico, pp. 146 - 157." type="book chapter" year="1995">Lieb 1995</bibRefCitation>
, 
<bibRefCitation author="Liner, E. A." box="[485,602,608,630]" pageId="84" pageNumber="85" pagination="1 - 60" refId="ref93458" refString="Liner, E. A. (2007) A checklist of the amphibians and reptiles of Mexico. Occasional Papers of the Museum of Natural Science, Louisiana State University, 80, 1 - 60." type="journal article" year="2007">Liner 2007</bibRefCitation>
, 
<bibRefCitation author="Flaschendrager, A. &amp; Wijffels, L. C. M." box="[616,952,608,630]" pageId="84" pageNumber="85" refId="ref90663" refString="Flaschendrager, A. &amp; Wijffels, L. C. M. (2009) Anolis. Im Biotop und Terrarium. Natur und Tier - Verlag, Munster, 319 pp." type="book" year="2009">Fläschendräger &amp; Wijffels 2009</bibRefCitation>
, 
<bibRefCitation author="Flores-Villela, O. &amp; Canseco-Marquez, L. &amp; Ochoa-Ochoa, L. M." box="[965,1229,608,630]" pageId="84" pageNumber="85" pagination="303 - 321" refId="ref90290" refString="Flores-Villela, O., Canseco-Marquez, L. &amp; Ochoa-Ochoa, L. M. (2010) Geographic distribution and conservation of the herpetofauna of northern Mexico. In: Wilson, L. D., Townsend, J. H. &amp; Johnson, J. D., (Eds.), Conservation of Mesoamerican Amphibians and Reptiles. Eagle Mountain Press, Eagle Mountain, Utah, pp. 303 - 321." type="book chapter" year="2010">
Flores-Villela 
<emphasis box="[1114,1168,608,630]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2010
</bibRefCitation>
, 
<bibRefCitation author="Urbina-Cardona, J. N. &amp; Flores-Villela, O." pageId="84" pageNumber="85" pagination="1031 - 1041" refId="ref97767" refString="Urbina-Cardona, J. N. &amp; Flores-Villela, O. (2010) Ecological-niche modeling and prioritization of conservation-area networks for Mexican herpetofauna. Conservation Biology, 24 (84), 1031 - 1041. http: // dx. doi. org / 10.1111 / j. 1523 - 1739.2009.01432. x" type="journal article" year="2010">Urbina-Cardona &amp; Flores-Villela 2010</bibRefCitation>
, 
<bibRefCitation author="Wilson, L. D. &amp; Townsend, J. H." box="[407,675,638,660]" pageId="84" pageNumber="85" pagination="761 - 812" refId="ref98308" refString="Wilson, L. D., Townsend, J. H. (2010) The herpetofauna of Mesoamerica: biodiversity significance, conservation status, and future challenges. In: Wilson, L. D., Townsend, J. H. &amp; Johnson, J. D., (Eds.), Conservation of Mesoamerican Amphibians and Reptiles. Eagle Mountain Press, Eagle Mountain, Utah, pp. 761 - 812." type="book chapter" year="2010">Wilson &amp; Townsend 2010</bibRefCitation>
, 
<bibRefCitation author="Wilson, L. D. &amp; Johnson, J. D. &amp; Mata-Silva, V." box="[685,873,638,660]" pageId="84" pageNumber="85" pagination="1 - 47" refId="ref98259" refString="Wilson, L. D. &amp; Johnson, J. D. &amp; Mata-Silva, V. (2013) A conservation reassessment of the reptiles of Mexico based on the EVS measure. Amphibian &amp; Reptile Conservation, 7 (1), 1 - 47." type="journal article" year="2013">
Wilson 
<emphasis box="[764,815,638,660]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2013
</bibRefCitation>
, 
<bibRefCitation author="Kohler, G. &amp; Gomez-Trejo-Perez, R. &amp; Petersen, C. B. P. &amp; Mendez de la Cruz, F." box="[884,1070,638,660]" pageId="84" pageNumber="85" pagination="453 - 468" refId="ref92525" refString="Kohler, G., Gomez-Trejo-Perez, R., Petersen, C. B. P. &amp; Mendez de la Cruz, F. (2014) A new species of pine anole from the Sierra Madre del Sur in Oaxaca, Mexico (Reptilia, Dactyloidae: Anolis). Zootaxa, 3753 (5), 453 - 468." type="journal article" year="2014">
Köhler 
<emphasis box="[960,1012,638,660]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2014
</bibRefCitation>
, Köhler 2014
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</paragraph>
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<subSubSection pageId="84" pageNumber="85" type="reference_group">
<paragraph blockId="84.[151,1436,546,722]" box="[151,605,669,691]" pageId="84" pageNumber="85">
<taxonomicName authority=": Gehlbach &amp; Colette 1957" authorityName=": Gehlbach &amp; Colette" authorityYear="1957" box="[151,605,669,691]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[151,332,669,691]" italics="true" pageId="84" pageNumber="85">Anolis nebuloides</emphasis>
: 
<bibRefCitation author="Gehlbach, F. &amp; Colette, B. B." box="[345,605,669,691]" pageId="84" pageNumber="85" pagination="227 - 231" refId="ref90960" refString="Gehlbach, F. R &amp; Colette, B. B. (1957) A Contribution to the herpetofauna of the highlands of Oaxaca and Puebla, Mexico. Herpetologica, 13 (3), 227 - 231." type="journal article" year="1957">Gehlbach &amp; Colette 1957</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="84.[151,1436,546,722]" box="[151,1202,700,722]" pageId="84" pageNumber="85">
<taxonomicName authority=": Savage &amp; Guyer 1989" authorityName="Savage &amp; Guyer" authorityYear="1989" box="[151,583,700,722]" class="Squamata" family="Polychrotidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[151,350,700,721]" italics="true" pageId="84" pageNumber="85">Norops quercorum:</emphasis>
<bibRefCitation author="Savage, J. M. &amp; Guyer, C." box="[356,583,700,722]" pageId="84" pageNumber="85" pagination="105 - 116" refId="ref95903" refString="Savage, J. M. &amp; Guyer, C. (1989) Infrageneric classification and species composition of the anole genera, Anolis, Ctenonotus, Dactyloa, Norops, and Semiurus (Sauria: Iguanidae). Amphibia-Reptilia, 10 (2), 105 - 116. http: // dx. doi. org / 10.1163 / 156853889 x 00142" type="journal article" year="1989">Savage &amp; Guyer 1989</bibRefCitation>
</taxonomicName>
, 
<bibRefCitation author="Nicholson, K. E." box="[594,757,700,722]" pageId="84" pageNumber="85" pagination="93 - 120" refId="ref94419" refString="Nicholson, K. E. (2002) Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs, 16, 93 - 120. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2002) 016 [0093: paaato] 2.0. co; 2" type="journal article" year="2002">Nicholson 2002</bibRefCitation>
, Castiglia 
<emphasis box="[864,915,700,722]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2010, 
<bibRefCitation author="Nicholson, K. E. &amp; Crother, B. I. &amp; Guyer, C. &amp; Savage, J. M." box="[984,1202,700,722]" pageId="84" pageNumber="85" pagination="1 - 108" refId="ref94482" refString="Nicholson, K. E., Crother, B. I., Guyer, C. &amp; Savage, J. M. (2012) It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa, 3477, 1 - 108." type="journal article" year="2012">
Nicholson 
<emphasis box="[1094,1146,700,722]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="diagnosis">
<paragraph blockId="84.[151,1437,766,2029]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[151,266,766,791]" pageId="84" pageNumber="85">Diagnosis</emphasis>
. A small species (SVL in largest male 46.0 mm, largest female 
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) of the genus 
<taxonomicName authority="(sensu Poe 2004)" baseAuthorityName="sensu Poe" baseAuthorityYear="2004" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="genus">
