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<document id="A80FDC3274B7EED79168376972CDCBC9" ID-CLB-Dataset="282358" ID-DOI="10.5852/ejt.2023.904.2323" ID-GBIF-Dataset="f7020c08-27b9-4282-b5b5-2b4dffffaf5a" ID-ISSN="2118-9773" ID-Zenodo-Dep="10123238" ID-ZooBank="C2FCC15D-1DE5-4198-B867-EE4C582BA689" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1699522769833" checkinUser="plazi" docAuthor="Lindsay, Kate &amp; Marshall, Stephen A." docDate="2023" docId="F62BA7121598FF5EFDA4FEB9110AA8C8" docLanguage="en" docName="EJT.2023.904.1-189.pdf" docOrigin="European Journal of Taxonomy 904" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2323/10097" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Scipopus (Phaeopterina) narupa Lindsay &amp; Marshall 2023, sp. nov." docType="treatment" docUuid="5034DA1D-9B72-4D92-A75F-8455C572AE1D" docUuidSource="ZooBank" docVersion="13" lastPageNumber="144" masterDocId="0A12DF6A1515FFD1FF80FFB0102FAB49" masterDocTitle="A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae)" masterLastPageNumber="189" masterPageNumber="1" pageNumber="142" updateTime="1710871134729" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A77EB7FE56E4C92155A90F0343A3868A">A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="1BC33C4401F8A789A51039A78B86021E">Lindsay, Kate</mods:namePart>
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<mods:affiliation id="1AFDFE7736C3F5E854204D7CF725472B">School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="6096362F9C5CC09BAEA736066F9F7265" type="email">klindsay@uoguelph.ca</mods:nameIdentifier>
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<mods:namePart id="6FBD5F7C499D88DD96658088B57FD523">Marshall, Stephen A.</mods:namePart>
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<mods:affiliation id="0A0A23CDB2233B314F18F0D1397B810D">School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="83F907EE48E72B1DBB75C326FA4EE54E" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
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<mods:title id="FB1C874DAF3950A40757977EFE89BCD2">European Journal of Taxonomy</mods:title>
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<mods:date id="C6DFFDE7738724B18F886A92BA7FEF53">2023</mods:date>
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<treatment id="F62BA7121598FF5EFDA4FEB9110AA8C8" ID-DOI="http://doi.org/10.5281/zenodo.10406247" ID-GBIF-Taxon="219998626" ID-Zenodo-Dep="10406247" LSID="urn:lsid:zoobank.org:act:5034DA1D-9B72-4D92-A75F-8455C572AE1D" httpUri="http://treatment.plazi.org/id/F62BA7121598FF5EFDA4FEB9110AA8C8" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142">
<subSubSection id="3698458F1598FF5CFDA4FEB9143FAA6A" box="[548,1040,264,291]" pageId="141" pageNumber="142" type="nomenclature">
<paragraph id="7E3D16041598FF5CFDA4FEB9143FAA6A" blockId="141.[363,1225,264,361]" box="[548,1040,264,291]" pageId="141" pageNumber="142">
<heading id="2575A1681598FF5CFDA4FEB9143FAA6A" box="[548,1040,264,291]" centered="true" fontSize="11" level="2" pageId="141" pageNumber="142" reason="2">
<taxonomicName id="B9826D871598FF5CFDA4FEB91381AA6A" ID-CoL="C2NMB" authority="Lindsay &amp; Marshall, 2023" authorityName="Lindsay &amp; Marshall" authorityYear="2023" box="[548,942,264,291]" class="Insecta" family="Micropezidae" genus="Scipopus" kingdom="Animalia" order="Diptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="narupa" status="sp. nov." subGenus="Phaeopterina">
<emphasis id="4CF6CA161598FF5CFDA4FEB91381AA6A" bold="true" box="[548,942,264,291]" pageId="141" pageNumber="142">
<emphasis id="4CF6CA161598FF5CFDA4FEB912BEAA6A" bold="true" box="[548,657,265,291]" italics="true" pageId="141" pageNumber="142">Scipopus</emphasis>
(
<emphasis id="4CF6CA161598FF5CFD22FEB8136AAA6A" bold="true" box="[674,837,264,291]" italics="true" pageId="141" pageNumber="142">Phaeopterina</emphasis>
)
