treatments-xml/data/06/5B/60/065B60075771FFC8FF22FCF30172FDCF.xml

<document id="C12022DD1D0346A44888B0C3B81E5D38" ID-CLB-Dataset="31874" ID-DOI="10.11646/zootaxa.4329.4.4" ID-GBIF-Dataset="df8e24d8-b6f2-4006-a065-dbae51af1600" ID-ISSN="1175-5326" ID-Zenodo-Dep="1003034" ID-ZooBank="E44A66Be-9226-4769-9B02-Eb681B705E00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507274003446" checkinUser="plazi" docAuthor="Hernández-Corral, Jesús" docDate="2017" docId="065B60075771FFC8FF22FCF30172FDCF" docLanguage="en" docName="zootaxa.4329.4.4.pdf" docOrigin="Zootaxa 4329 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysdera arnedoi Hernández-Corral 2017, new species" docType="treatment" docVersion="10" lastPageNumber="380" masterDocId="FA62187F5770FFCDFFB5FFEA033EFFAF" masterDocTitle="The spider family Dysderidae in the Balearic Islands" masterLastPageNumber="391" masterPageNumber="375" pageNumber="376" updateTime="1698469381106" updateUser="ExternalLinkService">
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<mods:title id="5F42AED2D70AA9B81A273C97B1BF260F">The spider family Dysderidae in the Balearic Islands</mods:title>
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<mods:namePart id="2AC682E5A38DD9BC23AEF2E7DEA84CEE">Hernández-Corral, Jesús</mods:namePart>
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<paragraph id="8E4DD1115771FFCCFF22FCF30164FC9C" blockId="1.[151,602,793,853]" box="[151,602,793,819]" pageId="1" pageNumber="376">
<heading id="D505667D5771FFCCFF22FCF30164FC9C" bold="true" box="[151,602,793,819]" fontSize="11" level="1" pageId="1" pageNumber="376" reason="1">
<emphasis id="BC860D035771FFCCFF22FCF30164FC9C" bold="true" box="[151,602,793,819]" pageId="1" pageNumber="376">
<taxonomicName id="49F2AA925771FFCCFF22FCF302FCFC9C" ID-CoL="388YC" authority="Lissner" authorityName="Hernández-Corral" authorityYear="2017" box="[151,450,793,819]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035771FFCCFF22FCF30260FC9C" bold="true" box="[151,350,793,819]" italics="true" pageId="1" pageNumber="376">Dysdera arnedoi</emphasis>
Lissner
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785771FFCCFE7FFCF30164FC9C" box="[458,602,793,819]" pageId="1" pageNumber="376" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8E4DD1115771FFCCFF22FCD60249FCFB" blockId="1.[151,602,793,853]" box="[151,375,828,853]" pageId="1" pageNumber="376">
<figureCitation id="16C9CD945771FFCCFF22FCD603E2FCFB" box="[151,220,828,853]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1920,1943]" captionTargetBox="[215,1372,193,1894]" captionTargetId="figure@2.[215,1372,193,1894]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fig. 1. Dysdera arnedoi Lissner new species. Female, dorsalateral view. Figs 2 – 3. Dysdera crocata C. L. Koch (from St Elm). 2. Male, dorsal view; 3. Female, dorsal view; Figs 4 – 5. Dysdera lata ReUss, 1834; 4. Male, dorsal view; 5. Female, dorsal view. Fig. 6. Harpactea dufouri (Thorell, 1839); Male, dorsal view. Fig. 7. Parachtes riberai Bosmans new species; Male, dorsal view." httpUri="https://zenodo.org/record/1003036/files/figure.png" pageId="1" pageNumber="376">Figs 1</figureCitation>
, 
<figureCitation id="16C9CD945771FFCCFF5FFCD6021BFCFB" box="[234,293,828,852]" captionStart="FIGURES 8 – 16" captionStartId="3.[151,264,1870,1893]" captionTargetBox="[239,1336,222,1803]" captionTargetId="figure@3.[226,1344,193,1843]" captionTargetPageId="3" captionText="FIGURES 8 – 16. Figs 8 – 14. Dysdera arnedoi Lissner new species; 8. Prosoma, dorsal view; 9. Left chelicera, ventral view; 10. FemUr IV, dorsal view; 11. MetatarsUs and tarsUs I, lateral view; 12. Tarsal claw, pectination only drawn for one claw; 13. VUlva, dorsal view; 14. VUlva, ventral view. Figs 15 – 16. Harpactea dufouri (Thorell, 1873). 15. Male palp, retrolateral view (from Estellencs); 16. VUlva, dorsal view (from Palma, Castilla de San Carlos). Scale bars: 8: 2 mm, 9: 0.5 mm, 10, 11: 1 mm, 12: 0.1 mm, 13 – 16: 0.2 mm." httpUri="https://zenodo.org/record/1003038/files/figure.png" pageId="1" pageNumber="376">8–14</figureCitation>
, 
<figureCitation id="16C9CD945771FFCCFE85FCD60272FCFB" box="[304,332,828,852]" captionStart="FIGURES 27 – 30" captionStartId="10.[151,264,1896,1919]" captionTargetBox="[151,1436,193,1847]" captionTargetId="figure@10.[151,1436,193,1876]" captionTargetPageId="10" captionText="FIGURES 27 – 30. Figs 27 – 28. Dysdera balearica Thorell, 1873; 27. VUlva, ventral view; 28. Idem, dorsal view. Fig. 29. Dysdera lata ReUss, 1834. VUlva, dorsal view. Figs 30 – 32. Dysdera arnedoi new species. 30. VUlva, dorsal view with transversal bar still in place; 31. Idem, ventral view; 32. Idem, dorsal view. Scale bars 0.2 mm." httpUri="https://zenodo.org/record/1003044/files/figure.png" pageId="1" pageNumber="376">30</figureCitation>
–32
</paragraph>
</subSubSection>
<subSubSection id="C6E8829A5771FFCCFF22FC6E0188FC6F" pageId="1" pageNumber="376" type="materials_examined">
<paragraph id="8E4DD1115771FFCCFF22FC6E0188FC6F" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">
<materialsCitation id="3E9ADB4C5771FFCCFF22FC6E018CFC6F" ID-GBIF-Occurrence="1664833317" collectingDate="2014-04-23" collectionCode="NHMD" collectorName="J. Lissner" country="Spain" county="Cova Morella" elevation="505" latitude="39.84436" location="Majorca" longLatPrecision="1" longitude="2.9840279" municipality="Pollenca" pageId="1" pageNumber="376" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="51496FB35771FFCCFF22FC6E03ECFC32" box="[151,210,900,925]" pageId="1" pageNumber="376">Type</typeStatus>
material.
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<typeStatus id="51496FB35771FFCCFEFEFC6E028DFC32" box="[331,435,900,925]" pageId="1" pageNumber="376" type="holotype">Holotype</typeStatus>
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from 
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, 
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, 
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, 
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, cave (
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), 
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, 
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<date id="FA4CF7D15771FFCCFF22FC42022BFC6F" box="[151,277,936,960]" pageId="1" pageNumber="376" value="2014-04-23">23.IV.2014</date>
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, 
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leg.; deposited at 
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.