<emphasis box="[1237,1309,766,790]" italics="true" pageId="84" pageNumber="85">Anolis</emphasis>
(sensu 
<bibRefCitation author="Poe, S." pageId="84" pageNumber="85" pagination="37 - 89" refId="ref95161" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
)
</taxonomicName>
that differs from all Mexican and Central American anoles except 
<taxonomicName box="[1000,1129,805,829]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis box="[1000,1129,805,829]" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[1189,1398,806,829]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[1189,1398,806,829]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
by having a combination of (1) strongly keeled ventral scales; (2) usually a patch of three greatly enlarged supraocular scales; (3) 12–20 rows of slightly to moderately enlarged dorsal scales that are smaller than ventral scales; (4) short hind legs, longest toe of adpressed hind leg usually reaching to ear opening or to a point between ear opening and eye, ratio shank length/SVL 0.22–0.26; (5) circumnasal usually in contact with first supralabial; and (6) a large pink dewlap in males and a very small pinkish orange-red dewlap in females (
<figureCitation box="[1046,1131,998,1023]" captionStart="FIGURE 65" captionStartId="85.[151,250,1966,1988]" captionTargetBox="[151,1436,1085,1939]" captionTargetId="figure-444@85.[151,1436,1085,1939]" captionTargetPageId="85" captionText="FIGURE 65. Dewlaps of Anolis quercorum in life. (a) SMF 96451; (b) IBH 26974; (c) IBH 26977; (d) SMF 96458; (e) IBH 26983; (f) IBH 26975. (a–d) are males, (e–f) are females." figureDoi="http://doi.org/10.5281/zenodo.4931093" httpUri="https://zenodo.org/record/4931093/files/figure.png" pageId="84" pageNumber="85">Fig. 65</figureCitation>
). 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[1153,1355,998,1022]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[1153,1355,998,1022]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[213,340,1037,1061]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis box="[213,340,1037,1061]" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[397,603,1038,1061]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[397,603,1038,1061]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
in the ranges and average values of several morphometric and pholidotic characters (see 
<tableCitation box="[323,408,1076,1101]" captionStart="TABLE 9" captionStartId="68.[151,241,152,175]" captionTargetPageId="68" captionText="TABLE 9. Selected measurements, proportions and scale characters of the species related to Anolis quercorum. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FF84E77D7EC3FF6EFB86C6E3" pageId="84" pageNumber="85" tableUuid="56FE6606FF84E77D7EC3FF6EFB86C6E3">Table 9</tableCitation>
), most obvious in (1) number of middorsal scales in one head length; (2) ratio shank length/ HL; and (3) subdigital lamellae on Phalanges II–IV of Toe IV. Also, 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[916,1116,1115,1139]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[916,1116,1115,1139]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[1263,1389,1115,1139]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis box="[1263,1389,1115,1139]" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[151,360,1154,1177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[151,360,1154,1177]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
in hemipenis morphology (hemipenis distinctly bilobed in 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[1057,1215,1154,1177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[1057,1215,1154,1177]" italics="true" pageId="84" pageNumber="85">A. quercorum</emphasis>
</taxonomicName>
vs. unilobed in 
<taxonomicName class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and slightly bilobed in 
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[510,716,1192,1215]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[510,716,1192,1215]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
) and by substantial genetic distances (see above).
</paragraph>
</subSubSection>
<subSubSection lastPageId="87" lastPageNumber="88" pageId="84" pageNumber="85" type="description">
<paragraph blockId="84.[151,1437,766,2029]" lastBlockId="85.[151,1437,151,1066]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis bold="true" box="[199,345,1230,1255]" pageId="84" pageNumber="85">Description.</emphasis>
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[355,559,1230,1254]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[355,559,1230,1254]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
</taxonomicName>
is a small anole (maximum recorded SVL 46.0 mm in males, 
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in females); dorsal head scales in internasal region smooth to weakly keeled, occasionally strongly keeled, unicarinate, other dorsal head scales smooth to weakly keeled in prefrontal region, in parietal region smooth to rugose; shallow to deep prefrontal depression present, shallow parietal depression; 4–7 postrostrals; anterior nasal usually single, exceptionally divided, the lower scale in contact with rostral and first supralabial; 4–8 internasals; canthal ridge sharply defined; scales comprising supraorbital semicircles well defined, smooth to strongly keeled, largest scale in semicircles usually subequal or larger than largest supraocular scale; supraorbital semicircles usually broadly in contact, occasionally separated by a complete row of scales; 1–4 scales separating supraorbital semicircles and interparietal at narrowest point; interparietal well defined, greatly enlarged relative to adjacent scales, surrounded by scales of moderate size, longer than wide, usually subequal to ear opening; enlarged supraoculars usually a patch of 3 greatly enlarged scales in a single row (occasionally a patch of 4–7 moderately to greatly enlarged, somewhat irregular arranged scales in two rows) rugose to weakly keeled, separated from supraorbital semicircles by a complete row of small scales, or these scales narrowly in contact (broadly in contact in 
<accessionNumber box="[182,324,1733,1758]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96456" pageId="84" pageNumber="85">SMF 96456</accessionNumber>