<emphasis id="4CF6CA161598FF5CFCD6FEB91381AA6A" bold="true" box="[854,942,265,291]" italics="true" pageId="141" pageNumber="142">narupa</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57C5776D1598FF5CFC35FEB9143FAA6A" box="[949,1040,265,291]" pageId="141" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3698458F1598FF5CFEEBFE9C1374AA20" pageId="141" pageNumber="142" type="description">
<paragraph id="7E3D16041598FF5CFEEBFE9C14E6AA0E" blockId="141.[363,1225,264,361]" box="[363,1225,300,327]" pageId="141" pageNumber="142">
<uri id="0A131A061598FF5CFEEBFE9C14E6AA0E" box="[363,1225,300,327]" pageId="141" pageNumber="142">
urn:lsid:zoobank.org:act:
<uuid id="0A242CD11598FF5CFD0CFE9C14E6AA0E" box="[652,1225,300,327]" pageId="141" pageNumber="142">5034DA1D-9B72-4D92-A75F-8455C572AE1D</uuid>
</uri>
</paragraph>
<paragraph id="7E3D16041598FF5CFD58FEFF1374AA20" blockId="141.[363,1225,264,361]" box="[728,859,335,361]" pageId="141" pageNumber="142">
<figureCitation id="E6B90A811598FF5CFD58FEFF1374AA20" box="[728,859,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="142.[189,232,1787,1813]" captionStartId-1="143.[189,232,1961,1987]" captionTargetBox-0="[268,1319,265,1737]" captionTargetBox-1="[189,1398,996,1922]" captionTargetId-0="figure-122@142.[268,1319,265,1737]" captionTargetId-1="figure-207@143.[189,1398,996,1922]" captionTargetPageId-0="142" captionTargetPageId-1="143" captionText-0="Fig. 62. Scipopus (Phaeopterina) narupa sp. nov. A. Head, ♀, anterolateral view, Ecuador, holotype (QCAZ). B. Female spermathecae and associated structures, Ecuador, non-type (DEBU). C. Head and thorax, dorsal view, holotype (QCAZ). D. Habitus, holotype (QCAZ). E. Male terminalia, lateral view, Ecuador, non-type (DEBU).F. Male genital fork, ventral view, Ecuador, non-type (DEBU).G. Living,♀, Ecuador. Abbreviations: bdp = basal distiphallus; e = epandrium; h = hypandrium; p =phallapodeme; pb = phallic bulb; pd= paired spermathecal duct; ps = paired spermatheca; pss = paired spermathecal stem; sd = single spermathecal duct; ss = single spermatheca." captionText-1="Fig. 63. Scipopus (Phaeopterina) narupa sp. nov. A. Living, ♀, Ecuador. B. Feeding, with abdomen upright, ♀, Ecuador (photo: K. Lindsay)." figureDoi-0="http://doi.org/10.5281/zenodo.10123425" figureDoi-1="http://doi.org/10.5281/zenodo.10123431" httpUri-0="https://zenodo.org/record/10123425/files/figure.png" httpUri-1="https://zenodo.org/record/10123431/files/figure.png" pageId="141" pageNumber="142">Figs 6263</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3698458F1598FF5CFF3DFE20152DAAB5" pageId="141" pageNumber="142" type="diagnosis">
<paragraph id="7E3D16041598FF5CFF3DFE2011E6AAE3" blockId="141.[189,457,400,426]" box="[189,457,400,426]" pageId="141" pageNumber="142">
<heading id="2575A1681598FF5CFF3DFE2011E6AAE3" bold="true" box="[189,457,400,426]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="4CF6CA161598FF5CFF3DFE2011E6AAE3" bold="true" box="[189,457,400,426]" pageId="141" pageNumber="142">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFE70152DAAB5" blockId="141.[189,1398,447,509]" pageId="141" pageNumber="142">
<taxonomicName id="B9826D871598FF5CFF3DFE70126FAA90" authority="Lindsay &amp; Marshall, 2023" authorityName="Lindsay &amp; Marshall" authorityYear="2023" box="[189,576,447,474]" class="Insecta" family="Micropezidae" genus="Scipopus" kingdom="Animalia" order="Diptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="narupa" status="sp. nov." subGenus="Phaeopterina">
<emphasis id="4CF6CA161598FF5CFF3DFE701108AA93" box="[189,295,448,474]" italics="true" pageId="141" pageNumber="142">Scipopus</emphasis>
(
<emphasis id="4CF6CA161598FF5CFEB9FE0F11F6AA90" box="[313,473,447,473]" italics="true" pageId="141" pageNumber="142">Phaeopterina</emphasis>
)
<emphasis id="4CF6CA161598FF5CFE6BFE70126FAA90" box="[491,576,448,473]" italics="true" pageId="141" pageNumber="142">narupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57C5776D1598FF5CFDC9FE70128BAA93" box="[585,676,448,474]" pageId="141" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
is easily distinguished from consubgeners by the blue-black, wide epicephalon and the primarily dark brown or black body with dark brown or black legs.