</paragraph>
</subSubSection>
<subSubSection id="C6E8829A5771FFCCFF72FC260290FB4E" pageId="1" pageNumber="376" type="diagnosis">
<paragraph id="8E4DD1115771FFCCFF72FC260290FB4E" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">
<emphasis id="BC860D035771FFCCFF72FC26027FFC4A" bold="true" box="[199,321,972,997]" pageId="1" pageNumber="376">Diagnosis.</emphasis>
The vulva seems closest to those of its Majorcan congener 
<taxonomicName id="49F2AA925771FFCCFC5CFC260663FC4B" authority="Thorell, 1873" authorityName="Thorell" authorityYear="1873" box="[1001,1373,972,997]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="balearica">
<emphasis id="BC860D035771FFCCFC5CFC260785FC4B" box="[1001,1211,972,996]" italics="true" pageId="1" pageNumber="376">Dysdera balearica</emphasis>
<bibRefCitation id="EA63ACE05771FFCCFB71FC260663FC4B" author="Thorell" box="[1220,1373,972,997]" pageId="1" pageNumber="376" refString="Thorell, T. (1873) Remarks on synonyms of European Spiders. Part IV. Uppsala, pp. 375 - 645." type="book chapter" year="1873">Thorell, 1873</bibRefCitation>
</taxonomicName>
from which it differs by having 6 spines on Fe IV compared to 
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<taxonomicName id="49F2AA925771FFCCFC36FC1B072CFBA7" authorityName="Thorell" authorityYear="1873" box="[899,1042,1008,1032]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="balearica">
<emphasis id="BC860D035771FFCCFC36FC1B072CFBA7" box="[899,1042,1008,1032]" italics="true" pageId="1" pageNumber="376">D. balearica</emphasis>
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and by having the cheliceral teeth equidistant while the distal tooth in 
<taxonomicName id="49F2AA925771FFCCFD9AFBFF0180FB83" authorityName="Thorell" authorityYear="1873" box="[559,702,1044,1068]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="balearica">
<emphasis id="BC860D035771FFCCFD9AFBFF0180FB83" box="[559,702,1044,1068]" italics="true" pageId="1" pageNumber="376">D. balearica</emphasis>
</taxonomicName>
is well separated from the two basal ones. The new species also differs by possessing troglobiomorphic features, such as appendage elongation, slight depigmentation and eye reduction, absent in 
<taxonomicName id="49F2AA925771FFCCFECEFBB70137FBDB" authorityName="Thorell" authorityYear="1873" box="[379,521,1116,1140]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="balearica">
<emphasis id="BC860D035771FFCCFECEFBB70137FBDB" box="[379,521,1116,1140]" italics="true" pageId="1" pageNumber="376">D. balearica</emphasis>
</taxonomicName>
. The vulva of 
<taxonomicName id="49F2AA925771FFCCFD1AFBB70016FBDB" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[687,808,1116,1140]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035771FFCCFD1AFBB70016FBDB" box="[687,808,1116,1140]" italics="true" pageId="1" pageNumber="376">D. arnedoi</emphasis>
</taxonomicName>
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<emphasis id="BC860D035771FFCCFC8DFBB60049FBDA" bold="true" box="[824,887,1116,1141]" pageId="1" pageNumber="376">n. sp.</emphasis>
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resembles those of 
<taxonomicName id="49F2AA925771FFCCFBE2FBB706A6FBDA" authority="Alicata, 1964" authorityName="Alicata" authorityYear="1964" box="[1111,1432,1116,1141]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="apenninica">
<emphasis id="BC860D035771FFCCFBE2FBB707C9FBDB" box="[1111,1271,1117,1140]" italics="true" pageId="1" pageNumber="376">D. apenninica</emphasis>
<bibRefCitation id="EA63ACE05771FFCCFB48FBB606A6FBDA" author="Alicata" box="[1277,1432,1116,1141]" pageId="1" pageNumber="376" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata, 1964</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="49F2AA925771FFCCFF22FB6B015BFB36" authority="Alicata, 1964" authorityName="Alicata" authorityYear="1964" box="[151,613,1152,1177]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="subSpecies" species="apenninica" subSpecies="aprutiana">
<emphasis id="BC860D035771FFCCFF22FB6B0287FB37" box="[151,441,1153,1176]" italics="true" pageId="1" pageNumber="376">D. apenninica aprutiana</emphasis>
<bibRefCitation id="EA63ACE05771FFCCFE72FB6A015BFB36" author="Alicata" box="[455,613,1152,1177]" pageId="1" pageNumber="376" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata, 1964</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="49F2AA925771FFCCFD1CFB6B00E3FB36" authority="Alicata, 1964" authorityName="Alicata" authorityYear="1964" box="[681,989,1152,1177]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="BC860D035771FFCCFD1CFB6B0010FB37" box="[681,814,1152,1176]" italics="true" pageId="1" pageNumber="376">D. lagrecai</emphasis>
<bibRefCitation id="EA63ACE05771FFCCFC89FB6A00E3FB36" author="Alicata" box="[828,989,1152,1177]" pageId="1" pageNumber="376" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata, 1964</bibRefCitation>
</taxonomicName>
, but none of these species share the troglobiomorphic features found in 
<taxonomicName id="49F2AA925771FFCCFD92FB4F019FFB13" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[551,673,1188,1212]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035771FFCCFD92FB4F019FFB13" box="[551,673,1188,1212]" italics="true" pageId="1" pageNumber="376">D. arnedoi</emphasis>
</taxonomicName>
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<emphasis id="BC860D035771FFCCFD1DFB4E01D9FB12" bold="true" box="[680,743,1188,1213]" pageId="1" pageNumber="376">n. sp.</emphasis>
</taxonomicNameLabel>
Comparisons to other congeners of this large genus are made in the comments section.
</paragraph>
</subSubSection>
<subSubSection id="C6E8829A5771FFCCFF72FB06006BFA86" pageId="1" pageNumber="376" type="etymology">
<paragraph id="8E4DD1115771FFCCFF72FB06006BFA86" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">
<emphasis id="BC860D035771FFCCFF72FB060270FAAA" bold="true" box="[199,334,1260,1285]" pageId="1" pageNumber="376">Etymology.</emphasis>
The species is dedicated to Miquel Arnedo in recognition of his important studies on woodlouse spiders, in particular those on species from the 
<collectingRegion id="4C361FF35771FFCCFD12FAFA006CFA86" box="[679,850,1296,1321]" country="Spain" name="Canarias" pageId="1" pageNumber="376">Canary Islands</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C6E8829A5771FFC9FF72FADE02FEFF1F" lastPageId="4" lastPageNumber="379" pageId="1" pageNumber="376" type="description">
<paragraph id="8E4DD1115771FFCCFF72FADE02ABFADE" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">
<emphasis id="BC860D035771FFCCFF72FADE02F9FAE2" bold="true" box="[199,455,1332,1357]" pageId="1" pageNumber="376">Description. Female.</emphasis>
Measurements: Female (n=1): Total length 11.3; prosoma 4.1 long, 3.0 wide. Opisthosoma 7.3 long.
</paragraph>
<paragraph id="8E4DD1115771FFCCFF72FA960134F9AE" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">Colour: Cephalothorax orange red in live specimen fading to pale yellow in alcohol preserved specimen, head region slightly darker than thoracic region (fig. 1). Cephalothorax covered with fairly long dark hairs. Chelicerae slightly darker than head. Legs orange with leg I and II darker than III and IV. Abdomen pale covered with uniformly distributed short hairs.</paragraph>
<paragraph id="8E4DD1115771FFCCFF72F9E707C9F9C2" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">Eyes (fig. 8): Sub-equal, rather small for the size of the spider; AM diameter 0.16; PL 0.17; PM 0.14; AM near anterior rim of frontal border and separated from each other by a little more than a diameter of an eye; PM nearly touching and separated from PL by less than half a diameter of PM; width of eye group equals 0.61.</paragraph>
<paragraph id="8E4DD1115771FFCCFF72F99201E0F93F" blockId="1.[151,1437,900,2005]" box="[199,734,1656,1681]" pageId="1" pageNumber="376">Chelicerae: With three equidistant teeth (fig. 9).</paragraph>
<paragraph id="8E4DD1115771FFCCFF72F97707BCF91B" blockId="1.[151,1437,900,2005]" box="[199,1154,1692,1717]" pageId="1" pageNumber="376">Legs: This is a long-legged species (figs. 1, 11). Leg lengths are presented in Table 2.</paragraph>
<paragraph id="8E4DD1115771FFCCFF72F92B012DF8EA" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">Leg spination: Legs I–II spineless. All patellae and tarsi spineless. Fe III spineless, Fe IV with 6 dorsal spines distributed as in fig. 10. Five of these spines are in a row while the sixth is slightly displaced prolaterally. Spination of Ti III, IV and Mt III, IV is listed in table 3. Tarsal claws large, each with 12 well-developed pectinate teeth (fig. 12, 2 small teeth hidden by claw).</paragraph>
<paragraph id="8E4DD1115771FFCCFF72F8BA01CEF87A" blockId="1.[151,1437,900,2005]" pageId="1" pageNumber="376">Vulva (figs 13–14, 30–32). Spermathecae oval, dorsal arch of anterior diverticulum in dorsal view with an anterior structure; bursal valve and transversal bar only seen in image of freshly removed vulva (fig. 30), during digestion in KOH they became detached from the anterior diverticulum. The lateral arms of the dorsal arch of anterior diverticulum do not reach the transversal bar.</paragraph>
<caption id="DA8D81995772FFCFFF22F86A0297F85C" httpUri="https://zenodo.org/record/1003036/files/figure.png" pageId="2" pageNumber="377" targetBox="[215,1372,193,1894]" targetPageId="2">
<paragraph id="8E4DD1115772FFCFFF22F86A0297F85C" blockId="2.[151,1436,1920,2035]" pageId="2" pageNumber="377">
<emphasis id="BC860D035772FFCFFF22F86A0207F839" bold="true" box="[151,313,1920,1943]" pageId="2" pageNumber="377">FIGURES 1–7.</emphasis>
Fig. 1. 
<taxonomicName id="49F2AA925772FFCFFE33F86B01BFF838" authority="Lissner" authorityName="Hernández-Corral" authorityYear="2017" box="[390,641,1921,1943]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035772FFCFFE33F86B010FF838" box="[390,561,1921,1943]" italics="true" pageId="2" pageNumber="377">Dysdera arnedoi</emphasis>
Lissner
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785772FFCFFD32F868003DF838" box="[647,771,1921,1943]" pageId="2" pageNumber="377" rank="species">new species</taxonomicNameLabel>
. Female, dorsalateral view. Figs 2–3. 