); 2–3 scales between enlarged supraoculars and superciliaries; usually 2 occasionally 3 elongate superciliaries, anterior one longest, followed posteriorly by a series of 3–4 rounded or squarish keeled scales of moderate size; usually 3 enlarged canthals, the second canthal largest; 4–9 scales between second canthals; 6–10 scales between posterior canthals; loreal region slightly concave, 11–29 mostly keeled (some smooth to rugose) loreal scales in a maximum of 3–5 horizontal rows; 5–7 supralabials to level below center of eye; suboculars keeled, broadly in contact with supralabials (2–3 suboculars in contact with 2–4 supralabials); ear opening vertically oval; scales anterior to ear opening keeled, juxtaposed, about four times larger than granulars posterior to ear opening; 5–8 infralabials to level below center of eye; 3–5 postmentals (usually 4), outer pair slightly to distinctly larger than adjacent median postmental scales; 0–3(commonly 0 or 1) enlarged sublabials in contact with infralabials on each side; keeled granular scales present on chin and throat; male dewlap of large size extending onto chest; 7–8 horizontal gorgetal-sternal rows with 14–18 scales per row; modal number of marginal pairs 2–3;female dewlap very small or small; a nuchal crest and a dorsal ridge present in males; scales on middorsum weakly to strongly keeled, subimbricate with rounded posterior margins; 12–20 middorsal scale rows slightly to moderately enlarged, with a gradual transition to lateral scales; lateral scales keeled, more or less homogeneous; 48–67 dorsal scales along vertebral midline between levels of axilla and groin in males, 
<quantity box="[1155,1255,383,408]" metricMagnitude="0" metricUnit="m" metricValue="1.524" metricValueMax="1.8034" metricValueMin="1.2446" pageId="85" pageNumber="86" unit="in" value="60.0" valueMax="71.0" valueMin="49.0">49–71 in</quantity>
females; 22–36 dorsal scales along vertebral midline contained in one head length in males, 
<quantity box="[1031,1132,422,447]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" metricValueMax="9.652" metricValueMin="5.588" pageId="85" pageNumber="86" unit="in" value="30.0" valueMax="38.0" valueMin="22.0">22–38 in</quantity>
females; ventral scales on midsection larger than largest dorsal scales; scales on midventer usually strongly keeled, imbricate, usually mucronate, occasionally weakly keeled, subimbricate, non-mucronate; 33–52 ventral scales along midventral line between levels of axilla and groin in males, 
<quantity box="[644,742,538,563]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.2446000000000002" metricValueMin="0.8382000000000002" pageId="85" pageNumber="86" unit="in" value="41.0" valueMax="49.0" valueMin="33.0">33–49 in</quantity>
females; 22–40 ventral scales contained in one head length in males, 
<quantity box="[231,331,577,602]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="7.112" metricValueMin="4.572" pageId="85" pageNumber="86" unit="in" value="23.0" valueMax="28.0" valueMin="18.0">18–28 in</quantity>
females; 106–140 scales around midbody in males, 
<quantity box="[923,1051,577,602]" metricMagnitude="0" metricUnit="m" metricValue="2.8701999999999996" metricValueMax="3.2004" metricValueMin="2.54" pageId="85" pageNumber="86" unit="in" value="113.0" valueMax="126.0" valueMin="100.0">100–126 in</quantity>
females; tube-like axillary pocket absent; precloacal scales smooth to weakly keeled; males with a pair of greatly enlarged postcloacal scales; tail moderately compressed in cross section, tail height/tail width 
<quantity box="[848,990,654,679]" metricMagnitude="-2" metricUnit="m" metricValue="3.7338" metricValueMax="4.5466" metricValueMin="2.921" pageId="85" pageNumber="86" unit="in" value="1.47" valueMax="1.79" valueMin="1.15">1.15–1.79 in</quantity>
males, 
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females; all caudal scales strongly keeled, homogeneous, although an indistinct division in segments is discernible; dorsal medial caudal scale row hardly enlarged, strongly keeled, not forming a crest; scales on anterodorsal surface of brachium subimbricate to imbricate, strongly keeled, unicarinate, non-mucronate; scales on dorsal surface of antebrachium strongly keeled, unicarinate, subimbricate to imbricate, some slightly mucronate; 19–26 subdigital lamellae on Phalanges II–IV of Toe IV of hind limbs; 5–7 subdigital lamellae on distal phalanx of Toe IV of hind limbs; digital pads dilated, slightly more than twice the size of distal phalanx. In all specimens examined, the longest toe of the adpressed hind leg reaches to level of tympanum or to a point between shoulder and tympanum. Of 14 individuals, the longest toe of adpressed hind leg reaching almost to ear opening in 3.3%, to ear opening in 60.0%, and to a point slightly beyond ear opening in 10.0%. For variation in selected scalation and morphometric characters see 
<tableCitation box="[151,237,1041,1066]" captionStart="TABLE 9" captionStartId="68.[151,241,152,175]" captionTargetPageId="68" captionText="TABLE 9. Selected measurements, proportions and scale characters of the species related to Anolis quercorum. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FF84E77D7EC3FF6EFB86C6E3" pageId="85" pageNumber="86" tableUuid="56FE6606FF84E77D7EC3FF6EFB86C6E3">Table 9</tableCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931093" ID-Zenodo-Dep="4931093" httpUri="https://zenodo.org/record/4931093/files/figure.png" pageId="85" pageNumber="86" startId="85.[151,250,1966,1988]" targetBox="[151,1436,1085,1939]" targetPageId="85">
<paragraph blockId="85.[151,1436,1966,2019]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[151,288,1966,1988]" pageId="85" pageNumber="86">FIGURE 65.</emphasis>
Dewlaps of 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[420,602,1966,1988]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="85" pageNumber="86" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[420,602,1966,1988]" italics="true" pageId="85" pageNumber="86">Anolis quercorum</emphasis>
</taxonomicName>
in life. (a) SMF 96451; (b) IBH 26974; (c) IBH 26977; (d) SMF 96458; (e) IBH 26983; (f) IBH 26975. (a–d) are males, (e–f) are females.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931095" ID-Zenodo-Dep="4931095" httpUri="https://zenodo.org/record/4931095/files/figure.png" pageId="86" pageNumber="87" startId="86.[151,250,1521,1543]" targetBox="[155,1436,187,1493]" targetPageId="86">
<paragraph blockId="86.[151,1435,1521,1574]" pageId="86" pageNumber="87">
<emphasis bold="true" box="[151,289,1521,1543]" pageId="86" pageNumber="87">FIGURE 66.</emphasis>
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[296,478,1521,1543]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="86" pageNumber="87" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[296,478,1521,1543]" italics="true" pageId="86" pageNumber="87">Anolis quercorum</emphasis>
</taxonomicName>
in life. (a) SMF 96448; (b) SMF 96438; (c) SMF 96447; (d) IBH 26983; (e) IBH 26976; (f) SMF 96456. (a–c) are males, (d–f) are females.
</paragraph>
</caption>
<paragraph blockId="86.[151,1437,1644,2011]" pageId="86" pageNumber="87">
The coloration in life of an adult male from 
<quantity box="[710,769,1644,1669]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca (
<accessionNumber box="[998,1136,1644,1669]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96457" pageId="86" pageNumber="87">SMF 96457</accessionNumber>
) was recorded as follows: Dorsal ground color of body, head, limbs and tail Drab (19); limbs with Tawny Olive (17) bands; tail with Olive Brown (278) bands; ventral surfaces of body, limbs and tail Light Buff (2); dewlap Pale Purple (223) with Pale Pinkish Buff (3) gorgetals; iris Dark Drab (45).