</paragraph>
</subSubSection>
<subSubSection id="3698458F1598FF5CFF3DFD9413DCA925" box="[189,1011,548,621]" pageId="141" pageNumber="142" type="etymology">
<paragraph id="7E3D16041598FF5CFF3DFD94116BA977" blockId="141.[189,324,548,574]" box="[189,324,548,574]" pageId="141" pageNumber="142">
<heading id="2575A1681598FF5CFF3DFD94116BA977" bold="true" box="[189,324,548,574]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="4CF6CA161598FF5CFF3DFD94116BA977" bold="true" box="[189,324,548,574]" pageId="141" pageNumber="142">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFDE213DCA925" blockId="141.[189,1011,594,621]" box="[189,1011,594,621]" pageId="141" pageNumber="142">
The species name is a noun in apposition inspired by the
<typeStatus id="A139A8A61598FF5CFCDCFDE313A0A924" box="[860,911,595,621]" pageId="141" pageNumber="142">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="3698458F1598FF5CFF3DFD241186AFCC" pageId="141" pageNumber="142" type="materials_examined">
<paragraph id="7E3D16041598FF5CFF3DFD2411C3A9E7" blockId="141.[189,492,660,686]" box="[189,492,660,686]" pageId="141" pageNumber="142">
<heading id="2575A1681598FF5CFF3DFD2411C3A9E7" bold="true" box="[189,492,660,686]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="4CF6CA161598FF5CFF3DFD2411C3A9E7" bold="true" box="[189,492,660,686]" pageId="141" pageNumber="142">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D16041598FF5CFF6AFD731173A994" blockId="141.[189,1399,707,803]" box="[234,348,707,733]" pageId="141" pageNumber="142">
<emphasis id="4CF6CA161598FF5CFF6AFD731173A994" bold="true" box="[234,348,707,733]" pageId="141" pageNumber="142">
<typeStatus id="A139A8A61598FF5CFF6AFD731173A994" box="[234,348,707,733]" pageId="141" pageNumber="142" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFD5612E7A86A" blockId="141.[189,1399,707,803]" pageId="141" pageNumber="142">
<materialsCitation id="CEEA1C591598FF5CFF3DFD5612E7A86A" ID-GBIF-Occurrence="4465943626" collectingDate="2019-05-03" collectionCode="QCAZ" collectorName="K. Lindsay" country="Ecuador" elevation="1100" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" municipality="Narupayacu" pageId="141" pageNumber="142" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="holotype">
<collectingCountry id="069556941598FF5CFF3DFD561164A849" box="[189,331,742,768]" name="Ecuador" pageId="141" pageNumber="142">ECUADOR</collectingCountry>
<specimenCount id="6884DD8D1598FF5CFEE1FD5611A2A849" box="[353,397,742,768]" count="1" pageId="141" pageNumber="142" type="female">1 ♀</specimenCount>
;
<collectingRegion id="BC46D8E61598FF5CFE1BFD5611F2A849" box="[411,477,742,768]" country="Ecuador" name="Napo" pageId="141" pageNumber="142">Napo</collectingRegion>
,
<location id="7B5D40DF1598FF5CFE68FD561255A849" LSID="urn:lsid:plazi:treatment:F62BA7121598FF5EFDA4FEB9110AA8C8:7B5D40DF1598FF5CFE68FD561255A849" box="[488,634,742,768]" country="Ecuador" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" name="Narupayacu" pageId="141" pageNumber="142" stateProvince="Napo">Narupayacu</location>
;
<geoCoordinate id="1BB670C31598FF5CFD04FD541325A849" box="[644,778,740,768]" degrees="0" direction="north" minutes="43" orientation="latitude" pageId="141" pageNumber="142" precision="15" seconds="30" value="0.72499996">0°4330″ N</geoCoordinate>
,
<geoCoordinate id="1BB670C31598FF5CFC95FD541380A849" box="[789,943,740,768]" degrees="77" direction="west" minutes="46" orientation="longitude" pageId="141" pageNumber="142" precision="15" seconds="02" value="-77.76723">77°4602″ W</geoCoordinate>
;
<elevation id="F5AFF1371598FF5CFC3AFD561460A849" box="[954,1103,742,768]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="141" pageNumber="142" unit="m" value="1100.0">
<quantity id="B97ABBE11598FF5CFC3AFD56143FA849" box="[954,1040,742,768]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="141" pageNumber="142" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
;
<date id="0A3C30C41598FF5CFBDBFD5614C5A849" box="[1115,1258,742,768]" pageId="141" pageNumber="142" value="2019-05-03">
<collectingDate id="1A78C92C1598FF5CFBDBFD5614C5A849" box="[1115,1258,742,768]" pageId="141" pageNumber="142" value="2019-05-03">3 May 2019</collectingDate>
</date>
;
<collectorName id="D37773D21598FF5CFB75FD561558A849" box="[1269,1399,742,768]" pageId="141" pageNumber="142">K. Lindsay</collectorName>
leg.;
<collectionCode id="18938EC11598FF5CFF78FCB91167A86A" box="[248,328,777,803]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="141" pageNumber="142" type="Museum">QCAZ</collectionCode>
(photographed,