<taxonomicName id="49F2AA925772FFCFFBCBF86B06A2F838" authority="C. L. Koch" authorityName="C. L. Koch" authorityYear="1838" box="[1150,1436,1921,1943]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="crocata">
<emphasis id="BC860D035772FFCFFBCBF86B0618F838" box="[1150,1318,1921,1943]" italics="true" pageId="2" pageNumber="377">Dysdera crocata</emphasis>
C. L. Koch
</taxonomicName>
(from St Elm). 2. Male, dorsal view; 3. Female, dorsal view; Figs 4–5. 
<taxonomicName id="49F2AA925772FFCFFC30F87507A8F81A" authority="ReUss, 1834" authorityName="Reuss" authorityYear="1834" box="[901,1174,1951,1973]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="BC860D035772FFCFFC30F8750735F81A" box="[901,1035,1951,1973]" italics="true" pageId="2" pageNumber="377">Dysdera lata</emphasis>
ReUss, 1834
</taxonomicName>
; 4. Male, dorsal view; 5. Female, dorsal view. Fig. 6. 
<taxonomicName id="49F2AA925772FFCFFE76F8550015F87B" authority="(Thorell, 1839)" authorityName="Thorell" authorityYear="1839" baseAuthorityName="Thorell" baseAuthorityYear="1839" box="[451,811,1982,2004]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="dufouri">
<emphasis id="BC860D035772FFCFFE76F85501BDF87B" box="[451,643,1982,2004]" italics="true" pageId="2" pageNumber="377">Harpactea dufouri</emphasis>
(Thorell, 1839)
</taxonomicName>
; Male, dorsal view. Fig. 7. 
<taxonomicName id="49F2AA925772FFCFFBF8F8540657F87B" authority="Bosmans" authorityName="Hernández-Corral" authorityYear="2017" box="[1101,1385,1982,2004]" class="Arachnida" family="Dysderidae" genus="Parachtes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">
<emphasis id="BC860D035772FFCFFBF8F854063DF87B" box="[1101,1283,1982,2004]" italics="true" pageId="2" pageNumber="377">Parachtes riberai</emphasis>
Bosmans
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785772FFCFFAC7F85503DDF85C" pageId="2" pageNumber="377" rank="species">new species</taxonomicNameLabel>
; Male, dorsal view.
</paragraph>
</caption>
<caption id="DA8D81995773FFCEFF22F8A40287F870" httpUri="https://zenodo.org/record/1003038/files/figure.png" pageId="3" pageNumber="378" targetBox="[239,1336,222,1803]" targetPageId="3">
<paragraph id="8E4DD1115773FFCEFF22F8A40287F870" blockId="3.[151,1436,1870,2015]" pageId="3" pageNumber="378">
<emphasis id="BC860D035773FFCEFF22F8A40278F8CB" bold="true" box="[151,326,1870,1893]" pageId="3" pageNumber="378">FIGURES 8–16.</emphasis>
Figs 8–14. 
<taxonomicName id="49F2AA925773FFCEFE09F8A50189F8CA" authority="Lissner" authorityName="Hernández-Corral" authorityYear="2017" box="[444,695,1871,1893]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035773FFCEFE09F8A50159F8CA" box="[444,615,1871,1893]" italics="true" pageId="3" pageNumber="378">Dysdera arnedoi</emphasis>
Lissner
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785773FFCEFD09F8BA0006F8CA" box="[700,824,1871,1893]" pageId="3" pageNumber="378" rank="species">new species</taxonomicNameLabel>
; 8. Prosoma, dorsal view; 9. Left chelicera, ventral view; 10. FemUr IV, dorsal view; 11. MetatarsUs and tarsUs I, lateral view; 12. Tarsal claw, pectination only drawn for one claw; 13. VUlva, dorsal view; 14. VUlva, ventral view. Figs 15–16. 
<taxonomicName id="49F2AA925773FFCEFD53F8670772F80D" authority="(Thorell, 1873)" authorityName="Thorell" authorityYear="1873" baseAuthorityName="Thorell" baseAuthorityYear="1873" box="[742,1100,1932,1954]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="dufouri">
<emphasis id="BC860D035773FFCEFD53F867009BF80D" box="[742,933,1932,1954]" italics="true" pageId="3" pageNumber="378">Harpactea dufouri</emphasis>
(Thorell, 1873)
</taxonomicName>
. 15. Male palp, retrolateral view (from Estellencs); 16. VUlva, dorsal view (from Palma, Castilla de San Carlos). Scale bars: 8: 2 mm, 9: 0.5 mm, 10, 11: 1 mm, 12: 0.1 mm, 13–16: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8E4DD1115774FFC9FF72FF7D02FEFF1F" blockId="4.[151,1436,151,717]" box="[199,448,151,176]" pageId="4" pageNumber="379">The male is unknown.</paragraph>
</subSubSection>
<subSubSection id="C6E8829A5774FFC8FF72FF5603E1FE7F" lastPageId="5" lastPageNumber="380" pageId="4" pageNumber="379" type="discussion">
<paragraph id="8E4DD1115774FFC9FF72FF56001EFD62" blockId="4.[151,1436,151,717]" pageId="4" pageNumber="379">
<emphasis id="BC860D035774FFC9FF72FF560271FF7A" bold="true" box="[199,335,188,213]" pageId="4" pageNumber="379">Comments.</emphasis>
The only troglobiomorphic 
<taxonomicName id="49F2AA925774FFC9FD2EFF5601C4FF7B" authorityName="Latreille" authorityYear="1804" box="[667,762,188,212]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="BC860D035774FFC9FD2EFF5601C4FF7B" box="[667,762,188,212]" italics="true" pageId="4" pageNumber="379">Dysdera</emphasis>
</taxonomicName>
species known from Majorca so far. The species exhibits pronounced leg elongation, slight depigmentation and reduced width of eye group and eye size compared to epigean species. The new species was compared to the descriptions and illustrations of the 142 species presently known to occur in Europe (Nentwig 
<emphasis id="BC860D035774FFC9FD89FEC30148FEEF" box="[572,630,296,320]" italics="true" pageId="4" pageNumber="379">et al.</emphasis>
, 2016; Le 
<bibRefCitation id="EA63ACE05774FFC9FD40FEC20053FEEF" author="Peru" box="[757,877,296,320]" pageId="4" pageNumber="379" refString="Le PerU, B. (2011) The spiders of Europe, a synthesis of data: Uolume 1 Atypidae to Theridiidae. Mem Soc Linn Lyon, 522 pp." type="book" year="2011">
<collectingCountry id="F6E591815774FFC9FD40FEC20016FE90" box="[757,808,296,319]" name="Peru" pageId="4" pageNumber="379">Peru</collectingCountry>
2011
</bibRefCitation>
). The combination of characters possessed by 
<taxonomicName id="49F2AA925774FFC9FA34FEC303D0FECB" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FA34FEC303D0FECB" italics="true" pageId="4" pageNumber="379">D. arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FF41FEA6020CFECA" box="[244,306,332,357]" pageId="4" pageNumber="379" rank="species">
<emphasis id="BC860D035774FFC9FF41FEA6020CFECA" bold="true" box="[244,306,332,357]" pageId="4" pageNumber="379">n. sp.</emphasis>
</taxonomicNameLabel>
, in particular troglobiomorphism, leg spination, prosoma scabrousness, cheliceral dentation, shape of female genitalia and habitat occupied prevents targeting close relatives except for 
<taxonomicName id="49F2AA925774FFC9FB9DFE9B078BFE27" authorityName="Thorell" authorityYear="1873" box="[1064,1205,368,392]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="balearica">
<emphasis id="BC860D035774FFC9FB9DFE9B078BFE27" box="[1064,1205,368,392]" italics="true" pageId="4" pageNumber="379">D. balearica</emphasis>
</taxonomicName>
, as mentioned in the diagnosis section. In the process of confirming a new species was at hand the specimen was compared to descriptions of other cave dwelling members of the genus (
<taxonomicName id="49F2AA925774FFC9FCF3FE5207A8FE7F" authority="Fage, 1931" authorityName="Fage" authorityYear="1931" box="[838,1174,440,464]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="bicornis">
<emphasis id="BC860D035774FFC9FCF3FE520734FE7F" box="[838,1034,440,464]" italics="true" pageId="4" pageNumber="379">Dysdera bicornis</emphasis>
<bibRefCitation id="EA63ACE05774FFC9FBA1FE5207A8FE7F" author="Fage" box="[1044,1174,440,464]" pageId="4" pageNumber="379" refString="Fage, L. (1931) Biospeologica LV. Araneae, 5 ª Serie, precedee d'Un esai sUr l'evolUtion soUterraine et son determinisme. Archives de Zoologie Experimentale et Generale, 71, 91 - 291." type="journal article" year="1931">Fage, 1931</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="49F2AA925774FFC9FB11FE53026EFE5A" authority="Ribera &amp; Ferrandez, 1986" authorityName="Ribera &amp; Ferrandez" authorityYear="1986" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="BC860D035774FFC9FB11FE530616FE7F" box="[1188,1320,440,464]" italics="true" pageId="4" pageNumber="379">D. espanoli</emphasis>