</paragraph>
<paragraph blockId="86.[151,1437,1644,2011]" lastBlockId="87.[151,1437,151,343]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
The coloration in life of an adult female from 
<quantity box="[725,783,1786,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca (
<accessionNumber box="[1010,1140,1786,1811]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26982" pageId="86" pageNumber="87">IBH 26982</accessionNumber>
) was recorded as follows: Dorsal ground color Mikado Brown (42) with a suffusion of Cinnamon Brown (43) on head; vertebral line Dark Salmon Color (59) grading into Orange-Rufous (56) on tail and edged by Cinnamon Brown (43) on dorsum; ventral surface of head Light Buff (2) ventral surface of limbs Drab (19); ventral surface of tail Salmon Color (58); dewlap Spinel Pink (235) with Pale Pinkish Buff (3) gorgetals; iris Sprout´s Brown (47). The coloration in life of another adult female from 
<quantity box="[452,509,1952,1977]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca (
<accessionNumber box="[734,865,1952,1977]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26983" pageId="86" pageNumber="87">IBH 26983</accessionNumber>
; 
<figureCitation box="[877,972,1952,1977]" captionStart="FIGURE 66" captionStartId="86.[151,250,1521,1543]" captionTargetBox="[155,1436,187,1493]" captionTargetId="figure-253@86.[155,1440,187,1493]" captionTargetPageId="86" captionText="FIGURE 66. Anolis quercorum in life. (a) SMF 96448; (b) SMF 96438; (c) SMF 96447; (d) IBH 26983; (e) IBH 26976; (f) SMF 96456. (a–c) are males, (d–f) are females." figureDoi="http://doi.org/10.5281/zenodo.4931095" httpUri="https://zenodo.org/record/4931095/files/figure.png" pageId="86" pageNumber="87">Fig. 66d</figureCitation>
) was recorded as follows: Dorsal ground color Drab (19) with a weakly defined Medium Fawn Color (257) zigzag band; dorsal surface of limbs True Cinnamon (260) with Cinnamon Drab (259) bands; ventral surface of head and body Pale Buff (1); ventral surface of limbs Pale Pinkish Buff (3); ventral surface of tail Beige (254); iris Sprout´s Brown (47).
</paragraph>
<paragraph blockId="87.[151,1437,151,343]" pageId="87" pageNumber="88">
The almost completely everted hemipenis of 
<accessionNumber box="[727,863,218,243]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96439" pageId="87" pageNumber="88">SMF 96439</accessionNumber>
(
<figureCitation box="[882,968,218,243]" captionStart="FIGURE 70" captionStartId="90.[151,250,954,976]" captionTargetBox="[151,1436,483,928]" captionTargetId="figure-220@90.[151,1436,483,929]" captionTargetPageId="90" captionText="FIGURE 70. Hemipenis of Anolis quercorum (SMF 96439): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4931103" httpUri="https://zenodo.org/record/4931103/files/figure.png" pageId="87" pageNumber="88">Fig. 70</figureCitation>
) is a bilobate organ; sulcus spermaticus bordered by well developed sulcal lips and opening into two apical fields void of ornamentation, one on each lobe; a large finger-like asulcate processus and a weak asulcate ridge present; apex strongly calyculate, truncus with transverse folds.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931097" ID-Zenodo-Dep="4931097" box="[151,1324,1737,1759]" httpUri="https://zenodo.org/record/4931097/files/figure.png" pageId="87" pageNumber="88" startId="87.[151,250,1737,1759]" targetBox="[155,1436,403,1709]" targetPageId="87">
<paragraph blockId="87.[151,1324,1737,1759]" box="[151,1324,1737,1759]" pageId="87" pageNumber="88">
<emphasis bold="true" box="[151,288,1737,1759]" pageId="87" pageNumber="88">FIGURE 67.</emphasis>
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[293,475,1737,1759]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="87" pageNumber="88" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[293,475,1737,1759]" italics="true" pageId="87" pageNumber="88">Anolis quercorum</emphasis>
</taxonomicName>
: head scalation in (a,c,e) KU 176051; (b,d,f) SMF 96444. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931099" ID-Zenodo-Dep="4931099" httpUri="https://zenodo.org/record/4931099/files/figure.png" pageId="88" pageNumber="89" startId="88.[151,250,1951,1973]" targetBox="[155,1436,181,1930]" targetPageId="88">
<paragraph blockId="88.[151,1436,1951,2035]" pageId="88" pageNumber="89">
<emphasis bold="true" box="[151,287,1951,1973]" pageId="88" pageNumber="89">FIGURE 68.</emphasis>
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[293,474,1951,1973]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="88" pageNumber="89" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[293,474,1951,1973]" italics="true" pageId="88" pageNumber="89">Anolis quercorum</emphasis>
</taxonomicName>
: nasal region in (a) SMF 96441; (b) KU 176051; superciliary region in (c) SMF 96444; (d) KU 176051; chin region in (e) SMF 96444; (f) SMF 96441; cloacal region in (g) IBH 26973; (h) KU 176051. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931101" ID-Zenodo-Dep="4931101" httpUri="https://zenodo.org/record/4931101/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,250,1522,1544]" targetBox="[151,1436,192,1498]" targetPageId="89">
<paragraph blockId="89.[151,1436,1522,1575]" pageId="89" pageNumber="90">
<emphasis bold="true" box="[151,290,1522,1544]" pageId="89" pageNumber="90">FIGURE 69.</emphasis>
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[298,482,1522,1544]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[298,482,1522,1544]" italics="true" pageId="89" pageNumber="90">Anolis quercorum</emphasis>
</taxonomicName>
: dorsal scales in (a) KU 176051; (b) SMF 96444; flank scales in (c) KU 176051; (d) SMF 96444; ventral scales in (e) KU 176051; (f) SMF 96444. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<subSubSection pageId="89" pageNumber="90" type="biology_ecology">
<paragraph blockId="89.[151,1436,1619,2036]" pageId="89" pageNumber="90">
<emphasis bold="true" box="[199,475,1619,1644]" pageId="89" pageNumber="90">Natural History Notes:</emphasis>
The habitat of 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[652,854,1620,1644]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[652,854,1620,1644]" italics="true" pageId="89" pageNumber="90">Anolis quercorum</emphasis>
</taxonomicName>
is leaf litter in oak forest. Here this typical ground anole can be found in amazing population densities. In the vicinity of the 
<typeStatus box="[1003,1051,1656,1680]" pageId="89" pageNumber="90">type</typeStatus>
locality of this species (
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SE Cuesta Blanca, 
<geoCoordinate box="[340,479,1690,1715]" degrees="17.33065" direction="north" orientation="latitude" pageId="89" pageNumber="90" precision="1" value="17.33065">17.33065°N</geoCoordinate>
, 
<geoCoordinate box="[497,654,1690,1715]" degrees="97.158420" direction="west" orientation="longitude" pageId="89" pageNumber="90" precision="1" value="-97.15842">97.158420°W</geoCoordinate>
, 
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), we saw at least 50 individuals within one hour of searching in a relatively small area (&lt;1 ha). When disturbed, these lizards darted a meter or so across the leaf litter and then usually froze and were well camouflaged against the vivid background colors.