<figureCitation id="E6B90A811598FF5CFD89FCB91293A86A" box="[521,700,777,803]" captionStart="Fig" captionStartId="142.[189,232,1787,1813]" captionTargetBox="[268,1319,265,1737]" captionTargetId="figure-122@142.[268,1319,265,1737]" captionTargetPageId="142" captionText="Fig. 62. Scipopus (Phaeopterina) narupa sp. nov. A. Head, ♀, anterolateral view, Ecuador, holotype (QCAZ). B. Female spermathecae and associated structures, Ecuador, non-type (DEBU). C. Head and thorax, dorsal view, holotype (QCAZ). D. Habitus, holotype (QCAZ). E. Male terminalia, lateral view, Ecuador, non-type (DEBU).F. Male genital fork, ventral view, Ecuador, non-type (DEBU).G. Living,♀, Ecuador. Abbreviations: bdp = basal distiphallus; e = epandrium; h = hypandrium; p =phallapodeme; pb = phallic bulb; pd= paired spermathecal duct; ps = paired spermatheca; pss = paired spermathecal stem; sd = single spermathecal duct; ss = single spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10123425" httpUri="https://zenodo.org/record/10123425/files/figure.png" pageId="141" pageNumber="142">Fig. 62A, CD</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="7E3D16041598FF5CFF6AFCFA1148A82D" blockId="141.[189,1035,842,904]" box="[234,359,842,868]" pageId="141" pageNumber="142">
<emphasis id="4CF6CA161598FF5CFF6AFCFA1148A82D" bold="true" box="[234,359,842,868]" pageId="141" pageNumber="142">
<typeStatus id="A139A8A61598FF5CFF6AFCFA1148A82D" box="[234,359,842,868]" pageId="141" pageNumber="142" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFCDE1424A8C1" blockId="141.[189,1035,842,904]" box="[189,1035,875,904]" pageId="141" pageNumber="142">
<materialsCitation id="CEEA1C591598FF5CFF3DFCDE1427A8C1" ID-GBIF-Occurrence="4465943613" box="[189,1032,875,904]" collectingDate="2019-05-03" collectionCode="QCAZ" collectorName="K. Lindsay" country="Ecuador" elevation="1100" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" municipality="Narupayacu" pageId="141" pageNumber="142" specimenCount="12" specimenCount-female="11" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<collectingCountry id="069556941598FF5CFF3DFCDE1164A8C1" box="[189,331,878,904]" name="Ecuador" pageId="141" pageNumber="142">ECUADOR</collectingCountry>
• 1 ♂,
<specimenCount id="6884DD8D1598FF5CFE1EFCDE11C0A8C1" box="[414,495,878,904]" count="11" pageId="141" pageNumber="142" type="female">11 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="18938EC11598FF5CFC34FCDE1427A8C1" box="[948,1032,878,904]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="141" pageNumber="142" type="Museum">QCAZ</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFC1F11D6A880" blockId="141.[189,505,943,969]" box="[189,505,943,969]" pageId="141" pageNumber="142">
<heading id="2575A1681598FF5CFF3DFC1F11D6A880" bold="true" box="[189,505,943,969]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="4CF6CA161598FF5CFF3DFC1F11D6A880" bold="true" box="[189,505,943,969]" pageId="141" pageNumber="142">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFC6E1186AFCC" blockId="141.[189,1398,987,1157]" pageId="141" pageNumber="142">
<materialsCitation id="CEEA1C591598FF5CFF3DFC6E1145AF52" ID-GBIF-Occurrence="4465943587" collectingDate="2019-05-03" collectionCode="DEBU" collectorName="K. Lindsay" country="Ecuador" elevation="1100" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" municipality="Narupayacu" pageId="141" pageNumber="142" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Napo" typeStatus="paratype">
<collectingCountry id="069556941598FF5CFF3DFC6E1164A8B1" box="[189,331,990,1016]" name="Ecuador" pageId="141" pageNumber="142">ECUADOR</collectingCountry>
• 2 ♂♂,
<specimenCount id="6884DD8D1598FF5CFE28FC6E11C7A8B1" box="[424,488,990,1016]" count="4" pageId="141" pageNumber="142" type="female">4 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="18938EC11598FF5CFC16FC6E13C9A8B1" box="[918,998,990,1016]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="141" pageNumber="142">DEBU</collectionCode>
(♂,
<specimenCount id="6884DD8D1598FF5CFB95FC6E1404A8B1" box="[1045,1067,990,1016]" count="1" pageId="141" pageNumber="142" type="female"></specimenCount>
dissected and photographed,
<figureCitation id="E6B90A811598FF5CFF3DFBB1114CAF52" box="[189,355,1025,1051]" captionStart="Fig" captionStartId="142.[189,232,1787,1813]" captionTargetBox="[268,1319,265,1737]" captionTargetId="figure-122@142.[268,1319,265,1737]" captionTargetPageId="142" captionText="Fig. 62. Scipopus (Phaeopterina) narupa sp. nov. A. Head, ♀, anterolateral view, Ecuador, holotype (QCAZ). B. Female spermathecae and associated structures, Ecuador, non-type (DEBU). C. Head and thorax, dorsal view, holotype (QCAZ). D. Habitus, holotype (QCAZ). E. Male terminalia, lateral view, Ecuador, non-type (DEBU).F. Male genital fork, ventral view, Ecuador, non-type (DEBU).G. Living,♀, Ecuador. Abbreviations: bdp = basal distiphallus; e = epandrium; h = hypandrium; p =phallapodeme; pb = phallic bulb; pd= paired spermathecal duct; ps = paired spermatheca; pss = paired spermathecal stem; sd = single spermathecal duct; ss = single spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10123425" httpUri="https://zenodo.org/record/10123425/files/figure.png" pageId="141" pageNumber="142">Fig. 62B, EF</figureCitation>