Ribera &amp; Ferrández, 1986
</taxonomicName>
, 
<taxonomicName id="49F2AA925774FFC9FEEEFE3701A6FE5B" authority="Caporiacco, 1941" authorityName="Caporiacco" authorityYear="1941" box="[347,664,476,501]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="pavani">
<emphasis id="BC860D035774FFC9FEEEFE3702F7FE5B" box="[347,457,477,500]" italics="true" pageId="4" pageNumber="379">D. pavani</emphasis>
Caporiacco, 1941
</taxonomicName>
, 
<taxonomicName id="49F2AA925774FFC9FD11FE3700F4FE5B" authority="Ribera, 2004" authorityName="Ribera" authorityYear="2004" box="[676,970,476,501]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="valentina">
<emphasis id="BC860D035774FFC9FD11FE370011FE5B" box="[676,815,476,500]" italics="true" pageId="4" pageNumber="379">D. valentina</emphasis>
Ribera, 2004
</taxonomicName>
, 
<taxonomicName id="49F2AA925774FFC9FC60FE37065AFE5A" authority="Ribera &amp; Ferrandez, 1986" authorityName="Ribera &amp; Ferrandez" authorityYear="1986" box="[981,1380,476,501]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="BC860D035774FFC9FC60FE370709FE5B" box="[981,1079,477,500]" italics="true" pageId="4" pageNumber="379">D. vivesi</emphasis>
Ribera &amp; Ferrández, 1986
</taxonomicName>
) and species with the most similar vulva (
<taxonomicName id="49F2AA925774FFC9FD83FDEB0005FDB7" authority="Simon, 1882" authorityName="Simon" authorityYear="1882" box="[566,827,511,536]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="nubila">
<emphasis id="BC860D035774FFC9FD83FDEB019FFDB7" box="[566,673,512,536]" italics="true" pageId="4" pageNumber="379">D. nubila</emphasis>
Simon, 1882
</taxonomicName>
, 
<taxonomicName id="49F2AA925774FFC9FCF2FDEB0769FDB7" authority="Alicata, 1973" authorityName="Alicata" authorityYear="1973" box="[839,1111,511,536]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="osellai">
<emphasis id="BC860D035774FFC9FCF2FDEB008BFDB7" box="[839,949,512,536]" italics="true" pageId="4" pageNumber="379">D. osellai</emphasis>
Alicata, 1973
</taxonomicName>
, 
<taxonomicName id="49F2AA925774FFC9FBD6FDEB06AEFDB7" authority="Simon, 1907" authorityName="Simon" authorityYear="1907" box="[1123,1424,511,536]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="subnubila">
<emphasis id="BC860D035774FFC9FBD6FDEB07CAFDB7" box="[1123,1268,512,536]" italics="true" pageId="4" pageNumber="379">D. subnubila</emphasis>
Simon, 1907
</taxonomicName>
). Some of the distinctive characters separating these species from 
<taxonomicName id="49F2AA925774FFC9FCC3FDCF00CFFD93" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[886,1009,548,572]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FCC3FDCF00CFFD93" box="[886,1009,548,572]" italics="true" pageId="4" pageNumber="379">D. arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FC4FFDCE0704FD92" box="[1018,1082,548,573]" pageId="4" pageNumber="379" rank="species">
<emphasis id="BC860D035774FFC9FC4FFDCE0704FD92" bold="true" box="[1018,1082,548,573]" pageId="4" pageNumber="379">n. sp.</emphasis>
</taxonomicNameLabel>
are presented below (
<tableCitation id="C370E4AA5774FFC9FA83FDCE06B5FD93" box="[1334,1419,548,573]" captionStart="TABLE 1" captionStartId="4.[151,239,760,783]" captionText="TABLE 1. Characteristics separating Dysdera arnedoi n. sp. from selected congeners." pageId="4" pageNumber="379">Table 1</tableCitation>
). 
<taxonomicName id="49F2AA925774FFC9FF22FDA2011FFDCF" authority="Alicata, 1964" authorityName="Alicata" authorityYear="1964" box="[151,545,583,608]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="apenninica">
<emphasis id="BC860D035774FFC9FF22FDA20243FDF0" box="[151,381,584,608]" italics="true" pageId="4" pageNumber="379">Dysdera apenninica</emphasis>
<bibRefCitation id="EA63ACE05774FFC9FE32FDAD011FFDCF" author="Alicata" box="[391,545,583,608]" pageId="4" pageNumber="379" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata, 1964</bibRefCitation>
</taxonomicName>
and the subspecies 
<taxonomicName id="49F2AA925774FFC9FCB8FDA307F3FDCF" authority="Alicata, 1964" authorityName="Alicata" authorityYear="1964" box="[781,1229,583,608]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="subSpecies" species="apenninica" subSpecies="aprutiana">
<emphasis id="BC860D035774FFC9FCB8FDA30717FDF0" box="[781,1065,584,608]" italics="true" pageId="4" pageNumber="379">D. apenninica aprutiana</emphasis>
<bibRefCitation id="EA63ACE05774FFC9FB87FDAD07F3FDCF" author="Alicata" box="[1074,1229,583,608]" pageId="4" pageNumber="379" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata, 1964</bibRefCitation>
</taxonomicName>
(both endemic to central 
<collectingCountry id="F6E591815774FFC9FF5BFD86021DFD2A" box="[238,291,620,645]" name="Italy" pageId="4" pageNumber="379">Italy</collectingCountry>
) and 
<taxonomicName id="49F2AA925774FFC9FEDFFD8701ADFD2A" authority="Alicata, 1964" authorityName="Alicata" authorityYear="1964" box="[362,659,620,645]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="BC860D035774FFC9FEDFFD8702D2FD2B" box="[362,492,620,644]" italics="true" pageId="4" pageNumber="379">D. lagrecai</emphasis>
<bibRefCitation id="EA63ACE05774FFC9FE4DFD8601ADFD2A" author="Alicata" box="[504,659,620,645]" pageId="4" pageNumber="379" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata, 1964</bibRefCitation>
</taxonomicName>
(endemic to 
<collectingRegion id="4C361FF35774FFC9FC86FD860048FD2A" box="[819,886,620,645]" country="Italy" name="Sicilia" pageId="4" pageNumber="379">Sicily</collectingRegion>
) may have similar vulvae according to outline illustrations in 
<bibRefCitation id="EA63ACE05774FFC9FEF6FD6502D4FD07" author="Alicata" box="[323,490,655,680]" pageId="4" pageNumber="379" refString="Alicata, P. (1964) Le Dysdera del grUppo andreinii della faUna italiana (Araneae, Dysderidae). Atti della Accademia gioenia di scienze naturali in Catania, 16 (6), 67 - 86." type="journal article" year="1964">Alicata (1964)</bibRefCitation>
. Both species are epigean and confined to high ground (above 
<quantity id="490A7CF45774FFC9FB75FD7A061FFD07" box="[1216,1313,656,680]" metricMagnitude="0" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="379" unit="m" value="1.3">1.300 m</quantity>
), therefore they are not considered conspecific with 
<taxonomicName id="49F2AA925774FFC9FDD7FD5F01E5FD63" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[610,731,692,716]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FDD7FD5F01E5FD63" box="[610,731,692,716]" italics="true" pageId="4" pageNumber="379">D. arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FD57FD5E001EFD62" box="[738,800,692,717]" pageId="4" pageNumber="379" rank="species">
<emphasis id="BC860D035774FFC9FD57FD5E001EFD62" bold="true" box="[738,800,692,717]" pageId="4" pageNumber="379">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<caption id="DA8D81995774FFC9FF22FD12070FFCA0" box="[151,1073,759,783]" pageId="4" pageNumber="379">
<paragraph id="8E4DD1115774FFC9FF22FD12070FFCA0" blockId="4.[151,1073,759,783]" box="[151,1073,759,783]" pageId="4" pageNumber="379">
<emphasis id="BC860D035774FFC9FF22FD12023DFCA0" bold="true" box="[151,259,760,783]" pageId="4" pageNumber="379">TABLE 1</emphasis>
. Characteristics separating 
<taxonomicName id="49F2AA925774FFC9FD9FFD1D01E0FCA0" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[554,734,759,783]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FD9FFD1D01E0FCA0" box="[554,734,759,783]" italics="true" pageId="4" pageNumber="379">Dysdera arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FD51FD12001FFCA0" box="[740,801,760,783]" pageId="4" pageNumber="379" rank="species">
<emphasis id="BC860D035774FFC9FD51FD12001FFCA0" bold="true" box="[740,801,760,783]" pageId="4" pageNumber="379">n. sp.</emphasis>
</taxonomicNameLabel>
from selected congeners.