</paragraph>
</subSubSection>
<subSubSection pageId="89" pageNumber="90" type="distribution">
<paragraph blockId="89.[151,1436,1619,2036]" pageId="89" pageNumber="90">
<emphasis bold="true" box="[199,723,1797,1822]" pageId="89" pageNumber="90">Geographic Distribution and Conservation</emphasis>
. As currently known, 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[988,1192,1797,1821]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[988,1192,1797,1821]" italics="true" pageId="89" pageNumber="90">Anolis quercorum</emphasis>
</taxonomicName>
is distributed on the Pacific versant of the western central and northwestern portions of the Mexican State of 
<collectingRegion box="[1196,1280,1832,1857]" country="Mexico" name="Oaxaca" pageId="89" pageNumber="90">Oaxaca</collectingRegion>
at elevations between 1616 and 
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(
<figureCitation box="[497,581,1868,1893]" captionStart="FIGURE 71" captionStartId="90.[151,250,1975,1997]" captionTargetBox="[286,1307,1067,1956]" captionTargetId="figure-229@90.[275,1317,1056,1964]" captionTargetPageId="90" captionText="FIGURE 71. Map indicating collecting localities of species related to Anolis quercorum. Each symbol can represent one or more adjacent localities. Black circles: Anolis carlliebi; white circles: A. quercorum; white squares: A. sacamecatensis." figureDoi="http://doi.org/10.5281/zenodo.4931105" httpUri="https://zenodo.org/record/4931105/files/figure.png" pageId="89" pageNumber="90">Fig. 71</figureCitation>
). Given its usual abundance wherever this species occurs, it seems justified to classify 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[272,425,1904,1927]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[272,425,1904,1927]" italics="true" pageId="89" pageNumber="90">A. quercorum</emphasis>
</taxonomicName>
as Least Concern based on the IUCN Red List Categories and Criteria (
<bibRefCitation author="IUCN" box="[1231,1369,1903,1928]" pageId="89" pageNumber="90" refId="ref91603" refString="IUCN (2012) IUCN Red List of Threatened Species. Version 2012.2. Available from: http: // www. iucnredlist. org (accessed 11 January 2013)" type="url" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="materials_examined">
<paragraph blockId="89.[151,1436,1619,2036]" lastBlockId="90.[151,1437,151,429]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis bold="true" box="[199,482,1939,1964]" pageId="89" pageNumber="90">Specimens examined—</emphasis>
<materialsCitation collectionCode="UTA" country="Mexico" elevation="2250" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
<emphasis bold="true" box="[482,575,1939,1964]" pageId="89" pageNumber="90">
<collectingCountry box="[482,571,1939,1964]" name="Mexico" pageId="89" pageNumber="90">Mexico</collectingCountry>
:
</emphasis>
<collectingRegion box="[586,674,1939,1964]" country="Mexico" name="Oaxaca" pageId="89" pageNumber="90">Oaxaca</collectingRegion>
: 0.9 mi N (by road) 
<collectorName box="[925,1049,1939,1964]" pageId="89" pageNumber="90">
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF99E7607DC9F865FC65C19D" box="[925,950,1939,1964]" country="Mexico" name="El" pageId="89" pageNumber="90" stateProvince="Oaxaca">El</location>
Cerezal
</collectorName>
, 
<collectorName box="[1064,1274,1939,1964]" pageId="89" pageNumber="90">Sierra Aloapaneca</collectorName>
(N of city of Oaxaca) 
<collectorName box="[258,377,1975,1999]" pageId="89" pageNumber="90">Mex Hwy</collectorName>
175: 
<collectionCode box="[454,509,1974,1999]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="89" pageNumber="90">UTA</collectionCode>
R-8421;
</materialsCitation>
<materialsCitation collectionCode="UTA" country="Mexico" elevation="2250" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
<locationDeviation box="[629,923,1974,1999]" location="Cuesta Blanca" pageId="89" pageNumber="90">2 km SE Cuesta Blanca</locationDeviation>
, 2250 m: 
<specimenCode box="[1053,1229,1974,1999]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="89" pageNumber="90">IBH 26973–76</specimenCode>
, 
<specimenCode box="[1248,1362,1974,1999]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="89" pageNumber="90">26982–83</specimenCode>
, 
<specimenCode collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">SMF 96441–55</specimenCode>
, 
<specimenCode box="[277,391,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">96457–58</specimenCode>
, 
<specimenCode box="[403,517,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">96747–51</specimenCode>
, 
<specimenCode box="[528,601,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">98021</specimenCode>
, 
<specimenCode box="[613,687,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">98735</specimenCode>
;
</materialsCitation>
<materialsCitation box="[697,1183,2011,2036]" collectionCode="UTA" country="Mexico" elevation="2250" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
1.7 mi SE 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF99E7607D7BF82DFC63C1C5" box="[815,944,2011,2036]" country="Mexico" name="Nochixtlan" pageId="89" pageNumber="90">Nochixtlán</location>
: 
<collectionCode box="[954,1014,2011,2036]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="89" pageNumber="90" type="Museum">MCZ</collectionCode>
R-133097–98;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="UTA" country="Mexico" elevation="2250" lastPageId="90" lastPageNumber="91" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
26 km SE 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF99E7607B48F82DFA4BC1C5" box="[1308,1432,2011,2036]" country="Mexico" name="Nochistlan" pageId="89" pageNumber="90">Nochistlán</location>
, 2.5 km NW 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637F73FF61FE1CC681" box="[295,463,151,176]" name="Cuesta Blanca" pageId="90" pageNumber="91">Cuesta Blanca</location>
: 
<specimenCode box="[475,654,151,176]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="90" pageNumber="91" type="University or college">
<accessionNumber box="[475,610,151,176]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/KU176051" pageId="90" pageNumber="91">KU 176051</accessionNumber>
–57
</specimenCode>
;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[667,1101,151,176]" collectionCode="KU" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
3.5 WSW 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637D41FF61FCA8C681" box="[789,891,151,176]" name="Tlaxiaco" pageId="90" pageNumber="91">Tlaxiaco</location>
: 
<collectionCode box="[903,958,151,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-3568–71;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
32 mi NW Oaxaca, 2164 m: 
<specimenCode box="[151,332,188,213]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="90" pageNumber="91" type="University or college">UMMZ 115057</specimenCode>
;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[342,1048,188,213]" collectionCode="KU" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
32.5 mi NW Oaxaca, 2134 m: 
<specimenCode box="[686,863,188,213]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="90" pageNumber="91" type="University or college">UMMZ 112636</specimenCode>
(
<specimenCount box="[879,1036,188,213]" pageId="90" pageNumber="91" type="generic">12 specimens</specimenCount>
);
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
3.1 km NW 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637AFEFF4AFAEBC6E4" box="[1194,1336,188,213]" name="Llano Verde" pageId="90" pageNumber="91" stateProvince="Oaxaca">Llano Verde</location>
: 
<collectionCode box="[1346,1401,188,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-11502–03;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[277,817,223,248]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
5.6 km E 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637FD5FF29FDC3C6C9" box="[385,528,223,248]" name="Llano Verde" pageId="90" pageNumber="91" stateProvince="Oaxaca">Llano Verde</location>
: 
<accessionNumber box="[538,672,223,248]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/KU193181" pageId="90" pageNumber="91">
<collectionCode box="[538,578,224,247]" country="USA" name="Biodiversity Institute, University of Kansas" pageId="90" pageNumber="91" type="University or college">KU</collectionCode>