)
</materialsCitation>
<materialsCitation id="CEEA1C591598FF5CFE01FBB1115AAF77" ID-GBIF-Occurrence="4465943544" collectingDate="2019-05-03" collectionCode="DEBU" collectorName="K. Lindsay" country="Ecuador" elevation="1100" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" municipality="Narupayacu" pageId="141" pageNumber="142" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="6884DD8D1598FF5CFE01FBB11183AF52" box="[385,428,1025,1051]" count="1" pageId="141" pageNumber="142" type="female">1 ♀</specimenCount>
; same collection data as for holotype; debu0399728/MYCRO810-20 sequenced for CO15;
<collectionCode id="18938EC11598FF5CFEA5FB94115AAF77" box="[293,373,1060,1086]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="141" pageNumber="142">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="CEEA1C591598FF5CFE0AFB941158AF2B" ID-GBIF-Occurrence="4465943355" collectingDate="2019-05-03" collectionCode="DEBU" collectorName="K. Lindsay" country="Ecuador" elevation="1100" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" municipality="Narupayacu" pageId="141" pageNumber="142" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="6884DD8D1598FF5CFE0AFB94119AAF77" box="[394,437,1060,1086]" count="1" pageId="141" pageNumber="142" type="female">1 ♀</specimenCount>
; same collection data as for holotype; debu0399675/MYCRO811-20 sequenced for CO15;
<collectionCode id="18938EC11598FF5CFEA7FBF81158AF2B" box="[295,375,1096,1122]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="141" pageNumber="142">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="CEEA1C591598FF5CFE10FBF7118AAFCC" ID-GBIF-Occurrence="4465943578" collectingDate="2019-05-03" collectionCode="DEBU" collectorName="K. Lindsay" country="Ecuador" elevation="1100" latitude="0.72499996" longLatPrecision="21" longitude="-77.76723" municipality="Narupayacu" pageId="141" pageNumber="142" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="6884DD8D1598FF5CFE10FBF71192AF2B" box="[400,445,1095,1122]" count="1" pageId="141" pageNumber="142" type="female">1 ♀</specimenCount>
; same collection data as for holotype; debu00399732/MYCRO816-20 sequenced for CO15;
<collectionCode id="18938EC11598FF5CFED1FBDB118AAFCC" box="[337,421,1131,1157]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="141" pageNumber="142">DEBU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3698458F1598FF5EFF3DFB1B14D7A971" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142" type="description">
<paragraph id="7E3D16041598FF5CFF3DFB1B1161AF8C" blockId="141.[189,334,1195,1221]" box="[189,334,1195,1221]" pageId="141" pageNumber="142">
<heading id="2575A1681598FF5CFF3DFB1B1161AF8C" bold="true" box="[189,334,1195,1221]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="4CF6CA161598FF5CFF3DFB1B1161AF8C" bold="true" box="[189,334,1195,1221]" pageId="141" pageNumber="142">Description</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFB6B1187AFBC" blockId="141.[189,424,1242,1269]" box="[189,424,1242,1269]" pageId="141" pageNumber="142">
<smallCapsWord id="78DB80D81598FF5CFF3DFB6B113AAFBC" baselines="1262,1262" box="[189,277,1243,1269]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Length" pageId="141" pageNumber="142">LENGTH</smallCapsWord>
.
<quantity id="B97ABBE11598FF5CFEA3FB6A118BAFBC" box="[291,420,1242,1269]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="141" pageNumber="142" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
.
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFAAC14FAAE8B" blockId="141.[189,1399,1307,1475]" pageId="141" pageNumber="142">
<smallCapsWord id="78DB80D81598FF5CFF3DFAAC10D3AE7F" baselines="1327,1327" box="[189,252,1308,1334]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="141" pageNumber="142">HEAD</smallCapsWord>
. Palpus orange, pale microtrichose and black setulose, narrow (length 4.5× height). Clypeus orange, width ~2.0 × height, entirely silvery microtrichose. Frontal vitta dull, orange, microtrichose, posteriorly tapered. Epicephalon black, shiny, bare, wide (width ⅔ or more of upper frontal vittal width at inner verticals), clearly delineated from upper frontal vitta. One of the two usual lower fronto-orbital setae absent, postocellar seta present or absent, all other head chaetotaxy well-developed.