</paragraph>
</caption>
<paragraph id="8E4DD1115774FFC9FF2AFCC001CDFAD3" pageId="4" pageNumber="379">
<table id="FCF223B157740032FF22FCC006B3FAD3" box="[151,1421,810,1404]" gridcols="3" gridrows="10" pageId="4" pageNumber="379">
<tr id="30C2D35357740032FF22FCC006B3FCEF" box="[151,1421,810,832]" gridrow="0" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FCC00204FCEF" box="[151,314,810,832]" gridcol="0" gridrow="0" pageId="4" pageNumber="379">Species</th>
<th id="7313BA2F57740032FEF4FCC00058FCEF" box="[321,870,810,832]" gridcol="1" gridrow="0" pageId="4" pageNumber="379">Characters compared</th>
<th id="7313BA2F57740032FCC9FCC006B3FCEF" box="[892,1421,810,832]" gridcol="2" gridrow="0" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FCC9FCC00719FCEF" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[892,1063,810,832]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FCC9FCC00719FCEF" box="[892,1063,810,832]" italics="true" pageId="4" pageNumber="379">Dysdera arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FB98FCC10759FCEF" box="[1069,1127,811,832]" pageId="4" pageNumber="379" rank="species">
<emphasis id="BC860D035774FFC9FB98FCC10759FCEF" bold="true" box="[1069,1127,811,832]" pageId="4" pageNumber="379">n. sp.</emphasis>
</taxonomicNameLabel>
</th>
</tr>
<tr id="30C2D35357740032FF22FCBE06B3FCC5" box="[151,1421,852,874]" gridrow="1" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FCBE0204FCC5" box="[151,314,852,874]" gridcol="0" gridrow="1" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFCBF022FFCC5" authorityName="Fage" authorityYear="1931" box="[159,273,852,874]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="bicornis">
<emphasis id="BC860D035774FFC9FF2AFCBF022FFCC5" box="[159,273,852,874]" italics="true" pageId="4" pageNumber="379">D. bicornis</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FCBE0058FCC5" box="[321,870,852,874]" gridcol="1" gridrow="1" pageId="4" pageNumber="379">Fe I–III with spines, chelicer with 4 teeth</td>
<td id="7313BA2F57740032FCC9FCBE06B3FCC5" box="[892,1421,852,874]" gridcol="2" gridrow="1" pageId="4" pageNumber="379">Fe I–III withoUt spines, chelicer with 3 teeth</td>
</tr>
<tr id="30C2D35357740032FF22FC9706B3FC3C" box="[151,1421,893,915]" gridrow="2" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FC970204FC3C" box="[151,314,893,915]" gridcol="0" gridrow="2" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFC94022AFC3C" authorityName="Ribera &amp; Ferrandez" authorityYear="1986" box="[159,276,893,915]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="BC860D035774FFC9FF2AFC94022AFC3C" box="[159,276,893,915]" italics="true" pageId="4" pageNumber="379">D. espanoli</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FC970058FC3C" box="[321,870,893,915]" gridcol="1" gridrow="2" pageId="4" pageNumber="379">Fe I–III spinose</td>
<td id="7313BA2F57740032FCC9FC9706B3FC3C" box="[892,1421,893,915]" gridcol="2" gridrow="2" pageId="4" pageNumber="379">Fe I–III withoUt spines</td>
</tr>
<tr id="30C2D35357740032FF22FC4C06B3FC74" box="[151,1421,934,987]" gridrow="3" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FC4C0204FC74" box="[151,314,934,987]" gridcol="0" gridrow="3" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFC42023CFC12" authorityName="Caporiacco" authorityYear="1941" box="[159,258,936,957]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="pavani">
<emphasis id="BC860D035774FFC9FF2AFC42023CFC12" box="[159,258,936,957]" italics="true" pageId="4" pageNumber="379">D. pavani</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FC4C0058FC74" box="[321,870,934,987]" gridcol="1" gridrow="3" pageId="4" pageNumber="379">Prosoma with pUnctUres, all legs spineless</td>
<td id="7313BA2F57740032FCC9FC4C06B3FC74" box="[892,1421,934,987]" gridcol="2" gridrow="3" pageId="4" pageNumber="379">Prosoma withoUt pUnctUres, some leg segments with spines</td>
</tr>
<tr id="30C2D35357740032FF22FC0406B3FBAA" box="[151,1421,1006,1029]" gridrow="4" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FC040204FBAA" box="[151,314,1006,1029]" gridcol="0" gridrow="4" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFC1A0222FBAB" authorityName="Ribera" authorityYear="2004" box="[159,284,1006,1029]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="valentina">
<emphasis id="BC860D035774FFC9FF2AFC1A0222FBAB" box="[159,284,1006,1029]" italics="true" pageId="4" pageNumber="379">D. valentina</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FC040058FBAA" box="[321,870,1006,1029]" gridcol="1" gridrow="4" pageId="4" pageNumber="379">Chelicer with two teeth, leg III spineless</td>
<td id="7313BA2F57740032FCC9FC0406B3FBAA" box="[892,1421,1006,1029]" gridcol="2" gridrow="4" pageId="4" pageNumber="379">Chelicer with three teeth, leg III with many spines</td>
</tr>
<tr id="30C2D35357740032FF22FBF206B3FB81" box="[151,1421,1048,1070]" gridrow="5" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FBF20204FB81" box="[151,314,1048,1070]" gridcol="0" gridrow="5" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFBF303C6FB81" authorityName="Ribera &amp; Ferrandez" authorityYear="1986" box="[159,248,1049,1070]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="BC860D035774FFC9FF2AFBF303C6FB81" box="[159,248,1049,1070]" italics="true" pageId="4" pageNumber="379">D. vivesi</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FBF20058FB81" box="[321,870,1048,1070]" gridcol="1" gridrow="5" pageId="4" pageNumber="379">All femora with nUmeroUs spines</td>
<td id="7313BA2F57740032FCC9FBF206B3FB81" box="[892,1421,1048,1070]" gridcol="2" gridrow="5" pageId="4" pageNumber="379">Only femora IV with spines</td>
</tr>
<tr id="30C2D35357740032FF22FBAB06B3FBD9" box="[151,1421,1089,1142]" gridrow="6" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FBAB0204FBD9" box="[151,314,1089,1142]" gridcol="0" gridrow="6" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFBA803C1FBF8" authorityName="Simon" authorityYear="1882" box="[159,255,1089,1111]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="nubila">
<emphasis id="BC860D035774FFC9FF2AFBA803C1FBF8" box="[159,255,1089,1111]" italics="true" pageId="4" pageNumber="379">D. nubila</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FBAB0058FBD9" box="[321,870,1089,1142]" gridcol="1" gridrow="6" pageId="4" pageNumber="379">FemUr I with spines, femUr IV with 2–3 spines, in litter</td>
<td id="7313BA2F57740032FCC9FBAB06B3FBD9" box="[892,1421,1089,1142]" gridcol="2" gridrow="6" pageId="4" pageNumber="379">FemUr I spineless, femUr IV with 6 spines, in caves</td>
</tr>
<tr id="30C2D35357740032FF22FB6306B3FB11" box="[151,1421,1161,1214]" gridrow="7" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FB630204FB11" box="[151,314,1161,1214]" gridcol="0" gridrow="7" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFB60023CFB30" authorityName="Alicata" authorityYear="1973" box="[159,258,1161,1183]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="osellai">