193181
</accessionNumber>
, 
<specimenCode box="[683,813,223,248]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="90" pageNumber="91" type="University or college">193188–89</specimenCode>
;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[824,1337,223,248]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
12.8 km W 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637DE8FF29FB98C6C9" box="[956,1099,223,248]" name="Llano Verde" pageId="90" pageNumber="91" stateProvince="Oaxaca">Llano Verde</location>
: 
<collectionCode box="[1109,1169,223,248]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-171142–44;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
52 mi N Oaxaca on 
<collectingCountry box="[282,344,260,283]" name="Mexico" pageId="90" pageNumber="91">MEX</collectingCountry>
190 
<locationDeviation box="[405,648,260,285]" location="El Cumbre" pageId="90" pageNumber="91">2.4 mi N El Cumbre</locationDeviation>
: 
<specimenCode box="[660,841,260,285]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="90" pageNumber="91" type="University or college">UMMZ 131485</specimenCode>
(
<specimenCount box="[858,1039,260,285]" pageId="90" pageNumber="91" type="generic">nine specimens</specimenCount>
);
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637A70FEF2FB1AC72C" box="[1060,1225,260,285]" country="Mexico" name="Hotel Victoria" pageId="90" pageNumber="91" stateProvince="Oaxaca">Hotel Victoria</location>
, 1616 m: 
<collectionCode box="[1343,1398,260,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-25789;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[239,561,295,320]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
Oaxaca: 
<specimenCode box="[341,557,295,320]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="90" pageNumber="91" type="Museum">FMNH 108789-92</specimenCode>
;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[572,1207,295,320]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
Oaxaca 
<collectingMunicipality box="[666,814,295,320]" pageId="90" pageNumber="91">Motor Court</collectingMunicipality>
: Oaxaca 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637DCEFED1FC1DC771" box="[922,974,295,320]" country="Mexico" municipality="Motor Court" name="City" pageId="90" pageNumber="91" stateProvince="Oaxaca">City</location>
: 
<collectionCode box="[989,1049,295,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-86374–75;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
grounds of Oaxaca 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637EC3FEBAFEC6C754" box="[151,277,332,357]" country="Mexico" name="Port Motel" pageId="90" pageNumber="91" stateProvince="Oaxaca">Port Motel</location>
: 
<collectionCode box="[289,349,332,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-79562–64;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[515,1121,332,357]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
3 mi N 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637C08FEBAFD3CC754" box="[604,751,332,357]" country="Mexico" name="La Heradura" pageId="90" pageNumber="91" stateProvince="Oaxaca">La Heradura</location>
: 
<collectionCode box="[762,823,332,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-177946–49, 54954–58;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637A3DFEBAFA4BC755" box="[1129,1432,332,357]" country="Mexico" name="Tamazulapan del Progreso" pageId="90" pageNumber="91" stateProvince="Oaxaca">Tamazulapán del Progreso</location>
, 1990 m: 
<accessionNumber box="[251,526,367,392]" isEnumeration="true" pageId="90" pageNumber="91">IBH 26977, SMF 96438</accessionNumber>
–40, 
<specimenCode box="[580,654,367,392]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="90" pageNumber="91">96456</specimenCode>
;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[665,1085,367,392]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637CCDFE99FC88C7B9" box="[665,859,367,392]" country="Mexico" name="Cerro San Felipe" pageId="90" pageNumber="91" stateProvince="Oaxaca">Cerro San Felipe</location>
: 
<collectionCode box="[870,931,367,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-93671–72;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
15 km NW (on road) 
<collectingMunicipality pageId="90" pageNumber="91">San Juan del Estado</collectingMunicipality>
, distrito 
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637F25FE62FE71C79C" box="[369,418,404,429]" country="Mexico" municipality="San Juan del Estado" name="Etla" pageId="90" pageNumber="91" stateProvince="Oaxaca">Etla</location>
: 
<specimenCode box="[429,585,404,429]" collectionCode="MVZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" lsid="urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="90" pageNumber="91" type="Museum">MVZ 144181</specimenCode>
;
</materialsCitation>
<materialsCitation accessionNumber="KU 176051" box="[597,1132,404,429]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637C01FE62FDBDC79C" box="[597,622,404,429]" country="Mexico" name="El" pageId="90" pageNumber="91" stateProvince="Oaxaca">El</location>
Tejocote, 2078 m: 
<collectionCode box="[838,893,404,429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-25788, 52818–23.
</materialsCitation>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931103" ID-Zenodo-Dep="4931103" httpUri="https://zenodo.org/record/4931103/files/figure.png" pageId="90" pageNumber="91" startId="90.[151,250,954,976]" targetBox="[151,1436,483,928]" targetPageId="90">
<paragraph blockId="90.[151,1437,954,1006]" pageId="90" pageNumber="91">
<emphasis bold="true" box="[151,290,954,976]" pageId="90" pageNumber="91">FIGURE 70.</emphasis>
Hemipenis of 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[446,629,954,976]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[446,629,954,976]" italics="true" pageId="90" pageNumber="91">Anolis quercorum</emphasis>
</taxonomicName>
(SMF 96439): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931105" ID-Zenodo-Dep="4931105" httpUri="https://zenodo.org/record/4931105/files/figure.png" pageId="90" pageNumber="91" startId="90.[151,250,1975,1997]" targetBox="[286,1307,1067,1956]" targetPageId="90">
<paragraph blockId="90.[151,1436,1975,2028]" pageId="90" pageNumber="91">
<emphasis bold="true" box="[151,289,1975,1997]" pageId="90" pageNumber="91">FIGURE 71.</emphasis>
Map indicating collecting localities of species related to 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[886,1069,1975,1997]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[886,1069,1975,1997]" italics="true" pageId="90" pageNumber="91">Anolis quercorum</emphasis>
</taxonomicName>
. Each symbol can represent one or more adjacent localities. Black circles: 
<taxonomicName box="[548,706,2006,2028]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="carlliebi">
<emphasis box="[548,706,2006,2028]" italics="true" pageId="90" pageNumber="91">Anolis carlliebi</emphasis>
</taxonomicName>
; white circles: 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[861,999,2006,2028]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="quercorum">
<emphasis box="[861,999,2006,2028]" italics="true" pageId="90" pageNumber="91">A. quercorum</emphasis>
</taxonomicName>
; white squares: 
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[1164,1349,2006,2028]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[1164,1349,2006,2028]" italics="true" pageId="90" pageNumber="91">A. sacamecatensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931107" ID-Zenodo-Dep="4931107" httpUri="https://zenodo.org/record/4931107/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,1526,1548]" targetBox="[151,1436,195,1502]" targetPageId="91">
<paragraph blockId="91.[151,1435,1526,1610]" pageId="91" pageNumber="92">
<emphasis bold="true" box="[151,290,1526,1548]" pageId="91" pageNumber="92">FIGURE 72.</emphasis>
Syntypes of 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[430,660,1526,1548]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[430,660,1526,1548]" italics="true" pageId="91" pageNumber="92">Anolis boulengerianus</emphasis>
</taxonomicName>
: (a) MNHN 6554 dorsal view; (b) MNHN 6554 ventral view; (c) MNHN 1994.1671 ventral view; (d) MNHN 1994.1671 lateral view; (e) MNHN 1994.1670 dorsal view; (f) MNHN 1994.1670 ventral view.