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DFA5A133DADD8" blockId="141.[188,1400,1514,1681]" pageId="141" pageNumber="142">
<smallCapsWord id="78DB80D81598FF5CFF3DFA5A1137AD4D" baselines="1533,1533" box="[189,280,1514,1540]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="141" pageNumber="142">THORAX</smallCapsWord>
. Scutum black with a wide, silvery-blue median sheen. Female cervical sclerite relatively flat. Postpronotal lobe black, with dense tuft of setulae on anterior corner. Notopleuron black with silvery-blue microtrichia outlining anterior notopleural seta and extending onto ventral margin. Thorax black with blue sheen, entirely silvery microtrichose. Legs black; fore and hind tarsomere 1 black with ventral golden fringe. Wing evenly black-brown infuscate.
</paragraph>
<paragraph id="7E3D16041598FF5CFF3DF90812D3AD9A" blockId="141.[189,764,1720,1747]" box="[189,764,1720,1747]" pageId="141" pageNumber="142">
<smallCapsWord id="78DB80D81598FF5CFF3DF9081104AD9B" baselines="1739,1740" box="[189,299,1720,1746]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="141" pageNumber="142">ABDOMEN</smallCapsWord>
(J+
<specimenCount id="6884DD8D1598FF5CFEE6F9081155AD9B" box="[358,378,1720,1746]" count="1" pageId="141" pageNumber="142" type="female"></specimenCount>
). T1 with fine, long, white setae.
</paragraph>
<paragraph id="7E3D16041598FF5EFF3DF94A1100AA0F" blockId="141.[189,1399,1785,2023]" lastBlockId="143.[189,1398,265,326]" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142">
<smallCapsWord id="78DB80D81598FF5CFF3DF94A113BAC5D" baselines="1805,1805" box="[189,276,1786,1812]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="141" pageNumber="142">FEMALE</smallCapsWord>
<smallCapsWord id="78DB80D81598FF5CFE9EF94F11A9AC5B" baselines="1805" box="[286,390,1791,1810]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="141" pageNumber="142">ABDOMEN</smallCapsWord>
. Pleuron pale grey, dorsal and ventral third of P25 dark brown or grey, leaving an indistinct pale convex median line on pleuron (
<figureCitation id="E6B90A811598FF5CFD72F8AD1370AC7E" box="[754,863,1821,1847]" captionStart="Fig" captionStartId="142.[189,232,1787,1813]" captionTargetBox="[268,1319,265,1737]" captionTargetId="figure-122@142.[268,1319,265,1737]" captionTargetPageId="142" captionText="Fig. 62. Scipopus (Phaeopterina) narupa sp. nov. A. Head, ♀, anterolateral view, Ecuador, holotype (QCAZ). B. Female spermathecae and associated structures, Ecuador, non-type (DEBU). C. Head and thorax, dorsal view, holotype (QCAZ). D. Habitus, holotype (QCAZ). E. Male terminalia, lateral view, Ecuador, non-type (DEBU).F. Male genital fork, ventral view, Ecuador, non-type (DEBU).G. Living,♀, Ecuador. Abbreviations: bdp = basal distiphallus; e = epandrium; h = hypandrium; p =phallapodeme; pb = phallic bulb; pd= paired spermathecal duct; ps = paired spermatheca; pss = paired spermathecal stem; sd = single spermathecal duct; ss = single spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10123425" httpUri="https://zenodo.org/record/10123425/files/figure.png" pageId="141" pageNumber="142">Fig. 62G</figureCitation>
). T1+2 ~1.8 × length of T3; posterior margin of T2 ~1.4× as wide as T1. Oviscape black, medium-length black setulose and white microtrichose on anterior ½, ~3.0× length of T6. Spermathecal ducts arising independently from bursa copulatrix. Paired spermathecal duct short (4.0 × length of paired spermathecae), narrow basally, swollen distally. Paired spermathecal stems long, narrow, inornate. Paired spermathecae nearly crescent-shaped, tapered apically. Single spermathecal duct narrow, same diameter as paired duct at base, same length as paired duct, swollen apically. Single spermatheca large (larger than paired), elongate, cup-shaped, with sparse tubercles.