<emphasis id="BC860D035774FFC9FF2AFB60023CFB30" box="[159,258,1161,1183]" italics="true" pageId="4" pageNumber="379">D. osellai</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FB630058FB11" box="[321,870,1161,1214]" gridcol="1" gridrow="7" pageId="4" pageNumber="379">FemUr I, III with spines, spermatheca recUrv in dorsal view</td>
<td id="7313BA2F57740032FCC9FB6306B3FB11" box="[892,1421,1161,1214]" gridcol="2" gridrow="7" pageId="4" pageNumber="379">FemUr I, III spineless, spermatheca straight in dorsal view</td>
</tr>
<tr id="30C2D35357740032FF22FB3B06B3FA8B" box="[151,1421,1233,1316]" gridrow="8" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FB3B0204FA8B" box="[151,314,1233,1316]" gridcol="0" gridrow="8" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FF2AFB38021CFB48" authorityName="Simon" authorityYear="1907" box="[159,290,1233,1255]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="subnubila">
<emphasis id="BC860D035774FFC9FF2AFB38021CFB48" box="[159,290,1233,1255]" italics="true" pageId="4" pageNumber="379">D. subnubila</emphasis>
</taxonomicName>
</th>
<td id="7313BA2F57740032FEF4FB3B0058FA8B" box="[321,870,1233,1316]" gridcol="1" gridrow="8" pageId="4" pageNumber="379">FemUr I with spines, spermatheca recUrv in dorsal view, dorsal arch of anterior diverticUlUm longer than wide</td>
<td id="7313BA2F57740032FCC9FB3B06B3FA8B" box="[892,1421,1233,1316]" gridcol="2" gridrow="8" pageId="4" pageNumber="379">FemUr I spineless, spermatheca straight in dorsal view, dorsal arch of anterior diverticUlUm wider than long</td>
</tr>
<tr id="30C2D35357740032FF22FA8E06B3FAD3" box="[151,1421,1380,1404]" gridrow="9" pageId="4" pageNumber="379" rowspan-2="1">
<th id="7313BA2F57740032FF22FA8E0204FAD3" box="[151,314,1380,1404]" gridcol="0" gridrow="9" pageId="4" pageNumber="379">
<emphasis id="BC860D035774FFC9FF22FA8F0237FAD3" bold="true" box="[151,265,1381,1404]" pageId="4" pageNumber="379">TABLE 2.</emphasis>
Leg
</th>
<td id="7313BA2F57740032FEF4FA8E0058FAD3" box="[321,870,1380,1404]" gridcol="1" gridrow="9" pageId="4" pageNumber="379">
measurements of 
<taxonomicName id="49F2AA925774FFC9FE49FA8E018EFAD3" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[508,688,1380,1404]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FE49FA8E018EFAD3" box="[508,688,1380,1404]" italics="true" pageId="4" pageNumber="379">Dysdera arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FD02FA8F01CDFAD3" box="[695,755,1381,1404]" pageId="4" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8E4DD1115774FFC9FF2AFA7301AFF904" pageId="4" pageNumber="379">
<table id="FCF223B157740032FF22FA720666F904" box="[151,1368,1432,1707]" gridcols="9" gridrows="6" pageId="4" pageNumber="379">
<tr id="30C2D35357740032FF22FA720666FA01" box="[151,1368,1432,1454]" gridrow="0" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FA7203D1FA01" box="[151,239,1432,1454]" gridcol="0" gridrow="0" pageId="4" pageNumber="379">Leg</th>
<th id="7313BA2F57740032FF43FA720249FA01" box="[246,375,1432,1454]" gridcol="1" gridrow="0" pageId="4" pageNumber="379">Co</th>
<th id="7313BA2F57740032FEC8FA7202CAFA01" box="[381,500,1432,1454]" gridcol="2" gridrow="0" pageId="4" pageNumber="379">Tr</th>
<th id="7313BA2F57740032FE4EFA7201AFFA01" box="[507,657,1432,1454]" gridcol="3" gridrow="0" pageId="4" pageNumber="379">Fe</th>
<th id="7313BA2F57740032FD7AFA7201C5FA01" box="[719,763,1432,1454]" gridcol="4" gridrow="0" pageId="4" pageNumber="379">Pa</th>
<th id="7313BA2F57740032FCEEFA7200B9FA01" box="[859,903,1432,1454]" gridcol="5" gridrow="0" pageId="4" pageNumber="379">Ti</th>
<th id="7313BA2F57740032FC6DFA72073AFA01" box="[984,1028,1432,1454]" gridcol="6" gridrow="0" pageId="4" pageNumber="379">Mt</th>
<th id="7313BA2F57740032FB21FA7207FEFA01" box="[1172,1216,1432,1454]" gridcol="7" gridrow="0" pageId="4" pageNumber="379">Ta</th>
<th id="7313BA2F57740032FAAAFA720666FA01" box="[1311,1368,1432,1454]" gridcol="8" gridrow="0" pageId="4" pageNumber="379">Total</th>
</tr>
<tr id="30C2D35357740032FF22FA2B0666FA78" box="[151,1368,1473,1495]" gridrow="1" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FA2B03D1FA78" box="[151,239,1473,1495]" gridcol="0" gridrow="1" pageId="4" pageNumber="379">I</th>
<td id="7313BA2F57740032FF43FA2B0249FA78" box="[246,375,1473,1495]" gridcol="1" gridrow="1" pageId="4" pageNumber="379">1.93</td>
<td id="7313BA2F57740032FEC8FA2B02CAFA78" box="[381,500,1473,1495]" gridcol="2" gridrow="1" pageId="4" pageNumber="379">0.53</td>
<td id="7313BA2F57740032FE4EFA2B01AFFA78" box="[507,657,1473,1495]" gridcol="3" gridrow="1" pageId="4" pageNumber="379">4.19</td>
<td id="7313BA2F57740032FD7AFA2B01C5FA78" box="[719,763,1473,1495]" gridcol="4" gridrow="1" pageId="4" pageNumber="379">2.55</td>
<td id="7313BA2F57740032FCEEFA2B00B9FA78" box="[859,903,1473,1495]" gridcol="5" gridrow="1" pageId="4" pageNumber="379">3.66</td>
<td id="7313BA2F57740032FC6DFA2B073AFA78" box="[984,1028,1473,1495]" gridcol="6" gridrow="1" pageId="4" pageNumber="379">3.45</td>
<td id="7313BA2F57740032FB21FA2B07FEFA78" box="[1172,1216,1473,1495]" gridcol="7" gridrow="1" pageId="4" pageNumber="379">0.79</td>
<td id="7313BA2F57740032FAAAFA2B0666FA78" box="[1311,1368,1473,1495]" gridcol="8" gridrow="1" pageId="4" pageNumber="379">17.10</td>
</tr>
<tr id="30C2D35357740032FF22FA000666F9AF" box="[151,1368,1514,1536]" gridrow="2" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22FA0003D1F9AF" box="[151,239,1514,1536]" gridcol="0" gridrow="2" pageId="4" pageNumber="379">II</th>
<td id="7313BA2F57740032FF43FA000249F9AF" box="[246,375,1514,1536]" gridcol="1" gridrow="2" pageId="4" pageNumber="379">1.72</td>
<td id="7313BA2F57740032FEC8FA0002CAF9AF" box="[381,500,1514,1536]" gridcol="2" gridrow="2" pageId="4" pageNumber="379">0.41</td>
<td id="7313BA2F57740032FE4EFA0001AFF9AF" box="[507,657,1514,1536]" gridcol="3" gridrow="2" pageId="4" pageNumber="379">3.80</td>
<td id="7313BA2F57740032FD7AFA0001C5F9AF" box="[719,763,1514,1536]" gridcol="4" gridrow="2" pageId="4" pageNumber="379">2.27</td>
<td id="7313BA2F57740032FCEEFA0000B9F9AF" box="[859,903,1514,1536]" gridcol="5" gridrow="2" pageId="4" pageNumber="379">3.35</td>
<td id="7313BA2F57740032FC6DFA00073AF9AF" box="[984,1028,1514,1536]" gridcol="6" gridrow="2" pageId="4" pageNumber="379">3.28</td>
<td id="7313BA2F57740032FB21FA0007FEF9AF" box="[1172,1216,1514,1536]" gridcol="7" gridrow="2" pageId="4" pageNumber="379">0.76</td>
<td id="7313BA2F57740032FAAAFA000666F9AF" box="[1311,1368,1514,1536]" gridcol="8" gridrow="2" pageId="4" pageNumber="379">15.59</td>
</tr>
<tr id="30C2D35357740032FF22F9FE0666F985" box="[151,1368,1556,1578]" gridrow="3" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22F9FE03D1F985" box="[151,239,1556,1578]" gridcol="0" gridrow="3" pageId="4" pageNumber="379">III</th>
<td id="7313BA2F57740032FF43F9FE0249F985" box="[246,375,1556,1578]" gridcol="1" gridrow="3" pageId="4" pageNumber="379">1.10</td>