</paragraph>
</caption>
<subSubSection lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92" type="discussion">
<paragraph blockId="91.[151,1165,1710,1737]" box="[151,1165,1710,1737]" pageId="91" pageNumber="92">
<emphasis bold="true" box="[151,1165,1710,1737]" pageId="91" pageNumber="92">
Taxonomy of the Mexican anoles related to 
<taxonomicName authority="Thominot 1887" authorityName="Thominot" authorityYear="1887" box="[693,1165,1710,1737]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis bold="true" box="[693,968,1710,1736]" italics="true" pageId="91" pageNumber="92">Anolis boulengerianus</emphasis>
<bibRefCitation author="Thominot, A." box="[975,1165,1711,1737]" pageId="91" pageNumber="92" pagination="182 - 190" refId="ref97703" refString="Thominot, A. (1887) Description de trois especes nouvelles d' Anolis et d'un amphisbaenien. Bulletin de la Societe Philomatique de Paris, Series 7, 11, 182 - 190." type="journal article" year="1887">Thominot 1887</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph blockId="91.[151,1436,1781,2022]" lastBlockId="92.[151,1436,151,284]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
<bibRefCitation author="Thominot, A." box="[151,345,1781,1806]" pageId="91" pageNumber="92" pagination="182 - 190" refId="ref97703" refString="Thominot, A. (1887) Description de trois especes nouvelles d' Anolis et d'un amphisbaenien. Bulletin de la Societe Philomatique de Paris, Series 7, 11, 182 - 190." type="journal article" year="1887">Thominot (1887)</bibRefCitation>
described 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[470,725,1781,1805]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[470,725,1781,1805]" italics="true" pageId="91" pageNumber="92">Anolis boulengerianus</emphasis>
</taxonomicName>
based on 
<specimenCount box="[844,1026,1781,1806]" pageId="91" pageNumber="92" type="generic">three specimens</specimenCount>
(now MNHN 6554, 1994.1670–71; 
<figureCitation box="[151,236,1817,1842]" captionStart="FIGURE 72" captionStartId="91.[151,250,1526,1548]" captionTargetBox="[151,1436,195,1502]" captionTargetId="figure-187@91.[151,1436,195,1502]" captionTargetPageId="91" captionText="FIGURE 72. Syntypes of Anolis boulengerianus: (a) MNHN 6554 dorsal view; (b) MNHN 6554 ventral view; (c) MNHN 1994.1671 ventral view; (d) MNHN 1994.1671 lateral view; (e) MNHN 1994.1670 dorsal view; (f) MNHN 1994.1670 ventral view." figureDoi="http://doi.org/10.5281/zenodo.4931107" httpUri="https://zenodo.org/record/4931107/files/figure.png" pageId="91" pageNumber="92">Fig. 72</figureCitation>
and 
<figureCitation box="[303,334,1817,1842]" captionStart="FIGURE 73" captionStartId="92.[151,250,1244,1266]" captionTargetBox="[151,1436,353,1217]" captionTargetId="figure-386@92.[151,1436,353,1217]" captionTargetPageId="92" captionText="FIGURE 73. Syntypes of Anolis boulengerianus: (a) MNHN 6554 dorsal view of head; (b) MNHN 6554 lateral view of head; (c) MNHN 6554 ventral view of head; (d) MNHN 6554 details of dorsum." figureDoi="http://doi.org/10.5281/zenodo.4931109" httpUri="https://zenodo.org/record/4931109/files/figure.png" pageId="91" pageNumber="92">73</figureCitation>
), examined by GK, from “Téhuantépec (
<collectingCountry box="[829,937,1817,1842]" name="Mexico" pageId="91" pageNumber="92">Mexique</collectingCountry>
).” For most of the time since its original description, this nominal species had remained in the synonymy of 
<taxonomicName baseAuthorityName="Wiegmann" baseAuthorityYear="1834" box="[927,1073,1853,1877]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="nebulosus">
<emphasis box="[927,1073,1853,1877]" italics="true" pageId="91" pageNumber="92">A. nebulosus</emphasis>
</taxonomicName>
or 
<taxonomicName authorityName=": Gehlbach &amp; Colette" authorityYear="1957" box="[1115,1269,1853,1877]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[1115,1269,1853,1877]" italics="true" pageId="91" pageNumber="92">A. nebuloides</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation author="Barbour, T." pageId="91" pageNumber="92" pagination="121 - 155" refId="ref87861" refString="Barbour, T. (1934) The anoles. II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology at Harvard College, 77, 121 - 155." type="journal article" year="1934">Barbour 1934</bibRefCitation>
, 
<bibRefCitation author="Smith, H. M. &amp; Taylor, E. H." box="[221,495,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 253" refId="ref97125" refString="Smith, H. M. &amp; Taylor, E. H. (1950 a) An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bulletin of the United States National Museum, 199, 1 - 253." type="journal article" year="1950">
Smith and Taylor 1950 
<bibRefCitation author="Peters, J. A." box="[482,495,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 37" refId="ref94820" refString="Peters, J. A. (1954) The amphibians and reptiles of the coast and coastal sierra of Michoacan, Mexico. Occasional Papers of the Museum of Zoology University of Michigan, 554, 1 - 37." type="journal article" year="1954">a</bibRefCitation>
</bibRefCitation>
, Guibé 1954, 
<bibRefCitation author="Brygoo, E. R." box="[653,805,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 112" refId="ref88294" refString="Brygoo, E. R. (1989) Les types d'Iguanides (Reptiles, Sauriens) du Museum national d'Histoire naturelle Catalogue critique. Bulletin Museum National d'Histoire Naturelle, 4, Series 11, Section A, No. 3 (Supplement), 1 - 112." type="book chapter" year="1989">Brygoo 1989</bibRefCitation>
). 