</paragraph>
<caption id="2AFD468C159BFF5FFF3DF94B13A0ACA1" ID-DOI="http://doi.org/10.5281/zenodo.10123425" ID-Zenodo-Dep="10123425" httpUri="https://zenodo.org/record/10123425/files/figure.png" pageId="142" pageNumber="143" startId="142.[189,232,1787,1813]" targetBox="[268,1319,265,1737]" targetPageId="142">
<paragraph id="7E3D1604159BFF5FFF3DF94B13A0ACA1" blockId="142.[189,1401,1787,2025]" pageId="142" pageNumber="143">
<emphasis id="4CF6CA16159BFF5FFF3DF94B1133AC5C" bold="true" box="[189,284,1787,1813]" pageId="142" pageNumber="143">Fig. 62.</emphasis>
<taxonomicName id="B9826D87159BFF5FFEA6F94B1283AC5C" authority="Lindsay &amp; Marshall, 2023" authorityName="Lindsay &amp; Marshall" authorityYear="2023" box="[294,684,1787,1813]" class="Insecta" family="Micropezidae" genus="Scipopus" kingdom="Animalia" order="Diptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="narupa" status="sp. nov." subGenus="Phaeopterina">
<emphasis id="4CF6CA16159BFF5FFEA6F94B11BFAC5C" box="[294,400,1787,1813]" italics="true" pageId="142" pageNumber="143">Scipopus</emphasis>
(
<emphasis id="4CF6CA16159BFF5FFE24F94B126BAC5C" box="[420,580,1787,1813]" italics="true" pageId="142" pageNumber="143">Phaeopterina</emphasis>
)
<emphasis id="4CF6CA16159BFF5FFDD7F94C1283AC5C" box="[599,684,1788,1813]" italics="true" pageId="142" pageNumber="143">narupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57C5776D159BFF5FFD36F94C133CAC5F" box="[694,787,1788,1814]" pageId="142" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="4CF6CA16159BFF5FFC9BF94B131FAC5C" bold="true" box="[795,816,1787,1813]" pageId="142" pageNumber="143">A</emphasis>
. Head, ♀, anterolateral view, Ecuador, holotype (QCAZ).
<emphasis id="4CF6CA16159BFF5FFEB0F8AE116BAC71" bold="true" box="[304,324,1822,1848]" pageId="142" pageNumber="143">B</emphasis>
. Female spermathecae and associated structures, Ecuador, non-type (DEBU).
<emphasis id="4CF6CA16159BFF5FFB62F8AE14D9AC71" bold="true" box="[1250,1270,1822,1848]" pageId="142" pageNumber="143">C</emphasis>
. Head and thorax, dorsal view, holotype (QCAZ).
<emphasis id="4CF6CA16159BFF5FFD0DF8F2128DAC15" bold="true" box="[653,674,1858,1884]" pageId="142" pageNumber="143">D</emphasis>
. Habitus, holotype (QCAZ).
<emphasis id="4CF6CA16159BFF5FFC7BF8F21421AC15" bold="true" box="[1019,1038,1858,1884]" pageId="142" pageNumber="143">E</emphasis>
. Male terminalia, lateral view, Ecuador, non-type (DEBU).
<emphasis id="4CF6CA16159BFF5FFD84F8D51239AC36" bold="true" box="[516,534,1893,1919]" pageId="142" pageNumber="143">F</emphasis>
. Male genital fork, ventral view, Ecuador, non-type (DEBU).
<emphasis id="4CF6CA16159BFF5FFB59F8D414DFAC37" bold="true" box="[1241,1264,1892,1918]" pageId="142" pageNumber="143">G</emphasis>
. Living,
<emphasis id="4CF6CA16159BFF5FFAD7F8D51542ACC8" bold="true" box="[1367,1389,1893,1921]" pageId="142" pageNumber="143"></emphasis>
, Ecuador. Abbreviations: bdp = basal distiphallus; e = epandrium; h = hypandrium; p =phallapodeme; pb = phallic bulb; pd= paired spermathecal duct; ps = paired spermatheca; pss = paired spermathecal stem; sd = single spermathecal duct; ss = single spermatheca.
</paragraph>
</caption>
<paragraph id="7E3D1604159AFF5EFF3DFEDE14D7A971" blockId="143.[189,1399,365,568]" pageId="143" pageNumber="144">
<smallCapsWord id="78DB80D8159AFF5EFF3DFEDE10D0AAC1" baselines="385,385" box="[189,255,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="143" pageNumber="144">MALE</smallCapsWord>
<smallCapsWord id="78DB80D8159AFF5EFE88FEC3115FAACF" baselines="385" box="[264,368,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="143" pageNumber="144">ABDOMEN</smallCapsWord>
. Pleuron pale grey, dorsal half of P25 dark brown. T1+2 ~2.0 × length of T3. Genital fork 2.5× as long as T6, arms converging; inner basal process
<emphasis id="4CF6CA16159AFF5EFC2EFE2213EDAAE7" bold="true" box="[942,962,402,430]" pageId="143" pageNumber="144">¹/6</emphasis>
the length of arm. Epandrium stout, length ~1.5× height, posteroventral margin short setose. Basiphallus frame-like. Basal distiphallus long (≈ length of epandrium). Phallic bulb large, elongate (length 3.0× height), irregular and complex, with multiple chambers, either terminating in a tube-like opening or a very short, wide distal distiphallus. Phallapodeme very broad and expanded apically. Anterior hypandrium broad and expanded.