<td id="7313BA2F57740032FEC8F9FE02CAF985" box="[381,500,1556,1578]" gridcol="2" gridrow="3" pageId="4" pageNumber="379">0.40</td>
<td id="7313BA2F57740032FE4EF9FE01AFF985" box="[507,657,1556,1578]" gridcol="3" gridrow="3" pageId="4" pageNumber="379">3.06</td>
<td id="7313BA2F57740032FD7AF9FE01C5F985" box="[719,763,1556,1578]" gridcol="4" gridrow="3" pageId="4" pageNumber="379">1.60</td>
<td id="7313BA2F57740032FCEEF9FE00B9F985" box="[859,903,1556,1578]" gridcol="5" gridrow="3" pageId="4" pageNumber="379">2.23</td>
<td id="7313BA2F57740032FC6DF9FE073AF985" box="[984,1028,1556,1578]" gridcol="6" gridrow="3" pageId="4" pageNumber="379">2.85</td>
<td id="7313BA2F57740032FB21F9FE07FEF985" box="[1172,1216,1556,1578]" gridcol="7" gridrow="3" pageId="4" pageNumber="379">0.75</td>
<td id="7313BA2F57740032FAAAF9FE0666F985" box="[1311,1368,1556,1578]" gridcol="8" gridrow="3" pageId="4" pageNumber="379">11.99</td>
</tr>
<tr id="30C2D35357740032FF22F9D70666F9FC" box="[151,1368,1597,1619]" gridrow="4" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF22F9D703D1F9FC" box="[151,239,1597,1619]" gridcol="0" gridrow="4" pageId="4" pageNumber="379">IV</th>
<td id="7313BA2F57740032FF43F9D70249F9FC" box="[246,375,1597,1619]" gridcol="1" gridrow="4" pageId="4" pageNumber="379">1.42</td>
<td id="7313BA2F57740032FEC8F9D702CAF9FC" box="[381,500,1597,1619]" gridcol="2" gridrow="4" pageId="4" pageNumber="379">0.48</td>
<td id="7313BA2F57740032FE4EF9D701AFF9FC" box="[507,657,1597,1619]" gridcol="3" gridrow="4" pageId="4" pageNumber="379">4.08</td>
<td id="7313BA2F57740032FD7AF9D701C5F9FC" box="[719,763,1597,1619]" gridcol="4" gridrow="4" pageId="4" pageNumber="379">2.09</td>
<td id="7313BA2F57740032FCEEF9D700B9F9FC" box="[859,903,1597,1619]" gridcol="5" gridrow="4" pageId="4" pageNumber="379">3.23</td>
<td id="7313BA2F57740032FC6DF9D7073AF9FC" box="[984,1028,1597,1619]" gridcol="6" gridrow="4" pageId="4" pageNumber="379">3.89</td>
<td id="7313BA2F57740032FB21F9D707FEF9FC" box="[1172,1216,1597,1619]" gridcol="7" gridrow="4" pageId="4" pageNumber="379">0.86</td>
<td id="7313BA2F57740032FAAAF9D70666F9FC" box="[1311,1368,1597,1619]" gridcol="8" gridrow="4" pageId="4" pageNumber="379">16.05</td>
</tr>
<tr id="30C2D35357740032FF22F9790666F904" box="[151,1368,1683,1707]" gridrow="5" pageId="4" pageNumber="379" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="7313BA2F57740032FF22F97903D1F904" box="[151,239,1683,1707]" gridcol="0" gridrow="5" pageId="4" pageNumber="379">
<emphasis id="BC860D035774FFC9FF22F97E03D1F904" bold="true" box="[151,239,1684,1707]" pageId="4" pageNumber="379">TABLE</emphasis>
</th>
<td id="7313BA2F57740032FF43F9790249F904" box="[246,375,1683,1707]" gridcol="1" gridrow="5" pageId="4" pageNumber="379">
<emphasis id="BC860D035774FFC9FF43F97E0237F904" bold="true" box="[246,265,1684,1707]" pageId="4" pageNumber="379">3.</emphasis>
Spination
</td>
<td id="7313BA2F57740032FEC8F97902CAF904" box="[381,500,1683,1707]" gridcol="2" gridrow="5" pageId="4" pageNumber="379">
of 
<taxonomicName id="49F2AA925774FFC9FE2FF97902CAF904" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[410,500,1683,1707]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="BC860D035774FFC9FE2FF97902CAF904" box="[410,500,1683,1707]" italics="true" pageId="4" pageNumber="379">Dysdera</emphasis>
</taxonomicName>
</td>
<td id="7313BA2F57740032FE4EF97901AFF904" box="[507,657,1683,1707]" gridcol="3" gridrow="5" pageId="4" pageNumber="379">
<taxonomicName id="49F2AA925774FFC9FE4EF9790170F904" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[507,590,1683,1707]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035774FFC9FE4EF9790170F904" box="[507,590,1683,1707]" italics="true" pageId="4" pageNumber="379">arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785774FFC9FDE0F97E01AFF904" box="[597,657,1684,1707]" pageId="4" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8E4DD1115774FFC9FF2AF92D062EF880" pageId="4" pageNumber="379">
<table id="FCF223B157740032FF2AF92C062EF880" box="[159,1296,1734,1839]" gridcols="4" gridrows="3" pageId="4" pageNumber="379">
<tr id="30C2D35357740032FF2AF92C062EF973" box="[159,1296,1734,1756]" gridrow="0" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF2AF92C03F9F973" box="[159,199,1734,1756]" gridcol="0" gridrow="0" pageId="4" pageNumber="379">Leg</th>
<th id="7313BA2F57740032FEA0F92C0283F973" box="[277,445,1734,1756]" gridcol="1" gridrow="0" pageId="4" pageNumber="379">Fe</th>
<th id="7313BA2F57740032FDA2F92C0016F973" box="[535,808,1734,1756]" gridcol="2" gridrow="0" pageId="4" pageNumber="379">Ti</th>
<th id="7313BA2F57740032FC03F92C062EF973" box="[950,1296,1734,1756]" gridcol="3" gridrow="0" pageId="4" pageNumber="379">Mt</th>
</tr>
<tr id="30C2D35357740032FF2AF91A062EF8A9" box="[159,1296,1776,1798]" gridrow="1" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF2AF91A03F9F8A9" box="[159,199,1776,1798]" gridcol="0" gridrow="1" pageId="4" pageNumber="379">III</th>
<td id="7313BA2F57740032FEA0F91A0283F8A9" box="[277,445,1776,1798]" gridcol="1" gridrow="1" pageId="4" pageNumber="379">-</td>
<td id="7313BA2F57740032FDA2F91A0016F8A9" box="[535,808,1776,1798]" gridcol="2" gridrow="1" pageId="4" pageNumber="379">(plrlpv)(plrlpv)(pvrv)</td>
<td id="7313BA2F57740032FC03F91A062EF8A9" box="[950,1296,1776,1798]" gridcol="3" gridrow="1" pageId="4" pageNumber="379">(pl)(plrlrv)(plrl)(pvrv)(pl)(pl*)</td>
</tr>
<tr id="30C2D35357740032FF2AF8F3062EF880" box="[159,1296,1817,1839]" gridrow="2" pageId="4" pageNumber="379">
<th id="7313BA2F57740032FF2AF8F303F9F880" box="[159,199,1817,1839]" gridcol="0" gridrow="2" pageId="4" pageNumber="379">IV</th>
<td id="7313BA2F57740032FEA0F8F30283F880" box="[277,445,1817,1839]" gridcol="1" gridrow="2" pageId="4" pageNumber="379">(d)(dpl)(d)(d)(d)</td>
<td id="7313BA2F57740032FDA2F8F30016F880" box="[535,808,1817,1839]" gridcol="2" gridrow="2" pageId="4" pageNumber="379">(plrlpvvrv)(plrlpvrv)(pvrv)</td>
<td id="7313BA2F57740032FC03F8F3062EF880" box="[950,1296,1817,1839]" gridcol="3" gridrow="2" pageId="4" pageNumber="379">(plrl)(pvrv)(plrl)(pvrv)(plrl)(pvrv)</td>
</tr>
</table>
</paragraph>
<paragraph id="8E4DD1115774FFC8FF72F88503E1FE7F" blockId="4.[151,1436,1903,2037]" lastBlockId="5.[151,1437,151,608]" lastPageId="5" lastPageNumber="380" pageId="4" pageNumber="379">
Adaptation to a cave life often leads to elongation of appendages (Kuntner 
<emphasis id="BC860D035774FFC9FB9DF89B0757F827" box="[1064,1129,1904,1928]" italics="true" pageId="4" pageNumber="379">et al.,</emphasis>
1999). The Mt I/ prosoma length ratio is proposed here as a way to quantify the degree of troglobiomorphism in 
<taxonomicName id="49F2AA925774FFC9FBEEF87E0784F803" authorityName="Hernández-Corral" authorityYear="2017" box="[1115,1210,1940,1964]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="BC860D035774FFC9FBEEF87E0784F803" box="[1115,1210,1940,1964]" italics="true" pageId="4" pageNumber="379">Dysdera</emphasis>
</taxonomicName>
, the higher the ratio the higher the degree of troglobiomorphism. 