<bibRefCitation author="Davis, W. B." box="[824,972,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 6" refId="ref89150" refString="Davis, W. B. (1954) Three new anoles from Mexico. Herpetologica, 10 (1), 1 - 6." type="journal article" year="1954">Davis (1954)</bibRefCitation>
described 
<taxonomicName authorityName="Davis" authorityYear="1954" box="[1094,1257,1889,1913]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="subocularis">
<emphasis box="[1094,1257,1889,1913]" italics="true" pageId="91" pageNumber="92">A. subocularis</emphasis>
</taxonomicName>
from “one mile southwest of Tierra Colorado, 
<quantity box="[494,570,1925,1950]" metricMagnitude="2" metricUnit="m" metricValue="2.7432" pageId="91" pageNumber="92" unit="ft" value="900.0">900 ft.</quantity>
, 
<collectingRegion box="[582,685,1925,1950]" country="Mexico" name="Guerrero" pageId="91" pageNumber="92">Guerrero</collectingRegion>
” (
<typeStatus box="[711,811,1925,1950]" pageId="91" pageNumber="92">holotype</typeStatus>
now TCWC 8675, 
<figureCitation box="[1032,1115,1925,1950]" captionStart="FIGURE 74" captionStartId="93.[151,250,1552,1574]" captionTargetBox="[155,1436,202,1508]" captionTargetId="figure-67@93.[155,1440,202,1508]" captionTargetPageId="93" captionText="FIGURE 74. Holotype of Anolis subocularis (TCWC 8675): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a–c) and 1.0 mm in (d–f), respectively." figureDoi="http://doi.org/10.5281/zenodo.4931111" httpUri="https://zenodo.org/record/4931111/files/figure.png" pageId="91" pageNumber="92">Fig. 74</figureCitation>
), examined by GK. In 1978, Fitch described 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[329,466,1961,1985]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="isthmicus">
<emphasis box="[329,466,1961,1985]" italics="true" pageId="91" pageNumber="92">A. isthmicus</emphasis>
</taxonomicName>
based on specimens from “ 
<quantity box="[774,868,1961,1986]" metricMagnitude="4" metricUnit="m" metricValue="1.95" pageId="91" pageNumber="92" unit="km" value="19.5">19.5 km</quantity>
WNW Tehuantepec, 
<collectingRegion box="[1110,1197,1961,1986]" country="Mexico" name="Oaxaca" pageId="91" pageNumber="92">Oaxaca</collectingRegion>
, 
<collectingCountry box="[1207,1295,1961,1986]" name="Mexico" pageId="91" pageNumber="92">Mexico</collectingCountry>
” and nearby localities. In his comparisons, he differentiated it from a selection of anole species from southern 
<collectingCountry box="[1256,1344,1997,2022]" name="Mexico" pageId="91" pageNumber="92">Mexico</collectingCountry>
, but did not include 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[282,487,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[282,487,152,176]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
in his considerations. Recently, 
<bibRefCitation author="Nieto Montes de Oca, A. &amp; Kohler, G. &amp; Feria-Ortiz, M." box="[848,1230,151,176]" pageId="92" pageNumber="93" pagination="125 - 133" refId="ref94533" refString="Nieto Montes de Oca, A., Kohler, G. &amp; Feria-Ortiz, M. (2014) Anolis boulengerianus Thominot 1887, a senior synonym of Anolis isthmicus Fitch 1978 (Squamata: Dactyloidae). Zootaxa, 3794 (1), 125 - 133. http: // dx. doi. org / 10.11646 / zootaxa. 3794.1.5" type="journal article" year="2014">
Nieto Montes de Oca 
<emphasis box="[1093,1149,152,176]" italics="true" pageId="92" pageNumber="93">et al.</emphasis>
(2014)
</bibRefCitation>
demonstrated that the type material of 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[375,580,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[375,580,188,212]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
is conspecific with that of 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[883,1020,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis box="[883,1020,188,212]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
and therefore synonymized the latter with 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[208,413,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[208,413,224,248]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
. Because of priority, 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[654,860,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[654,860,224,248]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
is the valid name for the species previously known as 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[181,318,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis box="[181,318,260,284]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931109" ID-Zenodo-Dep="4931109" httpUri="https://zenodo.org/record/4931109/files/figure.png" pageId="92" pageNumber="93" startId="92.[151,250,1244,1266]" targetBox="[151,1436,353,1217]" targetPageId="92">
<paragraph blockId="92.[151,1436,1244,1296]" pageId="92" pageNumber="93">
<emphasis bold="true" box="[151,288,1244,1266]" pageId="92" pageNumber="93">FIGURE 73.</emphasis>
Syntypes of 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[420,649,1244,1266]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[420,649,1244,1266]" italics="true" pageId="92" pageNumber="93">Anolis boulengerianus</emphasis>
</taxonomicName>
: (a) MNHN 6554 dorsal view of head; (b) MNHN 6554 lateral view of head; (c) MNHN 6554 ventral view of head; (d) MNHN 6554 details of dorsum.
</paragraph>
</caption>
<paragraph blockId="92.[151,1437,1366,1895]" pageId="92" pageNumber="93">
For our analysis, we defined three OTUs in this complex: OTU1 is formed by specimens from south-central and southeastern 
<collectingRegion box="[346,451,1402,1427]" country="Mexico" name="Guerrero" pageId="92" pageNumber="93">Guerrero</collectingRegion>
; OTU2 by specimens from the coastal plain between Huatulco and Puerto Escondido in south-central 
<collectingRegion box="[305,393,1438,1463]" country="Mexico" name="Oaxaca" pageId="92" pageNumber="93">Oaxaca</collectingRegion>
; and OTU 3 by specimens from the hills west and northwest of Tehuantepec in southeastern 
<collectingRegion box="[151,239,1474,1499]" country="Mexico" name="Oaxaca" pageId="92" pageNumber="93">Oaxaca</collectingRegion>
. The analysis of the CO1 and 16s gene fragments revealed the existence of three genetic clusters (
<figureCitation box="[1340,1424,1474,1499]" captionStart="FIGURE 75" captionStartId="94.[151,250,1931,1953]" captionTargetBox="[251,1331,200,1904]" captionTargetId="figure-82@94.[344,1242,195,1227]" captionTargetPageId="94" captionText="FIGURE 75. Phylogenetic tree of specimens of the species related to Anolis boulengerianus from a maximum-likelihood analysis of DNA sequences of two mitochondrial genes: (a) 16S rRNA and (b) CO1. A scale bar is indicated. The numbers at nodes are bootstrap values. The tree is rooted with the species Anolis macrinii. For designation of OTUs see text." figureDoi="http://doi.org/10.5281/zenodo.4931113" httpUri="https://zenodo.org/record/4931113/files/figure.png" pageId="92" pageNumber="93">Fig. 75</figureCitation>
). Genetic distances between our OTUs 2 and 3 are low in both genes (5.7–7.7% in CO1; 
<quantity box="[1129,1242,1510,1535]" metricMagnitude="-2" metricUnit="m" metricValue="5.460999999999999" metricValueMax="7.111999999999999" metricValueMin="3.81" pageId="92" pageNumber="93" unit="in" value="2.15" valueMax="2.8" valueMin="1.5">1.5–2.8 in</quantity>
16s). In contrast, our OTU1 has a genetic distance to OTU 2 and 3, respectively, of 13.9–17.6 (CO1) and 6.9–7.8 (16s), respectively. We observed differences in male dewlap coloration between OTUs 2 and 3: in OTU 2, the male dewlap is pinkish red without paler areas around gorgetals whereas in OTU 3 the male dewlap is orange yellow with paler areas around gorgetals. Also, the eastern populations (our OTU 3) occur in a much more arid habitat than the western populations (our OTU 2). These findings are evidence for the existence of three species in this complex. Our OTU 1 contains the 
<typeStatus box="[319,367,1727,1751]" pageId="92" pageNumber="93">type</typeStatus>
locality of 
<taxonomicName authorityName="Davis" authorityYear="1954" box="[502,666,1726,1750]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="subocularis">
<emphasis box="[502,666,1726,1750]" italics="true" pageId="92" pageNumber="93">A. subocularis</emphasis>
</taxonomicName>
and therefore this name has to be applied to this species. The 
<typeStatus box="[1388,1436,1727,1751]" pageId="92" pageNumber="93">type</typeStatus>
localities of 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[289,493,1763,1787]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[289,493,1763,1787]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[548,684,1763,1787]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis box="[548,684,1763,1787]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
, respectively, are both contained in our OTU3; thus the older name (i.e., 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[212,420,1798,1822]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[212,420,1798,1822]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
) is the valid name for this species and the nominal species 
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[1121,1261,1798,1822]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis box="[1121,1261,1798,1822]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
remains in the synonymy of 
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[307,514,1835,1859]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[307,514,1835,1859]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
. No scientific name is available for our OTU 2, and we therefore describe it as a new species below.
</paragraph>
</subSubSection>
</treatment>
</document>