</paragraph>
</subSubSection>
<subSubSection id="3698458F159AFF5EFF3DFDEF1304A858" pageId="143" pageNumber="144" type="discussion">
<paragraph id="7E3D1604159AFF5EFF3DFDEF1100A930" blockId="143.[189,303,607,633]" box="[189,303,607,633]" pageId="143" pageNumber="144">
<heading id="2575A168159AFF5EFF3DFDEF1100A930" bold="true" box="[189,303,607,633]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
<emphasis id="4CF6CA16159AFF5EFF3DFDEF1100A930" bold="true" box="[189,303,607,633]" pageId="143" pageNumber="144">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D1604159AFF5EFF3DFD3E1304A858" blockId="143.[189,1399,653,786]" pageId="143" pageNumber="144">
We observed
<taxonomicName id="B9826D87159AFF5EFEDCFD3E123DA9EE" authority="Lindsay &amp; Marshall, 2023" authorityName="Lindsay &amp; Marshall" authorityYear="2023" box="[348,530,653,680]" class="Insecta" family="Micropezidae" genus="Scipopus" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="5" phylum="Arthropoda" rank="species" species="narupa" status="sp. nov." subGenus="Phaeopterina">
<emphasis id="4CF6CA16159AFF5EFEDCFD3E115DA9E1" box="[348,370,654,680]" italics="true" pageId="143" pageNumber="144">S.</emphasis>
(
<emphasis id="4CF6CA16159AFF5EFE04FD3D118AA9EE" box="[388,421,653,679]" italics="true" pageId="143" pageNumber="144">Ph</emphasis>
.)
<emphasis id="4CF6CA16159AFF5EFE3DFD3E123DA9EE" box="[445,530,654,679]" italics="true" pageId="143" pageNumber="144">narupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57C5776D159AFF5EFD9AFD3E125BA9E1" box="[538,628,654,680]" pageId="143" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
feeding at a dung bait in Narupayacu,
<collectingCountry id="06955694159AFF5EFBBEFD3D148FA9EE" box="[1086,1184,653,679]" name="Ecuador" pageId="143" pageNumber="144">Ecuador</collectingCountry>
in a characteristic posture previously noted in
<emphasis id="4CF6CA16159AFF5EFD8AFD01120FA982" box="[522,544,689,715]" italics="true" pageId="143" pageNumber="144">S.</emphasis>
<taxonomicName id="B9826D87159AFF5EFDA8FD0112CAA982" authorityName="Hennig" authorityYear="1935" box="[552,741,689,715]" class="Insecta" family="Micropezidae" genus="Pseudeurybata" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="stigmatica" subGenus="Phaeopterina">
(
<emphasis id="4CF6CA16159AFF5EFDB1FD01127DA982" box="[561,594,689,715]" italics="true" pageId="143" pageNumber="144">Ph</emphasis>
.)
<emphasis id="4CF6CA16159AFF5EFDEBFD0212CAA982" box="[619,741,690,715]" italics="true" pageId="143" pageNumber="144">stigmatica</emphasis>
</taxonomicName>
, with the abdomen almost perpendicular to the feeding surface (
<figureCitation id="E6B90A81159AFF5EFEA3FD6411BFA9A7" box="[291,400,724,750]" captionStart="Fig" captionStartId="143.[189,232,1961,1987]" captionTargetBox="[189,1398,996,1922]" captionTargetId="figure-207@143.[189,1398,996,1922]" captionTargetPageId="143" captionText="Fig. 63. Scipopus (Phaeopterina) narupa sp. nov. A. Living, ♀, Ecuador. B. Feeding, with abdomen upright, ♀, Ecuador (photo: K. Lindsay)." figureDoi="http://doi.org/10.5281/zenodo.10123431" httpUri="https://zenodo.org/record/10123431/files/figure.png" pageId="143" pageNumber="144">Fig. 63B</figureCitation>
). A female of this species has been photographed with an associated pseudoscorpion (https://www.inaturalist.org/observations/31649795).
</paragraph>
</subSubSection>
<subSubSection id="3698458F159AFF5EFF3DFC88110AA8C8" box="[189,342,824,897]" pageId="143" pageNumber="144" type="distribution">
<paragraph id="7E3D1604159AFF5EFF3DFC881179A81B" blockId="143.[189,342,824,850]" box="[189,342,824,850]" pageId="143" pageNumber="144">
<heading id="2575A168159AFF5EFF3DFC881179A81B" bold="true" box="[189,342,824,850]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
<emphasis id="4CF6CA16159AFF5EFF3DFC881179A81B" bold="true" box="[189,342,824,850]" pageId="143" pageNumber="144">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="7E3D1604159AFF5EFF3DFCD7110AA8C8" blockId="143.[189,293,871,897]" box="[189,293,871,897]" pageId="143" pageNumber="144">
<collectingCountry id="06955694159AFF5EFF3DFCD7110EA8C8" box="[189,289,871,897]" name="Ecuador" pageId="143" pageNumber="144">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>