<taxonomicName id="49F2AA925774FFC9FD24F85201CEF87F" authorityName="Hernández-Corral" authorityYear="2017" box="[657,752,1976,2000]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="BC860D035774FFC9FD24F85201CEF87F" box="[657,752,1976,2000]" italics="true" pageId="4" pageNumber="379">Dysdera</emphasis>
</taxonomicName>
has radiated into many species in the 
<collectingRegion id="4C361FF35774FFC9FB2BF852067BF87F" box="[1182,1349,1975,2000]" country="Spain" name="Canarias" pageId="4" pageNumber="379">Canary Islands</collectingRegion>
to both epigean and hypogean environments. Using data available in 
<bibRefCitation id="EA63ACE05774FFC9FCF7F836076DF85B" author="Arnedo" box="[834,1107,2012,2037]" pageId="4" pageNumber="379" refString="Arnedo, M. &amp; Ribera, C. (1999). Radiation of the spider genUs Dysdera (Araneae, Dysderidae) in the Canary Islands: the island of Tenerife. Journal of Arachnology, 27 (3), 604 - 662." type="journal article" year="1999">Arnedo &amp; Ribera (1999)</bibRefCitation>
Mt I/ prosoma length ratio of females average 0.59 (range 0.42–0.76) in epigean species (
<taxonomicName id="49F2AA925775FFC8FC89FF7307F0FF1F" authority="Arnedo &amp; Ribera, 1999" authorityName="Arnedo &amp; Ribera" authorityYear="1999" box="[828,1230,151,176]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="guayota">
<emphasis id="BC860D035775FFC8FC89FF730086FF1F" box="[828,952,153,176]" italics="true" pageId="5" pageNumber="380">D. guayota</emphasis>
<bibRefCitation id="EA63ACE05775FFC8FC75FF7D07F0FF1F" author="Arnedo" box="[960,1230,151,176]" pageId="5" pageNumber="380" refString="Arnedo, M. &amp; Ribera, C. (1999). Radiation of the spider genUs Dysdera (Araneae, Dysderidae) in the Canary Islands: the island of Tenerife. Journal of Arachnology, 27 (3), 604 - 662." type="journal article" year="1999">Arnedo &amp; Ribera, 1999</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FB6FFF7303ECFF7B" authority="Simon, 1883" authorityName="Simon" authorityYear="1883" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="BC860D035775FFC8FB6FFF73067BFF1F" box="[1242,1349,153,176]" italics="true" pageId="5" pageNumber="380">D. macra</emphasis>
Simon, 1883
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FF69FF570109FF7B" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[220,567,188,213]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="breviseta">
<emphasis id="BC860D035775FFC8FF69FF57025AFF7B" box="[220,356,188,212]" italics="true" pageId="5" pageNumber="380">D. breviseta</emphasis>
Wunderlich, 1992
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FDF7FF570081FF7B" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[578,959,188,213]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="BC860D035775FFC8FDF7FF5701D5FF7B" box="[578,747,189,212]" italics="true" pageId="5" pageNumber="380">D. minutissima</emphasis>
Wunderlich, 1992
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FC7CFF57077EFF7B" authorityName="C. L. Koch" authorityYear="1838" box="[969,1088,189,212]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="crocata">
<emphasis id="BC860D035775FFC8FC7CFF57077EFF7B" box="[969,1088,189,212]" italics="true" pageId="5" pageNumber="380">D. crocata</emphasis>
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FBFBFF57064BFF7B" authority="Simon, 1883" authorityName="Simon" authorityYear="1883" box="[1102,1397,188,213]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="cribellata">
<emphasis id="BC860D035775FFC8FBFBFF5707E3FF7B" box="[1102,1245,188,212]" italics="true" pageId="5" pageNumber="380">D. cribellata</emphasis>
Simon, 1883
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FA34FF57012AFF57" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="montanetensis">
<emphasis id="BC860D035775FFC8FA34FF570209FF58" italics="true" pageId="5" pageNumber="380">D. montanetensis</emphasis>
Wunderlich, 1992
</taxonomicName>
) and 0.76 (range 0.51–1.21) in hypogean, more or less eyeless species (
<taxonomicName id="49F2AA925775FFC8FA34FF0B02E5FEB3" authority="Wunderlich, 1991" authorityName="Wunderlich" authorityYear="1991" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="chioensis">
<emphasis id="BC860D035775FFC8FA34FF0B03C1FEB3" italics="true" pageId="5" pageNumber="380">D. chioensis</emphasis>
Wunderlich, 1991
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FE59FEEF00ABFEB2" authority="Ribera &amp; Blasco, 1986" authorityName="Ribera &amp; Blasco" authorityYear="1986" box="[492,917,260,285]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="esquiveli">
<emphasis id="BC860D035775FFC8FE59FEEF0146FEB3" box="[492,632,260,284]" italics="true" pageId="5" pageNumber="380">D. esquiveli</emphasis>
Ribera &amp; Blasco, 1986
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FC13FEEF064EFEB3" authority="Arnedo &amp; Ribera, 1999" authorityName="Arnedo &amp; Ribera" authorityYear="1999" box="[934,1392,260,285]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="hernandezi">
<emphasis id="BC860D035775FFC8FC13FEEF0774FEB3" box="[934,1098,260,284]" italics="true" pageId="5" pageNumber="380">D. hernandezi</emphasis>
<bibRefCitation id="EA63ACE05775FFC8FBE3FEEE064EFEB3" author="Arnedo" box="[1110,1392,260,285]" pageId="5" pageNumber="380" refString="Arnedo, M. &amp; Ribera, C. (1999). Radiation of the spider genUs Dysdera (Araneae, Dysderidae) in the Canary Islands: the island of Tenerife. Journal of Arachnology, 27 (3), 604 - 662." type="journal article" year="1999">Arnedo &amp; Ribera, 1999</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FA34FEEF0190FEEF" authority="Ribera, Ferrandez &amp; Blasco, 1985" authorityName="Ribera, Ferrandez &amp; Blasco" authorityYear="1985" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="ambulotenta">
<emphasis id="BC860D035775FFC8FA34FEEF021DFEEF" italics="true" pageId="5" pageNumber="380">D. ambulotenta</emphasis>
Ribera, Ferrández &amp; Blasco, 1985
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FD0FFEC3077CFEEF" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[698,1090,295,320]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="labradaensis">
<emphasis id="BC860D035775FFC8FD0FFEC30050FEEF" box="[698,878,296,320]" italics="true" pageId="5" pageNumber="380">D. labradaensis</emphasis>
Wunderlich, 1992
</taxonomicName>
, 
<taxonomicName id="49F2AA925775FFC8FBF8FEC30213FECA" authority="Ferrandez &amp; Blasco, 1985" authorityName="Ribera, Ferrandez &amp; Blasco" authorityYear="1985" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="unguimmanis">
<emphasis id="BC860D035775FFC8FBF8FEC30639FE90" box="[1101,1287,296,320]" italics="true" pageId="5" pageNumber="380">D. unguimmanis</emphasis>
Ferrández &amp; Blasco, 1985
</taxonomicName>
). The Mt I/ prosoma length ratio of the female 
<taxonomicName id="49F2AA925775FFC8FC82FEA7008EFECB" authority="Hernández-Corral, 2017" authorityName="Hernández-Corral" authorityYear="2017" box="[823,944,332,356]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="arnedoi" status="sp. nov.">
<emphasis id="BC860D035775FFC8FC82FEA7008EFECB" box="[823,944,332,356]" italics="true" pageId="5" pageNumber="380">D. arnedoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7B5B0785775FFC8FC02FEA600CBFECA" box="[951,1013,332,357]" pageId="5" pageNumber="380" rank="species">
<emphasis id="BC860D035775FFC8FC02FEA600CBFECA" bold="true" box="[951,1013,332,357]" pageId="5" pageNumber="380">n. sp.</emphasis>
</taxonomicNameLabel>
is 0.83, thus grouping well within the hypogean species. The ratio is only surpassed by 
<taxonomicName id="49F2AA925775FFC8FD65FE9B0767FE27" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[720,1113,367,392]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="labradaensis">
<emphasis id="BC860D035775FFC8FD65FE9B00B8FE27" box="[720,902,368,392]" italics="true" pageId="5" pageNumber="380">D. labradaensis</emphasis>
Wunderlich, 1992
</taxonomicName>
(0.87) and 
<taxonomicName id="49F2AA925775FFC8FB54FE9B0119FE02" authority="Ribera, Ferrandez &amp; Blasco, 1985" authorityName="Ribera, Ferrandez &amp; Blasco" authorityYear="1985" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="species" species="unguimmanis">
<emphasis id="BC860D035775FFC8FB54FE9B06A2FE28" box="[1249,1436,368,392]" italics="true" pageId="5" pageNumber="380">D. unguimmanis</emphasis>
Ribera, Ferrández &amp; Blasco, 1985
</taxonomicName>
(1.21), the latter an extreme troglobiomorphic 
<taxonomicName id="49F2AA925775FFC8FBE3FE7E03E5FE7F" authority="(Arnedo &amp; Ribera, 1999)" baseAuthorityName="Arnedo &amp; Ribera" baseAuthorityYear="1999" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="BC860D035775FFC8FBE3FE7E078BFE03" box="[1110,1205,404,428]" italics="true" pageId="5" pageNumber="380">Dysdera</emphasis>
(
<bibRefCitation id="EA63ACE05775FFC8FB7DFE7E03EDFE7F" author="Arnedo" pageId="5" pageNumber="380" refString="Arnedo, M. &amp; Ribera, C. (1999). Radiation of the spider genUs Dysdera (Araneae, Dysderidae) in the Canary Islands: the island of Tenerife. Journal of Arachnology, 27 (3), 604 - 662." type="journal article" year="1999">Arnedo &amp; Ribera, 1999</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C6E8829A5775FFC8FF72FE360172FDCF" pageId="5" pageNumber="380" type="distribution">
<paragraph id="8E4DD1115775FFC8FF72FE360172FDCF" blockId="5.[151,1437,151,608]" pageId="5" pageNumber="380">
<emphasis id="BC860D035775FFC8FF72FE36025EFE5A" bold="true" box="[199,352,476,501]" pageId="5" pageNumber="380">Distribution.</emphasis>
A Majorcan endemic known from the Cova Morella cave near Pollença. The species possibly also occurs in the Coves de Campanet caves situated 
<quantity id="490A7CF45775FFC8FCB9FE150060FDB7" box="[780,862,511,536]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="380" unit="km" value="5.6">5.6 km</quantity>
to the south of Cova Morella, because the DNA barcode sequence of an immature specimen from the latter cave is very similar to the one obtained from the new species (Miquel Arnedo, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>